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<document id="0DBEAB755581DCBBB43017B66B858431" ID-DOI="10.11646/zootaxa.3937.1.1" ID-GBIF-Dataset="f0657878-554d-4991-9b89-30195335323e" ID-ISSN="1175-5326" ID-Zenodo-Dep="237131" ID-ZooBank="30026A41-4F7F-487F-8BE2-E7065FC3BD5F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461017696232" checkinUser="plazi" docAuthor="Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András &amp; Asami, Takahiro" docDate="2015" docId="314487958F40B314FF7DFC9BFBEAF93D" docLanguage="en" docName="zt03937p049.pdf" docOrigin="Zootaxa 3937 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Vargapupa biheli Pall-Gergely, n. sp." docType="treatment" docVersion="6" lastPageNumber="42" masterDocId="CD7DFFED8F69B33DFFEAFFD9FFC4FFE5" masterDocTitle="Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae)" masterLastPageNumber="49" masterPageNumber="1" pageNumber="42" updateTime="1698569286847" updateUser="plazi">
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<mods:title id="0433252CADAC1B328F679B030EC83082">Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae)</mods:title>
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<mods:namePart id="473EA9D21A967EE8B8DFA36F11696802">Páll-Gergely, Barna</mods:namePart>
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<mods:namePart id="E3D7042F308C1C9CD5681DF2BAAA914D">Fehér, Zoltán</mods:namePart>
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<mods:namePart id="3F11299BDDE07516A996626200A0C69A">Hunyadi, András</mods:namePart>
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<mods:namePart id="B5350DCA09C0958C12CF2B42D50AE12C">Asami, Takahiro</mods:namePart>
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<mods:number id="3C6A606E397797906EDF51D317B51327">3937</mods:number>
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<treatment id="314487958F40B314FF7DFC9BFBEAF93D" ID-DOI="http://doi.org/10.5281/zenodo.6121433" ID-GBIF-Taxon="114925946" ID-Zenodo-Dep="6121433" LSID="urn:lsid:plazi:treatment:314487958F40B314FF7DFC9BFBEAF93D" httpUri="http://treatment.plazi.org/id/314487958F40B314FF7DFC9BFBEAF93D" lastPageNumber="42" pageId="41" pageNumber="42">
<subSubSection id="F1F765088F40B314FF7DFC9BFDACFC26" pageId="41" pageNumber="42" type="nomenclature">
<paragraph id="B95236838F40B314FF7DFC9BFD9FFCB9" blockId="41.[151,603,834,894]" box="[151,603,834,860]" pageId="41" pageNumber="42">
<heading id="E21A81EF8F40B314FF7DFC9BFD9FFCB9" bold="true" box="[151,603,834,860]" fontSize="11" level="1" pageId="41" pageNumber="42" reason="1">
<emphasis id="8B99EA918F40B314FF7DFC9BFD9FFCB9" bold="true" box="[151,603,834,860]" pageId="41" pageNumber="42">
<taxonomicName id="7EED4D008F40B314FF7DFC9BFDC8FCB9" authority="Pall-Gergely" authorityName="Pall-Gergely" box="[151,524,834,860]" class="Gastropoda" family="Cyclophoridae" genus="Vargapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="biheli" status="sp. nov.">
<emphasis id="8B99EA918F40B314FF7DFC9BFEA3FCB9" bold="true" box="[151,359,834,860]" italics="true" pageId="41" pageNumber="42">Vargapupa biheli</emphasis>
Páll-Gergely
</taxonomicName>
,
<taxonomicNameLabel id="90AA57EA8F40B314FDF3FC9BFD9FFCB9" box="[537,603,834,860]" pageId="41" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B95236838F40B314FF7DFCBCFF30FC9B" blockId="41.[151,603,834,894]" box="[151,244,869,894]" pageId="41" pageNumber="42">
<figureCitation id="21D62A068F40B314FF7DFCBCFF1CFC9B" box="[151,216,869,894]" captionStart="FIGURE 8. A" captionStartId="17.[151,250,1838,1860]" captionTargetBox="[200,1365,195,1798]" captionTargetId="figure@17.[200,1366,195,1799]" captionTargetPageId="17" captionText="FIGURE 8. A. Csomapupa luyorensis (Godwin-Austen, 1917), syntype, NHMUK 1903.7.1.3377; B. Cs. grandis (Godwin- Austen, 1876), syntype, NHMUK 1903.7.1.3438; C. Vargapupa oharai n. sp., holotype, HNHM 98837; D. Vargapupa biheli n. sp., holotype, MNHN 2012 27020; E. Nodopomatias oakesi (Godwin-Austen, 1917), syntype, NHMUK 1903.7.1.3086; F. Nodopomatias sibbumensis (Godwin-Austen, 1917), syntype, NHMUK 1903.7. 1.3146. Photos: A B, E, F: H. Taylor; C D: B. Páll-Gergely. Scale bars = 5 mm; the smaller one refers to the photos of the whole shell, the larger one refers to the photos showing the lateral and basal side of the shells." httpUri="https://zenodo.org/record/237139/files/figure.png" pageId="41" pageNumber="42">Fig. 8</figureCitation>
D.
</paragraph>
<paragraph id="B95236838F40B314FF7DFC74FDACFC26" blockId="41.[151,616,941,963]" box="[151,616,941,963]" pageId="41" pageNumber="42">
<taxonomicName id="7EED4D008F40B314FF7DFC74FE45FC26" box="[151,385,941,963]" class="Gastropoda" family="Pupinidae" genus="Pseudopomatias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="fulvus">
<emphasis id="8B99EA918F40B314FF7DFC74FE45FC26" box="[151,385,941,963]" italics="true" pageId="41" pageNumber="42">Pseudopomatias fulvus</emphasis>
</taxonomicName>
<bibRefCitation id="DD7C4B728F40B314FE71FC74FDDDFC26" author="Saurin" box="[411,537,941,963]" pageId="41" pageNumber="42" refString="Saurin, E. (1953) Coquilles nouvelles de l'Indochine. Journal de Conchyliologie, 93 (4), 113 - 120." type="journal article" year="1953">Saurin 1953</bibRefCitation>
: p. 113.
</paragraph>
</subSubSection>
<subSubSection id="F1F765088F40B314FF7DFC36FBABFBED" box="[151,1135,1007,1032]" pageId="41" pageNumber="42" type="diagnosis">
<paragraph id="B95236838F40B314FF7DFC36FBABFBED" blockId="41.[151,1437,1007,1752]" box="[151,1135,1007,1032]" pageId="41" pageNumber="42">
<emphasis id="8B99EA918F40B314FF7DFC36FED5FBED" bold="true" box="[151,273,1007,1032]" pageId="41" pageNumber="42">Diagnosis.</emphasis>
A medium sized, slender turriform species with a well developed basal keel.
</paragraph>
</subSubSection>
<subSubSection id="F1F765088F40B314FF2DFBCDFA9BFA38" pageId="41" pageNumber="42" type="description">
<paragraph id="B95236838F40B314FF2DFBCDFE91FA95" blockId="41.[151,1437,1007,1752]" pageId="41" pageNumber="42">
<emphasis id="8B99EA918F40B314FF2DFBCDFE9DFBC8" bold="true" box="[199,345,1044,1069]" pageId="41" pageNumber="42">Description.</emphasis>
Shell turriform, widest at its base; only corroded shell material was available, therefore the original shell colour is unknown; there are 88.75, regularly increasing, rather flat or moderately bulging whorls; protoconch rather smooth with signs of small regular riblets; suture deeper on the upper whorls, especially at the protoconch; the coarse riblets are regular on the whole shell; sometimes on the last whorl they are less regular; keel blunt; the ribs of the body whorl are visible here too, especially close to the end of the keel; there are deep sutures on both sides of the keel; the keel starts from the back side of the most basal point of the peristome, and ends on the ventrolateral side; keel gradually becomes lower towards its end; keel visible inside the aperture as a shallow canal; aperture round with not angled columellar-parietal transition and slightly angled parietal-palatal transition; the lower incision (at the basal pont of the peristome) is the starting point of the keel; apertural rim thickened and slightly reflexed.
</paragraph>
<paragraph id="B95236838F40B314FF2DFAA5FC85FA71" blockId="41.[151,1437,1007,1752]" box="[199,833,1404,1429]" pageId="41" pageNumber="42">
<emphasis id="8B99EA918F40B314FF2DFAA5FE27FA70" bold="true" box="[199,483,1404,1429]" pageId="41" pageNumber="42">Measurements (in mm).</emphasis>
H: 8.910.5, D: 3.53.9 (n=3).
</paragraph>
<paragraph id="B95236838F40B314FF2DFA46FA9BFA38" blockId="41.[151,1437,1007,1752]" pageId="41" pageNumber="42">
<emphasis id="8B99EA918F40B314FF2DFA46FE0AFA5D" bold="true" box="[199,462,1439,1464]" pageId="41" pageNumber="42">Differential diagnosis.</emphasis>
<taxonomicName id="7EED4D008F40B314FE3EFA78FD51FA5D" box="[468,661,1439,1464]" class="Gastropoda" family="Cyclophoridae" genus="Vargapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="biheli" status="sp. nov.">
<emphasis id="8B99EA918F40B314FE3EFA78FD51FA5D" box="[468,661,1439,1464]" italics="true" pageId="41" pageNumber="42">Vargapupa biheli</emphasis>
</taxonomicName>
<emphasis id="8B99EA918F40B314FD76FA46FD1FFA5D" bold="true" box="[668,731,1439,1464]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="90AA57EA8F40B314FD76FA46FD1FFA5D" box="[668,731,1439,1464]" pageId="41" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is much smaller than
<taxonomicName id="7EED4D008F40B314FC39FA78FBFEFA5D" box="[979,1082,1439,1464]" class="Gastropoda" family="Cyclophoridae" genus="Vargapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="oharai" status="sp. nov.">
<emphasis id="8B99EA918F40B314FC39FA78FBFEFA5D" box="[979,1082,1439,1464]" italics="true" pageId="41" pageNumber="42">V. oharai</emphasis>
</taxonomicName>
<emphasis id="8B99EA918F40B314FBAAFA46FBBBFA5D" bold="true" box="[1088,1151,1439,1464]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="90AA57EA8F40B314FBAAFA46FBBBFA5D" box="[1088,1151,1439,1464]" pageId="41" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, has narrower umbilicus, more rounded aperture and only one basal keel. The sutures situated on both side of the main keel are deeper.
</paragraph>
</subSubSection>
<subSubSection id="F1F765088F40B314FF2DFA3EFC76F9AD" pageId="41" pageNumber="42" type="materials_examined">
<paragraph id="B95236838F40B314FF2DFA3EFDE6F9C1" blockId="41.[151,1437,1007,1752]" pageId="41" pageNumber="42">
<emphasis id="8B99EA918F40B314FF2DFA3EFEF2F9E5" bold="true" box="[199,310,1511,1536]" pageId="41" pageNumber="42">Material.</emphasis>
<collectingCountry id="C1FA76138F40B314FED4FA31FEB3FA1A" box="[318,375,1512,1535]" name="Laos" pageId="41" pageNumber="42">Laos</collectingCountry>
, Tran Ninh Province, Pa Hia, Pah Xieng Tong, coll. Saurin,
<collectionCode id="DFFCAE468F40B314FBC7FA31FB46FA1A" LSID="urn:lsid:biocol.org:col:34988" box="[1069,1154,1512,1535]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="41" pageNumber="42">MNHN</collectionCode>
201227020 (
<typeStatus id="665688218F40B314FAC2FA3EFA54F9E5" box="[1320,1424,1511,1536]" pageId="41" pageNumber="42" type="holotype">holotype</typeStatus>
),
<collectionCode id="DFFCAE468F40B314FF7DF9D4FF28F9C1" LSID="urn:lsid:biocol.org:col:34988" box="[151,236,1549,1572]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="41" pageNumber="42">MNHN</collectionCode>
201227021/9 (
<typeStatus id="665688218F40B314FE4EF9D5FDD2F9C1" box="[420,534,1548,1572]" pageId="41" pageNumber="42" type="paratype">paratypes</typeStatus>
).
</paragraph>
<paragraph id="B95236838F40B314FF2DF9F6FC76F9AD" blockId="41.[151,1437,1007,1752]" box="[199,946,1583,1608]" pageId="41" pageNumber="42">
<emphasis id="8B99EA918F40B314FF2DF9F6FEA1F9AD" bold="true" box="[199,357,1583,1608]" pageId="41" pageNumber="42">
<typeStatus id="665688218F40B314FF2DF9F6FEC6F9AD" box="[199,258,1583,1608]" pageId="41" pageNumber="42">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="C1FA76138F40B314FE86F9E9FE61F9A2" box="[364,421,1584,1607]" name="Laos" pageId="41" pageNumber="42">Laos</collectingCountry>
, Tran Ninh Province, Pa Hia, Pah Xieng Tong.
</paragraph>
</subSubSection>
<subSubSection id="F1F765088F40B314FF2DF98DFE3DF950" pageId="41" pageNumber="42" type="etymology">
<paragraph id="B95236838F40B314FF2DF98DFE3DF950" blockId="41.[151,1437,1007,1752]" pageId="41" pageNumber="42">
<emphasis id="8B99EA918F40B314FF2DF98DFE8AF988" bold="true" box="[199,334,1620,1645]" pageId="41" pageNumber="42">Etymology.</emphasis>
The new species is named after the Biodiversity Heritage Library (www.biodiversitylibrary.org) to thank the multitude of rare literature made available to us. The name “
<taxonomicName id="7EED4D008F40B314FC4CF9AEFC2EF975" box="[934,1002,1655,1680]" class="Gastropoda" family="Cyclophoridae" genus="Vargapupa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="biheli">biheli</taxonomicName>
” is an acronym derived from the name
<emphasis id="8B99EA918F40B314FF7DF945FF70F950" bold="true" box="[151,180,1692,1717]" pageId="41" pageNumber="42">
<collectingCountry id="C1FA76138F40B314FF7DF945FF70F950" box="[151,180,1692,1717]" name="Belgium" pageId="41" pageNumber="42">BI</collectingCountry>
</emphasis>
odiversity
<emphasis id="8B99EA918F40B314FEC1F945FE97F950" bold="true" box="[299,339,1692,1717]" pageId="41" pageNumber="42">HE</emphasis>
ritage
<emphasis id="8B99EA918F40B314FE71F945FE7CF950" bold="true" box="[411,440,1692,1717]" pageId="41" pageNumber="42">LI</emphasis>
brary.
</paragraph>
</subSubSection>
<subSubSection id="F1F765088F40B314FF2DF966FBEAF93D" box="[199,1070,1727,1752]" pageId="41" pageNumber="42" type="distribution">
<paragraph id="B95236838F40B314FF2DF966FBEAF93D" blockId="41.[151,1437,1007,1752]" box="[199,1070,1727,1752]" pageId="41" pageNumber="42">
<emphasis id="8B99EA918F40B314FF2DF966FE9DF93D" bold="true" box="[199,345,1727,1752]" pageId="41" pageNumber="42">Distribution</emphasis>
(
<figureCitation id="21D62A068F40B314FE82F966FE78F93D" box="[360,444,1727,1752]" captionStart="FIGURE 15" captionStartId="38.[151,250,1020,1042]" captionTargetBox="[151,1436,195,999]" captionTargetId="figure@38.[151,1437,195,999]" captionTargetPageId="38" captionText="FIGURE 15. Distribution of Pseudopomatias and Vargapupa n. gen. species. Empty triangle, top up: Vargapupa biheli n. sp.; filled triangle, top down: Vargapupa oharai n. sp.; empty square: Pseudopomatias maasseni n. sp., open circle: P. sophiae n. sp., closed circle: P. linanprietoae n. sp." httpUri="https://zenodo.org/record/237146/files/figure.png" pageId="41" pageNumber="42">Fig. 15</figureCitation>
): The new species is known from the
<typeStatus id="665688218F40B314FC8EF919FC50F93D" box="[868,916,1728,1752]" pageId="41" pageNumber="42">type</typeStatus>
locality only.
</paragraph>
</subSubSection>
</treatment>
</document>