208 lines
34 KiB
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208 lines
34 KiB
XML
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<mods:title id="AFE3857DA6D18778D8FB1D924069FBE5">Review Of The Genus Paraploderus Herman, 1970 (Coleoptera: Staphylinidae: Oxytelinae)</mods:title>
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<mods:namePart id="84E491AE89D2F9A170E575E9A533D5A4">Makranczy, György</mods:namePart>
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<mods:title id="ADBBB4F0D58EA1047828968465177D3D">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
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<mods:date id="602DC3A747B4735AAB9C5DA2FD391C54">2016</mods:date>
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<treatment id="631587C1FFC9FFB47D9730EA46634B19" LSID="urn:lsid:plazi:treatment:631587C1FFC9FFB47D9730EA46634B19" httpUri="http://treatment.plazi.org/id/631587C1FFC9FFB47D9730EA46634B19" lastPageId="8" lastPageNumber="67" pageId="2" pageNumber="61">
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<subSubSection id="A3A6655CFFC9FFBE7D9730EA414E4A61" pageId="2" pageNumber="61" type="nomenclature">
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<paragraph id="EB0336D7FFC9FFBE7D9730EA410C4A52" blockId="2.[411,814,1002,1028]" box="[411,814,1002,1028]" pageId="2" pageNumber="61">
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<heading id="B04B81BBFFC9FFBE7D9730EA410C4A52" box="[411,814,1002,1028]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="61" reason="2">
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<treatmentCitationGroup id="CBAC11F9FFC9FFBE7D9730EA410C4A52" box="[411,814,1002,1028]" pageId="2" pageNumber="61">
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<treatmentCitation id="6A1D10C6FFC9FFBE7D9730EA410C4A52" author="HeRmAN, L. H." box="[411,814,1002,1028]" page="400" pageId="2" pageNumber="61" year="1970">
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<taxonomicName id="2CBC4D54FFC9FFBE7D9730EA410C4A52" ID-CoL="8XNN6" authority="Herman, 1970: 400" authorityName="HERMAN" authorityYear="1970" box="[411,814,1002,1028]" class="Insecta" family="Staphylinidae" genus="Paraploderus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">
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<emphasis id="D9C8EAC5FFC9FFBE7D9730EA40154A52" box="[411,567,1002,1028]" italics="true" pageId="2" pageNumber="61">Paraploderus</emphasis>
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<bibRefCitation id="8F2D4B26FFC9FFBE7E3230EA410C4A52" author="HeRmAN, L. H." box="[574,814,1002,1028]" pageId="2" pageNumber="61" pagination="343 - 454" refId="ref18546" refString="HeRmAN, L. H. (1970) Phylogenyandreclassificationofthegeneraoftherove-beetlesubfamilyOxytelinaeoftheWorld (Coleoptera, Staphylinidae). BulletinoftheAmerican Museum of Natural History 142 (5): 343 - 454." type="journal article" year="1970">Herman, 1970: 400</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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</treatmentCitationGroup>
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</heading>
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</paragraph>
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<paragraph id="EB0336D7FFC9FFBE7C843722414E4A61" blockId="2.[136,876,1058,1080]" box="[136,876,1058,1080]" pageId="2" pageNumber="61">
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Typespecies:
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<emphasis id="D9C8EAC5FFC9FFBE7D103722402F4A61" box="[284,525,1058,1079]" italics="true" pageId="2" pageNumber="61">Aploderusspeculiventris</emphasis>
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Fauvel (byoriginaldesignation).
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</paragraph>
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</subSubSection>
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<subSubSection id="A3A6655CFFC9FFB47CC9375C46634B19" lastPageId="8" lastPageNumber="67" pageId="2" pageNumber="61" type="diagnosis">
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<paragraph id="EB0336D7FFC9FFBE7CC9375C41A64BBF" blockId="2.[136,1089,1116,1513]" pageId="2" pageNumber="61">
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Diagnosis –
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<taxonomicName id="2CBC4D54FFC9FFBE7D54375C43CE4A20" authorityName="HERMAN" authorityYear="1970" box="[344,492,1116,1142]" class="Insecta" family="Staphylinidae" genus="Paraploderus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="genus">
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<emphasis id="D9C8EAC5FFC9FFBE7D54375C43CE4A20" box="[344,492,1116,1142]" italics="true" pageId="2" pageNumber="61">Paraploderus</emphasis>
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</taxonomicName>
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canberecognizedbyitsrathershinyandsparselysetosebodywithweaksclerotization, abdomenandlegssomewhattranslucentbutoftenalsopartsofforebody. Mostspecieshavevaryingdegreeof punctationatleastontheforebody, butpuncturesarewithindefiniteedges, mostlycontributingtobodysurfaceperceivedasuneven. Headisrathershort, broadandtrapezoid, allspecieshavebulgingtemples, evenlycurved, though eye/templeratiovariesasalsowhetherheadisbroadestateyeortemples. Neckisdelimitedbyadistincttransversalgroove (occipitalgroove). Abdomenisfusiform, sterniteIXabsent, abdominaldefensiveglandopenings (in the form of a slot) are in unique position in tergites IX along the sides of tergite X. ThepeculiarformoftergiteXmakestheabdominalapexreminiscentofa horizontalfish-tail (slightlyincisedanddepressedinmiddle).
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</paragraph>
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<paragraph id="EB0336D7FFC8FFBF7CE733BE431A4C51" blockId="3.[174,1127,190,1594]" pageId="3" pageNumber="62">Description – Smalltomediumsized (1.8–4.0 mm). Bodyrathershiny andoftenslightlytranslucent. Forebodysparsely, ratherfinelypubescent, mostsetaeshort, onlyfewdarkerandslightlylongerinthefollowingpositions: atinneredgeofsupraantennalprominences, upperanterioredgeof eye, innermarginofeye, dorsalsideoftemplebehindeye, middleofimpressionsonpronotaldisc, pairofsetaeinproximityatandslightlybehindante- riorpronotalcornerandoneslightlybeforeposteriorcorner (oftenashorter onebehindmiddleofside); darkersetaeonouterelytraledgesoshortthat almostinvisibleindorsalview, fromlateralviewmoreapparentforbeing moreerect.</paragraph>
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<paragraph id="EB0336D7FFC8FFBF7CE7310E4645486C" blockId="3.[174,1127,190,1594]" pageId="3" pageNumber="62">Head. Clypeus (Fig. 1) ratherlarge (longandbroad), rectangular, mostlyshinyandslightlyconvex, delimitedbyadistinct, impressedepistomal suture, lessmarkedonsides; anteriormarginofclypeustruncatewiththin marginalbeadanteriorlyandlaterally. Dorsumofheadusuallywithstrong, butindistinct-borderedandscattered (unevenlydistributed) punctures, rest ofsurfaceshiny. Supraantennalprominenceswelldevelopedbutmoderately elevatedfromplaneofvertex; latterdelimitedfromthembyapairoflongitudinalimpressionsstartingfromoccipitalgrooveandmoreorlessvanishinganteriorly. Compoundeyesvarylessinsizethaninotherrelatedgenera, alwaysbulging, mediumroughlyfacetedonalmostentiresurface, however withashinyrimsurroundingitposteriorlyandformingasteplikeridgebeforetemples. Gularsuturesconfluentanteriorly; suturesnarrowlyseparate atanteriorneck, sharplydivergentfrommiddletoposteriorneck. Genalarea (temples) moresculptured, graduallycondensingpuncturesoftenturntoa ruggedsurfaceonthelateralparts. Baseofheadverystronglyconstrictedto formawelldefinedneck, delimitedbystronglyimpressedoccipitalgroove; latterwithsidepartsturningontouppersurfaceofheadinslightlyanterior direction, middlepartsometimesslightlyseparated. Antennaslightlyincrassateapically, firstthreesegmentsquiteelongate (articles 2 and 3 similarin length, togetherslightlyshorterthanscape), furtherarticleswithbasaldish, sculpturedandunlikerestofbodystronglysetose. Labrum (Fig. 2) divided inlongitudinalmidline, anterioredgerathertruncate, lateralportionsbearing membranouslobeswithmesialmarginfimbriate. Mandible (Fig. 4) denticulate, apexbifurcate, afurther, smallertoothbetweenapexandaknob-like tooth (anteriorlydemarcatedbyincision) atapical 1/3; prosthecaextending frominneredge, almostaslongashalfthelengthofentiremandible, composedofratherlongandslenderprocesses. Maxilla (Fig. 9) withcardosemi- triangular, stipeslikeperfectright-angledtriangle, laciniawithslenderlobe, galealarger, bothwithdensesetationatapex. Maxillarypalpuswithfirstseg- mentverysmallandring- like, secondlargeandapicallyincrassate, thirdeven larger (andalittlelonger), moreoval, aboutaswideatbaseasatapex, fourth almostaslongassecondbutstronglynarrowingtowardsmiddleandapically</paragraph>
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<paragraph id="EB0336D7FFCFFFB87C8436EC46634848" blockId="4.[136,1089,1516,1596]" pageId="4" pageNumber="63">
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<emphasis id="D9C8EAC5FFCFFFB87C8436EC42C44857" bold="true" box="[136,230,1516,1538]" pageId="4" pageNumber="63">Figs 1–7.</emphasis>
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<taxonomicName id="2CBC4D54FFCFFFB87CE736EC438A4854" authorityName="Makranczy" authorityYear="2016" box="[235,424,1516,1538]" class="Insecta" family="Staphylinidae" genus="Paraploderus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="wittei" status="sp. nov.">
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<emphasis id="D9C8EAC5FFCFFFB87CE736EC438A4854" box="[235,424,1516,1538]" italics="true" pageId="4" pageNumber="63">Paraploderus wittei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C2FB57BEFFCFFFB87DA136ED43C44854" box="[429,486,1517,1538]" pageId="4" pageNumber="63" rank="species">sp. n.</taxonomicNameLabel>
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1 = head capsule, 2 = labrum, 3 = mentum, 4 = mandibles, 5 = prothorax, 6 = scutellum, 7 = median pterothoracic parts. Scales:
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<quantity id="2C449B32FFCFFFB87F7F350941ED4848" box="[883,975,1545,1566]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="63" unit="mm" value="0.04">0.04 mm</quantity>
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for Fig. 3,
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</paragraph>
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<paragraph id="EB0336D7FFCFFFB87D1C35264198486D" blockId="4.[136,1089,1516,1596]" box="[272,954,1574,1596]" pageId="4" pageNumber="63">
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<quantity id="2C449B32FFCFFFB87D1C35264354486D" box="[272,374,1574,1596]" metricMagnitude="-5" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="63" unit="mm" value="0.045">0.045 mm</quantity>
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for Fig. 2,
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<quantity id="2C449B32FFCFFFB87DEA35264062486D" box="[486,576,1574,1596]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="63" unit="mm" value="0.06">0.06 mm</quantity>
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for Figs 4, 6,
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<quantity id="2C449B32FFCFFFB87EC135264139486D" box="[717,795,1574,1595]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="63" unit="mm" value="0.1">0.1 mm</quantity>
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for Figs 1, 5, 7.
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</paragraph>
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<paragraph id="EB0336D7FFCEFFB97CA233BE40364FB3" blockId="5.[174,1127,190,1594]" pageId="5" pageNumber="64">
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slender. Inlabium, mentum (Fig. 3) stronglytransversallyrectangular. Hy- popharynx (Fig. 8) withasetofbulboussetaenearapicaledge. Labialpalps three-segmentedsegments 1 and 3 almostofsamelength, segment 1 twiceas thickassegment 3, segment 2 slightlylongerthaneitherofothers, itsthick- nessintermediatebetweenothertwo. Apicalarticlewithgroupofshortsen- sillaeatapex. Plate-likearmatureinhypopharynx – introducedinMAKRANCZY (2006) andappliedinrecentworks (
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<bibRefCitation id="8F2D4B26FFCEFFB97E7B3288417B4FF7" author="MAKRANCZY, GY" box="[631,857,392,418]" pageId="5" pageNumber="64" pagination="77 - 133" refId="ref18692" refString="MAKRANCZY, GY. (2014 a) ReviewofthegenusParosusSharp, 1887 (Coleoptera: Staphylini- dae, Oxytelinae). Revue suisse de Zoologie 121 (1): 77 - 133." type="journal article" year="2014">
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MAKRANCZY 2014
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<emphasis id="D9C8EAC5FFCEFFB97F413288417B4FF7" box="[845,857,392,417]" italics="true" pageId="5" pageNumber="64">a</emphasis>
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</bibRefCitation>
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,
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<emphasis id="D9C8EAC5FFCEFFB97F6D3287414C4FF7" box="[865,878,391,417]" italics="true" pageId="5" pageNumber="64">b</emphasis>
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,
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<emphasis id="D9C8EAC5FFCEFFB97F79328841A24FF7" box="[885,896,392,417]" italics="true" pageId="5" pageNumber="64">c</emphasis>
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) notonlyasage- nericcharacterbutonediagnosticforspeciesgroupsinlargergenera – fully developed but rather simple.
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</paragraph>
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<paragraph id="EB0336D7FFCEFFB97CE732EC41934AF1" blockId="5.[174,1127,190,1594]" pageId="5" pageNumber="64">
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Thorax. Pronotum (Fig. 5) widestnearapex, lateralmarginsslightlycon- vergentposteriorly, bothanteriorandposteriorcornersrounded, lateralmar- ginsoftenslightlyconcaveanteriadposteriorcorners. Procoxalfissureabsent, protrochantinconcealed. Postprocoxallobeabsent, prohypomeronbroadand stronglydeflexed. Prosternalprocessascurvedkeelontransverserectangular furcasternum, shortlyextendingbetweenprocoxae. Scutellum (Fig. 6) with basallytransverserhomboidshaped, apicallymorearcuateimpression. Impressionanteriorlyconnectedtoproscutumbyamidlongitudinalinternal ridge (externallyasadarkline, ascratch-likeimpressiononsurface). Mesosternalprocess (Fig. 7) spiniform, metendosterniteslender, furcalarmsdeveloped, anteriorarmsbarelydiscernible. – Elytra. Discofelytrawithindistinctly borderedpuncturesarrangedmoreorlessrandomly. Elytralepipleuralridge present, well-markedexceptextremeanterior, slightlyfadingatouterposte- riorcorners. Suturemarkedbyunpunctured, slightlyelevatedmargin. – Legs. Tibiaewithmoderatelydevelopedmid-tibialspurs, distincttibialspinesabsentexceptatapex. Tarsomeres 3–3–3, basalarticlelongerthansecond, being threeformersegmentsfusedasevidentfromtheirapicalringsofsetaestill observable, apicalarticlelong, slender, slightlycurvedandonlywithacouple ofsetae. Formationoftarsallobeoutofapicalsetaeofsecondtarsomereonly evidentinhindleg; thisventralmodificationcharacteristicofmostoxytelines, butstronglyexpressedonlyinthe
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<taxonomicName id="2CBC4D54FFCEFFB97E57378C40FC4AF0" box="[603,734,1164,1190]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="64" phylum="Arthropoda" rank="genus">
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<emphasis id="D9C8EAC5FFCEFFB97E57378C40FC4AF0" box="[603,734,1164,1190]" italics="true" pageId="5" pageNumber="64">Carpelimus</emphasis>
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</taxonomicName>
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groupofgenera.
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</paragraph>
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<paragraph id="EB0336D7FFCEFFB97CE737AE4644486F" blockId="5.[174,1127,190,1594]" pageId="5" pageNumber="64">Abdomen. Intersegmentalmembranewithoutbrickwallpattern, second sternitefullydeveloped, firststernitecompletelyabsent. Abdomenspindleshaped, somewhatflatteneddorsally, twopairsoflateroscleritespresent. Whenabdomenlightcoloured, hindmarginoftergalfold (basalridge) onter- gitesIII–VIIwithadarkerline. Tergalbasolateralridgespresent, alsodarker marked, nolongitudinalmarksinmidline. Fimbriatehindmarginontergite VIIpresent, sterniteVIIwithoutmodifications. TergiteXsimilarinbothsexes, dorsoventrallyflattened, withmedialdepression, slightapicalemargination formingtwopeculiar‘ lobes’atapex; apexfimbriate, butasimilartransversal fimbriatelinerunningparalleltoapexataboutapical 1/4. Secondarysexual differencesslight, confinedtosomewhatdifferentapicalformationofster- nitesVIIIbetweenmalesandfemales. – Maleterminalia. TergiteVIIIwithout</paragraph>
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<paragraph id="EB0336D7FFCDFFBA7C8A36D4461B4816" blockId="6.[134,1087,1492,1601]" pageId="6" pageNumber="65">
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<emphasis id="D9C8EAC5FFCDFFBA7C8A36D442CC4BBF" bold="true" box="[134,238,1492,1514]" pageId="6" pageNumber="65">Figs 8–12.</emphasis>
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8–9 =
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<taxonomicName id="2CBC4D54FFCDFFBA7D3E36D443CE4BBC" authorityName="Makranczy" authorityYear="2016" box="[306,492,1492,1514]" class="Insecta" family="Staphylinidae" genus="Paraploderus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="wittei" status="sp. nov.">
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<emphasis id="D9C8EAC5FFCDFFBA7D3E36D443CE4BBC" box="[306,492,1492,1514]" italics="true" pageId="6" pageNumber="65">Paraploderus wittei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C2FB57BEFFCDFFBA7DFF36D540094BBC" box="[499,555,1493,1514]" pageId="6" pageNumber="65" rank="species">sp. n.</taxonomicNameLabel>
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8 = labium, 9 = maxilla. 10 =
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<taxonomicName id="2CBC4D54FFCDFFBA7F5136D5429C4851" authority="(Cameron, 1950)" baseAuthorityName="Cameron" baseAuthorityYear="1950" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="notabilis">
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<emphasis id="D9C8EAC5FFCDFFBA7F5136D541EB4BBF" box="[861,969,1492,1514]" italics="true" pageId="6" pageNumber="65">P. notabilis</emphasis>
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(Cameron, 1950)
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</taxonomicName>
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male tergites IX and X, ventral view. 11 =
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<taxonomicName id="2CBC4D54FFCDFFBA7E6736F241804851" authority="(Cameron, 1951)" baseAuthorityName="Cameron" baseAuthorityYear="1951" box="[619,930,1521,1543]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="nigronitens">
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<emphasis id="D9C8EAC5FFCDFFBA7E6736F240D14851" box="[619,755,1522,1543]" italics="true" pageId="6" pageNumber="65">P. nigronitens</emphasis>
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(Cameron, 1951)
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</taxonomicName>
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male tergite IX and X, dorsal view (defensive gland opening magnified 2×). 12 =
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<taxonomicName id="2CBC4D54FFCDFFBA7F16350E41594875" authorityName="Makranczy" authorityYear="2016" box="[794,891,1550,1571]" class="Insecta" family="Staphylinidae" genus="Paraploderus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
|
||
<emphasis id="D9C8EAC5FFCDFFBA7F16350E41594875" box="[794,891,1550,1571]" italics="true" pageId="6" pageNumber="65">P. grandis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C2FB57BEFFCDFFBA7F73350E41974875" box="[895,949,1550,1571]" pageId="6" pageNumber="65" rank="species">sp. n.</taxonomicNameLabel>
|
||
male sternite IX. Scales:
|
||
<quantity id="2C449B32FFCDFFBA7CF5352B43644816" box="[249,326,1579,1600]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="65" unit="mm" value="0.1">0.1 mm</quantity>
|
||
for Fig. 9,
|
||
<quantity id="2C449B32FFCDFFBA7DBF352B402E4816" box="[435,524,1579,1600]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="65" unit="mm" value="0.12">0.12 mm</quantity>
|
||
for Fig. 8,
|
||
<quantity id="2C449B32FFCDFFBA7E75352B40F04816" box="[633,722,1579,1600]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="65" unit="mm" value="0.17">0.17 mm</quantity>
|
||
for Fig. 12,
|
||
<quantity id="2C449B32FFCDFFBA7F5D352B41BC4816" box="[849,926,1579,1600]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="65" unit="mm" value="0.2">0.2 mm</quantity>
|
||
for Figs 10–11.
|
||
</paragraph>
|
||
<paragraph id="EB0336D7FFCCFFBB7CA233BE41374DCC" blockId="7.[174,1127,190,1527]" pageId="7" pageNumber="66">
|
||
peculiarmodification, apextruncate; TergiteIXpaired, withslenderbutnot tooelongate ‚ventralstruts’, platelaterallystronglyfusedtotergiteX (Fig. 11), formingacompactstructure (Fig. 10), defensiveglandopeningslongitudinal, slightlyobliqueintergiteIX, near, butdiscretefromplatefusionline. Ster- niteIXveryweaklydeveloped, entirelylostsetation, desclerotizedandmembranous, absolutelyinconspicuous (itis, however, moreapparent, spindleshaped, slightlyasymmetricanddenselyporousin
|
||
<emphasis id="D9C8EAC5FFCCFFBB7F3A3287460A4FF7" box="[822,1064,391,417]" italics="true" pageId="7" pageNumber="66">Paraploderusgrandis</emphasis>
|
||
, Fig. 12). – Malegenitalia. Aedeagustrilobed, medianlobebulb-like, scleritesand sclerotizedpartsconfinedmostlytoapicalpart: withonepairedblade-like openingprocessesandinbetweenoneunpairedappendage – latterreminiscenttoandperhapsprecursorofapicomedialhookin
|
||
<taxonomicName id="2CBC4D54FFCCFFBB7F58310E42C44C1C" authority="Gravenhorst, 1802" authorityName="Gravenhorst" authorityYear="1802" class="Insecta" family="Staphylinidae" genus="Oxytelus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D9C8EAC5FFCCFFBB7F58310E419E4C7E" box="[852,956,526,552]" italics="true" pageId="7" pageNumber="66">Oxytelus</emphasis>
|
||
Gravenhorst, 1802
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="8F2D4B26FFCCFFBB7CFB313043F34C1F" author="MAKRANCZY, GY" box="[247,465,559,586]" pageId="7" pageNumber="66" pagination="29 - 119" refId="ref18634" refString="MAKRANCZY, GY. (2006) Systematicsandphylogeneticrelationshipsofthegenerainthe Carpelimusgroup (Coleoptera: Staphylinidae: Oxytelinae). Annaleshistorico-naturales Musei nationalis hungarici 98: 29 - 119." type="journal article" year="2006">MAKRANCZY 2006</bibRefCitation>
|
||
, etc.) andmorerecentlydiscoveredwithin
|
||
<taxonomicName id="2CBC4D54FFCCFFBB7FF1312F434E4C3A" authority="Thomson, 1859" authorityName="Thomson" authorityYear="1859" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D9C8EAC5FFCCFFBB7FF1312F46444C1F" box="[1021,1126,559,585]" italics="true" pageId="7" pageNumber="66">Anotylus</emphasis>
|
||
Thomson, 1859
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="8F2D4B26FFCCFFBB7D713151407B4C3D" author="MAKRANCZY, GY" box="[381,601,593,619]" pageId="7" pageNumber="66" pagination="187 - 195" refId="ref18661" refString="MAKRANCZY, GY. (2013) DescriptionofanewAnotylusspeciesfromMadagascarandthe femaleofAnotylusflavipennisfromLaos (Coleoptera: Staphylinidae: Oxytelinae). Acta Zoologica Academiae Scientiarum Hungaricae 59 (3): 187 - 195." type="journal article" year="2013">MAKRANCZY 2013</bibRefCitation>
|
||
). Apicalpartwithavariouslywide, scle- rotized, reverseU-shapedsclerite (lengthofmedianlobeinkeymeasured fromapexofthissclerite), othersmuchlessconspicuous, moremembranous. Parameresextremelysetoseonmuchoflength (usuallyapical 2/3; uniquetrait inthesubfamily). Sizeofaedeagusvaryingoftendisproportionallytodiffer- encesinbodysize. – Femaleterminaliaverysimilartothatofmale, butter- gitesIXmissingelongate‘ventralstrut’. – Femalegenitalia. Spermathecascle- rotized, receptacledividedintoasmall, slendertubularportionandamore bulkybladder- likepartsometimesconstrictednearmiddle. Nosclerotized spermathecalglandobservable, umbilicusabsent.
|
||
</paragraph>
|
||
<paragraph id="EB0336D7FFCCFFBB7CE730A143EA4AD3" blockId="7.[174,1127,190,1527]" pageId="7" pageNumber="66">
|
||
Diversity – Only 6 nameshavebeenproposedforspeciesthatwerelater includedinortransferredto
|
||
<taxonomicName id="2CBC4D54FFCCFFBB7E1130C340934D8B" authorityName="HERMAN" authorityYear="1970" box="[541,689,963,989]" class="Insecta" family="Staphylinidae" genus="Paraploderus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D9C8EAC5FFCCFFBB7E1130C340934D8B" box="[541,689,963,989]" italics="true" pageId="7" pageNumber="66">Paraploderus</emphasis>
|
||
</taxonomicName>
|
||
, twoofthemhavenowfalleninto synonymy; withthecurrentcontributionthecountrisesto 15. Otherspecies may exist but most of them are hard to distinguish due to the rather simple anduniformbodybuiltandgenitaliawithveryfewdiagnostictraits. Itis believedthatthedifferencesbetweenspeciesarerathersubtleandsubjectto considerablevariation.
|
||
</paragraph>
|
||
<paragraph id="EB0336D7FFCCFFBB7CE7378C40144B19" blockId="7.[174,1127,190,1527]" pageId="7" pageNumber="66">
|
||
Distribution – Localitiesoftheexaminedmateriallayinforestedareas acrossalmosttheentireAfrotropicalandOrientalregions, althoughrecords areratherscattered; presenceofmaterialindicatesintensivecollectingactivity usingtheappropriatemethodstofind
|
||
<taxonomicName id="2CBC4D54FFCCFFBB7E9D37F141074B5D" authorityName="HERMAN" authorityYear="1970" box="[657,805,1265,1291]" class="Insecta" family="Staphylinidae" genus="Paraploderus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D9C8EAC5FFCCFFBB7E9D37F141074B5D" box="[657,805,1265,1291]" italics="true" pageId="7" pageNumber="66">Paraploderus</emphasis>
|
||
</taxonomicName>
|
||
ratherthanadistribution pattern. Onlyonespeciesisrecognizedtobeboundtohigheraltitudesand havesomewhatreducedwings.
|
||
</paragraph>
|
||
<paragraph id="EB0336D7FFCCFFBB7CE7365643EB4BA0" blockId="7.[174,1127,190,1527]" pageId="7" pageNumber="66">Naturalhistory – Availableinformationpreservedonspecimenlabels andcollectingeventlistsindicatesthatthesespeciesliveinwet, decaying plantmaterials, primarilyleaflitter. Preferredhabitatsareforested, oftenriv- erbanksorwaterfalls. Mostfrequentlyspecimenswerecollectedbysifting, but many flew to light.</paragraph>
|
||
<paragraph id="EB0336D7FFC3FFB47CC933BE46634B19" blockId="8.[136,1089,190,1359]" pageId="8" pageNumber="67">
|
||
The mature (L
|
||
<subScript id="77383492FFC3FFB47D7633D043A04E88" attach="left" box="[378,386,208,222]" fontSize="6" pageId="8" pageNumber="67">3</subScript>
|
||
) larva (of
|
||
<taxonomicName id="2CBC4D54FFC3FFB47E0433BE412D4E8E" authorityName="Makranczy" authorityYear="2016" box="[520,783,190,216]" class="Insecta" family="Staphylinidae" genus="Paraploderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="rufescens">
|
||
<emphasis id="D9C8EAC5FFC3FFB47E0433BE412D4E8E" box="[520,783,190,216]" italics="true" pageId="8" pageNumber="67">Paraploderus rufescens</emphasis>
|
||
</taxonomicName>
|
||
) – Length (in mm): 4.20 [ew = 0.50] (Fig. 79) and 3.45 [ew = 0.46] (specimen on Figs 105–116); the third larvalspecimenisinterpretedasLl
|
||
<subScript id="77383492FFC3FFB47E4F321440694F74" attach="both" box="[579,587,276,290]" fontSize="6" pageId="8" pageNumber="67">2</subScript>
|
||
arva, length 2.83 [ew = 0.48] (Fig. 78).
|
||
<emphasis id="D9C8EAC5FFC3FFB47C84322342E14F6B" box="[136,195,291,317]" italics="true" pageId="8" pageNumber="67">Head</emphasis>
|
||
. Headcapsule (Fig. 105) dorso-ventrallyflattened, roundedbutslightly transverse, asinglelargestemmaoneachside, supraantennalprominences transverselyelongate. Stemma (
|
||
<figureCitation id="73872A52FFC3FFB47E013266404F4FD7" box="[525,621,358,385]" captionStart="Figs 111–116" captionStartId="55.[174,219,1504,1526]" captionTargetBox="[176,1116,190,1470]" captionTargetId="figure-13@55.[174,1118,188,1484]" captionTargetPageId="55" captionText="Figs 111–116. Larva (L) of Paraploderus rufescens sp. n. 111 = abdominal segments IX–X, 3 112 = maxilla, 113 = mandible, 114 = antenna, 115 = stemma, lateral view, 116 = lateral view of 1st tergite, pleurite and spiracle. Scales: 0.1 mm for Fig. 114, 0.12 mm for Figs 115–116," figureDoi="http://doi.org/10.5281/zenodo.12625303" httpUri="https://zenodo.org/record/12625303/files/figure.png" pageId="8" pageNumber="67">Fig. 115</figureCitation>
|
||
) asclearsphericalcalotte, dark (black- ish) pigmentvariouslymovedawayfromthelensinpreservedspecimens (ar- tifactofpreservation). Dorsumofheadwithfrontalsetae (frontaldorsal = fd, frontallateral = fl, frontalmarginal = fm), epicranialsetae (epicranialdorsal = ed, epicraniallateral = el, epicranialmarginal = em) temporal (t) andlateral (l) setaeplustwopairsofposteriorsetae (p). Antenna (
|
||
<figureCitation id="73872A52FFC3FFB47F0C310E41424C7E" box="[768,864,526,552]" captionStart="Figs 111–116" captionStartId="55.[174,219,1504,1526]" captionTargetBox="[176,1116,190,1470]" captionTargetId="figure-13@55.[174,1118,188,1484]" captionTargetPageId="55" captionText="Figs 111–116. Larva (L) of Paraploderus rufescens sp. n. 111 = abdominal segments IX–X, 3 112 = maxilla, 113 = mandible, 114 = antenna, 115 = stemma, lateral view, 116 = lateral view of 1st tergite, pleurite and spiracle. Scales: 0.1 mm for Fig. 114, 0.12 mm for Figs 115–116," figureDoi="http://doi.org/10.5281/zenodo.12625303" httpUri="https://zenodo.org/record/12625303/files/figure.png" pageId="8" pageNumber="67">Fig. 114</figureCitation>
|
||
) threesegmented, threesensoryappendages (sa) onpenultimate, solenidia (so) onapical. La- brum (Fig. 108) trapeziform, halfaslongaswide, frontalmarginwithapair ofstout, spine-likesetae. Mandible (
|
||
<figureCitation id="73872A52FFC3FFB47E443173408B4CDB" box="[584,681,627,653]" captionStart="Figs 111–116" captionStartId="55.[174,219,1504,1526]" captionTargetBox="[176,1116,190,1470]" captionTargetId="figure-13@55.[174,1118,188,1484]" captionTargetPageId="55" captionText="Figs 111–116. Larva (L) of Paraploderus rufescens sp. n. 111 = abdominal segments IX–X, 3 112 = maxilla, 113 = mandible, 114 = antenna, 115 = stemma, lateral view, 116 = lateral view of 1st tergite, pleurite and spiracle. Scales: 0.1 mm for Fig. 114, 0.12 mm for Figs 115–116," figureDoi="http://doi.org/10.5281/zenodo.12625303" httpUri="https://zenodo.org/record/12625303/files/figure.png" pageId="8" pageNumber="67">Fig. 113</figureCitation>
|
||
) bifid, narrowtriangularshaped butthickatbase, graduallynarrowingtowardsbifurcateapex; amorebasal toothondorsalsideofmandibleobservableinonespecimenonly: eitheran aberrationorpossiblyaremnantofthemolarlobeofmoreprimitivestaphyli- noids. Maxilla (
|
||
<figureCitation id="73872A52FFC3FFB47D4531F943894D45" box="[329,427,761,788]" captionStart="Figs 111–116" captionStartId="55.[174,219,1504,1526]" captionTargetBox="[176,1116,190,1470]" captionTargetId="figure-13@55.[174,1118,188,1484]" captionTargetPageId="55" captionText="Figs 111–116. Larva (L) of Paraploderus rufescens sp. n. 111 = abdominal segments IX–X, 3 112 = maxilla, 113 = mandible, 114 = antenna, 115 = stemma, lateral view, 116 = lateral view of 1st tergite, pleurite and spiracle. Scales: 0.1 mm for Fig. 114, 0.12 mm for Figs 115–116," figureDoi="http://doi.org/10.5281/zenodo.12625303" httpUri="https://zenodo.org/record/12625303/files/figure.png" pageId="8" pageNumber="67">Fig. 112</figureCitation>
|
||
) withtriangularcardo, moderatelywidestipes, mala withctenidiumonadoralmarginsomewhatinterruptedinmiddle. Labium (Fig. 109) withmentumsaddle-shaped, prementumquiteelongate, ligula shortandsmall, labialpalpstwosegmented.
|
||
<emphasis id="D9C8EAC5FFC3FFB47EB3305E41324D2E" box="[703,784,862,888]" italics="true" pageId="8" pageNumber="67">Thorax</emphasis>
|
||
. Tergiteswithshortand longsetaearrangedmoreorlessalongtransversallines; prothoraxasinFig. 106, mesothoraxasinFig. 107, metanotumwithsetationidenticaltothatof mesonotum. Pronotaldiscalsetaeinrows (a- e) (terminologyexplainedin StANieC (2003)). Legswithfivearticles. Anteriorleg (Fig. 110) withproximal pairoftibialsetaeuneveninsize (onestout, otherthinandlonger), meso-and metatibiaewithsetaeeveninsizewithinpair. Tarsunguluswithtwosmall setaeinbasalposition, oneoneachside.
|
||
<emphasis id="D9C8EAC5FFC3FFB47E70374940CA4A35" box="[636,744,1097,1123]" italics="true" pageId="8" pageNumber="67">Abdomen</emphasis>
|
||
. SegmentsI–VIIIcomposed oftergitesandsternites. Firsttergumwithasmall, island-likesclerite (pleurite) separatedlaterally (
|
||
<figureCitation id="73872A52FFC3FFB47DCF378C40084AF1" box="[451,554,1164,1191]" captionStart="Figs 111–116" captionStartId="55.[174,219,1504,1526]" captionTargetBox="[176,1116,190,1470]" captionTargetId="figure-13@55.[174,1118,188,1484]" captionTargetPageId="55" captionText="Figs 111–116. Larva (L) of Paraploderus rufescens sp. n. 111 = abdominal segments IX–X, 3 112 = maxilla, 113 = mandible, 114 = antenna, 115 = stemma, lateral view, 116 = lateral view of 1st tergite, pleurite and spiracle. Scales: 0.1 mm for Fig. 114, 0.12 mm for Figs 115–116," figureDoi="http://doi.org/10.5281/zenodo.12625303" httpUri="https://zenodo.org/record/12625303/files/figure.png" pageId="8" pageNumber="67">Fig. 116</figureCitation>
|
||
), weaklysclerotizedbutmarkedbythree setae, spiraclessituatedbetweenpleuriteandtergite, separatefromboth. Seg- mentsII–VIIIwithspiraclesintergites. AbdominalsegmentsIXandX (
|
||
<figureCitation id="73872A52FFC3FFB4781D37CF42914B5D" captionStart="Figs 111–116" captionStartId="55.[174,219,1504,1526]" captionTargetBox="[176,1116,190,1470]" captionTargetId="figure-13@55.[174,1118,188,1484]" captionTargetPageId="55" captionText="Figs 111–116. Larva (L) of Paraploderus rufescens sp. n. 111 = abdominal segments IX–X, 3 112 = maxilla, 113 = mandible, 114 = antenna, 115 = stemma, lateral view, 116 = lateral view of 1st tergite, pleurite and spiracle. Scales: 0.1 mm for Fig. 114, 0.12 mm for Figs 115–116," figureDoi="http://doi.org/10.5281/zenodo.12625303" httpUri="https://zenodo.org/record/12625303/files/figure.png" pageId="8" pageNumber="67">Fig. 111</figureCitation>
|
||
) formingtergosternite (dorsalandventralscleritesfused). Urogomphi onearticled, blackanddistallynarrowed, extendedbeyondsegmentX. Anal lobesnotevertedinexaminedspecimensbutwithoutconspicuousstructures.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |