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<document id="CC95D2F7B6107EBDAFADB2F0BF4BC754" ID-DOI="10.11646/zootaxa.3937.1.1" ID-GBIF-Dataset="f0657878-554d-4991-9b89-30195335323e" ID-ISSN="1175-5326" ID-Zenodo-Dep="237131" ID-ZooBank="30026A41-4F7F-487F-8BE2-E7065FC3BD5F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461017696232" checkinUser="plazi" docAuthor="Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András &amp; Asami, Takahiro" docDate="2015" docId="314487958F40B317FF7DF8F5FEC5FC09" docLanguage="en" docName="zt03937p049.pdf" docOrigin="Zootaxa 3937 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Vargapupa oharai Pall-Gergely, n. sp." docType="treatment" docVersion="6" lastPageNumber="43" masterDocId="CD7DFFED8F69B33DFFEAFFD9FFC4FFE5" masterDocTitle="Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae)" masterLastPageNumber="49" masterPageNumber="1" pageNumber="42" updateTime="1698569286847" updateUser="plazi">
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<mods:title id="109881F07E26496BD1B304CD8F12F481">Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae)</mods:title>
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<mods:namePart id="C6CB8B90F5338C64B7732C289DC7F85D">Páll-Gergely, Barna</mods:namePart>
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<mods:namePart id="A17948481BB07A44F55978D34E85230E">Fehér, Zoltán</mods:namePart>
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<mods:namePart id="6A6AB8FA183C27F65E92C457837C1A64">Hunyadi, András</mods:namePart>
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<mods:namePart id="CA7F1757FF3D93F652ABEBE4E756FE1F">Asami, Takahiro</mods:namePart>
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<mods:title id="53994A8407CED0C63C1AAD0F37CF8785">Zootaxa</mods:title>
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<mods:date id="D7F870E00C403491CF5495C6BBC31CBB">2015</mods:date>
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<mods:number id="C78654B7C5C3505C1BC50B99E55459F2">3937</mods:number>
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<mods:identifier id="F5ED289E5CEC16BF691127A91562BBA9" type="DOI">10.11646/zootaxa.3937.1.1</mods:identifier>
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<mods:identifier id="94CD998487918BBD590F0684A75EE841" type="Zenodo-Dep">237131</mods:identifier>
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<treatment id="314487958F40B317FF7DF8F5FEC5FC09" ID-DOI="http://doi.org/10.5281/zenodo.6121435" ID-GBIF-Taxon="114925942" ID-Zenodo-Dep="6121435" LSID="urn:lsid:plazi:treatment:314487958F40B317FF7DF8F5FEC5FC09" httpUri="http://treatment.plazi.org/id/314487958F40B317FF7DF8F5FEC5FC09" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
<subSubSection id="F1F765088F40B314FF7DF8F5FF37F882" pageId="41" pageNumber="42" type="nomenclature">
<paragraph id="B95236838F40B314FF7DF8F5FDA3F8A3" blockId="41.[151,615,1836,1895]" box="[151,615,1836,1862]" pageId="41" pageNumber="42">
<heading id="E21A81EF8F40B314FF7DF8F5FDA3F8A3" bold="true" box="[151,615,1836,1862]" fontSize="11" level="1" pageId="41" pageNumber="42" reason="1">
<emphasis id="8B99EA918F40B314FF7DF8F5FDA3F8A3" bold="true" box="[151,615,1836,1862]" pageId="41" pageNumber="42">
<taxonomicName id="7EED4D008F40B314FF7DF8F5FDD3F8A3" authority="Pall-Gergely" authorityName="Pall-Gergely" box="[151,535,1836,1862]" class="Gastropoda" family="Cyclophoridae" genus="Vargapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="oharai" status="sp. nov.">
<emphasis id="8B99EA918F40B314FF7DF8F5FEB6F8A3" bold="true" box="[151,370,1836,1862]" italics="true" pageId="41" pageNumber="42">Vargapupa oharai</emphasis>
Páll-Gergely
</taxonomicName>
,
<taxonomicNameLabel id="90AA57EA8F40B314FDCEF8F5FDA3F8A3" box="[548,615,1836,1862]" pageId="41" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B95236838F40B314FF7DF897FF37F882" blockId="41.[151,615,1836,1895]" box="[151,243,1870,1895]" pageId="41" pageNumber="42">
<figureCitation id="21D62A068F40B314FF7DF897FF1CF882" box="[151,216,1870,1895]" captionStart="FIGURE 8. A" captionStartId="17.[151,250,1838,1860]" captionTargetBox="[200,1365,195,1798]" captionTargetId="figure@17.[200,1366,195,1799]" captionTargetPageId="17" captionText="FIGURE 8. A. Csomapupa luyorensis (Godwin-Austen, 1917), syntype, NHMUK 1903.7.1.3377; B. Cs. grandis (Godwin- Austen, 1876), syntype, NHMUK 1903.7.1.3438; C. Vargapupa oharai n. sp., holotype, HNHM 98837; D. Vargapupa biheli n. sp., holotype, MNHN 2012 27020; E. Nodopomatias oakesi (Godwin-Austen, 1917), syntype, NHMUK 1903.7.1.3086; F. Nodopomatias sibbumensis (Godwin-Austen, 1917), syntype, NHMUK 1903.7. 1.3146. Photos: A B, E, F: H. Taylor; C D: B. Páll-Gergely. Scale bars = 5 mm; the smaller one refers to the photos of the whole shell, the larger one refers to the photos showing the lateral and basal side of the shells." httpUri="https://zenodo.org/record/237139/files/figure.png" pageId="41" pageNumber="42">Fig. 8</figureCitation>
C.
</paragraph>
</subSubSection>
<subSubSection id="F1F765088F40B314FF7DF84FFA92F84A" box="[151,1366,1942,1967]" pageId="41" pageNumber="42" type="diagnosis">
<paragraph id="B95236838F40B314FF7DF84FFA92F84A" blockId="41.[151,1436,1942,2003]" box="[151,1366,1942,1967]" pageId="41" pageNumber="42">
<emphasis id="8B99EA918F40B314FF7DF84FFED5F84A" bold="true" box="[151,273,1942,1967]" pageId="41" pageNumber="42">Diagnosis.</emphasis>
A very large, slender turriform, brownish species with a well-developed and a weaker basal keel.
</paragraph>
</subSubSection>
<subSubSection id="F1F765088F40B317FF2DF863FC95FDFD" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="description">
<paragraph id="B95236838F40B317FF2DF863FA83FE35" blockId="41.[151,1436,1942,2003]" lastBlockId="42.[151,1437,151,1004]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
<emphasis id="8B99EA918F40B314FF2DF863FE9DF836" bold="true" box="[199,345,1978,2003]" pageId="41" pageNumber="42">Description.</emphasis>
Shell brownish corneous or yellowish, slender turriform, widest at its base; there are 10.25 (n=2) regularly increasing whorls; first whorl of the protoconch rather smooth, the second whorl is finely ribbed; suture deeper on the upper whorls, especially at the protoconch; the coarse riblets are regular on the whole shell; keel blunt, the ribs of the body whorl are visible here too, especially close to the end of the keel (close to the columellarparietal transition of the peristome); there are rather shallow sutures on both sides of the keel where it joins the shell surface of the last whorl; keel starts from the back side of the most basal point apertural rim, and ends on the ventrolateral side; keel gradually becomes lower towards its end; there is a second, lower keel between the main keel and the umbilicus; it also starts from the back side of the apertural rim; main keel visible inside the aperture as a shallow canal; aperture round with angled columellar-parietal and parietal-palatal transitions; the lower incision (basal part of the aperture) is the starting point of the keel; apertural rim thickened and very much reflexed.
</paragraph>
<paragraph id="B95236838F43B317FF2DFE05FC8BFE11" blockId="42.[151,1437,151,1004]" box="[199,847,476,501]" pageId="42" pageNumber="43">
<emphasis id="8B99EA918F43B317FF2DFE05FE27FE10" bold="true" box="[199,483,476,501]" pageId="42" pageNumber="43">Measurements (in mm).</emphasis>
H: 14.716.1, D: 5.35.6 (n=2).
</paragraph>
<paragraph id="B95236838F43B317FF2DFE26FC95FDFD" blockId="42.[151,1437,151,1004]" box="[199,849,511,536]" pageId="42" pageNumber="43">
<emphasis id="8B99EA918F43B317FF2DFE26FE0AFDFD" bold="true" box="[199,462,511,536]" pageId="42" pageNumber="43">Differential diagnosis.</emphasis>
See under
<taxonomicName id="7EED4D008F43B317FDA0FDD8FCCFFDFD" box="[586,779,511,536]" class="Gastropoda" family="Cyclophoridae" genus="Vargapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="42" pageNumber="43" phylum="Mollusca" rank="species" species="biheli" status="sp. nov.">
<emphasis id="8B99EA918F43B317FDA0FDD8FCCFFDFD" box="[586,779,511,536]" italics="true" pageId="42" pageNumber="43">Vargapupa biheli</emphasis>
</taxonomicName>
<emphasis id="8B99EA918F43B317FCF8FE26FC95FDFD" bold="true" box="[786,849,511,536]" pageId="42" pageNumber="43">
<taxonomicNameLabel id="90AA57EA8F43B317FCF8FE26FC95FDFD" box="[786,849,511,536]" pageId="42" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="F1F765088F43B317FF2DFDFDFDCEFCDD" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="B95236838F43B317FF2DFDFDFF3BFD15" blockId="42.[151,1437,151,1004]" pageId="42" pageNumber="43">
<emphasis id="8B99EA918F43B317FF2DFDFDFEF2FDD8" bold="true" box="[199,310,548,573]" pageId="42" pageNumber="43">Material.</emphasis>
<collectingCountry id="C1FA76138F43B317FEA9FDFDFE61FDD8" box="[323,421,548,573]" name="Vietnam" pageId="42" pageNumber="43">Vietnam</collectingCountry>
, Ninh Binh Province, Cuc Phuong National Park, Cave of Prehistoric
<collectingCountry id="C1FA76138F43B317FAEAFDFCFAF2FDD9" box="[1280,1334,549,572]" name="Isle of Man" pageId="42" pageNumber="43">Man</collectingCountry>
,
<quantity id="7E159B668F43B317FAADFDFDFA5CFDD9" box="[1351,1432,548,572]" metricMagnitude="2" metricUnit="m" metricValue="2.37" pageId="42" pageNumber="43" unit="m" value="237.0">237 m</quantity>
,
<geoCoordinate id="DCD950448F43B317FF7DFD9EFE87FD85" box="[151,323,583,608]" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="20.292812">20°17.56874'N</geoCoordinate>
,
<geoCoordinate id="DCD950448F43B317FEBBFD9EFDCCFD85" box="[337,520,583,608]" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="105.66687">105°40.01229'E</geoCoordinate>
, leg. Ohara, K., Okubo, K., Otani, J. U.,
<date id="CD5310438F43B317FC03FD9EFBA8FD85" box="[1001,1132,583,608]" pageId="42" pageNumber="43" value="2007-11-16">16.11.2007</date>
.,
<typeStatus id="665688218F43B317FB94FD9EFB22FD85" box="[1150,1254,583,608]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
<collectionCode id="DFFCAE468F43B317FB1AFD91FA81FDBA" LSID="urn:lsid:biocol.org:col:34991" box="[1264,1349,584,607]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="42" pageNumber="43">HNHM</collectionCode>
98837, OK/2, JUO/2; same data, leg. Okubo, K. &amp; Otani, J. U.,
<date id="CD5310438F43B317FCBFFDB5FC1DFD61" box="[853,985,620,644]" pageId="42" pageNumber="43" value="2007-12-04">12.04.2007</date>
., JUO/1, PGB/1;
<collectingCountry id="C1FA76138F43B317FB47FDB5FACBFD60" box="[1197,1295,620,645]" name="Vietnam" pageId="42" pageNumber="43">Vietnam</collectingCountry>
, Ninh Binh Province, Cuc Phuong National Park, excoll Jose Ahuir Galindo, PGB/1;
<collectingCountry id="C1FA76138F43B317FC38FD56FBF0FD4D" box="[978,1076,655,680]" name="Vietnam" pageId="42" pageNumber="43">Vietnam</collectingCountry>
, Thanh Hóa Prov., Pu Luong N. P., surroundings of village Am,
<geoCoordinate id="DCD950448F43B317FDC8FD6DFD76FD28" box="[546,690,692,717]" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="20.46975">20°28.185'N</geoCoordinate>
,
<geoCoordinate id="DCD950448F43B317FD2CFD6DFCA5FD28" box="[710,865,692,717]" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="105.22182">105°13.309'E</geoCoordinate>
, collected by native people, 0
<date id="CD5310438F43B317FB3FFD6DFA8EFD28" box="[1237,1354,692,717]" pageId="42" pageNumber="43" value="2010-05-10">5.10.2010</date>
., HE/1
<typeStatus id="665688218F43B317FF7DFD01FF3FFD15" box="[151,251,728,752]" pageId="42" pageNumber="43" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="B95236838F43B317FF2DFD25FDCEFCDD" blockId="42.[151,1437,151,1004]" pageId="42" pageNumber="43">
<emphasis id="8B99EA918F43B317FF2DFD25FEACFCF0" bold="true" box="[199,360,764,789]" pageId="42" pageNumber="43">
<typeStatus id="665688218F43B317FF2DFD25FEC6FCF0" box="[199,258,764,789]" pageId="42" pageNumber="43">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="C1FA76138F43B317FE98FD25FE10FCF0" box="[370,468,764,789]" name="Vietnam" pageId="42" pageNumber="43">Vietnam</collectingCountry>
, Ninh Binh Province, Cuc Phuong National Park, Cave of Prehistoric
<collectingCountry id="C1FA76138F43B317FAEDFD24FAF9FCF1" box="[1287,1341,765,788]" name="Isle of Man" pageId="42" pageNumber="43">Man</collectingCountry>
,
<quantity id="7E159B668F43B317FAA1FD25FA53FCF1" box="[1355,1431,764,789]" metricMagnitude="2" metricUnit="m" metricValue="1.46" pageId="42" pageNumber="43" unit="m" value="146.0">146 m</quantity>
,
<geoCoordinate id="DCD950448F43B317FF7DFCC6FE87FCDD" box="[151,323,799,824]" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="20.258974">20°15.53843'N</geoCoordinate>
,
<geoCoordinate id="DCD950448F43B317FEA4FCC6FDC2FCDD" box="[334,518,799,824]" direction="east" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="105.70649">105°42.38950'E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="F1F765088F43B317FF2DFC9DFBEEFCB8" box="[199,1066,836,861]" pageId="42" pageNumber="43" type="etymology">
<paragraph id="B95236838F43B317FF2DFC9DFBEEFCB8" blockId="42.[151,1437,151,1004]" box="[199,1066,836,861]" pageId="42" pageNumber="43">
<emphasis id="8B99EA918F43B317FF2DFC9DFE8AFCB8" bold="true" box="[199,334,836,861]" pageId="42" pageNumber="43">Etymology.</emphasis>
Named after our friend, Kenji Ohara, who provided the material.
</paragraph>
</subSubSection>
<subSubSection id="F1F765088F43B317FF2DFCBEFD41FC40" pageId="42" pageNumber="43" type="distribution">
<paragraph id="B95236838F43B317FF2DFCBEFD41FC40" blockId="42.[151,1437,151,1004]" pageId="42" pageNumber="43">
<emphasis id="8B99EA918F43B317FF2DFCBEFE9DFC65" bold="true" box="[199,345,871,896]" pageId="42" pageNumber="43">Distribution</emphasis>
(
<figureCitation id="21D62A068F43B317FE9AFCBEFE0FFC65" box="[368,459,871,896]" captionStart="FIGURE 15" captionStartId="38.[151,250,1020,1042]" captionTargetBox="[151,1436,195,999]" captionTargetId="figure@38.[151,1437,195,999]" captionTargetPageId="38" captionText="FIGURE 15. Distribution of Pseudopomatias and Vargapupa n. gen. species. Empty triangle, top up: Vargapupa biheli n. sp.; filled triangle, top down: Vargapupa oharai n. sp.; empty square: Pseudopomatias maasseni n. sp., open circle: P. sophiae n. sp., closed circle: P. linanprietoae n. sp." httpUri="https://zenodo.org/record/237146/files/figure.png" pageId="42" pageNumber="43">Fig. 15</figureCitation>
): The new species is known from two localities in Northern
<collectingCountry id="C1FA76138F43B317FB2FFCBEFAECFC65" box="[1221,1320,871,896]" name="Vietnam" pageId="42" pageNumber="43">Vietnam</collectingCountry>
. There is approximately
<quantity id="7E159B668F43B317FED5FC55FE43FC40" box="[319,391,908,933]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="42" pageNumber="43" unit="km" value="60.0">60 km</quantity>
between the two sites.
</paragraph>
</subSubSection>
<subSubSection id="F1F765088F43B317FF2DFC76FEC5FC09" pageId="42" pageNumber="43" type="discussion">
<paragraph id="B95236838F43B317FF2DFC76FEC5FC09" blockId="42.[151,1437,151,1004]" pageId="42" pageNumber="43">
<emphasis id="8B99EA918F43B317FF2DFC76FEFFFC2D" bold="true" box="[199,315,943,968]" pageId="42" pageNumber="43">Remarks.</emphasis>
In the Cuc Phuong National Park
<taxonomicName id="7EED4D008F43B317FD53FC68FC40FC2D" box="[697,900,943,968]" class="Gastropoda" family="Cyclophoridae" genus="Vargapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="42" pageNumber="43" phylum="Mollusca" rank="species" species="oharai" status="sp. nov.">
<emphasis id="8B99EA918F43B317FD53FC68FC40FC2D" box="[697,900,943,968]" italics="true" pageId="42" pageNumber="43">Vargapupa oharai</emphasis>
</taxonomicName>
<emphasis id="8B99EA918F43B317FC61FC76FC0DFC2D" bold="true" box="[907,969,943,968]" pageId="42" pageNumber="43">
<taxonomicNameLabel id="90AA57EA8F43B317FC61FC76FC0DFC2D" box="[907,969,943,968]" pageId="42" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
lives sympatrically with
<taxonomicName id="7EED4D008F43B317FB09FC76FF3FFC09" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="42" pageNumber="43" phylum="Mollusca" rank="species" species="amoenus">
<emphasis id="8B99EA918F43B317FB09FC76FF3FFC09" italics="true" pageId="42" pageNumber="43">Pseudopomatias amoenus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>