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<mods:title id="3F227A1FCA045A9DF44B3717FD00AFAF">Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae)</mods:title>
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<treatment id="314487958F79B32DFF7DFF4EFB14FB91" ID-DOI="http://doi.org/10.5281/zenodo.6121385" ID-GBIF-Taxon="159607938" ID-Zenodo-Dep="6121385" LSID="urn:lsid:plazi:treatment:314487958F79B32DFF7DFF4EFB14FB91" httpUri="http://treatment.plazi.org/id/314487958F79B32DFF7DFF4EFB14FB91" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection id="F1F765088F79B32DFF7DFF4EFF28FF36" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="B95236838F79B32DFF7DFF4EFD75FF57" blockId="16.[151,689,151,211]" box="[151,689,151,178]" pageId="16" pageNumber="17">
<heading id="E21A81EF8F79B32DFF7DFF4EFD75FF57" bold="true" box="[151,689,151,178]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<taxonomicName id="7EED4D008F79B32DFF7DFF4EFD75FF57" ID-CoL="3245Q" authority="Godwin-Austen, 1876" authorityName="Godwin-Austen" authorityYear="1876" box="[151,689,151,178]" class="Gastropoda" family="Pupinidae" genus="Csomapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="grandis">
<emphasis id="8B99EA918F79B32DFF7DFF4EFD75FF57" bold="true" box="[151,689,151,178]" pageId="16" pageNumber="17">
<emphasis id="8B99EA918F79B32DFF7DFF4EFE4CFF54" bold="true" box="[151,392,151,177]" italics="true" pageId="16" pageNumber="17">Csomapupa grandis</emphasis>
(
<bibRefCitation id="DD7C4B728F79B32DFE7DFF41FD6DFF57" author="Godwin-Austen" box="[407,681,152,178]" pageId="16" pageNumber="17" refString="Godwin-Austen, H. H. (1876) On the Cyclostomacea of the Dafla Hills, Assam. The Journal of the Asiatic Society of Bengal, 45 (2), 171 - 184." type="journal article" year="1876">Godwin-Austen, 1876</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="B95236838F79B32DFF7DFF63FF28FF36" blockId="16.[151,689,151,211]" box="[151,236,186,211]" pageId="16" pageNumber="17">
<figureCitation id="21D62A068F79B32DFF7DFF63FF13FF36" box="[151,215,186,211]" captionStart="FIGURE 8. A" captionStartId="17.[151,250,1838,1860]" captionTargetBox="[200,1365,195,1798]" captionTargetId="figure@17.[200,1366,195,1799]" captionTargetPageId="17" captionText="FIGURE 8. A. Csomapupa luyorensis (Godwin-Austen, 1917), syntype, NHMUK 1903.7.1.3377; B. Cs. grandis (Godwin- Austen, 1876), syntype, NHMUK 1903.7.1.3438; C. Vargapupa oharai n. sp., holotype, HNHM 98837; D. Vargapupa biheli n. sp., holotype, MNHN 2012 27020; E. Nodopomatias oakesi (Godwin-Austen, 1917), syntype, NHMUK 1903.7.1.3086; F. Nodopomatias sibbumensis (Godwin-Austen, 1917), syntype, NHMUK 1903.7. 1.3146. Photos: A B, E, F: H. Taylor; C D: B. Páll-Gergely. Scale bars = 5 mm; the smaller one refers to the photos of the whole shell, the larger one refers to the photos showing the lateral and basal side of the shells." httpUri="https://zenodo.org/record/237139/files/figure.png" pageId="16" pageNumber="17">Fig. 8</figureCitation>
B
</paragraph>
</subSubSection>
<subSubSection id="F1F765088F79B32DFF7DFEDAFD63FE0A" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="B95236838F79B32DFF7DFEDAFD41FEFD" blockId="16.[151,792,258,495]" box="[151,645,258,280]" pageId="16" pageNumber="17">
<emphasis id="8B99EA918F79B32DFF7DFEDAFE8EFEFD" box="[151,330,258,280]" italics="true" pageId="16" pageNumber="17">
Pomatias
<taxonomicName id="7EED4D008F79B32DFF17FEDBFE8EFEFD" ID-CoL="3245Q" authority="Godwin-Austen, 1876" authorityName="Godwin-Austen" authorityYear="1876" box="[253,330,258,280]" class="Gastropoda" family="Pupinidae" genus="Csomapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="grandis">grandis</taxonomicName>
</emphasis>
<bibRefCitation id="DD7C4B728F79B32DFEBAFEDBFDF0FEFD" author="Godwin-Austen" box="[336,564,258,280]" pageId="16" pageNumber="17" refString="Godwin-Austen, H. H. (1876) On the Cyclostomacea of the Dafla Hills, Assam. The Journal of the Asiatic Society of Bengal, 45 (2), 171 - 184." type="journal article" year="1876">Godwin-Austen, 1876</bibRefCitation>
: p. 181.
</paragraph>
<paragraph id="B95236838F79B32DFF7DFEFBFDEBFED2" blockId="16.[151,792,258,495]" box="[151,559,289,311]" pageId="16" pageNumber="17">
<emphasis id="8B99EA918F79B32DFF7DFEFBFE8EFED2" box="[151,330,289,311]" italics="true" pageId="16" pageNumber="17">
Pomatias
<taxonomicName id="7EED4D008F79B32DFF17FEF8FE8EFED2" ID-CoL="3245Q" box="[253,330,289,311]" class="Gastropoda" family="Pupinidae" genus="Csomapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="grandis">grandis</taxonomicName>
</emphasis>
<bibRefCitation id="DD7C4B728F79B32DFE8EFEF8FE1BFED2" author="Nevill" box="[356,479,289,311]" pageId="16" pageNumber="17" refString="Nevill, G. (1878) Hand list of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda. Pulmonata and Prosobranchia- Neurobranchia. Order of the Trustees, Calcutta, 338 pp. http: // dx. doi. org / 10.5962 / bhl. title. 6561" type="book" year="1878">Nevill 1878</bibRefCitation>
: p. 255.
</paragraph>
<paragraph id="B95236838F79B32DFF7DFE99FDB8FE57" blockId="16.[151,792,258,495]" pageId="16" pageNumber="17">
<taxonomicName id="7EED4D008F79B32DFF7DFE99FE56FEB3" ID-CoL="7SVGW" box="[151,402,320,342]" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="grandis">
<emphasis id="8B99EA918F79B32DFF7DFE99FE56FEB3" box="[151,402,320,342]" italics="true" pageId="16" pageNumber="17">Pseudopomatias grandis</emphasis>
</taxonomicName>
<bibRefCitation id="DD7C4B728F79B32DFE41FE99FDA6FEB3" author="Mollendorff" box="[427,610,320,342]" pageId="16" pageNumber="17" refString="Mollendorff, O. F. von (1885) Diagnoses specierum novarum sinensium. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 11 - 12, 161 - 170." type="book chapter" year="1885">Möllendorff 1885</bibRefCitation>
: p. 164.
<taxonomicName id="7EED4D008F79B32DFF7DFE87FE56FE91" ID-CoL="7SVGW" box="[151,402,350,372]" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="grandis">
<emphasis id="8B99EA918F79B32DFF7DFE87FE56FE91" box="[151,402,350,372]" italics="true" pageId="16" pageNumber="17">Pseudopomatias grandis</emphasis>
</taxonomicName>
<bibRefCitation id="DD7C4B728F79B32DFE41FE87FDA6FE91" author="Mollendorff" box="[427,610,350,372]" pageId="16" pageNumber="17" refString="Mollendorff, O. F. von (1886) Materialen zur Fauna for China. Jahrbucher der Deutschen Malakozoologischen Gesellschaft, 13, 156 - 210." type="journal article" year="1886">Möllendorff 1886</bibRefCitation>
: p. 179.
<taxonomicName id="7EED4D008F79B32DFF7DFEA4FE56FE76" ID-CoL="7SVGW" box="[151,402,381,403]" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="grandis">
<emphasis id="8B99EA918F79B32DFF7DFEA4FE56FE76" box="[151,402,381,403]" italics="true" pageId="16" pageNumber="17">Pseudopomatias grandis</emphasis>
</taxonomicName>
<bibRefCitation id="DD7C4B728F79B32DFE41FEA4FD0CFE76" author="Kobelt" box="[427,712,381,403]" pageId="16" pageNumber="17" refString="Kobelt, W. &amp; Mollendorff, O. von (1897) Catalog der gegenvartig lebend bekannten Pneumonopomen. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 29, 73 - 88, 105 - 120, 137 - 152." type="journal article" year="1897">Kobelt &amp; Möllendorff 1897</bibRefCitation>
: p. 140.
<taxonomicName id="7EED4D008F79B32DFF7DFE45FE56FE57" ID-CoL="7SVGW" box="[151,402,412,434]" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="grandis">
<emphasis id="8B99EA918F79B32DFF7DFE45FE56FE57" box="[151,402,412,434]" italics="true" pageId="16" pageNumber="17">Pseudopomatias grandis</emphasis>
</taxonomicName>
<bibRefCitation id="DD7C4B728F79B32DFE41FE45FDE8FE57" author="Kobelt" box="[427,556,412,434]" pageId="16" pageNumber="17" refString="Kobelt, W. (1902) Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In Verbindung mit der Deutschen Zoologischen Gesellschaft herausgegeben von der Koniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R. Friedlander und Sohn, Berlin, 662 pp." type="book" year="1902">Kobelt 1902</bibRefCitation>
: p. 273.
</paragraph>
<paragraph id="B95236838F79B32DFF7DFE63FD63FE0A" blockId="16.[151,792,258,495]" pageId="16" pageNumber="17">
<taxonomicName id="7EED4D008F79B32DFF7DFE63FE56FE35" ID-CoL="7SVGW" box="[151,402,442,464]" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="grandis">
<emphasis id="8B99EA918F79B32DFF7DFE63FE56FE35" box="[151,402,442,464]" italics="true" pageId="16" pageNumber="17">Pseudopomatias grandis</emphasis>
</taxonomicName>
<bibRefCitation id="DD7C4B728F79B32DFE41FE63FD4DFE35" author="Godwin-Austen" box="[427,649,442,464]" pageId="16" pageNumber="17" refString="Godwin-Austen, H. H. (1917) Zoological results of the Abor Expedition, 1911 - 12. XLVII Mollusca, VII. Cyclophoridae (in part) Records of the Indian Museum, 8, 493 - 580." type="journal article" year="1917">Godwin-Austen 1917</bibRefCitation>
: p. 578.
<taxonomicName id="7EED4D008F79B32DFF7DFE00FE56FE0A" ID-CoL="7SVGW" box="[151,402,473,495]" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="grandis">
<emphasis id="8B99EA918F79B32DFF7DFE00FE56FE0A" box="[151,402,473,495]" italics="true" pageId="16" pageNumber="17">Pseudopomatias grandis</emphasis>
</taxonomicName>
<bibRefCitation id="DD7C4B728F79B32DFE41FE00FDDAFE0A" author="Gude" box="[427,542,473,495]" pageId="16" pageNumber="17" refString="Gude, G. K. (1921) The Fauna of British India including Ceylon and Burma. Mollusca. - III. Land operculates (Cyclophoridae, Truncatellidae, Assimineidae, Helicinidae). Taylor and Francis, London, 386 pp. http: // dx. doi. org / 10.5962 / bhl. title. 12890" type="book" year="1921">Gude 1921</bibRefCitation>
: pp 158159.
</paragraph>
</subSubSection>
<subSubSection id="F1F765088F79B32DFF7DFDC5FDB4FDBD" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="B95236838F79B32DFF7DFDC5FDB4FDBD" blockId="16.[151,1437,540,1141]" pageId="16" pageNumber="17">
<emphasis id="8B99EA918F79B32DFF7DFDC5FED5FDD0" bold="true" box="[151,273,540,565]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
A large species with &quot;double&quot; suture, weak sculpture, elongated apical portion, and simple-looking but conspicuously widened peristome margin.
</paragraph>
</subSubSection>
<subSubSection id="F1F765088F79B32DFF2DFDBDFD1FFBC8" pageId="16" pageNumber="17" type="description">
<paragraph id="B95236838F79B32DFF2DFDBDFD00FC78" blockId="16.[151,1437,540,1141]" pageId="16" pageNumber="17">
<emphasis id="8B99EA918F79B32DFF2DFDBDFE9DFD98" bold="true" box="[199,345,612,637]" pageId="16" pageNumber="17">Description.</emphasis>
Shell yellowish, turriform, with the upper portion of the shell (first five whorls) being slightly concave; the 7.758.25 (n=3), rather flat whorls are separated by shallow suture; above the suture there is a groove running parallel with it; this is present on the whole shell except of above the last one to two whorls; protoconch of two whorls; the sculpture of the first whorl is finely granulated, glossy, after that it becomes regularly ribbed; teleoconch regularly, extremely finely ribbed with no spiral lines visible; the ribs are very low and are regular even behind the aperture; aperture rather rounded; the parietal-palatal transition is slightly angled; columellar-parietal transition not angled; peristome white, very much widened and reflexed, consists of two circles; the inner one is slimmer, the outer one is much wider, expanded and reflexed; the boundary between the two circles is hardly visible, the whole periostome seems to be simple.
</paragraph>
<paragraph id="B95236838F79B32DFF2DFC7EFCE1FC25" blockId="16.[151,1437,540,1141]" box="[199,805,935,960]" pageId="16" pageNumber="17">
<emphasis id="8B99EA918F79B32DFF2DFC7EFE27FC25" bold="true" box="[199,483,935,960]" pageId="16" pageNumber="17">Measurements (in mm).</emphasis>
H: 1213.3, D: 45.6 (n=3).
</paragraph>
<paragraph id="B95236838F79B32DFF2DFC15FD1FFBC8" blockId="16.[151,1437,540,1141]" pageId="16" pageNumber="17">
<emphasis id="8B99EA918F79B32DFF2DFC15FE0BFC00" bold="true" box="[199,463,972,997]" pageId="16" pageNumber="17">Differential diagnosis.</emphasis>
<taxonomicName id="7EED4D008F79B32DFE3CFC14FDB1FC00" box="[470,629,972,997]" class="Gastropoda" family="Pupinidae" genus="Csomapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="luyorensis">
<emphasis id="8B99EA918F79B32DFE3CFC14FDB1FC00" box="[470,629,972,997]" italics="true" pageId="16" pageNumber="17">Cs. luyorensis</emphasis>
</taxonomicName>
possesses a smaller shell, deeper suture and more conspicuous parallel groove above the suture. Moreover, the peristome of
<taxonomicName id="7EED4D008F79B32DFD00FC29FC4CFBED" box="[746,904,1007,1032]" class="Gastropoda" family="Pupinidae" genus="Csomapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="luyorensis">
<emphasis id="8B99EA918F79B32DFD00FC29FC4CFBED" box="[746,904,1007,1032]" italics="true" pageId="16" pageNumber="17">Cs. luyorensis</emphasis>
</taxonomicName>
is clearly separated into two circles, but that of
<taxonomicName id="7EED4D008F79B32DFF7DFBCCFEDCFBC8" box="[151,280,1044,1069]" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="grandis">
<emphasis id="8B99EA918F79B32DFF7DFBCCFEDCFBC8" box="[151,280,1044,1069]" italics="true" pageId="16" pageNumber="17">Cs. grandis</emphasis>
</taxonomicName>
seems to be rather simple, but widened.
</paragraph>
</subSubSection>
<subSubSection id="F1F765088F79B32DFF2DFBEEFA4DFBB5" box="[199,1417,1079,1104]" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="B95236838F79B32DFF2DFBEEFA4DFBB5" blockId="16.[151,1437,540,1141]" box="[199,1417,1079,1104]" pageId="16" pageNumber="17">
<emphasis id="8B99EA918F79B32DFF2DFBEEFEB1FBB5" bold="true" box="[199,373,1079,1104]" pageId="16" pageNumber="17">
<typeStatus id="665688218F79B32DFF2DFBEEFEC6FBB5" box="[199,258,1079,1104]" pageId="16" pageNumber="17">Type</typeStatus>
material.
</emphasis>
Dafla Hills, Assam,
<collectingCountry id="C1FA76138F79B32DFDB5FBEEFD5FFBB5" box="[607,667,1079,1104]" name="India" pageId="16" pageNumber="17">India</collectingCountry>
, Godwin-Austin Collection,
<collectionCode id="DFFCAE468F79B32DFC35FBE1FB8DFBAA" LSID="urn:lsid:biocol.org:col:34665" box="[991,1097,1080,1103]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="16" pageNumber="17">NHMUK</collectionCode>
1903.7.1.3438 (9
<typeStatus id="665688218F79B32DFAFCFBE1FAB9FBB5" box="[1302,1405,1080,1104]" pageId="16" pageNumber="17" type="syntype">syntypes</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="F1F765088F79B32DFF2DFB85FB14FB91" box="[199,1232,1116,1141]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="B95236838F79B32DFF2DFB85FB14FB91" blockId="16.[151,1437,540,1141]" box="[199,1232,1116,1141]" pageId="16" pageNumber="17">
<emphasis id="8B99EA918F79B32DFF2DFB85FEA4FB90" bold="true" box="[199,352,1116,1141]" pageId="16" pageNumber="17">Distribution.</emphasis>
<taxonomicName id="7EED4D008F79B32DFE8DFB84FE2CFB90" box="[359,488,1116,1141]" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="grandis">
<emphasis id="8B99EA918F79B32DFE8DFB84FE2CFB90" box="[359,488,1116,1141]" italics="true" pageId="16" pageNumber="17">Cs. grandis</emphasis>
</taxonomicName>
is known from the
<typeStatus id="665688218F79B32DFD29FB84FD37FB90" box="[707,755,1117,1141]" pageId="16" pageNumber="17">type</typeStatus>
locality only (see also
<figureCitation id="21D62A068F79B32DFC1CFB85FBF3FB91" box="[1014,1079,1116,1141]" captionStart="FIGURE 2" captionStartId="7.[151,250,1286,1308]" captionTargetBox="[151,1435,610,1264]" captionTargetId="figure@7.[151,1436,556,1265]" captionTargetPageId="7" captionText="FIGURE 2. Different areas of north-eastern India mentioned in the text. See also Table 3. 1: Sikkim, 2: Aka Hills, 3: Garo Hills, 4: Khasi Hills, 5: Dafla Hills, 6: Abor Hills, 7: Naga Hills." httpUri="https://zenodo.org/record/237133/files/figure.png" pageId="16" pageNumber="17">Fig. 2</figureCitation>
and
<tableCitation id="F46F03388F79B32DFB84FB85FB06FB91" box="[1134,1218,1116,1141]" captionStart="TABLE 3" captionStartId="6.[151,239,152,175]" captionTargetBox="[159,1404,202,1732]" captionTargetPageId="6" captionText="TABLE 3. Indian and Burmese localities mentioned in the paper. See also Figure 2." httpUri="http://table.plazi.org/id/ED92660B8F6FB33BFF7DFF41FBD0FF4A" pageId="16" pageNumber="17" tableUuid="ED92660B8F6FB33BFF7DFF41FBD0FF4A">Table 3</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>