285 lines
34 KiB
XML
285 lines
34 KiB
XML
<document ID-CLB-Dataset="24300" ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="FD38D22C887123E96F071BBED66BF240" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole glabra Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="C54726DF-9759-447D-AC25-87D471335D43" docUuidSource="ZooBank" docVersion="7" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1732649601836" updateUser="ExternalLinkService">
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<mods:mods id="7EBC88A44C0D0B4CD2DC605CF5F1D999" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BB801697AFE44C897BF6D226E57401F7">
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<mods:title id="6DF6790A6C8CB526E39A01B0BE3E2649">Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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<mods:name id="4F4ED5E7D3FAE0202195DEC2500713FF" type="personal">
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<mods:namePart id="56E7013B2A980DDB90E4C48087012472">Salata, Sebastian</mods:namePart>
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<mods:name id="168957C3F41EAF7ABF7BF93EA96E94B1" type="personal">
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<mods:namePart id="588087168426846D02775FCA2AB2D1CD">Fisher, Brian L.</mods:namePart>
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<mods:typeOfResource id="069C8F9B560499BC78BB36D66690C85B">text</mods:typeOfResource>
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<mods:title id="6032DAC490815174D09A2B0280648291">ZooKeys</mods:title>
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<mods:part id="26228A92E37DF6456197A6AA0FA5740A">
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<mods:date id="B12CA892EB980B7C28A2E8FF48D1FE52">2020</mods:date>
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<mods:detail id="2226855757F7FF88722E567F29280502" type="volume">
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<mods:number id="0602210EA91320B6629515F4003DF736">905</mods:number>
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<mods:start id="5884E509B368DCE565403BD624699B64">1</mods:start>
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<mods:end id="5AD66230A0428C68B61CDA8CA5CA716F">235</mods:end>
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<mods:location id="314512903DAEF0ADE5126917525C7217">
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<mods:url id="1663867F7FD59E7970037B42E713D22D">http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
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<mods:classification id="474E5DF8F735014F80B36899F92DB2EE">journal article</mods:classification>
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<mods:identifier id="1C547A1E7FD9727ADC3C854EEA2B1C61" type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
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<mods:identifier id="A257BBE795D5DB04757516DE9AC84616" type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
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<treatment id="FD38D22C887123E96F071BBED66BF240" ID-GBIF-Taxon="161777041" LSID="urn:lsid:zoobank.org:act:C54726DF-9759-447D-AC25-87D471335D43" httpUri="http://treatment.plazi.org/id/FD38D22C887123E96F071BBED66BF240" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="33B93E328F8C010F949106FE50AD87A2" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="C554EC2A7633490C02BE70FC489939CA" pageId="0" pageNumber="1">
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<taxonomicName id="DBF0AAAD40030ACA8ABFFEA952C379E1" ID-CoL="CML7D" LSID="http://zoobank.org/C54726DF-9759-447D-AC25-87D471335D43" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole glabra" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="glabra" status="sp. nov.">Pheidole glabra</taxonomicName>
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<taxonomicNameLabel id="2496683C0B8463A0A317878923864430" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="311267783A7065F353301AD02490A85E" captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Pheidole glabra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235034) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134271)." figureDoi="10.3897/zookeys.905.39592.figure68" httpUri="https://binary.pensoft.net/fig/373115" pageId="0" pageNumber="1">Figs 68A-F</figureCitation>
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<figureCitation id="BE2684D849658A001775C49D56A2DC7D" captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84T</figureCitation>
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<figureCitation id="611D38318700805E16822D726F80B3F6" captionStart="Figure 86" captionStartId="F86" captionText="Figure 86. Distribution. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure86" httpUri="https://binary.pensoft.net/fig/373133" pageId="0" pageNumber="1">, 86T</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="13420B0979127375F360A441135494E4" pageId="0" pageNumber="1" type="type material">
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<paragraph id="AEAEB7D7997F35F2FC2374470AE25501" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="4D1D9EE376B3F79770A775010DC18F9F" pageId="0" pageNumber="1">
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<emphasis id="B3F0137A9621BC146B839771C32071F9" bold="true" pageId="0" pageNumber="1">
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<emphasis id="889DEF4E2161CCDC4BBF9352A9D88C87" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Madagascar. •1 major worker; Toliara; Grand Lavasoa, 25.9 km W Tolagnaro;
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<geoCoordinate id="363BDF2BA81EAD1CF3C6C173F45CB961" degrees="25.08767" direction="south" orientation="latitude" precision="1" value="-25.08767">-25.08767</geoCoordinate>
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,
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<geoCoordinate id="471C92A1BEAAF17BB23849E400BA9A18" degrees="46.749" direction="east" orientation="longitude" precision="55" value="46.749">46.749</geoCoordinate>
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; alt. 450 m; 30 Nov 2006; B.L. Fisher et al. leg.; BLF15399, CASENT0122911 (CASC).
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<emphasis id="E47C881B80F59D45477FC3D556DE46F8" bold="true" pageId="0" pageNumber="1">
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<emphasis id="169D7C46C1A568DE5403D53307EAC719" italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
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.
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</emphasis>
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Madagascar. •1 w.; same data as for holotype; CASENT0235037 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection id="9D2F988E3938E6A756E1D53D2AA5447B" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="40BA3AE8D6FB44312A23028C9549FD03" pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph id="2F93B3636B77B381F5C2DEFFED5EF14E" pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis id="8F81C9B33EC35E5927C6944675166AA4" bold="true" italics="true" pageId="0" pageNumber="1">Toliara</emphasis>
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: •1w., 1s.; Manatantely, 8.9 km NW Tolagnaro;
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<geoCoordinate id="7C2E49A389C7CF47ACF97721C48ABD04" degrees="24.9815" direction="south" orientation="latitude" precision="5" value="-24.9815">-24.9815</geoCoordinate>
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,
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<geoCoordinate id="01A18E1F007F5ECFEE8C209E7B1BC69A" degrees="46.92567" direction="east" orientation="longitude" precision="1" value="46.92567">46.92567</geoCoordinate>
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; alt. 100 m; 27 Nov 2006; B.L. Fisher et al. leg.; CASENT0125808, CASENT0125809 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection id="1E36FE83CCFCA9FD63A74DC8FFC97F4E" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="4067BB6F3A26B28FD636F1F48888C9EE" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="80E1CDD9FE2EC399FBFB76FF833E00C3" pageId="0" pageNumber="1">
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<emphasis id="78FA90750E108756102A77843000621B" bold="true" pageId="0" pageNumber="1">
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<emphasis id="4EC607D0051CEE5DC10EB724194F1BEC" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
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.
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</emphasis>
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Head in full-face view square, anterior and posterior of eyes relatively straight; sides of the head with dense, short, erect pilosity; malar area with sparse to moderately dense longitudinal rugae, interspaces smooth to finely rugulose; inner hypostomal teeth distinct, big, thick, triangular, with rounded apex; outer hypostomal teeth slightly but distinctly smaller and thin, with rounded tips, and wide base; propodeal spines moderately long, with base slightly wider than top; first gastral tergite shiny and finely shagreened; body brownish black to black.
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<emphasis id="40D61AAD9793DDF4D3D800D1DE9331EF" bold="true" pageId="0" pageNumber="1">
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<emphasis id="D18A69384C1772F7723C67930406FC53" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
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.
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</emphasis>
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Whole head with sparse foveolae, frons and malar area with few additional, indistinct, longitudinal rugae, genae and centre of frons with sculpture reduced to absent; promesonotum low, slightly convex, with relatively steep posterior declivity; mesosoma foveolate, reduced on dorsal surface of pronotum, with katepisternum, anepisternum, promesonotal declivity, and lateral surfaces of propodeum smooth; propodeal spines very short, triangular; body dark brown.
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</paragraph>
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</subSubSection>
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<subSubSection id="698EB8409AB00BB0A09DC7AAB30D1AE1" pageId="0" pageNumber="1" type="description">
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<paragraph id="E94BA0D6CAA897F2BFAD9500F44499B6" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="096718052A87645EA8C0DB3D12B7CDDA" pageId="0" pageNumber="1">
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<emphasis id="9AA46D8D87E0B29AE836D30C8616215C" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis id="ADA693C21EC3F5EBD5BB9759F7DA9FD0" italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 1): HL: 1.38; HW: 1.33; SL: 0.58; EL: 0.15; WL: 0.9; PSL: 0.2; MTL: 0.56; PNW: 0.63; PTW: 0.17; PPW: 0.61; CI: 96.3; SI: 43.8; PSLI: 14.1; PPI: 27.9; PNI: 47.1; MTI: 42.2.
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<emphasis id="6B7FDE01930E7873CE425487B5A89535" bold="true" pageId="0" pageNumber="1">
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<emphasis id="F015E666E7DB83009FFCC2F62969FAB7" italics="true" pageId="0" pageNumber="1">Head</emphasis>
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.
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</emphasis>
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In full-face view square, anterior and posterior of eyes relatively straight (Fig.
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<figureCitation id="C79DF6A6D3D8DE434CB1DD64BDE3BDD4" captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Pheidole glabra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235034) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134271)." figureDoi="10.3897/zookeys.905.39592.figure68" httpUri="https://binary.pensoft.net/fig/373115" pageId="0" pageNumber="1">68B</figureCitation>
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). In lateral view sub-rectangular; ventral and dorsal faces relatively flat; dorsal face indistinctly depressed posteriorly, forming indistinct transverse depression between frons and occipital lobes; inner hypostomal teeth visible. Sides of the head with dense, short, erect pilosity; whole head with dense, short, suberect to erect pilosity. Antennal scrobes indistinct and not delimited by carinulae; scrobe surface shiny, with dense to sparse, fine, longitudinal to irregular rugae, interspaces smooth to rugulae. Occipital lobes with thick, sparse, irregular rugae, interspaces smooth; frons with thick, sparse to moderately sparse, longitudinal rugae, interspaces smooth; malar area with sparse to moderately dense longitudinal rugae, interspaces smooth to finely rugulose; genae with fine, thin to indistinct rugulae. Clypeus shiny and smooth, with thin, longitudinal rugulae on the lateral sides; median notch present, wide and shallow; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, slightly surpassing the mid-length of head; pilosity decumbent to erect (Fig.
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<figureCitation id="C8DB233A433FA9B7A32F8AE0ECCC27C4" captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Pheidole glabra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235034) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134271)." figureDoi="10.3897/zookeys.905.39592.figure68" httpUri="https://binary.pensoft.net/fig/373115" pageId="0" pageNumber="1">68B, D</figureCitation>
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). Inner hypostomal teeth distinct, big, thick, triangular, with rounded apex; outer hypostomal teeth slightly but distinctly smaller and thin, with rounded tips, and wide base (Fig.
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<figureCitation id="E5E8434A52B711B5D4C9636F11A6CC40" captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84T</figureCitation>
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).
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<emphasis id="358D12FC90666365180DCDA002D07001" bold="true" pageId="0" pageNumber="1">
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<emphasis id="E7911910DADC5A8D9E2772A260AD488E" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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.
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</emphasis>
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In lateral view, promesonotum short, angular, and relatively low, posterior mesonotum slightly steep, with indistinct tubercle-like projections; promesonotal groove absent; metanotal groove absent or indistinct; propodeal spines moderately long, with base slightly wider than top, apex rounded; humeral area with small and flat tubercles (Fig.
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<figureCitation id="0E855BFAC7C44A789CAC6D9700BDD039" captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Pheidole glabra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235034) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134271)." figureDoi="10.3897/zookeys.905.39592.figure68" httpUri="https://binary.pensoft.net/fig/373115" pageId="0" pageNumber="1">68D</figureCitation>
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). Surface shiny, with fine and sparse rugoreticulation, pronotum with sculpture reduced, sometimes with smooth notches on lateral sides and centre of its dorsal surface; katepisternum and lateral sides of propodeum at least partially smooth. Pilosity moderately dense, long, and erect (Fig.
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<figureCitation id="29E28DDE166EF06BF1ADAA43D9142A5C" captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Pheidole glabra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235034) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134271)." figureDoi="10.3897/zookeys.905.39592.figure68" httpUri="https://binary.pensoft.net/fig/373115" pageId="0" pageNumber="1">68D, F</figureCitation>
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).
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<emphasis id="23841A19549E5368F4559E4B568ABD3A" bold="true" pageId="0" pageNumber="1">
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<emphasis id="03E26A42863A122D587DE7A616833816" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Shiny, with dense and fine rugulae; peduncle short, with small, rounded, horizontal lobes on its basal part; node moderately high and narrow, with convex apex, in rear view node slightly dorsoventrally concave; pilosity moderately and erect, long to moderately long (Fig.
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<figureCitation id="F88ACBD152C6E5BEA7E061EB933EB562" captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Pheidole glabra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235034) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134271)." figureDoi="10.3897/zookeys.905.39592.figure68" httpUri="https://binary.pensoft.net/fig/373115" pageId="0" pageNumber="1">68D, F</figureCitation>
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).
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<emphasis id="68B59ACAF5BDEBDAB7F5F3AC6FFB0A14" bold="true" pageId="0" pageNumber="1">
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<emphasis id="BC05B922F13B822EAF8CAC02B86C9C8A" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
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.
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</emphasis>
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Shiny, with fine and dense rugulae; short and rounded; in dorsal view sides with relatively long, acute, narrow and triangular projections; pilosity long, moderately dense, and erect (Fig.
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<figureCitation id="27C5E5CA4C931829A3ADC7F495464E8A" captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Pheidole glabra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235034) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134271)." figureDoi="10.3897/zookeys.905.39592.figure68" httpUri="https://binary.pensoft.net/fig/373115" pageId="0" pageNumber="1">68D, F</figureCitation>
|
|
).
|
|
<emphasis id="4F313976FAAB62C38F67DE66ADDA1F40" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="C3A8EAA3DE50CA976E7DC0BCAD122B50" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
First gastral tergite shiny and finely shagreened; pilosity moderately dense, moderately long, and erect (Fig.
|
|
<figureCitation id="79C87D79C4D9C2228F2E955326459A56" captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Pheidole glabra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235034) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134271)." figureDoi="10.3897/zookeys.905.39592.figure68" httpUri="https://binary.pensoft.net/fig/373115" pageId="0" pageNumber="1">68D, F</figureCitation>
|
|
).
|
|
<emphasis id="18D5AF7199B9EF12C37F507E64A7DA76" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="A1F3C1ED07ECB6E1F6BBF7BC865443C4" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
|
.
|
|
</emphasis>
|
|
Unicolourous, brownish black to black; legs yellow to brown (Fig.
|
|
<figureCitation id="4D104233B85CCC3CD0230A37A246DEB4" captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Pheidole glabra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235034) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134271)." figureDoi="10.3897/zookeys.905.39592.figure68" httpUri="https://binary.pensoft.net/fig/373115" pageId="0" pageNumber="1">68D, F</figureCitation>
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|
).
|
|
</paragraph>
|
|
<caption id="C0A63269CF96F53D4835176DCB2D6CAB" doi="10.3897/zookeys.905.39592.figure68" httpUri="https://binary.pensoft.net/fig/373115" pageId="0" pageNumber="1" start="Figure 68" startId="F68">
|
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<paragraph id="A17E10645E99CFED77F100D69EA191DD" pageId="0" pageNumber="1">
|
|
<emphasis id="9150F2F8703BDF0E2F2A77D049F01DDF" bold="true" pageId="0" pageNumber="1">Figure 68.</emphasis>
|
|
<taxonomicName id="A429BFF805C59E72E3338ABF40887ECD" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole glabra" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="glabra">
|
|
<emphasis id="FA64E82063F99829CCFB6EDECE39E5F7" italics="true" pageId="0" pageNumber="1">Pheidole glabra</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., full-face view (
|
|
<emphasis id="0750FF970F8B1139CC824EB736C2893C" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
), profile (
|
|
<emphasis id="1267C506BA6E27C675723F5E744423E3" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
), and dorsal view (
|
|
<emphasis id="DF25BC1C7FDCDFEAAAF8A0F0DEF2A8B3" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
|
) of paratype minor worker (CASENT0235034) and full-face view (
|
|
<emphasis id="8EDD42F17CA2015AE24882FCDF8F5561" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
), profile (
|
|
<emphasis id="49C27FF6671F8E7EF0CD535AEC756170" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
), and dorsal view (
|
|
<emphasis id="A0CCC6002F13432AB5B722B19D1F0680" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
|
) of holotype major worker (CASENT0134271).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="40F72CBEF2799FFE6C3CAA0666768A35" pageId="0" pageNumber="1">
|
|
<emphasis id="398574C0E05FA6BA2F29313004D837B1" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
|
Measurements (
|
|
<emphasis id="F70449791CD2F8F7DABEE53A771E57A1" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
|
= 1): HL: 0.53; HW: 0.47; SL: 0.52; EL: 0.1; WL: 0.59; PSL: 0.1; MTL: 0.4; PNW: 0.32; PTW: 0.07; PPW: 0.11; CI: 88.4; SI: 111.4; PSLI: 18.0; PPI: 64.3; PNI: 68.9; MTI: 84.1.
|
|
<emphasis id="BD2E16E93B3F0007660C44EBD4283F46" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="D9F8F3D66EFA21EA0139091732AD8011" italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
|
.
|
|
</emphasis>
|
|
Occipital margin straight or indistinctly concave; occipital carina indistinct, weakly developed (Fig.
|
|
<figureCitation id="045E94BA23B8CCA3C268557151531AA2" captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Pheidole glabra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235034) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134271)." figureDoi="10.3897/zookeys.905.39592.figure68" httpUri="https://binary.pensoft.net/fig/373115" pageId="0" pageNumber="1">68A</figureCitation>
|
|
). Pilosity moderately dense, long, erect. Whole head with sparse foveolae, frons and malar area with few additional, indistinct, longitudinal rugae, genae and centre of frons with sculpture reduced to absent. Clypeus shiny, rugoreticulate; median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by one-fifth of its length; pilosity erect (Fig.
|
|
<figureCitation id="59C5D6DD4A2D4E536F350DECAB495956" captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Pheidole glabra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235034) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134271)." figureDoi="10.3897/zookeys.905.39592.figure68" httpUri="https://binary.pensoft.net/fig/373115" pageId="0" pageNumber="1">68A, C</figureCitation>
|
|
).
|
|
<emphasis id="34F93639472F05E7FCEFE3FC2BF39FB2" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="AA246C131DF87546B6100E3AC2610D3F" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
|
.
|
|
</emphasis>
|
|
In lateral view, promesonotum low, slightly convex, with relatively steep posterior declivity; promesonotal groove absent; metanotal groove indistinct; propodeal spines very short, triangular, apex acute (Fig.
|
|
<figureCitation id="CB4F877B206C233B8F98A432AB8F4BC5" captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Pheidole glabra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235034) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134271)." figureDoi="10.3897/zookeys.905.39592.figure68" httpUri="https://binary.pensoft.net/fig/373115" pageId="0" pageNumber="1">68C</figureCitation>
|
|
). Sculpture foveolate, reduced on dorsal surface of pronotum, with katepisternum, anepisternum, promesonotal declivity, and lateral surfaces of propodeum smooth. Pilosity sparse, long, and erect (Fig.
|
|
<figureCitation id="F9330C112A8C08A01D40477568BF41F0" captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Pheidole glabra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235034) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134271)." figureDoi="10.3897/zookeys.905.39592.figure68" httpUri="https://binary.pensoft.net/fig/373115" pageId="0" pageNumber="1">68C, E</figureCitation>
|
|
).
|
|
<emphasis id="6C8206CB3070796EBFC01A1EFD5DACB2" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="D3DE938DB06B17C91C8B02743E3CD31A" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Peduncle short and thin with ventral face slightly convex; node low, globular, and small; with few long, erect setae (Fig.
|
|
<figureCitation id="5B2E01D76D3E6424AAC6E256B226A150" captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Pheidole glabra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235034) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134271)." figureDoi="10.3897/zookeys.905.39592.figure68" httpUri="https://binary.pensoft.net/fig/373115" pageId="0" pageNumber="1">68C, E</figureCitation>
|
|
).
|
|
<emphasis id="B108A1DDAE7EEB0D2551B893C20E78FB" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="A75646C52782E656FE35995FBD4DEE46" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Moderately long, low, and convex; with few short, erect setae (Fig.
|
|
<figureCitation id="DC7CB5E78F8F648783CE0A8C84CB7A3A" captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Pheidole glabra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235034) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134271)." figureDoi="10.3897/zookeys.905.39592.figure68" httpUri="https://binary.pensoft.net/fig/373115" pageId="0" pageNumber="1">68C, E</figureCitation>
|
|
).
|
|
<emphasis id="72968E41E817AEED446E973785FCE431" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="3A9EAE8A6826B3A2616C5E050D03C590" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
With moderately sparse, erect pilosity (Fig.
|
|
<figureCitation id="E192C382067BAE66EED0F427DB751414" captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Pheidole glabra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235034) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134271)." figureDoi="10.3897/zookeys.905.39592.figure68" httpUri="https://binary.pensoft.net/fig/373115" pageId="0" pageNumber="1">68C, E</figureCitation>
|
|
).
|
|
<emphasis id="9DE0737CA5C9282A310B36CE5F81104D" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="93B58F39F922AA94866F8BB2DD834F2B" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
|
.
|
|
</emphasis>
|
|
Unicolourous, dark brown (Fig.
|
|
<figureCitation id="A6FFCB9250A031F70E5BA0D75C34D6AA" captionStart="Figure 68" captionStartId="F68" captionText="Figure 68. Pheidole glabra sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235034) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0134271)." figureDoi="10.3897/zookeys.905.39592.figure68" httpUri="https://binary.pensoft.net/fig/373115" pageId="0" pageNumber="1">68C, E</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="BA2CCA80A0AA7D70AC49F2C92A146F04" pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph id="6903A58A4083DA28D9313FD860B65AF3" pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph id="2723BEB20EF8EBFDD1FC962DE0F8A2F3" pageId="0" pageNumber="1">Latin for smooth, in reference to reduced sculpture of minor workers.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="49FC5842E66042F215E2D9ADE17C0083" pageId="0" pageNumber="1" type="biology_ecology">
|
|
<paragraph id="EE4E57428B374246D40676054EBC3FDA" pageId="0" pageNumber="1">Biology.</paragraph>
|
|
<paragraph id="D98D9F5E8B12FB04DBA27065EAA89F95" pageId="0" pageNumber="1">The species was collected at 450 m in elevation, in rainforest. Nest was located in rotten log.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="6FAEAA7DD3BA2169DE8933A84FAF4F43" pageId="0" pageNumber="1" type="comments">
|
|
<paragraph id="9637596C2495317F59EB4108B980AEE9" pageId="0" pageNumber="1">Comments.</paragraph>
|
|
<paragraph id="E5A17E43DEC52CBA458C6699BC0D5AD0" pageId="0" pageNumber="1">
|
|
This species is most similar to
|
|
<taxonomicName id="CF30ADB59B3582F639C76DBF6E4930FC" lsidName="P. petax" pageId="0" pageNumber="1" rank="species" species="petax">
|
|
<emphasis id="DA644C7622991783BDE13C12DF846040" italics="true" pageId="0" pageNumber="1">P. petax</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="DC62E7288A060815A88B8DBC3E01653A" lsidName="P. brevipilosa" pageId="0" pageNumber="1" rank="species" species="brevipilosa">
|
|
<emphasis id="5B3EABB342E7F0E56847032F7A618D81" italics="true" pageId="0" pageNumber="1">P. brevipilosa</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
<emphasis id="373427820C5C4A26A3B28384D143AD3E" bold="true" pageId="0" pageNumber="1">
|
|
<emphasis id="C9A8ACC0F5BBE203E8CA1C9B7CE4B16F" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
<taxonomicName id="C7C844EE185CDC3E401C69C9AF460E4E" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole glabra" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="glabra">
|
|
<emphasis id="24C97F66F7C9896935DD83993FE49ADD" italics="true" pageId="0" pageNumber="1">Pheidole glabra</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. can be distinguished from
|
|
<taxonomicName id="11EA8AAB3D1D203C22E25E95DC7E1758" lsidName="P. petax" pageId="0" pageNumber="1" rank="species" species="petax">
|
|
<emphasis id="C7905CECB0AFC561224309D4A3D1A3C8" italics="true" pageId="0" pageNumber="1">P. petax</emphasis>
|
|
</taxonomicName>
|
|
by presence of erect pilosity on sides of head and lateral sides of head, and square head shape in full-face view; from
|
|
<taxonomicName id="F866934989745EE8BF6211FA2F1DCFA0" lsidName="P. brevipilosa" pageId="0" pageNumber="1" rank="species" species="brevipilosa">
|
|
<emphasis id="1BC3FBCE71529F53CD109F6A3447CE84" italics="true" pageId="0" pageNumber="1">P. brevipilosa</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by absence of foveolate sculpture on head, well-developed outer hypostomal teeth, pronotum with reduced sculpture and presence of smooth notches on its lateral and dorsal surfaces, and propodeum with presence of smooth notches on lateral sides
|
|
<emphasis id="1282F2D3E450FD944C454A32384F0DD1" bold="true" pageId="0" pageNumber="1">
|
|
.
|
|
<emphasis id="1CAE262A85353B836E2F1AEA5D1883C7" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
<taxonomicName id="8A6DBEF7A0CBC8893F245C577D32EC93" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole glabra" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="glabra">
|
|
<emphasis id="067B05D412572E107263CC6E107E69B9" italics="true" pageId="0" pageNumber="1">Pheidole glabra</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. can be distinguished from
|
|
<taxonomicName id="1074E7E7886887AC8CE1BAB4DC471367" lsidName="P. petax" pageId="0" pageNumber="1" rank="species" species="petax">
|
|
<emphasis id="D8D4E21CB8AD03315717CE6706F4BF06" italics="true" pageId="0" pageNumber="1">P. petax</emphasis>
|
|
</taxonomicName>
|
|
by dark brown body colouration, presence of additional indistinct, longitudinal rugae on frons and malar area, clypeus never foveolate, and reduced sculpture on frons, dorsal surface of pronotum, promesonotal declivity, and lateral surfaces of propodeum; from
|
|
<taxonomicName id="C640EB04076541A7B9F4DFCD98F5DDBD" lsidName="P. brevipilosa" pageId="0" pageNumber="1" rank="species" species="brevipilosa">
|
|
<emphasis id="05E43AFD21AC28C16A09796FD18F8022" italics="true" pageId="0" pageNumber="1">P. brevipilosa</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by denser and longer pilosity on head, and frons, dorsal surface of pronotum, promesonotal declivity, and lateral surfaces of propodeum with sculpture reduced or absent.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |