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<mods:title id="9B9CEE609887031C73955859C069B4CE">Synopsis of the pelidnotine scarabs (Coleoptera, Scarabaeidae, Rutelinae, Rutelini) and annotated catalog of the species and subspecies</mods:title>
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<mods:affiliation id="F86918A910EFE428B2D7483F8EEA882A">Department of Entomology and Nematology, University of Florida Building 1881 Natural Drive Area, Steinmetz Hall, Box 110620, Gainesville, FL 32611 - 0620, USA</mods:affiliation>
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<mods:namePart id="55908A136764DDB41D0232B404C4B87B">Jameson, Mary L.</mods:namePart>
<mods:affiliation id="E36D182F055E77419CE195391E72262E">Department of Biological Sciences, Wichita State University 1845 Fairmount, Box 26, Wichita, KS 67260 - 0026, USA</mods:affiliation>
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<mods:affiliation id="E8A2462DFBA08E5620239306DD4C2F33">Musee des Confluences, Centre de Conservation et d'Etude des Collections, 13 A Rue Bancel, F- 69007 Lyon, France</mods:affiliation>
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<mods:namePart id="17B5AF1AF768B1141371B94DC1D55587">Smith, Andrew B. T.</mods:namePart>
<mods:affiliation id="F58B1F29C190A42396D1A996B644021E">Research Division, Canadian Museum of Nature, P. O. Box 3443, Station D, Ottawa, Ontario, K 1 P 6 P 4, Canada</mods:affiliation>
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<treatment id="C6D3E092A1BB26254D21140C29E342EB" ID-GBIF-Taxon="128344759" LSID="urn:lsid:plazi:treatment:C6D3E092A1BB26254D21140C29E342EB" httpUri="http://treatment.plazi.org/id/C6D3E092A1BB26254D21140C29E342EB" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName id="3C326D20CD20412A15DA7A14A17FDCEA" ID-CoL="3MLXL" LSID="C6D3E092-A1BB-2625-4D21-140C29E342EB" authority="(Laporte, 1840)" baseAuthorityName="Laporte" baseAuthorityYear="1840" class="Insecta" family="Scarabaeidae" genus="Hoplopelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopelidnota metallica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="metallica">Hoplopelidnota metallica (Laporte, 1840)</taxonomicName>
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<paragraph id="2241C1F82A2EF433C43BE1E261D5A877" pageId="0" pageNumber="1">
<taxonomicName id="CF8633E436A722FF179DD57FBDAF1D7B" authorityName="Laporte" authorityYear="1840" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota metallica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="metallica">Pelidnota metallica</taxonomicName>
Laporte, 1840: 122 [original combination].
</paragraph>
<paragraph id="5760E14CA2F42D56080C676F299C8BC1" pageId="0" pageNumber="1">
<taxonomicName id="99225D844C261A8B364EF7C5FD655DBA" authorityName="Bates" authorityYear="1904" class="Insecta" family="Scarabaeidae" genus="Hoplopelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopelidnota candezei" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="candezei">Hoplopelidnota candezei</taxonomicName>
F. Bates [syn. by
<bibRefCitation id="05D9EEFD739B4D150A395AD83F4D6A69" author="Machatschke, JW" journalOrPublisher="Coleopterorum Catalogus Supplementa" pageId="0" pageNumber="1" pagination="1 - 361" refId="B188" refString="Machatschke, JW, 1972. Scarabaeoidea: Melolonthidae, Rutelinae. Coleopterorum Catalogus Supplementa 66: 1 - 361" title="Scarabaeoidea: Melolonthidae, Rutelinae." volume="66" year="1972">Machatschke 1972</bibRefCitation>
: 12].
</paragraph>
<paragraph id="AA1EBD494507171B0B9A74D77FF769D7" pageId="0" pageNumber="1">
<taxonomicName id="1803087E9F0FBB81EE127417393BDCEF" baseAuthorityName="Laporte" baseAuthorityYear="1840" class="Insecta" family="Scarabaeidae" genus="Hoplopelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopelidnota metallica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="metallica">Hoplopelidnota metallica</taxonomicName>
(Laporte) [revised species status by
<bibRefCitation id="26122DADD167F640F865E030BF96FB8B" author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B359" refString="Soula, M, 2008. Les Coleopteres du Nouveau Monde. Volume 2: Rutelini 2. Revision des Pelidnotina 2. Revision des genres: Parhoplognathus, Chipita n. gen., Heteropelidnota, Homothermon, Hoplopelidnota, Mesomerodon, Mecopelidnota, Patatra n. gen. (Coleoptera: Scarabaeidae, Rutelinae, Rutelini). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 2: Rutelini 2. Revision des Pelidnotina 2. Revision des genres: Parhoplognathus, Chipita n. gen., Heteropelidnota, Homothermon, Hoplopelidnota, Mesomerodon, Mecopelidnota, Patatra n. gen. (Coleoptera: Scarabaeidae, Rutelinae, Rutelini). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2008">Soula 2008</bibRefCitation>
: 17].
</paragraph>
<paragraph id="70A8329CBF1E74CE00C30EBB0E7D73D7" pageId="0" pageNumber="1">
<taxonomicName id="D9AF6345E818F12006E2ABED946962BA" authorityName="Ohaus" authorityYear="1912" class="Insecta" family="Scarabaeidae" genus="Hoplopelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopelidnota armata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="armata">Hoplopelidnota armata</taxonomicName>
Ohaus, 1912
<emphasis id="0CD96CE8C0F9C204A5CDCFF8543F17ED" bold="true" pageId="0" pageNumber="1">synonym.</emphasis>
</paragraph>
<paragraph id="9B562537ABBE28BB84429D0DCF6F0A99" pageId="0" pageNumber="1">
<taxonomicName id="36D036AA53FF2B3BC5C79FE259CDC55C" authorityName="Ohaus" authorityYear="1912" class="Insecta" family="Scarabaeidae" genus="Hoplopelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopelidnota armata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="armata">Hoplopelidnota armata</taxonomicName>
Ohaus, 1912: 309 [original combination; sometimes erroneously referred to as
<taxonomicName id="11C2247BE4EED2DE36576FBF38ED592F" family="Scarabaeidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="H. armata" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="armata">
<emphasis id="E5ACCA0BA332B5AE41D1417455B0E8E4" italics="true" pageId="0" pageNumber="1">H. armata</emphasis>
</taxonomicName>
F. Bates].
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<taxonomicName id="6464CD4232046045BAF1F58A6A67F411" baseAuthorityName="Laporte" baseAuthorityYear="1840" class="Insecta" family="Scarabaeidae" genus="Hoplopelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopelidnota metallica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="metallica">Hoplopelidnota metallica</taxonomicName>
(Laporte) [syn. by
<bibRefCitation id="FD936386A3787D060B1BAD9A6A6EB0B1" DOI="https://doi.org/10.1649/0010-065X-67.3.377" author="Moore, MR" journalOrPublisher="The Coleopterists Bulletin" pageId="0" pageNumber="1" pagination="377 - 387" refId="B226" refString="Moore, MR, Jameson, ML, 2013. Taxonomic and nomenclatural changes in the pelidnotine scarabs (Coleoptera: Scarabaeidae: Rutelinae: Rutelini). The Coleopterists Bulletin 67: 377 - 387, DOI: https://doi.org/10.1649/0010-065X-67.3.377" title="Taxonomic and nomenclatural changes in the pelidnotine scarabs (Coleoptera: Scarabaeidae: Rutelinae: Rutelini)." url="https://doi.org/10.1649/0010-065X-67.3.377" volume="67" year="2013">Moore and Jameson 2013</bibRefCitation>
: 381].
</paragraph>
<paragraph id="C8D74879F9E96BC056908C6BE22A92C4" pageId="0" pageNumber="1">
<taxonomicName id="9EBE8542C0B93FDFF3DDBF334892CD73" authorityName="Bates" authorityYear="1904" class="Insecta" family="Scarabaeidae" genus="Hoplopelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopelidnota candezei" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="candezei">Hoplopelidnota candezei</taxonomicName>
F. Bates, 1904
<emphasis id="B895501383ED50BF22C03E74F55E90EE" bold="true" pageId="0" pageNumber="1">synonym.</emphasis>
</paragraph>
<paragraph id="B93D39E00BA4F2170D64987FA72E545C" pageId="0" pageNumber="1">
<taxonomicName id="CAB430F413508DAA99246B2F94C1248B" authorityName="Bates" authorityYear="1904" class="Insecta" family="Scarabaeidae" genus="Hoplopelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopelidnota candezei" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="candezei">Hoplopelidnota candezei</taxonomicName>
F. Bates, 1904: 274-275 [original combination].
</paragraph>
<paragraph id="55D416055C6371DBFC240E7B5A14196A" pageId="0" pageNumber="1">
<taxonomicName id="B300ED577BD24D22C7ED8D91BA44948E" baseAuthorityName="Laporte" baseAuthorityYear="1840" class="Insecta" family="Scarabaeidae" genus="Hoplopelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopelidnota metallica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="metallica">Hoplopelidnota metallica</taxonomicName>
(Laporte) [syn. by
<bibRefCitation id="0B26883AF778F364068CD60D7B2FAAE9" author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B359" refString="Soula, M, 2008. Les Coleopteres du Nouveau Monde. Volume 2: Rutelini 2. Revision des Pelidnotina 2. Revision des genres: Parhoplognathus, Chipita n. gen., Heteropelidnota, Homothermon, Hoplopelidnota, Mesomerodon, Mecopelidnota, Patatra n. gen. (Coleoptera: Scarabaeidae, Rutelinae, Rutelini). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 2: Rutelini 2. Revision des Pelidnotina 2. Revision des genres: Parhoplognathus, Chipita n. gen., Heteropelidnota, Homothermon, Hoplopelidnota, Mesomerodon, Mecopelidnota, Patatra n. gen. (Coleoptera: Scarabaeidae, Rutelinae, Rutelini). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2008">Soula 2008</bibRefCitation>
: 17].
</paragraph>
</subSubSection>
<subSubSection id="CAD354654405FDB0C2090AC026783ACB" pageId="0" pageNumber="1" type="distribution">
<paragraph id="B5D7CB88714D809E61A4DA10E717328B" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="4285DAB3FA589E3E664CBC077AE21476" pageId="0" pageNumber="1">
BRAZIL: Territorio de Amapa (Serra Navia). FRENCH GUIANA: Cayenne (
<bibRefCitation id="C2380BF8425CCE7412D21C82C455A0BC" author="Laporte, [= Castelnau] FLNCde" journalOrPublisher="Dumenil, Paris" pageId="0" pageNumber="1" refId="B163" refString="Laporte, [=Castelnau] FLNCde, 1840. Histoire naturelle des insectes Coleopteres; avec une introduction renfermant l'anatomie et la physiologie des animaux articules, par M. Brulle. Tome premier. Histoire naturelle des animaux articules, annelides, crustaces, arachnides, myriapodes et insectes Tome troisieme. P. Dumenil, Paris" title="Histoire naturelle des insectes Coleopteres; avec une introduction renfermant l'anatomie et la physiologie des animaux articules, par M. Brulle. Tome premier. Histoire naturelle des animaux articules, annelides, crustaces, arachnides, myriapodes et insectes Tome troisieme. P." year="1840">Laporte 1840</bibRefCitation>
, F.
<bibRefCitation id="85139946823306FF85908C44B6BAA8E5" DOI="https://doi.org/10.1111/j.1365-2311.1904.tb02746.x" author="Bates, F" journalOrPublisher="Transactions of the Entomological Society of London" pageId="0" pageNumber="1" pagination="249 - 276" refId="B7" refString="Bates, F, 1904. A revision of the sub-family Pelidnotinae of the coleopterous family Rutelidae, with descriptions of new genera and species. Transactions of the Entomological Society of London 1904: 249 - 276, DOI: https://doi.org/10.1111/j.1365-2311.1904.tb02746.x" title="A revision of the sub-family Pelidnotinae of the coleopterous family Rutelidae, with descriptions of new genera and species." url="https://doi.org/10.1111/j.1365-2311.1904.tb02746.x" volume="1904" year="1904">Bates 1904</bibRefCitation>
,
<bibRefCitation id="696E14316E604D98163335152B33D936" author="Ohaus, F" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="0" pageNumber="1" pagination="273 - 319" refId="B273" refString="Ohaus, F, 1912. Beitraege zur Kenntnis der Ruteliden, X. Stettiner Entomologische Zeitung 73: 273 - 319" title="Beitraege zur Kenntnis der Ruteliden, X." volume="73" year="1912">Ohaus 1912</bibRefCitation>
,
<bibRefCitation id="FDD7A3B302DC853A5BA458985E1700D8" author="Ohaus, F" journalOrPublisher="Coleopterorum Catalogus" pageId="0" pageNumber="1" pagination="1 - 241" refId="B278" refString="Ohaus, F, 1918. Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae. Coleopterorum Catalogus 20: 1 - 241" title="Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae." volume="20" year="1918">1918</bibRefCitation>
,
<bibRefCitation id="88E43030883E90F1EE7C2CBE9E33FE99" author="Ohaus, F" journalOrPublisher="Mitteilungen der Deutschen Entomologischen Gesellschaft" pageId="0" pageNumber="1" refId="B286" refString="Ohaus, F, 1934b. Coleopteralamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T.1: Tribus Rutelini. Genera Insectorum Fasc. 199A: 1-172." title="Coleopteralamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T. 1: Tribus Rutelini. Genera Insectorum Fasc. 199 A: 1 - 172." year="1934 b">1934b</bibRefCitation>
,
<bibRefCitation id="BE3ED54513F717CC747E1C0889D97DA1" author="Blackwelder, RE" journalOrPublisher="United States National Museum Bulletin" pageId="0" pageNumber="1" pagination="189 - 265" refId="B14" refString="Blackwelder, RE, 1944. Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America. United States National Museum Bulletin 185: 189 - 265" title="Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
,
<bibRefCitation id="A4A598327BA14ECFC1EE4AF50FE61A83" author="Machatschke, JW" journalOrPublisher="Coleopterorum Catalogus Supplementa" pageId="0" pageNumber="1" pagination="1 - 361" refId="B188" refString="Machatschke, JW, 1972. Scarabaeoidea: Melolonthidae, Rutelinae. Coleopterorum Catalogus Supplementa 66: 1 - 361" title="Scarabaeoidea: Melolonthidae, Rutelinae." volume="66" year="1972">Machatschke 1972</bibRefCitation>
,
<bibRefCitation id="34CBB0E1F1A72EECE5EA47554D0E5E64" author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B359" refString="Soula, M, 2008. Les Coleopteres du Nouveau Monde. Volume 2: Rutelini 2. Revision des Pelidnotina 2. Revision des genres: Parhoplognathus, Chipita n. gen., Heteropelidnota, Homothermon, Hoplopelidnota, Mesomerodon, Mecopelidnota, Patatra n. gen. (Coleoptera: Scarabaeidae, Rutelinae, Rutelini). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 2: Rutelini 2. Revision des Pelidnotina 2. Revision des genres: Parhoplognathus, Chipita n. gen., Heteropelidnota, Homothermon, Hoplopelidnota, Mesomerodon, Mecopelidnota, Patatra n. gen. (Coleoptera: Scarabaeidae, Rutelinae, Rutelini). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2008">Soula 2008</bibRefCitation>
,
<bibRefCitation id="854AAAD547C9667FC6FBF16348263B3D" author="Soula, M" editor="Touroult, J" journalOrPublisher="Tome II. Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;, Paris" pageId="0" pageNumber="1" pagination="50 - 61" refId="B363" refString="Soula, M, 2010c. Les Rutelinae: presentation des tribus et genres de Guyane (Coleoptera, Scarabaeidae). In: Touroult, J, Ed., Contribution a l'Etude des Coleopteres de Guyane. Tome II. Supplement au Bulletin de liaison d'ACOREP-France &quot;Le Coleopteriste&quot;, Paris: 50 - 61" title="Les Rutelinae: presentation des tribus et genres de Guyane (Coleoptera, Scarabaeidae)." volumeTitle="Contribution a l'Etude des Coleopteres de Guyane." year="2010 c">2010c</bibRefCitation>
). GUYANA: Essequibo River, Moraballi Creek. VENEZUELA: Amazonas (Rio Negro).
</paragraph>
</subSubSection>
<subSubSection id="EC1E429B223E26C28D7D01CA823D596B" pageId="0" pageNumber="1" type="types">
<paragraph id="5A5A2B673C71A72E79BE0B0F49AF93FC" pageId="0" pageNumber="1">Types.</paragraph>
<paragraph id="5B02D8083EA1D9D7EBC577FCC1947692" pageId="0" pageNumber="1">
The following specimen is deposited at CCECL. 1 invalid ♂ neotype (Fig.
<figureCitation id="8A42F96B89E4551C5333D3F899246CB0" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Hoplopelidnota metallica (Laporte) invalid neotype male from CCECL. A Dorsal habitus B Male parameres, caudal view C Specimen labels." figureDoi="10.3897/zookeys.666.9191.figure36" httpUri="https://binary.pensoft.net/fig/130464" pageId="0" pageNumber="1">36</figureCitation>
): &quot;pk 23 p. de Belizon G. F. 8/91 coll. - SOULA [obverse] pk 23//
<normalizedToken id="32DD01820820E04B39B49B7FC0133089" originalValue="Néotype">Neotype</normalizedToken>
2007
<taxonomicName id="98BDF659352CAA2AFB2EFE228672EE70" authorityName="Laporte" authorityYear="1840" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota metallica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="0E79A438CE49E85275C29D8280FF97C7" italics="true" pageId="0" pageNumber="1">Pelidnota metallica</emphasis>
</taxonomicName>
Lap. Soula det.//
<taxonomicName id="7F3F9A521F6F07D3CF7A60F283DE71A9" baseAuthorityName="Laporte" baseAuthorityYear="1840" class="Insecta" family="Scarabaeidae" genus="Hoplopelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopelidnota metallica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="E0C45A8E87B0398BD3D5BECCDA164DD1" italics="true" pageId="0" pageNumber="1">Hoplopelidnota metallica</emphasis>
</taxonomicName>
(Lap.) M. SOULA 19 2007&quot; (47031033). Genitalia card-mounted underneath the invalid neotype. Box 4618690 SOULA.
</paragraph>
</subSubSection>
<subSubSection id="CCD5A5598B64FFFB7B67F95481700DB5" pageId="0" pageNumber="1" type="remarks">
<paragraph id="1589225D87550DF35FA6DBF4FECBFD26" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="545C764CEEDC04C02540EBFAB0173A53" pageId="0" pageNumber="1">
The classification of
<taxonomicName id="1E94926C0BAEFE82EA27763E7119EF26" baseAuthorityName="Laporte" baseAuthorityYear="1840" class="Insecta" family="Scarabaeidae" genus="Hoplopelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopelidnota metallica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="F1330C6AEBF902F3264F38CD4C499BE1" italics="true" pageId="0" pageNumber="1">Hoplopelidnota metallica</emphasis>
</taxonomicName>
(Figs
<figureCitation id="56E4F3AC32818197CADA77684DA61218" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Hoplopelidnota metallica (Laporte) invalid neotype male from CCECL. A Dorsal habitus B Male parameres, caudal view C Specimen labels." figureDoi="10.3897/zookeys.666.9191.figure36" httpUri="https://binary.pensoft.net/fig/130464" pageId="0" pageNumber="1">36</figureCitation>
,
<figureCitation id="F8AE197981361F2E958DC291C85B5CE0" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Hoplopelidnota metallica (Laporte) male from MSPC. A Dorsal habitus B Lateral habitus C Male genitalia, lateral view D Male parameres, caudal view." figureDoi="10.3897/zookeys.666.9191.figure37" httpUri="https://binary.pensoft.net/fig/130465" pageId="0" pageNumber="1">37</figureCitation>
) has been tumultuous (
<bibRefCitation id="29C95CFDCB8F7900C5F16FD8E7B238DC" DOI="https://doi.org/10.1649/0010-065X-67.3.377" author="Moore, MR" journalOrPublisher="The Coleopterists Bulletin" pageId="0" pageNumber="1" pagination="377 - 387" refId="B226" refString="Moore, MR, Jameson, ML, 2013. Taxonomic and nomenclatural changes in the pelidnotine scarabs (Coleoptera: Scarabaeidae: Rutelinae: Rutelini). The Coleopterists Bulletin 67: 377 - 387, DOI: https://doi.org/10.1649/0010-065X-67.3.377" title="Taxonomic and nomenclatural changes in the pelidnotine scarabs (Coleoptera: Scarabaeidae: Rutelinae: Rutelini)." url="https://doi.org/10.1649/0010-065X-67.3.377" volume="67" year="2013">Moore and Jameson 2013</bibRefCitation>
).
<bibRefCitation id="22BFE066A3E2368F239423D3C5C255B5" author="Laporte, [= Castelnau] FLNCde" journalOrPublisher="Dumenil, Paris" pageId="0" pageNumber="1" refId="B163" refString="Laporte, [=Castelnau] FLNCde, 1840. Histoire naturelle des insectes Coleopteres; avec une introduction renfermant l'anatomie et la physiologie des animaux articules, par M. Brulle. Tome premier. Histoire naturelle des animaux articules, annelides, crustaces, arachnides, myriapodes et insectes Tome troisieme. P. Dumenil, Paris" title="Histoire naturelle des insectes Coleopteres; avec une introduction renfermant l'anatomie et la physiologie des animaux articules, par M. Brulle. Tome premier. Histoire naturelle des animaux articules, annelides, crustaces, arachnides, myriapodes et insectes Tome troisieme. P." year="1840">Laporte (1840)</bibRefCitation>
named
<taxonomicName id="8A48E4DCA501B21789B41AA123E22717" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. metallica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="53C254859420896FD8A943FD3F896A34" italics="true" pageId="0" pageNumber="1">P. metallica</emphasis>
</taxonomicName>
, clearly indicating the unusual form of the elytral apex.
<bibRefCitation id="2C535D146D0DCF2B6070A7F7D0F62E9D" DOI="https://doi.org/10.1111/j.1365-2311.1904.tb02746.x" author="Bates, F" journalOrPublisher="Transactions of the Entomological Society of London" pageId="0" pageNumber="1" pagination="249 - 276" refId="B7" refString="Bates, F, 1904. A revision of the sub-family Pelidnotinae of the coleopterous family Rutelidae, with descriptions of new genera and species. Transactions of the Entomological Society of London 1904: 249 - 276, DOI: https://doi.org/10.1111/j.1365-2311.1904.tb02746.x" title="A revision of the sub-family Pelidnotinae of the coleopterous family Rutelidae, with descriptions of new genera and species." url="https://doi.org/10.1111/j.1365-2311.1904.tb02746.x" volume="1904" year="1904">
<normalizedToken id="5FCB401766A8C4D890E80F7AA32F18EF" originalValue="Bates’s">Bates's</normalizedToken>
(1904)
</bibRefCitation>
description of
<taxonomicName id="7DCE7C97239E80F53B444D8F06DA3E6D" authorityName="Bates" authorityYear="1904" class="Insecta" family="Scarabaeidae" genus="Hoplopelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopelidnota candezei" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="05AE476293762138D926B276AEDB5AFA" italics="true" pageId="0" pageNumber="1">Hoplopelidnota candezei</emphasis>
</taxonomicName>
overlooked the conspecific
<emphasis id="D808DF19985B99475DAF7B3D6D185F2B" italics="true" pageId="0" pageNumber="1">
<taxonomicName id="39466398EB1013B450EC58272BB98290" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. metallica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="metallica">P. metallica</taxonomicName>
.
<taxonomicName id="A20B651423DF27924A32E5AC3E33D534" authorityName="Bates" authorityYear="1904" class="Insecta" family="Scarabaeidae" genus="Hoplopelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopelidnota candezei" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="candezei">Hoplopelidnota candezei</taxonomicName>
</emphasis>
was based on a single male specimen that was labeled &quot;
<taxonomicName id="7A9150721272ED7A53FFD258696A38E2" authorityName="MacLeay" authorityYear="1819" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Pelidnota</taxonomicName>
<taxonomicName id="6ED71D9B0CC4D726F6E09D469CEDEA6A" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="armata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="armata">armata</taxonomicName>
&quot; by
<normalizedToken id="82CFC7564CC283D27C167C7A599EDDD5" originalValue="Candèze">Candeze</normalizedToken>
. The name &quot;
<taxonomicName id="BC7CE5BB54E53FBB2181B0BA6DB6F50E" family="Scarabaeidae" kingdom="Animalia" lsidName="H. armata" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="armata">H. armata</taxonomicName>
&quot;, however, had not been validly described and therefore was not available. However,
<bibRefCitation id="ABF13D458FEEF5BAA5CE75A3BABAE0D8" author="Ohaus, F" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="0" pageNumber="1" pagination="273 - 319" refId="B273" refString="Ohaus, F, 1912. Beitraege zur Kenntnis der Ruteliden, X. Stettiner Entomologische Zeitung 73: 273 - 319" title="Beitraege zur Kenntnis der Ruteliden, X." volume="73" year="1912">Ohaus (1912)</bibRefCitation>
provided a description of the female, he used the name &quot;
<taxonomicName id="7E225504BCF848A5F9DA63BEA7C63DED" authorityName="Ohaus" authorityYear="1912" class="Insecta" family="Scarabaeidae" genus="Hoplopelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopelidnota armata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="armata">Hoplopelidnota armata</taxonomicName>
&quot; (rather than
<taxonomicName id="51BB5C76897B1D43DC14029BDA753670" family="Scarabaeidae" kingdom="Animalia" lsidName="H. candezei" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="candezei">
<emphasis id="C67D2B4FABB8D3B16B857442C3FE69B0" italics="true" pageId="0" pageNumber="1">H. candezei</emphasis>
</taxonomicName>
). This act made the name
<taxonomicName id="BB0047CA9E56F7C8D81A07940EAC86BE" family="Scarabaeidae" kingdom="Animalia" lsidName="H. armata" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="armata">
<emphasis id="9E369419B33EB0E9D9098F76981FCD73" italics="true" pageId="0" pageNumber="1">H. armata</emphasis>
</taxonomicName>
an available name and a junior synonym of
<taxonomicName id="11A1C6D0D9469A2C664CC51F32490E72" family="Scarabaeidae" kingdom="Animalia" lsidName="H. metallica" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="metallica">
<emphasis id="E073A989ED89AEF5186979867DAC8293" italics="true" pageId="0" pageNumber="1">H. metallica</emphasis>
</taxonomicName>
. Most recently,
<bibRefCitation id="86FE100FEDF3BDE74EFE12D530B57DE7" author="Krajcik, M" journalOrPublisher="X Supplement 5, Plzen" pageId="0" pageNumber="1" refId="B156" refString="Krajcik, M, 2012. Checklist of the world Scarabaeoidea. Animma. X Supplement 5, Plzen" title="Checklist of the world Scarabaeoidea. Animma." year="2012">Krajcik (2012</bibRefCitation>
,
<bibRefCitation id="84B0D0FBEFA505C5F012B810CFFE3BD2" author="Krajcik, M" journalOrPublisher="49 - 54, Plzen" pageId="0" pageNumber="1" refId="B157" refString="Krajcik, M, 2013. Addenda to the checklist of world Scarabaeoidea 2012. Animma.X. No. 49-54, Plzen" title="Addenda to the checklist of world Scarabaeoidea 2012. Animma. X. No." year="2013">2013</bibRefCitation>
) considered
<taxonomicName id="4E99CFA7E5C8476F665E330ACF4F181A" family="Scarabaeidae" kingdom="Animalia" lsidName="H. candezei" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="candezei">
<emphasis id="B05E6F847C4E3E686B891F9D10ADD0D5" italics="true" pageId="0" pageNumber="1">H. candezei</emphasis>
</taxonomicName>
to be a subspecies of
<taxonomicName id="63F4BB3E57EB86C45E7BC52605D55B04" family="Scarabaeidae" kingdom="Animalia" lsidName="H. metallica" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="metallica">
<emphasis id="A86D2C48BEA7170612368E75512A4B2E" italics="true" pageId="0" pageNumber="1">H. metallica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="47F48983901EF0912EDD020A5EB9FF64" pageId="0" pageNumber="1">
<bibRefCitation id="DBB36C0C751ADD3F220F48F8171FCE3D" author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B359" refString="Soula, M, 2008. Les Coleopteres du Nouveau Monde. Volume 2: Rutelini 2. Revision des Pelidnotina 2. Revision des genres: Parhoplognathus, Chipita n. gen., Heteropelidnota, Homothermon, Hoplopelidnota, Mesomerodon, Mecopelidnota, Patatra n. gen. (Coleoptera: Scarabaeidae, Rutelinae, Rutelini). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 2: Rutelini 2. Revision des Pelidnotina 2. Revision des genres: Parhoplognathus, Chipita n. gen., Heteropelidnota, Homothermon, Hoplopelidnota, Mesomerodon, Mecopelidnota, Patatra n. gen. (Coleoptera: Scarabaeidae, Rutelinae, Rutelini). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2008">Soula (2008</bibRefCitation>
: 17-18) attempted to designate a neotype specimen for
<taxonomicName id="E29C25BF3BBA1F4A4037168DF28A8470" baseAuthorityName="Laporte" baseAuthorityYear="1840" class="Insecta" family="Scarabaeidae" genus="Hoplopelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopelidnota metallica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="6C20729A2FD81BAE9F8DA6EAE861D2EE" italics="true" pageId="0" pageNumber="1">Hoplopelidnota metallica</emphasis>
</taxonomicName>
. Soula stated that the neotype is in &quot;Collection Soula&quot;, but Article 75.3.7 (ICZN 1999) requires a statement that the &quot;neotype is, or immediately upon publication has become, the property of a recognized scientific or educational institution, cited by name, that maintains a research collection, with proper facilities for preserving name-bearing types, and that makes them accessible for study&quot;. Because
<normalizedToken id="44A5E88B68337F9ADBB2E280BB89B7A4" originalValue="Soula’s">Soula's</normalizedToken>
collection was private at the time of designation,
<normalizedToken id="0732698C57A24AF3128204760EBBA038" originalValue="Soula’s">Soula's</normalizedToken>
neotype is invalid.
</paragraph>
<paragraph id="AA840AA17760672F3D75ED3AB5712486" pageId="0" pageNumber="1">
<taxonomicName id="DC6227240BFB1D77C07B7F2007F47A72" baseAuthorityName="Laporte" baseAuthorityYear="1840" class="Insecta" family="Scarabaeidae" genus="Hoplopelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopelidnota metallica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="22C6939DF6D362760EC6A27EBFFFC17F" italics="true" pageId="0" pageNumber="1">Hoplopelidnota metallica</emphasis>
</taxonomicName>
is distributed in northern South America. Prior to this work,
<taxonomicName id="A2786AFDC6E095944C399D219ECBD634" family="Scarabaeidae" kingdom="Animalia" lsidName="H. metallica" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="metallica">
<emphasis id="88593E4C819F84151943F93EE756328A" italics="true" pageId="0" pageNumber="1">H. metallica</emphasis>
</taxonomicName>
was only recorded from French Guiana. In addition to French Guiana, we record the species from Guyana (Moraballi Creek, Essequibo River), Venezuela (Amazonas Dept., Rio Negro) and Brazil (Territorio de Amapa, Serra Navia). The species is rare in collections, and is apparently much more wide spread in northern South America than previous data would indicate. Specimens are recorded from 140 m elevation in March, April, July, August, and November.
</paragraph>
<caption id="FCC6DAC69CE480D68824A2B6904E1597" ID-Zenodo-Dep="982369" doi="10.3897/zookeys.666.9191.figure36" httpUri="https://binary.pensoft.net/fig/130464" pageId="0" pageNumber="1" start="Figure 36" startId="F36">
<paragraph id="CF284D64C7FDA9535B35AB492DC1DD9C" pageId="0" pageNumber="1">
<emphasis id="9D59C7F276C09F7CC43EA20F967CFDC2" bold="true" pageId="0" pageNumber="1">Figure 36.</emphasis>
<taxonomicName id="3E8CD4E699B187F0D929DE9FB878C603" baseAuthorityName="Laporte" baseAuthorityYear="1840" class="Insecta" family="Scarabaeidae" genus="Hoplopelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopelidnota metallica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="C1400E694908EC940AFCD85BD1680D50" italics="true" pageId="0" pageNumber="1">Hoplopelidnota metallica</emphasis>
</taxonomicName>
(Laporte) invalid neotype male from CCECL.
<emphasis id="2E714CBB0EC43908A839A82BD33A0C04" bold="true" pageId="0" pageNumber="1">A</emphasis>
Dorsal habitus
<emphasis id="B4D320C4D02CC5DBB14288EE8C936B85" bold="true" pageId="0" pageNumber="1">B</emphasis>
Male parameres, caudal view
<emphasis id="39A91362BFB938F3318D3F72BBC01E45" bold="true" pageId="0" pageNumber="1">C</emphasis>
Specimen labels.
</paragraph>
</caption>
<caption id="7245ACC258DFE6701CAE85655FAFA556" ID-Zenodo-Dep="982371" doi="10.3897/zookeys.666.9191.figure37" httpUri="https://binary.pensoft.net/fig/130465" pageId="0" pageNumber="1" start="Figure 37" startId="F37">
<paragraph id="915EDC74E98EA318CD38753E71A4B700" pageId="0" pageNumber="1">
<emphasis id="53B38D6E7D492DCADCCB06BCA7F56E9F" bold="true" pageId="0" pageNumber="1">Figure 37.</emphasis>
<taxonomicName id="7EE74B9E342FF6B9F1CE2C3A79F02EB1" baseAuthorityName="Laporte" baseAuthorityYear="1840" class="Insecta" family="Scarabaeidae" genus="Hoplopelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplopelidnota metallica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="25B25642EF0BE4A4F85A0BD08BEEFF37" italics="true" pageId="0" pageNumber="1">Hoplopelidnota metallica</emphasis>
</taxonomicName>
(Laporte) male from MSPC.
<emphasis id="0B40A5C3B18391DCB697CC05428D9B0D" bold="true" pageId="0" pageNumber="1">A</emphasis>
Dorsal habitus
<emphasis id="F1EEEA5478F53F26DA38E25E3C3938D8" bold="true" pageId="0" pageNumber="1">B</emphasis>
Lateral habitus
<emphasis id="0B52044CE0F3CC0C6B7F9B39B206ABE7" bold="true" pageId="0" pageNumber="1">C</emphasis>
Male genitalia, lateral view
<emphasis id="54D17B97F47E6D12BB9107234C368F79" bold="true" pageId="0" pageNumber="1">D</emphasis>
Male parameres, caudal view.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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