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<document id="5A0C8E5F3387B66E6B30C8A0BF0DB30A" ID-DOI="10.11646/zootaxa.4813.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3944139" ID-ZooBank="0ED34D69-0BC1-4E7D-A50D-6C0A31AB0374" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594719214043" checkinUser="plazi" docAuthor="Jordal, Bjarte H. &amp; Smith, Sarah M." docDate="2020" docId="9E3D1C0AFB38FFBBFF62F9B81A90FB99" docLanguage="en" docName="zootaxa.4813.1.1.pdf" docOrigin="Zootaxa 4813 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Scolytodes validus Jordal and Smith 2020, sp. nov." docType="treatment" docUuid="B10647EC-104A-4299-8ACF-D66EA8F7005B" docUuidSource="ZooBank" docVersion="5" lastPageNumber="56" masterDocId="62046472FB0EFF8CFFF5FFEA1918FF8A" masterDocTitle="Scolytodes Ferrari (Coleoptera, Scolytinae) from Ecuador: 40 new species, and a molecular phylogenetic guide to infer species differences" masterLastPageNumber="67" masterPageNumber="1" pageNumber="55" updateTime="1698781377542" updateUser="plazi">
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<mods:title id="135AA676D08E2EB3789174F2DE89B0BE">Scolytodes Ferrari (Coleoptera, Scolytinae) from Ecuador: 40 new species, and a molecular phylogenetic guide to infer species differences</mods:title>
</mods:titleInfo>
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<mods:namePart id="9F8A16230DC89E9711EA7CEAE75E759F">Jordal, Bjarte H.</mods:namePart>
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<mods:namePart id="20D87FB01014FDA52FACC4705E5FCB8E">Smith, Sarah M.</mods:namePart>
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<mods:title id="326B3F296D2697BDFA92C36512F35AB5">Zootaxa</mods:title>
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<mods:part id="4DCA1B908E98C53E765020911833C01B">
<mods:date id="323C0E561CE087132F4D4DC1A0E029B1">2020</mods:date>
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<mods:number id="9176C6728B36847EAC5E50D38F64F509">2020-07-14</mods:number>
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<mods:classification id="608820923EA86A620828193D4C2007FA">journal article</mods:classification>
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<treatment id="9E3D1C0AFB38FFBBFF62F9B81A90FB99" ID-DOI="http://doi.org/10.5281/zenodo.4338696" ID-Zenodo-Dep="4338696" LSID="urn:lsid:zoobank.org:act:B10647EC-104A-4299-8ACF-D66EA8F7005B" httpUri="http://treatment.plazi.org/id/9E3D1C0AFB38FFBBFF62F9B81A90FB99" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
<subSubSection id="5E8EFE97FB38FFBAFF62F9B81AD1F91A" pageId="54" pageNumber="55" type="nomenclature">
<paragraph id="162BAD1CFB38FFBAFF62F9B81BDAF9E7" blockId="54.[151,969,1618,1716]" box="[151,706,1618,1645]" pageId="54" pageNumber="55">
<heading id="4D631A70FB38FFBAFF62F9B81BDAF9E7" bold="true" box="[151,706,1618,1645]" fontSize="11" level="1" pageId="54" pageNumber="55" reason="1">
<emphasis id="24E0710EFB38FFBAFF62F9B81BDAF9E7" bold="true" box="[151,706,1618,1645]" pageId="54" pageNumber="55">
<taxonomicName id="D194D69FFB38FFBAFF62F9B81B40F9E6" authority="Jordal and Smith" authorityName="Jordal and Smith" authorityYear="2020" box="[151,600,1618,1645]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="validus" status="sp. nov.">
<emphasis id="24E0710EFB38FFBAFF62F9B8186CF9E7" bold="true" box="[151,372,1618,1645]" italics="true" pageId="54" pageNumber="55">Scolytodes validus</emphasis>
Jordal and Smith
</taxonomicName>
,
<taxonomicNameLabel id="3FD3CC75FB38FFBAFD91F9B91BDAF9E7" box="[612,706,1619,1645]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
<uuid id="623297C9FB38FFBAFE52F99F1AD1F91A" box="[423,969,1653,1680]" pageId="54" pageNumber="55">B10647EC-104A-4299-8ACF-D66EA8F7005B</uuid>
</uri>
</paragraph>
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<subSubSection id="5E8EFE97FB38FFBAFF62F9701862F93E" box="[151,378,1689,1716]" pageId="54" pageNumber="55" type="description">
<paragraph id="162BAD1CFB38FFBAFF62F9701862F93E" blockId="54.[151,969,1618,1716]" box="[151,378,1689,1716]" pageId="54" pageNumber="55">
(
<figureCitation id="8EAFB199FB38FFBAFF6AF970186AF93E" box="[159,370,1689,1716]" captionStart="FIGURES 118126" captionStartId="56.[151,264,1934,1959]" captionTargetBox="[162,1428,198,1909]" captionTargetId="figure@56.[152,1436,181,1909]" captionTargetPageId="56" captionText="FIGURES 118126. Dorsal, lateral and frontal views of the female holotype of Scolytodes arcuatus (118, 121, 124), Scolytodes validus (119, 122, 125), and Scolytodes sparsus (120, 123, 126)." figureDoi="http://doi.org/10.5281/zenodo.3944167" httpUri="https://zenodo.org/record/3944167/files/figure.png" pageId="54" pageNumber="55">Figs 119, 122, 125</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB38FFBAFF62F9081DF3F807" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="162BAD1CFB38FFBAFF62F9081DF3F807" blockId="54.[151,1437,1761,2041]" pageId="54" pageNumber="55">
<emphasis id="24E0710EFB38FFBAFF62F908185DF976" bold="true" box="[151,325,1761,1788]" pageId="54" pageNumber="55">Type material.</emphasis>
<materialsCitation id="A6FCA741FB38FFBAFEBEF9081A2BF8AA" collectingDate="1998-10" collectorName="T. L. Erwin" country="Ecuador" elevation="235" latitude="-0.6319445" location="Tiputini" longLatPrecision="21" longitude="-76.144165" pageId="54" pageNumber="55" specimenCount="1" typeStatus="holotype">
<emphasis id="24E0710EFB38FFBAFEBEF9081B06F976" bold="true" box="[331,542,1761,1788]" pageId="54" pageNumber="55">
<typeStatus id="C92F13BEFB38FFBAFEBEF90818A2F976" box="[331,442,1762,1788]" pageId="54" pageNumber="55" type="holotype">Holotype</typeStatus>
, female:
</emphasis>
<collectingCountry id="6E83ED8CFB38FFBAFDD0F9081B9FF976" box="[549,647,1762,1788]" name="Ecuador" pageId="54" pageNumber="55">Ecuador</collectingCountry>
:
<location id="134BFBC7FB38FFBAFD65F9081BF1F976" LSID="urn:lsid:plazi:treatment:9E3D1C0AFB38FFBBFF62F9B81A90FB99:134BFBC7FB38FFBAFD65F9081BF1F976" box="[656,745,1762,1788]" country="Ecuador" latitude="-0.6319445" longLatPrecision="21" longitude="-76.144165" name="Tiputini" pageId="54" pageNumber="55">Tiputini</location>
biodiversity station,
<quantity id="D16C00F9FB38FFBAFC21F90B1D4CF977" box="[980,1108,1761,1789]" metricMagnitude="2" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="54" pageNumber="55" unit="m" value="235.0" valueMax="250.0" valueMin="220.0">
<elevation id="9DB94A2FFB38FFBAFC21F90B1D4CF977" box="[980,1108,1761,1789]" metricMagnitude="2" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="54" pageNumber="55" unit="m" value="235.0" valueMax="250.0" valueMin="220.0">220250 m</elevation>
</quantity>
,
<date id="622A8BDCFB38FFBAFB95F9081DE3F976" box="[1120,1275,1762,1788]" pageId="54" pageNumber="55" value="1998-10">
<collectingDate id="726E7234FB38FFBAFB95F9081DE3F976" box="[1120,1275,1762,1788]" pageId="54" pageNumber="55" value="1998-10">October 1998</collectingDate>
</date>
,
<geoCoordinate id="73A0CBDBFB38FFBAFAF3F9081C8FF976" box="[1286,1431,1761,1788]" degrees="00" direction="south" minutes="37" orientation="latitude" pageId="54" pageNumber="55" precision="15" seconds="55" value="-0.6319445">00° 3755S</geoCoordinate>
,
<geoCoordinate id="73A0CBDBFB38FFBAFF62F8EC182EF8AA" box="[151,310,1798,1824]" degrees="76" direction="west" minutes="08" orientation="longitude" pageId="54" pageNumber="55" precision="15" seconds="39" value="-76.144165">76° 0839W</geoCoordinate>
,
<collectorName id="BB61C8CAFB38FFBAFEB0F8ED18DAF8AA" box="[325,450,1798,1825]" pageId="54" pageNumber="55">T.L. Erwin</collectorName>
et al collectors, indv. #000448
</materialsCitation>
.
<emphasis id="24E0710EFB38FFBAFCB4F8EF1AE3F8AA" bold="true" box="[833,1019,1797,1824]" pageId="54" pageNumber="55">
<typeStatus id="C92F13BEFB38FFBAFCB4F8EF1AB0F8AA" box="[833,936,1797,1824]" pageId="54" pageNumber="55" type="allotype">Allotype</typeStatus>
, male:
</emphasis>
same data as holotype.
<materialsCitation id="A6FCA741FB38FFBAFAEEF8EC1824F806" collectingDate="1995-10" collectionCode="USNM" collectorName="T. L. Erwin &amp; The" country="Ecuador" elevation="220" latitude="-0.65277773" location="Trans. Et." longLatPrecision="925" longitude="-76.433334" municipality="Res. Ethnica Waorani" pageId="54" pageNumber="55" specimenCount="3" stateProvince="Napo" typeStatus="paratype">
<emphasis id="24E0710EFB38FFBAFAEEF8EC1C84F8AA" bold="true" box="[1307,1436,1798,1824]" pageId="54" pageNumber="55">
<typeStatus id="C92F13BEFB38FFBAFAEEF8EC1C8FF8AA" box="[1307,1431,1798,1824]" pageId="54" pageNumber="55" type="paratype">Paratypes</typeStatus>
:
</emphasis>
same data as holotype (3);
<collectingRegion id="D45063FEFB38FFBAFE31F8C11B5CF8CF" box="[452,580,1835,1861]" country="Ecuador" name="Napo" pageId="54" pageNumber="55">Napo Prov.</collectingRegion>
,
<collectingMunicipality id="F64F3766FB38FFBAFDBAF8C11A5CF8CE" box="[591,836,1834,1861]" pageId="54" pageNumber="55">Res. Ethnica Waorani</collectingMunicipality>
,
<locationDeviation id="0799B43EFB38FFBAFCBAF8C01D90F8CE" box="[847,1160,1834,1860]" location="Onkone Gare Camp." pageId="54" pageNumber="55">
<quantity id="D16C00F9FB38FFBAFCBAF8C01A99F8CE" box="[847,897,1834,1860]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="54" pageNumber="55" unit="km" value="1.0">1km</quantity>
S Onkone Gare Camp.
</locationDeviation>
,
<location id="134BFBC7FB38FFBAFB67F8C11C19F8CF" LSID="urn:lsid:plazi:treatment:9E3D1C0AFB38FFBBFF62F9B81A90FB99:134BFBC7FB38FFBAFB67F8C11C19F8CF" box="[1170,1281,1835,1861]" latitude="-0.65277773" longLatPrecision="925" longitude="-76.433334" municipality="Res. Ethnica Waorani" name="Trans. Et." pageId="54" pageNumber="55" stateProvince="Napo">Trans. Et.</location>
,
<geoCoordinate id="73A0CBDBFB38FFBAFAF8F8C01C80F8CE" box="[1293,1432,1834,1860]" degrees="00" direction="south" minutes="39" orientation="latitude" pageId="54" pageNumber="55" precision="15" seconds="10" value="-0.65277773">00°3910S</geoCoordinate>
,
<geoCoordinate id="73A0CBDBFB38FFBAFF62F8A419E7F8E2" box="[151,255,1870,1896]" degrees="76" direction="west" minutes="26" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="-76.433334">76°26W</geoCoordinate>
,
<quantity id="D16C00F9FB38FFBAFEFFF8A41851F8E2" box="[266,329,1870,1896]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="54" pageNumber="55" unit="m" value="220.0">
<elevation id="9DB94A2FFB38FFBAFEFFF8A41851F8E2" box="[266,329,1870,1896]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="54" pageNumber="55" unit="m" value="220.0">220m</elevation>
</quantity>
elev.,
<date id="622A8BDCFB38FFBAFE67F8A41B36F8E2" box="[402,558,1869,1896]" pageId="54" pageNumber="55" value="1995-10">
<collectingDate id="726E7234FB38FFBAFE67F8A41B36F8E2" box="[402,558,1869,1896]" pageId="54" pageNumber="55" value="1995-10">October 1995</collectingDate>
</date>
,
<collectorName id="BB61C8CAFB38FFBAFDCCF8A51BAAF8E2" box="[569,690,1870,1897]" pageId="54" pageNumber="55">T.L. Erwin</collectorName>
<emphasis id="24E0710EFB38FFBAFD4CF8A51BE9F8E2" box="[697,753,1870,1896]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
collectors (2).
<collectorName id="BB61C8CAFB38FFBAFC6FF8A41ADEF8E2" box="[922,966,1870,1896]" pageId="54" pageNumber="55">The</collectorName>
<typeStatus id="C92F13BEFB38FFBAFC39F8A41D36F8E2" box="[972,1070,1870,1896]" pageId="54" pageNumber="55" type="holotype">holotype</typeStatus>
and
<typeStatus id="C92F13BEFB38FFBAFB90F8A41DA7F8E2" box="[1125,1215,1870,1896]" pageId="54" pageNumber="55" type="allotype">allotype</typeStatus>
temporarily held in trust at
<collectionCode id="708535D9FB38FFBAFF1EF8981824F806" box="[235,316,1906,1932]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="54" pageNumber="55" type="Museum">USNM</collectionCode>
</materialsCitation>
for
<collectionCode id="708535D9FB38FFBAFE9EF89818A7F806" box="[363,447,1906,1932]" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" pageId="54" pageNumber="55">MECN</collectionCode>
,
<materialsCitation id="A6FCA741FB38FFBAFE3FF8991D13F806" box="[458,1035,1906,1933]" collectionCode="QCAZ, MSUC" country="Ecuador" pageId="54" pageNumber="55" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
two
<typeStatus id="C92F13BEFB38FFBAFE0EF8991B7EF807" box="[507,614,1907,1933]" pageId="54" pageNumber="55" type="paratype">paratypes</typeStatus>
each deposited in
<collectionCode id="708535D9FB38FFBAFCC3F8981A9AF806" box="[822,898,1906,1932]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="54" pageNumber="55" type="Museum">QCAZ</collectionCode>
and
<collectionCode id="708535D9FB38FFBAFC4CF8981D13F806" box="[953,1035,1906,1932]" pageId="54" pageNumber="55">MSUC</collectionCode>
</materialsCitation>
, and one in
<collectionCode id="708535D9FB38FFBAFB67F8991DFFF807" box="[1170,1255,1907,1933]" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="54" pageNumber="55">ZMBN</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB38FFBBFF32F87F1866FF5D" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55" type="diagnosis">
<paragraph id="162BAD1CFB38FFBBFF32F87F1866FF5D" blockId="54.[151,1437,1761,2041]" lastBlockId="55.[151,1437,153,1043]" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
<emphasis id="24E0710EFB38FFBAFF32F87F185AF83A" bold="true" box="[199,322,1941,1968]" pageId="54" pageNumber="55">Diagnosis.</emphasis>
Interstriae 10 sharply carinate to near apex. Protibiae without additional mesal tooth, tooth 1 larger than tooth 2. Lateral carinae on pronotum anteriorly turned medially. A pair of broad, blunt carinae continue dorsad from epistoma to middle of frons. Related to
<taxonomicName id="D194D69FFB38FFBAFD64F8351A00F872" authorityName="Wood" authorityYear="1974" box="[657,792,2014,2041]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="costabilis">
<emphasis id="24E0710EFB38FFBAFD64F8351A00F872" box="[657,792,2014,2041]" italics="true" pageId="54" pageNumber="55">S. costabilis</emphasis>
</taxonomicName>
and
<taxonomicName id="D194D69FFB38FFBAFCBAF8351AD4F872" authorityName="Jordal and Smith" authorityYear="2020" box="[847,972,2015,2041]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="24E0710EFB38FFBAFCBAF8351AD4F872" box="[847,972,2015,2041]" italics="true" pageId="54" pageNumber="55">S. arcuatus</emphasis>
</taxonomicName>
but distinguished from both by the broad carinae in the frons, and further from
<taxonomicName id="D194D69FFB39FFBBFDC3FF731BA4FF39" authorityName="Wood" authorityYear="1974" box="[566,700,153,179]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="costabilis">
<emphasis id="24E0710EFB39FFBBFDC3FF731BA4FF39" box="[566,700,153,179]" italics="true" pageId="55" pageNumber="56">S. costabilis</emphasis>
</taxonomicName>
by the much stouter body shape, and from
<taxonomicName id="D194D69FFB39FFBBFB6DFF731C0CFF39" authorityName="Jordal and Smith" authorityYear="2020" box="[1176,1300,153,179]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="24E0710EFB39FFBBFB6DFF731C0CFF39" box="[1176,1300,153,179]" italics="true" pageId="55" pageNumber="56">S. arcuatus</emphasis>
</taxonomicName>
by the presence of elytral setae.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB39FFBBFF32FF0A1BB9FCD5" pageId="55" pageNumber="56" type="description">
<paragraph id="162BAD1CFB39FFBBFF32FF0A19CFFC9D" blockId="55.[151,1437,153,1043]" pageId="55" pageNumber="56">
<emphasis id="24E0710EFB39FFBBFF32FF0A18A2FF71" bold="true" box="[199,442,224,251]" pageId="55" pageNumber="56">Description, female.</emphasis>
Length
<quantity id="D16C00F9FB39FFBBFDEAFF0B1BB3FF71" box="[543,683,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="55" pageNumber="56" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.22.4 mm</quantity>
, 2.0 × as long as wide; colour dark brown.
<emphasis id="24E0710EFB39FFBBFB40FF0B1DE4FF71" bold="true" box="[1205,1276,225,251]" pageId="55" pageNumber="56">Head.</emphasis>
Eyes slightly emarginate, separated above by 3.33.5 × their width. Frons flattened and impressed between eyes from vertex to epistoma, a pair of broad, shiny carinae running dorsad from epistoma to the middle of frons, smooth and micropunctate between; vestiture consisting of long setae along outer margin of impressed area, from vertex to level of antennal insertion, setae curved towards the centre of frons. Antennal club pilose, sutures obscure. Funiculus 6-segmented.
<emphasis id="24E0710EFB39FFBBFEC1FE7F18B5FE25" bold="true" box="[308,429,405,431]" pageId="55" pageNumber="56">Pronotum</emphasis>
reticulate, dull, punctures spaced by 1 × their diameter, reaching anterior margin; anterior half with low asperities; lateral carinae anteriorly turned medially before interrupted. Vestiture consisting of 4 long, erect setae (2-0-2).
<emphasis id="24E0710EFB39FFBBFE98FE3718A1FE7D" bold="true" box="[365,441,477,503]" pageId="55" pageNumber="56">Elytra</emphasis>
shiny, smooth; striae more or less regular, not impressed, punctures large, deep, separated by 12 × their diameter; interstriae 56 × broader than striae, punctures almost as large as strial punctures, entirely confused. Interstriae 10 sharply raised to near apex. Vestiture consisting of 1530 erect setae of variable length, on odd-numbered interstriae, the majority on 7 and 9.
<emphasis id="24E0710EFB39FFBBFD17FDA31A39FDE9" bold="true" box="[738,801,585,611]" pageId="55" pageNumber="56">Legs.</emphasis>
Procoxae separated by 0.50.7 × the width of one procoxa. Mesocoxae separated by 0.70.8 × the width of a procoxa. Protibiae narrow, distal teeth 1 larger than 2, with 34 additional small spines or transverse rugae towards tibial base; protibial mucro short, blunt, curved laterally. Meso- and metatibiae with 78 socketed teeth on distal half.
<emphasis id="24E0710EFB39FFBBFCA1FD5F1D06FD45" bold="true" box="[852,1054,692,719]" pageId="55" pageNumber="56">Ventral vestiture</emphasis>
. Setae on mesanepisternum trifid or plumose, on metanepisternum simple, on metasternum mixed plumose, bi- or trifid setae, with scattered simple setae.
</paragraph>
<paragraph id="162BAD1CFB39FFBBFF32FCCB1BB9FCD5" blockId="55.[151,1437,153,1043]" pageId="55" pageNumber="56">
<emphasis id="24E0710EFB39FFBBFF32FCCB1813FCB1" bold="true" box="[199,267,801,827]" pageId="55" pageNumber="56">Male.</emphasis>
Similar to female, except frons convex, reticulated, with clusters of two or three contiguous punctures; eyes separated above by 2.93.0 × their width.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB39FFBBFF32FC831DB7FC09" box="[199,1199,872,899]" pageId="55" pageNumber="56" type="key">
<paragraph id="162BAD1CFB39FFBBFF32FC831DB7FC09" blockId="55.[151,1437,153,1043]" box="[199,1199,872,899]" pageId="55" pageNumber="56">
<emphasis id="24E0710EFB39FFBBFF32FC831885FC09" bold="true" box="[199,413,872,899]" pageId="55" pageNumber="56">
Key (
<bibRefCitation id="7205D0EDFB39FFBBFEF3FC821888FC09" author="Wood, S. L." box="[262,400,872,899]" pageId="55" pageNumber="56" refId="ref46297" refString="Wood, S. L. (2007) Bark and ambrosia beetles of South America (Coleoptera, Scolytidae). Brigham Young University Press, Provo, Utah, 900 pp." type="book" year="2007">Wood 2007</bibRefCitation>
).
</emphasis>
Keys to couplet 19,
<taxonomicName id="D194D69FFB39FFBBFD70FC831A14FC09" ID-CoL="4VT3V" authorityName="Wood" authorityYear="1974" box="[645,780,873,899]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="costabilis">
<emphasis id="24E0710EFB39FFBBFD70FC831A14FC09" box="[645,780,873,899]" italics="true" pageId="55" pageNumber="56">S. costabilis</emphasis>
</taxonomicName>
, but differs as noted in the diagnosis.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB39FFBBFF32FC6719C7FC41" pageId="55" pageNumber="56" type="etymology">
<paragraph id="162BAD1CFB39FFBBFF32FC6719C7FC41" blockId="55.[151,1437,153,1043]" pageId="55" pageNumber="56">
<emphasis id="24E0710EFB39FFBBFF32FC671856FC2D" bold="true" box="[199,334,909,935]" pageId="55" pageNumber="56">Etymology.</emphasis>
The Latin name
<taxonomicName id="D194D69FFB39FFBBFDFFFC671B43FC2D" authorityName="Jordal and Smith" authorityYear="2020" box="[522,603,909,935]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="validus">
<emphasis id="24E0710EFB39FFBBFDFFFC671B43FC2D" box="[522,603,909,935]" italics="true" pageId="55" pageNumber="56">validus</emphasis>
</taxonomicName>
is a masculine adjective, meaning strong or mighty, referring to stout body shape.
</paragraph>
</subSubSection>
<subSubSection id="5E8EFE97FB39FFBBFF32FC3E1A90FB99" pageId="55" pageNumber="56" type="biology_ecology">
<paragraph id="162BAD1CFB39FFBBFF32FC3E1A90FB99" blockId="55.[151,1437,153,1043]" pageId="55" pageNumber="56">
<emphasis id="24E0710EFB39FFBBFF32FC3E18E8FC65" bold="true" box="[199,496,980,1007]" pageId="55" pageNumber="56">Biology and distribution.</emphasis>
Only known from two nearby localities in the Ecuadorian Amazon. Specimens were collected by canopy fogging in the month of October (1995, 1998).
</paragraph>
</subSubSection>
</treatment>
</document>