treatments-xml/data/9D/64/C2/9D64C25EF658EC339FD19A0A1BEAFCFA.xml

<document id="24E3C4767CB5667EC0269CFE110830C7" ID-CLB-Dataset="54383" ID-DOI="10.11646/zootaxa.5301.4.4" ID-GBIF-Dataset="c767a139-ee8d-49fb-924f-76050006336f" ID-ISSN="1175-5326" ID-Zenodo-Dep="8036043" ID-ZooBank="F5B58CAD-6420-4A08-90A6-F7EAF2D54826" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1686722943637" checkinUser="plazi" docAuthor="Hernáez, Patricio, Villegas-Castro, Erick &amp; Boyko, Christopher B." docDate="2023" docId="9D64C25EF658EC339FD19A0A1BEAFCFA" docLanguage="en" docName="zootaxa.5301.4.4.pdf" docOrigin="Zootaxa 5301 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5301.4.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ione sarahae Hernáez &amp; Villegas-Castro &amp; Boyko 2023, n. sp." docType="treatment" docVersion="4" lastPageNumber="466" masterDocId="615DBA26F659EC369F469E391F63FFEF" masterDocTitle="A new species of parasitic isopod (Bopyroidea: Ionidae) infesting burrowing shrimp (Axiidea: Callianassidae) in the tropical eastern Pacific" masterLastPageNumber="468" masterPageNumber="461" pageNumber="462" updateTime="1699523626710" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D8A59AB878B48E080BF11D6BBBE2A640">A new species of parasitic isopod (Bopyroidea: Ionidae) infesting burrowing shrimp (Axiidea: Callianassidae) in the tropical eastern Pacific</mods:title>
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<mods:namePart id="7FC747D5017ABF93F179C55E3693F005">Hernáez, Patricio</mods:namePart>
<mods:affiliation id="96D6813D2856A197EC88698F70F19C3E">Universidad de Tarapacá - UTA, Facultad de Ciencias, Departamento de Biología, Centro de Estudios Marinos y Limnológicos, Av. General Velásquez, 1775, Arica, Chile.</mods:affiliation>
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<mods:namePart id="25AA27D88E8BD2F3FD0DACE3FE98181C">Villegas-Castro, Erick</mods:namePart>
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<mods:affiliation id="60F01B2F58803A0AED3888F40742EE8F">Universidad de Costa Rica - UCR, Centro de Investigaciones en Nutrición Animal, San José, Costa Rica. erick. villegascastro @ ucr. ac. cr; https: // orcid. org / 0000 - 0001 - 8583 - 5450</mods:affiliation>
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<mods:namePart id="23E814C3C78614AB615B28FAB67F432B">Boyko, Christopher B.</mods:namePart>
<mods:affiliation id="C2617EA75392E9EDEBAAAA105C488BF3">Department of Biology, Hofstra University, Hempstead, NY 11549, U. S. A. and Division of Invertebrate Zoology, American Museum of Natural History, Central Park West @ 79 Street, New York, NY, U. S. A.</mods:affiliation>
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<heading id="4E3AC424F658EC379FD19A0A1EE3FBA2" bold="true" box="[151,384,1074,1101]" fontSize="11" level="1" pageId="1" pageNumber="462" reason="1">
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<taxonomicName id="D2CD08CBF658EC379FD19A0A1E55FBA2" ID-CoL="8a6a1a01-3be5-4e7e-9262-dc242468320e" authority="Hernáez &amp; Villegas-Castro &amp; Boyko, 2023" authorityName="Hernáez &amp; Villegas-Castro &amp; Boyko" authorityYear="2023" box="[151,310,1074,1101]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="462" phylum="Arthropoda" rank="species" species="sarahae" status="sp. nov.">
<emphasis id="27B9AF5AF658EC379FD19A0A1E55FBA2" bold="true" box="[151,310,1074,1101]" italics="true" pageId="1" pageNumber="462">Ione sarahae</emphasis>
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<taxonomicNameLabel id="3C8A1221F658EC379E7B9A0A1EE3FBA2" box="[317,384,1075,1101]" pageId="1" pageNumber="462" rank="species">n. sp.</taxonomicNameLabel>
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(
<figureCitation id="8DF66FCDF658EC379FD99A6E1E62FB9E" box="[159,257,1111,1137]" captionStart="FIGURE 1" captionStartId="2.[151,250,1153,1177]" captionTargetBox="[183,1404,349,1127]" captionTargetId="figure-102@2.[183,1404,345,1130]" captionTargetPageId="2" captionText="FIGURE 1. Neotrypaea tabogensis (Sakai, 2005) A, male and female hosts, dorsal view; B, Ione sarahae n. sp. female and male, dorsal view (MZUCR 3789-01). Scale bars: A = 5 mm, B = 2 mm (female), 250 µm (male)." figureDoi="http://doi.org/10.5281/zenodo.8036045" httpUri="https://zenodo.org/record/8036045/files/figure.png" pageId="1" pageNumber="462">Figs. 1B</figureCitation>
, 
<figureCitation id="8DF66FCDF658EC379E4B9A6E1E5AFB9E" box="[269,313,1111,1137]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="3.[151,250,1276,1300]" captionStartId-1="4.[151,250,1575,1599]" captionStartId-2="5.[151,250,1977,2001]" captionTargetBox-0="[192,1435,192,1251]" captionTargetBox-1="[178,1414,185,1549]" captionTargetBox-2="[169,1421,828,1951]" captionTargetId-0="figure-18@3.[151,1435,181,1253]" captionTargetId-1="figure-26@4.[170,1416,181,1551]" captionTargetId-2="figure-356@5.[164,1423,813,1954]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 2. Ione sarahae n. sp., holotype, female, 6.6 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view. Scale bars: A, B = 2 mm." captionText-1="FIGURE 3. Ione sarahae n. sp., paratype, female, 4.3 mm TL (MZUCR 3791-01). A, right antennula; B, right antenna; C, barbula; D, right maxilliped; E, right pereopod 1; F, right pereopod 7; G, right oostegite 1, external view; H, right oostegite 1, internal view. Scale bars: A, B, E, F = 100 µm; C = 500 µm; G, H = 1 mm; D = 50 µm." captionText-2="FIGURE 4. Ione sarahae n. sp., allotype, male, 3.5 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view; C, left antennula; D, left antenna; E, left pereopod 1. Scale bars: A, B = 250 µm; C, D = 50 µm; E = 200 µm." figureDoi-0="http://doi.org/10.5281/zenodo.8036047" figureDoi-1="http://doi.org/10.5281/zenodo.8036049" figureDoi-2="http://doi.org/10.5281/zenodo.8036051" httpUri-0="https://zenodo.org/record/8036047/files/figure.png" httpUri-1="https://zenodo.org/record/8036049/files/figure.png" httpUri-2="https://zenodo.org/record/8036051/files/figure.png" pageId="1" pageNumber="462">2–4</figureCitation>
)
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<emphasis id="27B9AF5AF658EC379FD19AA61E24FB57" bold="true" box="[151,327,1182,1209]" pageId="1" pageNumber="462">Type material:</emphasis>
<materialsCitation id="A5A57915F658EC379E0B9AA61C75FB56" ID-GBIF-Occurrence="4130680302" box="[333,790,1183,1209]" collectingDate="2012-03-16" collectionCode="MZUCR" collectorName="P. Hernaez" country="Costa Rica" location="Mata de Limon" municipality="Mata de Limon" pageId="1" pageNumber="462" specimenCode="MZUCR 3789-01" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="27B9AF5AF658EC379E0B9AA61ED7FB56" box="[333,436,1183,1209]" italics="true" pageId="1" pageNumber="462">
<typeStatus id="CA76CDEAF658EC379E0B9AA61ED7FB56" box="[333,436,1183,1209]" pageId="1" pageNumber="462" type="holotype">Holotype</typeStatus>
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, ovigerous 
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, 
<quantity id="D235DEADF658EC379DD79AA61D85FB56" box="[657,742,1183,1209]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="1" pageNumber="462" unit="mm" value="6.6">6.6 mm</quantity>
TL;
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<materialsCitation id="A5A57915F658EC379C5B9AA61C67FAEE" ID-GBIF-Occurrence="4130680301" collectingDate="2012-03-16" collectionCode="MZUCR" collectorName="P. Hernaez" country="Costa Rica" location="Mata de Limon" municipality="Mata de Limon" pageId="1" pageNumber="462" specimenCode="MZUCR 3789-01" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
allotype, 
<specimenCount id="03CBB8C1F658EC379CC29AA61CDEFB56" box="[900,957,1183,1209]" pageId="1" pageNumber="462" type="male">male</specimenCount>
, 
<quantity id="D235DEADF658EC379C8E9AA71B7DFB56" box="[968,1054,1182,1209]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="462" unit="mm" value="3.5">3.5 mm</quantity>
TL; from right branchial chamber of female 
<taxonomicName id="D2CD08CBF658EC379E4F9AFA1D68FB32" baseAuthorityName="Sakai" baseAuthorityYear="2005" box="[265,523,1219,1245]" class="Malacostraca" family="Callianassidae" genus="Neotrypaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="462" phylum="Arthropoda" rank="species" species="tabogensis">
<emphasis id="27B9AF5AF658EC379E4F9AFA1D68FB32" box="[265,523,1219,1245]" italics="true" pageId="1" pageNumber="462">Neotrypaea tabogensis</emphasis>
</taxonomicName>
(
<quantity id="D235DEADF658EC379D5F9AFB1D12FB32" box="[537,625,1218,1245]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="462" unit="mm" value="5.3">5.3 mm</quantity>
CL), 
<collectingMunicipality id="F516E932F658EC379DF59AFA1C00FB32" box="[691,867,1219,1245]" pageId="1" pageNumber="462">Mata de Limón</collectingMunicipality>
, Pacific coast of 
<collectingCountry id="6DDA33D8F658EC379B679AFA1BFDFB32" box="[1057,1182,1219,1245]" name="Costa Rica" pageId="1" pageNumber="462">Costa Rica</collectingCountry>
, in burrows, intertidal, coll. 
<collectorName id="B838169EF658EC379F929ADE1E31FAEE" box="[212,338,1255,1281]" pageId="1" pageNumber="462">P. Hernáez</collectorName>
, 
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<collectingDate id="7137AC60F658EC379E249ADE1D75FAEE" box="[354,534,1255,1281]" pageId="1" pageNumber="462" value="2012-03-16">16 March 2012</collectingDate>
</date>
(
<specimenCode id="456BDB33F658EC379D6F9ADE1D9FFAEE" box="[553,764,1255,1281]" pageId="1" pageNumber="462">MZUCR 3789-01</specimenCode>
)
</materialsCitation>
. 
<typeStatus id="CA76CDEAF658EC379C559ADE1CE7FAEE" box="[787,900,1255,1281]" pageId="1" pageNumber="462" type="paratype">Paratypes</typeStatus>
: female, 
<quantity id="D235DEADF658EC379CB49ADE1B2FFAEE" box="[1010,1100,1255,1281]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="462" unit="mm" value="2.8">2.8 mm</quantity>
TL, without male, from left branchial chamber of male of 
<taxonomicName id="D2CD08CBF658EC379EA19B321DE3FACA" baseAuthorityName="Sakai" baseAuthorityYear="2005" box="[487,640,1291,1317]" class="Malacostraca" family="Callianassidae" genus="Neotrypaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="462" phylum="Arthropoda" rank="species" species="tabogensis">
<emphasis id="27B9AF5AF658EC379EA19B321DE3FACA" box="[487,640,1291,1317]" italics="true" pageId="1" pageNumber="462">N. tabogensis</emphasis>
</taxonomicName>
(
<quantity id="D235DEADF658EC379DC99B331D85FACA" box="[655,742,1290,1317]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="1" pageNumber="462" unit="mm" value="5.1">5.1 mm</quantity>
CL), Mata de 
<collectingRegion id="D709BDAAF658EC379CCF9B321CB4FACA" box="[905,983,1291,1317]" country="Costa Rica" name="Limon" pageId="1" pageNumber="462">Limón</collectingRegion>
, Pacific coast of 
<collectingCountry id="6DDA33D8F658EC379BD39B321A72FACA" box="[1173,1297,1291,1317]" name="Costa Rica" pageId="1" pageNumber="462">Costa Rica</collectingCountry>
, in burrows, intertidal, coll. P. Hernáez, 
<date id="61735588F658EC379E819B161D1CFAA6" box="[455,639,1327,1353]" pageId="1" pageNumber="462" value="2013-01-21">21 January 2013</date>
(
<collectionCode id="73DCEB8DF658EC379DCB9B161D90FAA6" box="[653,755,1327,1353]" pageId="1" pageNumber="462">MZUCR</collectionCode>
3790-01); female, 
<quantity id="D235DEADF658EC379C8F9B161B7DFAA6" box="[969,1054,1327,1353]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="1" pageNumber="462" unit="mm" value="4.3">4.3 mm</quantity>
TL, from right branchial chamber of male of 
<taxonomicName id="D2CD08CBF658EC379E529B6A1ECEFA82" baseAuthorityName="Sakai" baseAuthorityYear="2005" box="[276,429,1363,1389]" class="Malacostraca" family="Callianassidae" genus="Neotrypaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="462" phylum="Arthropoda" rank="species" species="tabogensis">
<emphasis id="27B9AF5AF658EC379E529B6A1ECEFA82" box="[276,429,1363,1389]" italics="true" pageId="1" pageNumber="462">N. tabogensis</emphasis>
</taxonomicName>
(
<quantity id="D235DEADF658EC379EFA9B6A1D76FA82" box="[444,533,1363,1389]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="1" pageNumber="462" unit="mm" value="4.4">4.4 mm</quantity>
CL), Mata de 
<collectingRegion id="D709BDAAF658EC379DFD9B6A1C6AFA82" box="[699,777,1363,1389]" country="Costa Rica" name="Limon" pageId="1" pageNumber="462">Limón</collectingRegion>
, Pacific coast of 
<collectingCountry id="6DDA33D8F658EC379C8D9B6A1B2BFA82" box="[971,1096,1363,1389]" name="Costa Rica" pageId="1" pageNumber="462">Costa Rica</collectingCountry>
, in burrows, intertidal, coll. P. Hernáez, 
<date id="61735588F658EC379E449B4E1EBBFA7E" box="[258,472,1399,1425]" pageId="1" pageNumber="462" value="2012-12-21">21 December 2012</date>
(
<collectionCode id="73DCEB8DF658EC379EA09B4E1D2FFA7E" box="[486,588,1399,1425]" pageId="1" pageNumber="462">MZUCR</collectionCode>
3791-01).
</paragraph>
<paragraph id="15727348F658EC379F819BA21B00FA5A" blockId="1.[151,1437,1182,2037]" box="[199,1123,1434,1461]" pageId="1" pageNumber="462">
<emphasis id="27B9AF5AF658EC379F819BA21E27FA5A" bold="true" box="[199,324,1435,1461]" pageId="1" pageNumber="462">Type host:</emphasis>
The burrowing shrimp 
<taxonomicName id="D2CD08CBF658EC379D0B9BA21C8EFA5A" authority="(Sakai, 2005)" baseAuthorityName="Sakai" baseAuthorityYear="2005" box="[589,1005,1434,1461]" class="Malacostraca" family="Callianassidae" genus="Neotrypaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="462" phylum="Arthropoda" rank="species" species="tabogensis">
<emphasis id="27B9AF5AF658EC379D0B9BA21C2CFA5A" box="[589,847,1435,1461]" italics="true" pageId="1" pageNumber="462">Neotrypaea tabogensis</emphasis>
(
<bibRefCitation id="715C0EB9F658EC379C189BA21C86FA5A" author="Sakai, K." box="[862,997,1434,1461]" pageId="1" pageNumber="462" pagination="1 - 285" refId="ref5794" refString="Sakai, K. (2005) Callianassoidea of the world (Decapoda: Thalassinidea). Crustaceana Monographs, 4, 1 - 285. https: // doi. org / 10.1163 / 9789047416890" type="journal article" year="2005">Sakai, 2005</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="8DF66FCDF658EC379CBA9BA21B36FA5A" box="[1020,1109,1435,1461]" captionStart="FIGURE 1" captionStartId="2.[151,250,1153,1177]" captionTargetBox="[183,1404,349,1127]" captionTargetId="figure-102@2.[183,1404,345,1130]" captionTargetPageId="2" captionText="FIGURE 1. Neotrypaea tabogensis (Sakai, 2005) A, male and female hosts, dorsal view; B, Ione sarahae n. sp. female and male, dorsal view (MZUCR 3789-01). Scale bars: A = 5 mm, B = 2 mm (female), 250 µm (male)." figureDoi="http://doi.org/10.5281/zenodo.8036045" httpUri="https://zenodo.org/record/8036045/files/figure.png" pageId="1" pageNumber="462">Fig. 1A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5DD720C3F658EC379F819B871B8AF88A" pageId="1" pageNumber="462" type="diagnosis">
<paragraph id="15727348F658EC379F819B871B8AF88A" blockId="1.[151,1437,1182,2037]" pageId="1" pageNumber="462">
<emphasis id="27B9AF5AF658EC379F819B871E27FA37" bold="true" box="[199,324,1470,1496]" pageId="1" pageNumber="462">Diagnosis:</emphasis>
<emphasis id="27B9AF5AF658EC379E0B9B861D76FA36" box="[333,533,1471,1497]" italics="true" pageId="1" pageNumber="462">Ovigerous female</emphasis>
with body shape obovate, asymmetrical, slight sinistral distortion up to 13° in adults, all segments of pereon distinct; head not fused with first pereomere, posterior margin nearly straight; antenna composed of five articles; barbula formed by four lamellae, posterior lamella with one tapered lobe on each side; oostegite 1 ornamented with small spines from base to middle region on outer surface; maxillipedal palp composed of three articles. Pereopods subequal in structure and size, dactyli progressively less pointed posteriorly, propodus not fused with carpus, carpus composed of two articles. Five pleomeres fused dorsally, laterally separated, with long strongly ramified featherlike lateral projections posteriorly; three pairs of biramous pleopods, reduced progressively in size posteriorly, pleopods 4 and 5 absent; uropods strongly segmented. 
<emphasis id="27B9AF5AF658EC379C8D98821B3BF93A" box="[971,1112,1723,1749]" italics="true" pageId="1" pageNumber="462">Mature male</emphasis>
with body elongated, clearly segmented dorsally and laterally, symmetrical; head separated from first pereomere; maxillipeds absent; antennula and antenna of 3 and 6 articles, respectively. Pereomeres separated dorsally, segmentation not visible ventrally, directed laterally. Pereopods subequal in size and length. Pleon shorter than pereon. Pleomeres progressively narrower posteriorly, distinctly separated; pleotelson truncated; uropods uniramous and digitiform.
</paragraph>
</subSubSection>
<subSubSection id="5DD720C3F658EC329F8199571C8BF8F7" lastPageId="4" lastPageNumber="465" pageId="1" pageNumber="462" type="description">
<paragraph id="15727348F658EC349F8199571A71FEAE" blockId="1.[151,1437,1182,2037]" lastBlockId="2.[151,1436,151,321]" lastPageId="2" lastPageNumber="463" pageId="1" pageNumber="462">
<emphasis id="27B9AF5AF658EC379F8199571DC9F867" bold="true" box="[199,682,1902,1929]" pageId="1" pageNumber="462">
Description: 
<typeStatus id="CA76CDEAF658EC379E2799561EADF866" box="[353,462,1903,1929]" pageId="1" pageNumber="462" type="holotype">Holotype</typeStatus>
ovigerous female (
</emphasis>
MZUCR 3789-01
<emphasis id="27B9AF5AF658EC379C3799571C19F867" bold="true" box="[881,890,1902,1928]" pageId="1" pageNumber="462">)</emphasis>
: 
<emphasis id="27B9AF5AF658EC379CCE99561B05F866" box="[904,1126,1903,1929]" italics="true" pageId="1" pageNumber="462">General body shape</emphasis>
(
<figureCitation id="8DF66FCDF658EC379B3399561BB4F866" box="[1141,1239,1903,1929]" captionStart="FIGURE 1" captionStartId="2.[151,250,1153,1177]" captionTargetBox="[183,1404,349,1127]" captionTargetId="figure-102@2.[183,1404,345,1130]" captionTargetPageId="2" captionText="FIGURE 1. Neotrypaea tabogensis (Sakai, 2005) A, male and female hosts, dorsal view; B, Ione sarahae n. sp. female and male, dorsal view (MZUCR 3789-01). Scale bars: A = 5 mm, B = 2 mm (female), 250 µm (male)." figureDoi="http://doi.org/10.5281/zenodo.8036045" httpUri="https://zenodo.org/record/8036045/files/figure.png" pageId="1" pageNumber="462">Figs. 1A</figureCitation>
, 
<figureCitation id="8DF66FCDF658EC379BA499561B92F866" box="[1250,1265,1903,1929]" captionStart="FIGURE 2" captionStartId="3.[151,250,1276,1300]" captionTargetBox="[192,1435,192,1251]" captionTargetId="figure-18@3.[151,1435,181,1253]" captionTargetPageId="3" captionText="FIGURE 2. Ione sarahae n. sp., holotype, female, 6.6 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view. Scale bars: A, B = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8036047" httpUri="https://zenodo.org/record/8036047/files/figure.png" pageId="1" pageNumber="462">2</figureCitation>
) obovate, body segments separate, asymmetrical, slight sinistral distortion up to 13° in adults; total length 
<quantity id="D235DEADF658EC379BD699AA1B8BF842" box="[1168,1256,1939,1965]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="1" pageNumber="462" unit="mm" value="6.6">6.6 mm</quantity>
, maximal width (across pereomere 3) 
<quantity id="D235DEADF658EC379ECB998E1E8BF83E" box="[397,488,1975,2001]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="1" pageNumber="462" unit="mm" value="3.4">3.4 mm</quantity>
. 
<emphasis id="27B9AF5AF658EC379EB2998E1D52F83E" box="[500,561,1975,2001]" italics="true" pageId="1" pageNumber="462">Head</emphasis>
with hexagonal outline, approximately 1.5 times wider than long (length 
<quantity id="D235DEADF658EC379A3F998E1FA6F81A" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="462" unit="mm" value="1.1">1.1 mm</quantity>
, width 
<quantity id="D235DEADF658EC379E5C99E21E16F81A" box="[282,373,2011,2037]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="462" unit="mm" value="1.7">1.7 mm</quantity>
), straight anteriorly, nearly rounded posteriorly, not fused with first pereomere, dorsal surface smooth, lateral margins produced into obtuse angle (112°) on each side. 
<emphasis id="27B9AF5AF65BEC349CF49EAE1B39FF5E" box="[946,1114,151,177]" italics="true" pageId="2" pageNumber="463">Frontal lamina</emphasis>
very long, extending beyond anterior margin of head, with one lateral leaflike projection on each side; eyes absent. 
<emphasis id="27B9AF5AF65BEC349BD49E821B81FF3B" box="[1170,1250,187,212]" italics="true" pageId="2" pageNumber="463">Pereon</emphasis>
broadest across pereomere 3, all pereomeres dorsally, ventrally and laterally distinct. 
<emphasis id="27B9AF5AF65BEC349CFD9EE61B2FFF16" box="[955,1100,223,249]" italics="true" pageId="2" pageNumber="463">Pereomere 1</emphasis>
curved forward around head reaching its lateral angles; pereomere 2 anterior margin curved forward, others progressively curved backward. All pereomeres without tergal projections. 
<emphasis id="27B9AF5AF65BEC349D0A9F1E1DBAFEAE" box="[588,729,295,321]" italics="true" pageId="2" pageNumber="463">Coxal plates</emphasis>
oval, not overlapping; dorsolateral bosses present.
</paragraph>
<caption id="41B223C0F65BEC349FD19AB81B14FB51" ID-DOI="http://doi.org/10.5281/zenodo.8036045" ID-Zenodo-Dep="8036045" httpUri="https://zenodo.org/record/8036045/files/figure.png" pageId="2" pageNumber="463" startId="2.[151,250,1153,1177]" targetBox="[183,1404,349,1127]" targetPageId="2" targetType="figure">
<paragraph id="15727348F65BEC349FD19AB81B14FB51" blockId="2.[151,1437,1153,1214]" pageId="2" pageNumber="463">
<emphasis id="27B9AF5AF65BEC349FD19AB81E76FB75" bold="true" box="[151,277,1153,1178]" pageId="2" pageNumber="463">FIGURE 1.</emphasis>
<taxonomicName id="D2CD08CBF65BEC349E5B9ABB1DFAFB75" authority="(Sakai, 2005)" baseAuthorityName="Sakai" baseAuthorityYear="2005" box="[285,665,1154,1178]" class="Malacostraca" family="Callianassidae" genus="Neotrypaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="463" phylum="Arthropoda" rank="species" species="tabogensis">
<emphasis id="27B9AF5AF65BEC349E5B9ABB1D64FB75" box="[285,519,1154,1178]" italics="true" pageId="2" pageNumber="463">Neotrypaea tabogensis</emphasis>
(
<bibRefCitation id="715C0EB9F65BEC349D509ABB1DF1FB75" author="Sakai, K." box="[534,658,1154,1178]" pageId="2" pageNumber="463" pagination="1 - 285" refId="ref5794" refString="Sakai, K. (2005) Callianassoidea of the world (Decapoda: Thalassinidea). Crustaceana Monographs, 4, 1 - 285. https: // doi. org / 10.1163 / 9789047416890" type="journal article" year="2005">Sakai, 2005</bibRefCitation>
)
</taxonomicName>
A, male and female hosts, dorsal view; B, 
<taxonomicName id="D2CD08CBF65BEC349B1C9ABB1B82FB75" authority="Hernáez &amp; Villegas-Castro &amp; Boyko, 2023" authorityName="Hernáez &amp; Villegas-Castro &amp; Boyko" authorityYear="2023" box="[1114,1249,1154,1178]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="463" phylum="Arthropoda" rank="species" species="sarahae" status="sp. nov.">
<emphasis id="27B9AF5AF65BEC349B1C9ABB1B82FB75" box="[1114,1249,1154,1178]" italics="true" pageId="2" pageNumber="463">Ione sarahae</emphasis>
</taxonomicName>
<emphasis id="27B9AF5AF65BEC349BAE9ABB1A40FB75" bold="true" box="[1256,1315,1154,1178]" pageId="2" pageNumber="463">
<taxonomicNameLabel id="3C8A1221F65BEC349BAE9ABB1A40FB75" box="[1256,1315,1154,1178]" pageId="2" pageNumber="463" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
female and male, dorsal view (MZUCR 3789-01). Scale bars: A = 5 mm, B = 2 mm (female), 250 µm (male).
</paragraph>
</caption>
<paragraph id="15727348F65BEC349F819ADE1E91FAA6" blockId="2.[151,1436,1255,2037]" pageId="2" pageNumber="463">
<emphasis id="27B9AF5AF65BEC349F819ADE1E59FAEE" box="[199,314,1255,1281]" italics="true" pageId="2" pageNumber="463">Antennula</emphasis>
(
<figureCitation id="8DF66FCDF65BEC349E0F9ADE1EC3FAEE" box="[329,416,1255,1281]" captionStart="FIGURE 3" captionStartId="4.[151,250,1575,1599]" captionTargetBox="[178,1414,185,1549]" captionTargetId="figure-26@4.[170,1416,181,1551]" captionTargetPageId="4" captionText="FIGURE 3. Ione sarahae n. sp., paratype, female, 4.3 mm TL (MZUCR 3791-01). A, right antennula; B, right antenna; C, barbula; D, right maxilliped; E, right pereopod 1; F, right pereopod 7; G, right oostegite 1, external view; H, right oostegite 1, internal view. Scale bars: A, B, E, F = 100 µm; C = 500 µm; G, H = 1 mm; D = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.8036049" httpUri="https://zenodo.org/record/8036049/files/figure.png" pageId="2" pageNumber="463">Fig. 3A</figureCitation>
) composed of three articles; basal article globose with sparse, short stiff setae distolaterally; second article subcylindrical with isolated stiff setae distally; distal article subcylindrical, shorter than second article, with tuft of short setae distally.
</paragraph>
<paragraph id="15727348F65BEC349F819B6A1E9FFA5A" blockId="2.[151,1436,1255,2037]" pageId="2" pageNumber="463">
<emphasis id="27B9AF5AF65BEC349F819B6A1E47FA82" box="[199,292,1363,1389]" italics="true" pageId="2" pageNumber="463">Antenna</emphasis>
(
<figureCitation id="8DF66FCDF65BEC349E739B6A1EE8FA82" box="[309,395,1363,1389]" captionStart="FIGURE 3" captionStartId="4.[151,250,1575,1599]" captionTargetBox="[178,1414,185,1549]" captionTargetId="figure-26@4.[170,1416,181,1551]" captionTargetPageId="4" captionText="FIGURE 3. Ione sarahae n. sp., paratype, female, 4.3 mm TL (MZUCR 3791-01). A, right antennula; B, right antenna; C, barbula; D, right maxilliped; E, right pereopod 1; F, right pereopod 7; G, right oostegite 1, external view; H, right oostegite 1, internal view. Scale bars: A, B, E, F = 100 µm; C = 500 µm; G, H = 1 mm; D = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.8036049" httpUri="https://zenodo.org/record/8036049/files/figure.png" pageId="2" pageNumber="463">Fig. 3B</figureCitation>
) composed of five articles; basal fifth article globose, not fused with fourth article, article 2 subquadrate, articles 3 and 4 cylindrical, with isolated setae distolaterally, distal article subcylindrical, shorter than former, with tuft of setae on tip.
</paragraph>
<paragraph id="15727348F65BEC349F819B861C2AFA12" blockId="2.[151,1436,1255,2037]" pageId="2" pageNumber="463">
<emphasis id="27B9AF5AF65BEC349F819B861E40FA36" box="[199,291,1471,1497]" italics="true" pageId="2" pageNumber="463">Barbula</emphasis>
(
<figureCitation id="8DF66FCDF65BEC349E739B861EEDFA36" box="[309,398,1471,1497]" captionStart="FIGURE 3" captionStartId="4.[151,250,1575,1599]" captionTargetBox="[178,1414,185,1549]" captionTargetId="figure-26@4.[170,1416,181,1551]" captionTargetPageId="4" captionText="FIGURE 3. Ione sarahae n. sp., paratype, female, 4.3 mm TL (MZUCR 3791-01). A, right antennula; B, right antenna; C, barbula; D, right maxilliped; E, right pereopod 1; F, right pereopod 7; G, right oostegite 1, external view; H, right oostegite 1, internal view. Scale bars: A, B, E, F = 100 µm; C = 500 µm; G, H = 1 mm; D = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.8036049" httpUri="https://zenodo.org/record/8036049/files/figure.png" pageId="2" pageNumber="463">Fig. 3C</figureCitation>
) with two pair of smooth lamellae: posterior lamella with one tapered lobe on each side, middle region flat; each lateral lamella with one tapered lobe.
</paragraph>
<paragraph id="15727348F65BEC349F81983E1C06F9AA" blockId="2.[151,1436,1255,2037]" pageId="2" pageNumber="463">
<emphasis id="27B9AF5AF65BEC349F81983E1E23F9CE" box="[199,320,1543,1569]" italics="true" pageId="2" pageNumber="463">Maxilliped</emphasis>
(
<figureCitation id="8DF66FCDF65BEC349E13983E1ED2F9CE" box="[341,433,1543,1569]" captionStart="FIGURE 3" captionStartId="4.[151,250,1575,1599]" captionTargetBox="[178,1414,185,1549]" captionTargetId="figure-26@4.[170,1416,181,1551]" captionTargetPageId="4" captionText="FIGURE 3. Ione sarahae n. sp., paratype, female, 4.3 mm TL (MZUCR 3791-01). A, right antennula; B, right antenna; C, barbula; D, right maxilliped; E, right pereopod 1; F, right pereopod 7; G, right oostegite 1, external view; H, right oostegite 1, internal view. Scale bars: A, B, E, F = 100 µm; C = 500 µm; G, H = 1 mm; D = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.8036049" httpUri="https://zenodo.org/record/8036049/files/figure.png" pageId="2" pageNumber="463">Fig. 3D</figureCitation>
) articulated, prominent basal process anterolaterally, distal article subpyramidal; palp composed of three articles, distal article with 1 or 2 setae on tip.
</paragraph>
<paragraph id="15727348F65BEC349F8198761EC1F93A" blockId="2.[151,1436,1255,2037]" pageId="2" pageNumber="463">
<emphasis id="27B9AF5AF65BEC349F8198761E5DF986" box="[199,318,1615,1641]" italics="true" pageId="2" pageNumber="463">Pereopods</emphasis>
(
<figureCitation id="8DF66FCDF65BEC349E0998761EA6F986" box="[335,453,1615,1641]" captionStart="FIGURE 3" captionStartId="4.[151,250,1575,1599]" captionTargetBox="[178,1414,185,1549]" captionTargetId="figure-26@4.[170,1416,181,1551]" captionTargetPageId="4" captionText="FIGURE 3. Ione sarahae n. sp., paratype, female, 4.3 mm TL (MZUCR 3791-01). A, right antennula; B, right antenna; C, barbula; D, right maxilliped; E, right pereopod 1; F, right pereopod 7; G, right oostegite 1, external view; H, right oostegite 1, internal view. Scale bars: A, B, E, F = 100 µm; C = 500 µm; G, H = 1 mm; D = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.8036049" httpUri="https://zenodo.org/record/8036049/files/figure.png" pageId="2" pageNumber="463">Fig. 3E, F</figureCitation>
) subequal in size and structure; ischium widening distally in lateral view (
<figureCitation id="8DF66FCDF65BEC349A5B98761A10F986" box="[1309,1395,1615,1641]" captionStart="FIGURE 3" captionStartId="4.[151,250,1575,1599]" captionTargetBox="[178,1414,185,1549]" captionTargetId="figure-26@4.[170,1416,181,1551]" captionTargetPageId="4" captionText="FIGURE 3. Ione sarahae n. sp., paratype, female, 4.3 mm TL (MZUCR 3791-01). A, right antennula; B, right antenna; C, barbula; D, right maxilliped; E, right pereopod 1; F, right pereopod 7; G, right oostegite 1, external view; H, right oostegite 1, internal view. Scale bars: A, B, E, F = 100 µm; C = 500 µm; G, H = 1 mm; D = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.8036049" httpUri="https://zenodo.org/record/8036049/files/figure.png" pageId="2" pageNumber="463">Fig. 3E</figureCitation>
) or subquadrate (
<figureCitation id="8DF66FCDF65BEC349E68984A1EE7F962" box="[302,388,1651,1677]" captionStart="FIGURE 3" captionStartId="4.[151,250,1575,1599]" captionTargetBox="[178,1414,185,1549]" captionTargetId="figure-26@4.[170,1416,181,1551]" captionTargetPageId="4" captionText="FIGURE 3. Ione sarahae n. sp., paratype, female, 4.3 mm TL (MZUCR 3791-01). A, right antennula; B, right antenna; C, barbula; D, right maxilliped; E, right pereopod 1; F, right pereopod 7; G, right oostegite 1, external view; H, right oostegite 1, internal view. Scale bars: A, B, E, F = 100 µm; C = 500 µm; G, H = 1 mm; D = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.8036049" httpUri="https://zenodo.org/record/8036049/files/figure.png" pageId="2" pageNumber="463">Fig. 3F</figureCitation>
), similar in length to merus; carpus two-segmented; propodus with anterior margin expended dorsally, not fused with carpus, inner margin with small crenulations used for attachment to host; dactyli progressively less pointed posteriorly.
</paragraph>
<paragraph id="15727348F65BEC349F8198E61E93F866" blockId="2.[151,1436,1255,2037]" pageId="2" pageNumber="463">
<taxonomicName id="D2CD08CBF65BEC349F8198E61E2BF917" box="[199,328,1759,1784]" class="Magnoliopsida" family="Fabaceae" genus="Marsupium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Fabales" pageId="2" pageNumber="463" phylum="Echinodermata" rank="genus">
<emphasis id="27B9AF5AF65BEC349F8198E61E2BF917" box="[199,328,1759,1784]" italics="true" pageId="2" pageNumber="463">Marsupium</emphasis>
</taxonomicName>
(
<figureCitation id="8DF66FCDF65BEC349E1198E61EDBF916" box="[343,440,1759,1785]" captionStart="FIGURE 2" captionStartId="3.[151,250,1276,1300]" captionTargetBox="[192,1435,192,1251]" captionTargetId="figure-18@3.[151,1435,181,1253]" captionTargetPageId="3" captionText="FIGURE 2. Ione sarahae n. sp., holotype, female, 6.6 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view. Scale bars: A, B = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8036047" httpUri="https://zenodo.org/record/8036047/files/figure.png" pageId="2" pageNumber="463">Figs. 2B</figureCitation>
, 
<figureCitation id="8DF66FCDF65BEC349E8598E61D64F916" box="[451,519,1759,1785]" captionStart="FIGURE 3" captionStartId="4.[151,250,1575,1599]" captionTargetBox="[178,1414,185,1549]" captionTargetId="figure-26@4.[170,1416,181,1551]" captionTargetPageId="4" captionText="FIGURE 3. Ione sarahae n. sp., paratype, female, 4.3 mm TL (MZUCR 3791-01). A, right antennula; B, right antenna; C, barbula; D, right maxilliped; E, right pereopod 1; F, right pereopod 7; G, right oostegite 1, external view; H, right oostegite 1, internal view. Scale bars: A, B, E, F = 100 µm; C = 500 µm; G, H = 1 mm; D = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.8036049" httpUri="https://zenodo.org/record/8036049/files/figure.png" pageId="2" pageNumber="463">3G, H</figureCitation>
) with five pairs of oostegites, inner surface concave in both segments, ornamented with small spines from base to middle region of outer surface, similar in shape; oostegites overlapping, enclosing brood pouch (
<figureCitation id="8DF66FCDF65BEC349E73991E1EEDF8AE" box="[309,398,1831,1857]" captionStart="FIGURE 2" captionStartId="3.[151,250,1276,1300]" captionTargetBox="[192,1435,192,1251]" captionTargetId="figure-18@3.[151,1435,181,1253]" captionTargetPageId="3" captionText="FIGURE 2. Ione sarahae n. sp., holotype, female, 6.6 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view. Scale bars: A, B = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8036047" httpUri="https://zenodo.org/record/8036047/files/figure.png" pageId="2" pageNumber="463">Fig. 2B</figureCitation>
); oostegite 1 smallest, each segment formed by two plates covering oral region completely; oostegites 2 and 3 largest; others similar in size. Oostegites 2–5 each with longitudinal carina, posterior margin of oostegite 5 covered with setae.
</paragraph>
<paragraph id="15727348F65BEC349F8199AA1CB5F81A" blockId="2.[151,1436,1255,2037]" pageId="2" pageNumber="463">
Length of pleon approximately 4.8 times smaller than head/pereon (
<figureCitation id="8DF66FCDF65BEC349C9199AA1B57F842" box="[983,1076,1939,1965]" captionStart="FIGURE 2" captionStartId="3.[151,250,1276,1300]" captionTargetBox="[192,1435,192,1251]" captionTargetId="figure-18@3.[151,1435,181,1253]" captionTargetPageId="3" captionText="FIGURE 2. Ione sarahae n. sp., holotype, female, 6.6 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view. Scale bars: A, B = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8036047" httpUri="https://zenodo.org/record/8036047/files/figure.png" pageId="2" pageNumber="463">Fig. 2A</figureCitation>
). Pleomeres 1-5 fused dorsally, laterally not separated, with long strongly ramified featherlike lateral plates posteriorly; ventral surface of pleomeres smooth; pleotelson subquadrate, approximately 1.4 times wider than long.
</paragraph>
<caption id="41B223C0F65AEC359FD19AC51EB0FAD7" ID-DOI="http://doi.org/10.5281/zenodo.8036047" ID-Zenodo-Dep="8036047" httpUri="https://zenodo.org/record/8036047/files/figure.png" pageId="3" pageNumber="464" startId="3.[151,250,1276,1300]" targetBox="[192,1435,192,1251]" targetPageId="3" targetType="figure">
<paragraph id="15727348F65AEC359FD19AC51EB0FAD7" blockId="3.[151,1436,1276,1337]" pageId="3" pageNumber="464">
<emphasis id="27B9AF5AF65AEC359FD19AC51E70FAFA" bold="true" box="[151,275,1276,1301]" pageId="3" pageNumber="464">FIGURE 2.</emphasis>
<taxonomicName id="D2CD08CBF65AEC359E5F9AC41EFCFAFA" authority="Hernáez &amp; Villegas-Castro &amp; Boyko, 2023" authorityName="Hernáez &amp; Villegas-Castro &amp; Boyko" authorityYear="2023" box="[281,415,1277,1301]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="464" phylum="Arthropoda" rank="species" species="sarahae" status="sp. nov.">
<emphasis id="27B9AF5AF65AEC359E5F9AC41EFCFAFA" box="[281,415,1277,1301]" italics="true" pageId="3" pageNumber="464">Ione sarahae</emphasis>
</taxonomicName>
<emphasis id="27B9AF5AF65AEC359EE39AC41EBDFAFA" bold="true" box="[421,478,1277,1301]" pageId="3" pageNumber="464">
<taxonomicNameLabel id="3C8A1221F65AEC359EE39AC41EBDFAFA" box="[421,478,1277,1301]" pageId="3" pageNumber="464" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, female, 6.6 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view. Scale bars: A, B = 2 mm.
</paragraph>
</caption>
<paragraph id="15727348F65AEC359F819B5E1E02FA26" blockId="3.[151,1437,1383,2021]" pageId="3" pageNumber="464">
Pleopods 1-3 biramous, reduced progressively in size posteriorly; pleopods 4 and 5 absent, endopod much larger than exopod (
<figureCitation id="8DF66FCDF65AEC359EC49BB21EBDFA4A" box="[386,478,1419,1445]" captionStart="FIGURE 2" captionStartId="3.[151,250,1276,1300]" captionTargetBox="[192,1435,192,1251]" captionTargetId="figure-18@3.[151,1435,181,1253]" captionTargetPageId="3" captionText="FIGURE 2. Ione sarahae n. sp., holotype, female, 6.6 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view. Scale bars: A, B = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8036047" httpUri="https://zenodo.org/record/8036047/files/figure.png" pageId="3" pageNumber="464">Fig. 2B</figureCitation>
). 
<emphasis id="27B9AF5AF65AEC359EB09BB21D34FA4A" box="[502,599,1419,1445]" italics="true" pageId="3" pageNumber="464">Uropods</emphasis>
(
<figureCitation id="8DF66FCDF65AEC359D2C9BB21D8BFA4A" box="[618,744,1419,1445]" captionStart="FIGURE 2" captionStartId="3.[151,250,1276,1300]" captionTargetBox="[192,1435,192,1251]" captionTargetId="figure-18@3.[151,1435,181,1253]" captionTargetPageId="3" captionText="FIGURE 2. Ione sarahae n. sp., holotype, female, 6.6 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view. Scale bars: A, B = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8036047" httpUri="https://zenodo.org/record/8036047/files/figure.png" pageId="3" pageNumber="464">Fig. 2A, B</figureCitation>
) strongly segmented, lanceolate and biramous, outer ramus strongly ramified.
</paragraph>
<paragraph id="15727348F65AEC359F819BEA1EACF992" blockId="3.[151,1437,1383,2021]" pageId="3" pageNumber="464">
<emphasis id="27B9AF5AF65AEC359F819BEA1EE6FA03" bold="true" box="[199,389,1490,1517]" pageId="3" pageNumber="464">
<typeStatus id="CA76CDEAF65AEC359F819BEA1E4FFA02" box="[199,300,1491,1517]" pageId="3" pageNumber="464" type="allotype">Allotype</typeStatus>
male (
</emphasis>
<specimenCode id="456BDB33F65AEC359EC39BEA1D32FA02" box="[389,593,1491,1517]" collectionCode="MZUCR" pageId="3" pageNumber="464">MZUCR 3789-01</specimenCode>
<emphasis id="27B9AF5AF65AEC359D179BEB1D00FA03" bold="true" box="[593,611,1490,1516]" pageId="3" pageNumber="464">):</emphasis>
<emphasis id="27B9AF5AF65AEC359D289BEA1DCBFA02" box="[622,680,1491,1517]" italics="true" pageId="3" pageNumber="464">Body</emphasis>
(
<figureCitation id="8DF66FCDF65AEC359DFD9BEA1C59FA02" box="[699,826,1491,1517]" captionStart="FIGURE 4" captionStartId="5.[151,250,1977,2001]" captionTargetBox="[169,1421,828,1951]" captionTargetId="figure-356@5.[164,1423,813,1954]" captionTargetPageId="5" captionText="FIGURE 4. Ione sarahae n. sp., allotype, male, 3.5 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view; C, left antennula; D, left antenna; E, left pereopod 1. Scale bars: A, B = 250 µm; C, D = 50 µm; E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.8036051" httpUri="https://zenodo.org/record/8036051/files/figure.png" pageId="3" pageNumber="464">Fig. 4A, B</figureCitation>
) symmetrical, with lanceolate outline, unpigmented, elongated, distinctly segmented dorsally and laterally, body boundaries not visible ventrally; total length 
<quantity id="D235DEADF65AEC359A3F9BCE1FA6F9DA" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="464" unit="mm" value="3.5">3.5 mm</quantity>
, maximal width at fourth pereomere, 
<quantity id="D235DEADF65AEC359DC598221D8DF9DA" box="[643,750,1563,1590]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="3" pageNumber="464" unit="mm" value="0.57">0.57 mm</quantity>
. 
<emphasis id="27B9AF5AF65AEC359DBA98221C5AF9DA" box="[764,825,1563,1589]" italics="true" pageId="3" pageNumber="464">Head</emphasis>
(
<figureCitation id="8DF66FCDF65AEC359C0A98221CA8F9DA" box="[844,971,1563,1589]" captionStart="FIGURE 4" captionStartId="5.[151,250,1977,2001]" captionTargetBox="[169,1421,828,1951]" captionTargetId="figure-356@5.[164,1423,813,1954]" captionTargetPageId="5" captionText="FIGURE 4. Ione sarahae n. sp., allotype, male, 3.5 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view; C, left antennula; D, left antenna; E, left pereopod 1. Scale bars: A, B = 250 µm; C, D = 50 µm; E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.8036051" httpUri="https://zenodo.org/record/8036051/files/figure.png" pageId="3" pageNumber="464">Fig. 4A, B</figureCitation>
) suboval, distinct from first pereomere, approximately 4 times as wide as long; anterolateral borders rounded and produced ventrally; eyes small, dark near to posterior margin of head.
</paragraph>
<paragraph id="15727348F65AEC359F8198BE1C57F906" blockId="3.[151,1437,1383,2021]" pageId="3" pageNumber="464">
<emphasis id="27B9AF5AF65AEC359F8198BE1EB2F94E" box="[199,465,1671,1697]" italics="true" pageId="3" pageNumber="464">Antennula and antenna</emphasis>
(
<figureCitation id="8DF66FCDF65AEC359EA298BE1D02F94E" box="[484,609,1671,1697]" captionStart="FIGURE 4" captionStartId="5.[151,250,1977,2001]" captionTargetBox="[169,1421,828,1951]" captionTargetId="figure-356@5.[164,1423,813,1954]" captionTargetPageId="5" captionText="FIGURE 4. Ione sarahae n. sp., allotype, male, 3.5 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view; C, left antennula; D, left antenna; E, left pereopod 1. Scale bars: A, B = 250 µm; C, D = 50 µm; E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.8036051" httpUri="https://zenodo.org/record/8036051/files/figure.png" pageId="3" pageNumber="464">Fig. 4C, D</figureCitation>
) of 3 and 6 articles, respectively; antennula much shorter than antenna, antenna approximately 2.6 times longer than antennula; antennula with small tuft of setae at tip of distal article; antennae (
<figureCitation id="8DF66FCDF65AEC359E4C98F61E01F906" box="[266,354,1743,1769]" captionStart="FIGURE 4" captionStartId="5.[151,250,1977,2001]" captionTargetBox="[169,1421,828,1951]" captionTargetId="figure-356@5.[164,1423,813,1954]" captionTargetPageId="5" captionText="FIGURE 4. Ione sarahae n. sp., allotype, male, 3.5 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view; C, left antennula; D, left antenna; E, left pereopod 1. Scale bars: A, B = 250 µm; C, D = 50 µm; E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.8036051" httpUri="https://zenodo.org/record/8036051/files/figure.png" pageId="3" pageNumber="464">Fig. 4A</figureCitation>
) extending beyond distal margin of head.
</paragraph>
<paragraph id="15727348F65AEC359F8198CA1EC1F8E2" blockId="3.[151,1437,1383,2021]" box="[199,418,1779,1805]" pageId="3" pageNumber="464">
<emphasis id="27B9AF5AF65AEC359F8198CA1E28F8E2" box="[199,331,1779,1805]" italics="true" pageId="3" pageNumber="464">Maxillipeds</emphasis>
absent.
</paragraph>
<paragraph id="15727348F65AEC359F8199211B23F8DE" blockId="3.[151,1437,1383,2021]" box="[199,1088,1815,1841]" pageId="3" pageNumber="464">
<emphasis id="27B9AF5AF65AEC359F8199211E28F8DE" box="[199,331,1816,1841]" italics="true" pageId="3" pageNumber="464">Pereomeres</emphasis>
(
<figureCitation id="8DF66FCDF65AEC359E1C992E1ED1F8DE" box="[346,434,1815,1841]" captionStart="FIGURE 4" captionStartId="5.[151,250,1977,2001]" captionTargetBox="[169,1421,828,1951]" captionTargetId="figure-356@5.[164,1423,813,1954]" captionTargetPageId="5" captionText="FIGURE 4. Ione sarahae n. sp., allotype, male, 3.5 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view; C, left antennula; D, left antenna; E, left pereopod 1. Scale bars: A, B = 250 µm; C, D = 50 µm; E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.8036051" httpUri="https://zenodo.org/record/8036051/files/figure.png" pageId="3" pageNumber="464">Fig. 4A</figureCitation>
) separated, directed laterally; midventral tubercles absent.
</paragraph>
<paragraph id="15727348F65AEC359F8199021E0BF80A" blockId="3.[151,1437,1383,2021]" pageId="3" pageNumber="464">
<emphasis id="27B9AF5AF65AEC359F8199021E5DF8BA" box="[199,318,1851,1877]" italics="true" pageId="3" pageNumber="464">Pereopods</emphasis>
(
<figureCitation id="8DF66FCDF65AEC359E0A99021EA3F8BA" box="[332,448,1851,1877]" captionStart="FIGURE 4" captionStartId="5.[151,250,1977,2001]" captionTargetBox="[169,1421,828,1951]" captionTargetId="figure-356@5.[164,1423,813,1954]" captionTargetPageId="5" captionText="FIGURE 4. Ione sarahae n. sp., allotype, male, 3.5 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view; C, left antennula; D, left antenna; E, left pereopod 1. Scale bars: A, B = 250 µm; C, D = 50 µm; E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.8036051" httpUri="https://zenodo.org/record/8036051/files/figure.png" pageId="3" pageNumber="464">Fig. 4B, E</figureCitation>
) subequal in size and length, not visible in dorsal view; pereopods 1–7 used for attaching to female. 
<emphasis id="27B9AF5AF65AEC359E4A99661EE8F896" box="[268,395,1887,1913]" italics="true" pageId="3" pageNumber="464">Pereopod 1</emphasis>
prehensile; length of ischium more than twice that of merus; merus slightly widening distally; carpus two-segmented, armed with sclerotized setae distolaterally; dorsal margin of propodus expanded anteriorly, inner margin irregularly crenulate used for attachment to female; dactyli strongly arcuate with each tip slightly curved downward.
</paragraph>
<caption id="41B223C0F65DEC329FD1981E1B69F967" ID-DOI="http://doi.org/10.5281/zenodo.8036049" ID-Zenodo-Dep="8036049" httpUri="https://zenodo.org/record/8036049/files/figure.png" pageId="4" pageNumber="465" startId="4.[151,250,1575,1599]" targetBox="[178,1414,185,1549]" targetPageId="4" targetType="figure">
<paragraph id="15727348F65DEC329FD1981E1B69F967" blockId="4.[151,1436,1575,1672]" pageId="4" pageNumber="465">
<emphasis id="27B9AF5AF65DEC329FD1981E1E76F9D0" bold="true" box="[151,277,1575,1599]" pageId="4" pageNumber="465">FIGURE 3.</emphasis>
<taxonomicName id="D2CD08CBF65DEC329E5898111EC6F9D0" authority="Hernáez &amp; Villegas-Castro &amp; Boyko, 2023" authorityName="Hernáez &amp; Villegas-Castro &amp; Boyko" authorityYear="2023" box="[286,421,1575,1599]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="sarahae" status="sp. nov.">
<emphasis id="27B9AF5AF65DEC329E5898111EC6F9D0" box="[286,421,1575,1599]" italics="true" pageId="4" pageNumber="465">Ione sarahae</emphasis>
</taxonomicName>
<emphasis id="27B9AF5AF65DEC329EEB981E1E8BF9D0" bold="true" box="[429,488,1575,1599]" pageId="4" pageNumber="465">
<taxonomicNameLabel id="3C8A1221F65DEC329EEB981E1E8BF9D0" box="[429,488,1575,1599]" pageId="4" pageNumber="465" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, paratype, female, 4.3 mm TL (MZUCR 3791-01). A, right antennula; B, right antenna; C, barbula; D, right maxilliped; E, right pereopod 1; F, right pereopod 7; G, right oostegite 1, external view; H, right oostegite 1, internal view. Scale bars: A, B, E, F = 100 µm; C = 500 µm; G, H = 1 mm; D = 50 µm.
</paragraph>
</caption>
<paragraph id="15727348F65DEC329F81988C1C8BF8F7" blockId="4.[151,1437,1717,2032]" pageId="4" pageNumber="465">
<emphasis id="27B9AF5AF65DEC329F81988C1E6BF920" box="[199,264,1717,1743]" italics="true" pageId="4" pageNumber="465">Pleon</emphasis>
(
<figureCitation id="8DF66FCDF65DEC329E5E988C1EECF920" box="[280,399,1717,1743]" captionStart="FIGURE 4" captionStartId="5.[151,250,1977,2001]" captionTargetBox="[169,1421,828,1951]" captionTargetId="figure-356@5.[164,1423,813,1954]" captionTargetPageId="5" captionText="FIGURE 4. Ione sarahae n. sp., allotype, male, 3.5 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view; C, left antennula; D, left antenna; E, left pereopod 1. Scale bars: A, B = 250 µm; C, D = 50 µm; E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.8036051" httpUri="https://zenodo.org/record/8036051/files/figure.png" pageId="4" pageNumber="465">Fig. 4A, B</figureCitation>
) shorter than pereon. Pleomeres 1-5 progressively narrower posteriorly, distinctly separated, lateral margins rounded; each pleomere with pair of digitiform and elongated lateral plates. Pleopods absent. Pleotelson truncated. 
<emphasis id="27B9AF5AF65DEC329ECF98C41E89F8F8" box="[393,490,1789,1815]" italics="true" pageId="4" pageNumber="465">Uropods</emphasis>
uniramous and digitiform; anal cone present.
</paragraph>
</subSubSection>
<subSubSection id="5DD720C3F65DEC329F8199181E39F8B0" pageId="4" pageNumber="465" type="distribution">
<paragraph id="15727348F65DEC329F8199181E39F8B0" blockId="4.[151,1437,1717,2032]" pageId="4" pageNumber="465">
<emphasis id="27B9AF5AF65DEC329F8199181E00F8D4" bold="true" box="[199,355,1825,1851]" pageId="4" pageNumber="465">Distribution:</emphasis>
Known only from the intertidal zone at the 
<typeStatus id="CA76CDEAF65DEC329C00991B1C15F8D3" box="[838,886,1826,1852]" pageId="4" pageNumber="465">type</typeStatus>
locality of Mata de 
<collectingRegion id="D709BDAAF65DEC329B1299181BC1F8D4" box="[1108,1186,1825,1851]" country="Costa Rica" name="Limon" pageId="4" pageNumber="465">Limón</collectingRegion>
, Pacific coast of 
<collectingCountry id="6DDA33D8F65DEC329A1A99181FAEF8B0" name="Costa Rica" pageId="4" pageNumber="465">Costa Rica</collectingCountry>
, 
<collectingCountry id="6DDA33D8F65DEC329F9F997C1E35F8B0" box="[217,342,1861,1887]" name="Costa Rica" pageId="4" pageNumber="465">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="5DD720C3F65DEC329F8199501C65F848" pageId="4" pageNumber="465" type="etymology">
<paragraph id="15727348F65DEC329F8199501C65F848" blockId="4.[151,1437,1717,2032]" pageId="4" pageNumber="465">
<emphasis id="27B9AF5AF65DEC329F8199501E31F86C" bold="true" box="[199,338,1897,1923]" pageId="4" pageNumber="465">Etymology:</emphasis>
The specific name 
<taxonomicName id="D2CD08CBF65DEC329D6E99501DE1F86C" authorityName="Hernáez &amp; Villegas-Castro &amp; Boyko" authorityYear="2023" box="[552,642,1897,1923]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="sarahae">
<emphasis id="27B9AF5AF65DEC329D6E99501DE1F86C" box="[552,642,1897,1923]" italics="true" pageId="4" pageNumber="465">sarahae</emphasis>
</taxonomicName>
is given to honor the young lady Sarah Villegas Abarca, daughter of the second author, who has filled her father's life with love.
</paragraph>
</subSubSection>
<subSubSection id="5DD720C3F65DEC339F8199881BEAFCFA" lastPageId="5" lastPageNumber="466" pageId="4" pageNumber="465" type="discussion">
<paragraph id="15727348F65DEC339F8199881E8FFE3E" blockId="4.[151,1437,1717,2032]" lastBlockId="5.[151,1437,151,789]" lastPageId="5" lastPageNumber="466" pageId="4" pageNumber="465">
<emphasis id="27B9AF5AF65DEC329F8199881E5EF824" bold="true" box="[199,317,1969,1995]" pageId="4" pageNumber="465">Remarks:</emphasis>
The new species described here is assigned to 
<taxonomicName id="D2CD08CBF65DEC329C08998B1CE3F824" authorityName="Latreille" authorityYear="1818" box="[846,896,1970,1995]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis id="27B9AF5AF65DEC329C08998B1CE3F824" box="[846,896,1970,1995]" italics="true" pageId="4" pageNumber="465">Ione</emphasis>
</taxonomicName>
, whose main diagnostic characters (cf. 
<bibRefCitation id="715C0EB9F65DEC329A7B99881FB1F800" author="Latreille, P. A." pageId="4" pageNumber="465" pagination="1 - 38" refId="ref5348" refString="Latreille, P. A. (1818) Crustaces, arachnides et insectes. In: Tableau encyclopedique et methodique des trois regnes de la nature. Agasse, Paris, 24, [1 - 4], pp. 1 - 38, pls. 268 - 397." type="book chapter" year="1818">Latreille 1818</bibRefCitation>
; see also 
<bibRefCitation id="715C0EB9F65DEC329E0299EC1D77F800" author="Boyko, C. B. &amp; Moss, J. &amp; Williams, J. D. &amp; Shields, J. D." box="[324,532,2005,2032]" pageId="4" pageNumber="465" pagination="495 - 506" refId="ref4365" refString="Boyko, C. B., Moss, J., Williams, J. D. &amp; Shields, J. D. (2013) A molecular phylogeny of Bopyroidea and Cryptoniscoidea (Crustacea: Isopoda). Systematics and Biodiversity, 11 (4), 495 - 506. https: // doi. org / 10.1080 / 14772000.2013.865679" type="journal article" year="2013">
Boyko 
<emphasis id="27B9AF5AF65DEC329EDE99EF1EAFF800" box="[408,460,2005,2031]" italics="true" pageId="4" pageNumber="465">et al</emphasis>
. 2013
</bibRefCitation>
and references therein) are readily recognized in the new species. They are as follows: female pleonal lateral plates with numerous filamentous rami, and biramous uropods (see 
<figureCitation id="8DF66FCDF65CEC339B989EAE1A5DFF5E" box="[1246,1342,151,177]" captionStart="FIGURE 1" captionStartId="2.[151,250,1153,1177]" captionTargetBox="[183,1404,349,1127]" captionTargetId="figure-102@2.[183,1404,345,1130]" captionTargetPageId="2" captionText="FIGURE 1. Neotrypaea tabogensis (Sakai, 2005) A, male and female hosts, dorsal view; B, Ione sarahae n. sp. female and male, dorsal view (MZUCR 3789-01). Scale bars: A = 5 mm, B = 2 mm (female), 250 µm (male)." figureDoi="http://doi.org/10.5281/zenodo.8036045" httpUri="https://zenodo.org/record/8036045/files/figure.png" pageId="5" pageNumber="466">Figs. 1B</figureCitation>
, 
<figureCitation id="8DF66FCDF65CEC339A0F9EAE1AEEFF5E" box="[1353,1421,151,177]" captionStart="FIGURE 2" captionStartId="3.[151,250,1276,1300]" captionTargetBox="[192,1435,192,1251]" captionTargetId="figure-18@3.[151,1435,181,1253]" captionTargetPageId="3" captionText="FIGURE 2. Ione sarahae n. sp., holotype, female, 6.6 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view. Scale bars: A, B = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8036047" httpUri="https://zenodo.org/record/8036047/files/figure.png" pageId="5" pageNumber="466">2A, B</figureCitation>
); male with five pairs of simple pleonal lateral plates, and uniramous uropods (see 
<figureCitation id="8DF66FCDF65CEC339B529E821B10FF3A" box="[1044,1139,187,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1153,1177]" captionTargetBox="[183,1404,349,1127]" captionTargetId="figure-102@2.[183,1404,345,1130]" captionTargetPageId="2" captionText="FIGURE 1. Neotrypaea tabogensis (Sakai, 2005) A, male and female hosts, dorsal view; B, Ione sarahae n. sp. female and male, dorsal view (MZUCR 3789-01). Scale bars: A = 5 mm, B = 2 mm (female), 250 µm (male)." figureDoi="http://doi.org/10.5281/zenodo.8036045" httpUri="https://zenodo.org/record/8036045/files/figure.png" pageId="5" pageNumber="466">Figs. 1B</figureCitation>
, 
<figureCitation id="8DF66FCDF65CEC339B399E821BA1FF3A" box="[1151,1218,187,213]" captionStart="FIGURE 4" captionStartId="5.[151,250,1977,2001]" captionTargetBox="[169,1421,828,1951]" captionTargetId="figure-356@5.[164,1423,813,1954]" captionTargetPageId="5" captionText="FIGURE 4. Ione sarahae n. sp., allotype, male, 3.5 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view; C, left antennula; D, left antenna; E, left pereopod 1. Scale bars: A, B = 250 µm; C, D = 50 µm; E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.8036051" httpUri="https://zenodo.org/record/8036051/files/figure.png" pageId="5" pageNumber="466">4A, B</figureCitation>
). 
<taxonomicName id="D2CD08CBF65CEC339B909E821A6BFF3B" authorityName="Latreille" authorityYear="1818" box="[1238,1288,187,212]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="genus">
<emphasis id="27B9AF5AF65CEC339B909E821A6BFF3B" box="[1238,1288,187,212]" italics="true" pageId="5" pageNumber="466">Ione</emphasis>
</taxonomicName>
now consists of seven species (
<bibRefCitation id="715C0EB9F65CEC339E1B9EE61D4FFF16" author="Boyko, C. B. &amp; Bruce, N. L. &amp; Hadfield, K. A. &amp; Merrin, K. L. &amp; Ota, Y. &amp; Poore, G. C. B. &amp; Taiti, S." box="[349,556,223,249]" pageId="5" pageNumber="466" refId="ref4257" refString="Boyko, C. B., Bruce, N. L., Hadfield, K. A., Merrin, K. L., Ota, Y., Poore, G. C. B. &amp; Taiti, S. (Eds.) (2008 onwards). World Marine, Freshwater and Terrestrial Isopod Crustaceans database. Ione Latreille, 1818. Accessed through: World Register of Marine Species. Available from: https: // www. marinespecies. org / aphia. php? p = taxdetails &amp; id = 118178 (accessed 21 December 2022)" type="url" year="2008">
Boyko 
<emphasis id="27B9AF5AF65CEC339EF79EE61E89FF16" box="[433,490,223,249]" italics="true" pageId="5" pageNumber="466">et al.</emphasis>
2008
</bibRefCitation>
; for details on synonymy, see also 
<bibRefCitation id="715C0EB9F65CEC339CF39EE61BE0FF16" author="Boyko, C. B. &amp; Williams, J. D. &amp; Shields, J. D." box="[949,1155,223,249]" pageId="5" pageNumber="466" pagination="251 - 301" refId="ref4429" refString="Boyko, C. B., Williams, J. D. &amp; Shields, J. D. (2017) Parasites (Isopoda: Epicaridea and Nematoda) from ghost and mud shrimp (Decapoda: Axiidea and Gebiidea) with descriptions of a new genus and a new species of bopyrid isopod and clarification of Pseudione Kossmann, 1881. Zootaxa, 4365 (3), 251 - 301. https: // doi. org / 10.11646 / zootaxa. 4365.3.1" type="journal article" year="2017">
Boyko 
<emphasis id="27B9AF5AF65CEC339B419EE61B5AFF16" box="[1031,1081,223,249]" italics="true" pageId="5" pageNumber="466">et al</emphasis>
. 2017
</bibRefCitation>
), 
<emphasis id="27B9AF5AF65CEC339BD19EE61BD5FF17" box="[1175,1206,223,248]" italics="true" pageId="5" pageNumber="466">viz</emphasis>
: 
<taxonomicName id="D2CD08CBF65CEC339B839EE61A57FF17" authorityName="Bate" authorityYear="1863" box="[1221,1332,223,248]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="27B9AF5AF65CEC339B839EE61A57FF17" box="[1221,1332,223,248]" italics="true" pageId="5" pageNumber="466">I. cornuta</emphasis>
</taxonomicName>
(parasite of 
<taxonomicName id="D2CD08CBF65CEC339FF09F3A1D45FEF2" authority="(Dana, 1852))" authorityName="" baseAuthorityName="Dana" baseAuthorityYear="1852" box="[182,550,258,285]" class="Malacostraca" family="Callianassidae" genus="Neotrypaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="27B9AF5AF65CEC339FF09F3A1E1DFEF3" box="[182,382,259,284]" italics="true" pageId="5" pageNumber="466">Neotrypaea gigas</emphasis>
(
<bibRefCitation id="715C0EB9F65CEC339ECB9F3A1D70FEF2" author="Dana, J. D." box="[397,531,258,285]" pageId="5" pageNumber="466" refId="ref4516" refString="Dana, J. D. (1852) United States Exploring Expedition. During the Years 1838, 1839, 1840, 1841, 1842. Under the Command of Charles Wilkes, U. S. N. Crustacea Part I. Vol. 13. C. Sherman, Philadelphia." type="journal volume" year="1852">Dana, 1852</bibRefCitation>
))
</taxonomicName>
, 
<taxonomicName id="D2CD08CBF65CEC339D779F3A1DE4FEF3" authorityName="Shiino" authorityYear="1964" box="[561,647,259,284]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="27B9AF5AF65CEC339D779F3A1DE4FEF3" box="[561,647,259,284]" italics="true" pageId="5" pageNumber="466">I. ovata</emphasis>
</taxonomicName>
(parasite of 
<taxonomicName id="D2CD08CBF65CEC339C519F3A1BB4FEF2" authority="(H. Milne Edwards, 1837))" authorityName="" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1837" box="[791,1239,259,285]" class="Malacostraca" family="Callianassidae" genus="Neotrypaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="uncinata">
<emphasis id="27B9AF5AF65CEC339C519F3A1CF9FEF3" box="[791,922,259,284]" italics="true" pageId="5" pageNumber="466">N. uncinata</emphasis>
(H. Milne Edwards, 1837))
</taxonomicName>
, 
<taxonomicName id="D2CD08CBF65CEC339BA49F3A1A36FEF2" authority="Hernáez &amp; Villegas-Castro &amp; Boyko, 2023" authorityName="Hernáez &amp; Villegas-Castro &amp; Boyko" authorityYear="2023" box="[1250,1365,259,285]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="sarahae" status="sp. nov.">
<emphasis id="27B9AF5AF65CEC339BA49F3A1A36FEF2" box="[1250,1365,259,285]" italics="true" pageId="5" pageNumber="466">I. sarahae</emphasis>
</taxonomicName>
<emphasis id="27B9AF5AF65CEC339A1A9F3A1AFFFEF2" bold="true" box="[1372,1436,259,285]" pageId="5" pageNumber="466">
<taxonomicNameLabel id="3C8A1221F65CEC339A1A9F3A1AFFFEF2" box="[1372,1436,259,285]" pageId="5" pageNumber="466" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(parasite of 
<taxonomicName id="D2CD08CBF65CEC339E659F1E1EDCFEAE" baseAuthorityName="Sakai" baseAuthorityYear="2005" box="[291,447,295,321]" class="Malacostraca" family="Callianassidae" genus="Neotrypaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="tabogensis">
<emphasis id="27B9AF5AF65CEC339E659F1E1EDCFEAE" box="[291,447,295,321]" italics="true" pageId="5" pageNumber="466">N. tabogensis</emphasis>
</taxonomicName>
), 
<taxonomicName id="D2CD08CBF65CEC339E9F9F1E1D14FEAF" authorityName="Markham" authorityYear="1995" box="[473,631,295,320]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="27B9AF5AF65CEC339E9F9F1E1D14FEAF" box="[473,631,295,320]" italics="true" pageId="5" pageNumber="466">I. taiwanensis</emphasis>
</taxonomicName>
(parasite of 
<taxonomicName id="D2CD08CBF65CEC339C4B9F1E1B94FEAE" authority="H. Milne Edwards, 1837" authorityName="H. Milne Edwards" authorityYear="1837" box="[781,1271,295,321]" class="Malacostraca" family="Callianideidae" genus="Callianidea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="typa">
<emphasis id="27B9AF5AF65CEC339C4B9F1E1CA8FEAF" box="[781,971,295,321]" italics="true" pageId="5" pageNumber="466">Callianidea typa</emphasis>
H. Milne Edwards, 1837
</taxonomicName>
), 
<taxonomicName id="D2CD08CBF65CEC339A4B9F1E1AFEFEAE" authorityName="Richardson" authorityYear="1904" box="[1293,1437,295,321]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="thompsoni">
<emphasis id="27B9AF5AF65CEC339A4B9F1E1AFEFEAE" box="[1293,1437,295,321]" italics="true" pageId="5" pageNumber="466">I. thompsoni</emphasis>
</taxonomicName>
(parasite of 
<taxonomicName id="D2CD08CBF65CEC339E639F721DB5FE8A" authority="(DeKay, 1844))" baseAuthorityName="DeKay" baseAuthorityYear="1844" box="[293,726,331,357]" class="Malacostraca" family="Callianassidae" genus="Gilvossius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="setimanus">
<emphasis id="27B9AF5AF65CEC339E639F721D77FE8B" box="[293,532,331,357]" italics="true" pageId="5" pageNumber="466">Gilvossius setimanus</emphasis>
(
<bibRefCitation id="715C0EB9F65CEC339D619F721DA0FE8A" author="DeKay, J. E." box="[551,707,331,357]" pageId="5" pageNumber="466" refId="ref4570" refString="DeKay, J. E. (1844) Zoology of New-York, or the New-York fauna; comprising detailed descriptions of all the animals hitherto observed within the state of New-York, with brief notices of those occasionally found near its borders, and accompanied by appropriate illustrations. Part VI. Crustacea. Carroll and Cook, Albany." type="book" year="1844">DeKay, 1844</bibRefCitation>
))
</taxonomicName>
, 
<taxonomicName id="D2CD08CBF65CEC339DA39F721C08FE8A" baseAuthorityName="Montagu" baseAuthorityYear="1808" box="[741,875,331,357]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="thoracica">
<emphasis id="27B9AF5AF65CEC339DA39F721C08FE8A" box="[741,875,331,357]" italics="true" pageId="5" pageNumber="466">I. thoracica</emphasis>
</taxonomicName>
(parasite of 
<taxonomicName id="D2CD08CBF65CEC339B429F721F86FE66" authority="(Montagu, 1808))" baseAuthorityName="Montagu" baseAuthorityYear="1808" class="Malacostraca" family="Callianassidae" genus="Callianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis id="27B9AF5AF65CEC339B429F721A7FFE8A" box="[1028,1308,331,357]" italics="true" pageId="5" pageNumber="466">Callianassa subterranea</emphasis>
(
<bibRefCitation id="715C0EB9F65CEC339A689F721FB2FE66" author="Montagu, G." pageId="5" pageNumber="466" pagination="81 - 114" refId="ref5604" refString="Montagu, G. (1808) Description of several marine animals found on the south coast of Devonshire. Transactions of the Linnean Society of London, 9, 81 - 114, pls. 2 - 8. https: // doi. org / 10.1111 / j. 1096 - 3642.1818. tb 00327. x" type="journal article" year="1808">Montagu, 1808</bibRefCitation>
))
</taxonomicName>
, and 
<taxonomicName id="D2CD08CBF65CEC339E669F561EF6FE66" authorityName="Bourdon" authorityYear="1976" box="[288,405,367,393]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="tubulata">
<emphasis id="27B9AF5AF65CEC339E669F561EF6FE66" box="[288,405,367,393]" italics="true" pageId="5" pageNumber="466">I. tubulata</emphasis>
</taxonomicName>
(host unknown). The new species described herein raises to two (with 
<taxonomicName id="D2CD08CBF65CEC339BF09F561A6FFE67" authorityName="Shiino" authorityYear="1964" box="[1206,1292,367,392]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="27B9AF5AF65CEC339BF09F561A6FFE67" box="[1206,1292,367,392]" italics="true" pageId="5" pageNumber="466">I. ovata</emphasis>
</taxonomicName>
) the number of eastern Pacific species of 
<taxonomicName id="D2CD08CBF65CEC339E939FAA1D64FE43" authorityName="Latreille" authorityYear="1818" box="[469,519,403,428]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="genus">
<emphasis id="27B9AF5AF65CEC339E939FAA1D64FE43" box="[469,519,403,428]" italics="true" pageId="5" pageNumber="466">Ione</emphasis>
</taxonomicName>
, and is the only species known from the Pacific coast of 
<collectingCountry id="6DDA33D8F65CEC339BC69FAA1B9AFE42" box="[1152,1273,403,429]" name="Costa Rica" pageId="5" pageNumber="466">Costa Rica</collectingCountry>
and the whole of the tropical Eastern Pacific.
</paragraph>
<paragraph id="15727348F65CEC339F819FE21BEAFCFA" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="466">
Morphologically, the new species differs from the six other congeners by the following combination of characters: (i) body of female slightly asymmetrical in 
<taxonomicName id="D2CD08CBF65CEC339C5A9FC61CD1FDF6" authority="Hernáez &amp; Villegas-Castro &amp; Boyko, 2023" authorityName="Hernáez &amp; Villegas-Castro &amp; Boyko" authorityYear="2023" box="[796,946,511,537]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="sarahae" status="sp. nov.">
<emphasis id="27B9AF5AF65CEC339C5A9FC61CD1FDF6" box="[796,946,511,537]" italics="true" pageId="5" pageNumber="466">Ione sarahae</emphasis>
</taxonomicName>
<emphasis id="27B9AF5AF65CEC339CFB9FC61C9CFDF6" bold="true" box="[957,1023,511,537]" pageId="5" pageNumber="466">
<taxonomicNameLabel id="3C8A1221F65CEC339CFB9FC61C9CFDF6" box="[957,1023,511,537]" pageId="5" pageNumber="466" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, but symmetrical in 
<taxonomicName id="D2CD08CBF65CEC339BB39FC61A2DFDF7" authorityName="Shiino" authorityYear="1964" box="[1269,1358,511,536]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="27B9AF5AF65CEC339BB39FC61A2DFDF7" box="[1269,1358,511,536]" italics="true" pageId="5" pageNumber="466">I. ovata</emphasis>
</taxonomicName>
and 
<taxonomicName id="D2CD08CBF65CEC339ACA9FC61F97FDD2" authorityName="Bourdon" authorityYear="1976" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="tubulata">
<emphasis id="27B9AF5AF65CEC339ACA9FC61F97FDD2" italics="true" pageId="5" pageNumber="466">I. tubulata</emphasis>
</taxonomicName>
; (ii) the posterior margin of the head of female is nearly straight in the new species, whereas it is evenly rounded in 
<taxonomicName id="D2CD08CBF65CEC339E5E9C7E1EC6FD8E" authorityName="Richardson" authorityYear="1904" box="[280,421,583,609]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="thompsoni">
<emphasis id="27B9AF5AF65CEC339E5E9C7E1EC6FD8E" box="[280,421,583,609]" italics="true" pageId="5" pageNumber="466">I. thompsoni</emphasis>
</taxonomicName>
, and 
<taxonomicName id="D2CD08CBF65CEC339EA29C7E1D04FD8E" baseAuthorityName="Montagu" baseAuthorityYear="1808" box="[484,615,583,609]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="thoracica">
<emphasis id="27B9AF5AF65CEC339EA29C7E1D04FD8E" box="[484,615,583,609]" italics="true" pageId="5" pageNumber="466">I. thoracica</emphasis>
</taxonomicName>
; (iii) the antenna of the female is composed of five articles in 
<taxonomicName id="D2CD08CBF65CEC339A6C9C7E1AFFFD8E" authority="Hernáez &amp; Villegas-Castro &amp; Boyko, 2023" authorityName="Hernáez &amp; Villegas-Castro &amp; Boyko" authorityYear="2023" box="[1322,1436,583,609]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="sarahae" status="sp. nov.">
<emphasis id="27B9AF5AF65CEC339A6C9C7E1A50FD8F" box="[1322,1331,583,608]" italics="true" pageId="5" pageNumber="466">I</emphasis>
. 
<emphasis id="27B9AF5AF65CEC339A049C7E1AFFFD8E" box="[1346,1436,583,609]" italics="true" pageId="5" pageNumber="466">sarahae</emphasis>
</taxonomicName>
<emphasis id="27B9AF5AF65CEC339FD19C521FB6FD6A" bold="true" box="[151,213,619,645]" pageId="5" pageNumber="466">
<taxonomicNameLabel id="3C8A1221F65CEC339FD19C521FB6FD6A" box="[151,213,619,645]" pageId="5" pageNumber="466" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, whereas it is composed of four or six articles in 
<taxonomicName id="D2CD08CBF65CEC339DBF9C521CE5FD6A" authorityName="Montagu" authorityYear="1808" box="[761,902,619,645]" class="Malacostraca" family="Oniscidae" genus="Oniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="thoracicus">
<emphasis id="27B9AF5AF65CEC339DBF9C521CE5FD6A" box="[761,902,619,645]" italics="true" pageId="5" pageNumber="466">I. thoracicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="D2CD08CBF65CEC339CFA9C521B35FD6B" authorityName="Markham" authorityYear="1995" box="[956,1110,619,644]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="27B9AF5AF65CEC339CFA9C521B35FD6B" box="[956,1110,619,644]" italics="true" pageId="5" pageNumber="466">I. taiwanensis</emphasis>
</taxonomicName>
, respectively; (iv) the frontal lamina of the female bears lateral leaflike projections in the new species, instead the posteriorly curved horn-shape projections present in 
<taxonomicName id="D2CD08CBF65CEC339EC89C8A1E9FFD23" authorityName="Bate" authorityYear="1863" box="[398,508,691,716]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="27B9AF5AF65CEC339EC89C8A1E9FFD23" box="[398,508,691,716]" italics="true" pageId="5" pageNumber="466">I. cornuta</emphasis>
</taxonomicName>
; (v) the pleotelson of the male is truncated in the new species, whereas it is distally tapered in both 
<taxonomicName id="D2CD08CBF65CEC339E0C9CEE1EBBFD1E" authorityName="Richardson" authorityYear="1904" box="[330,472,727,753]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="thompsoni">
<emphasis id="27B9AF5AF65CEC339E0C9CEE1EBBFD1E" box="[330,472,727,753]" italics="true" pageId="5" pageNumber="466">I. thompsoni</emphasis>
</taxonomicName>
and 
<taxonomicName id="D2CD08CBF65CEC339D579CEE1DF6FD1E" baseAuthorityName="Montagu" baseAuthorityYear="1808" box="[529,661,727,753]" class="Malacostraca" family="Ionidae" genus="Ione" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="thoracica">
<emphasis id="27B9AF5AF65CEC339D579CEE1DF6FD1E" box="[529,661,727,753]" italics="true" pageId="5" pageNumber="466">I. thoracica</emphasis>
</taxonomicName>
(
<bibRefCitation id="715C0EB9F65CEC339DE39CEE1C27FD1E" author="Bonnier, J." box="[677,836,727,753]" pageId="5" pageNumber="466" pagination="1 - 475" refId="ref4181" refString="Bonnier, J. (1900) Contribution a l'etude des epicarides. Les Bopyridae. Travaux de la Station Zoologique de Wimereux, 8, 1 - 475, pls. 1 - 41. https: // doi. org / 10.5962 / bhl. title. 10522" type="journal article" year="1900">Bonnier 1900</bibRefCitation>
, pl. 1, fig. 2, 6; 
<bibRefCitation id="715C0EB9F65CEC339CB89CEE1BA3FD1E" author="Richardson, H." box="[1022,1216,727,753]" pageId="5" pageNumber="466" pagination="1 - 89" refId="ref5701" refString="Richardson, H. (1904) Contributions to the natural history of the Isopoda. Proceedings of the United States National Museum, 27, 1 - 89. https: // doi. org / 10.5479 / si. 00963801.27 - 1369.657" type="journal article" year="1904">Richardson 1904</bibRefCitation>
, fig. 64, 68; 
<bibRefCitation id="715C0EB9F65CEC339A159CEE1FB1FCFA" author="Shiino, S. M." pageId="5" pageNumber="466" pagination="27 - 32" refId="ref5830" refString="Shiino, S. M. (1964) On two species of bopyrid isopods parasitic on Callianassa uncinata Milne-Edwards from Chile. Report of the Faculty of Fisheries. Prefectural University of Mie, 5 (1), 27 - 32." type="journal article" year="1964">Shiino 1964</bibRefCitation>
, fig. 2a; 
<bibRefCitation id="715C0EB9F65CEC339E729CC21EBBFCFA" author="Bourdon, R." box="[308,472,763,789]" pageId="5" pageNumber="466" pagination="353 - 392" refId="ref4231" refString="Bourdon, R. (1976) Epicarides de Madagascar. I. Bulletin du Museum National d'Histoire Naturelle, 371, 353 - 392." type="journal article" year="1976">Bourdon 1976</bibRefCitation>
, fig. 17; 
<bibRefCitation id="715C0EB9F65CEC339D7C9CC21D8AFCFA" author="Markham, J. C." box="[570,745,762,789]" pageId="5" pageNumber="466" pagination="83 - 89" refId="ref5505" refString="Markham, J. C. (1995) Two new species of Isopoda Bopyridae (Crustacea) infesting thalassinideans in the western Pacific. Raffles Bulletin of Zoology, 43 (1), 83 - 89." type="journal article" year="1995">Markham 1995</bibRefCitation>
, fig. 1c; 
<figureCitation id="8DF66FCDF65CEC339C0D9CC21CCEFCFA" box="[843,941,763,789]" captionStart="FIGURE 2" captionStartId="3.[151,250,1276,1300]" captionTargetBox="[192,1435,192,1251]" captionTargetId="figure-18@3.[151,1435,181,1253]" captionTargetPageId="3" captionText="FIGURE 2. Ione sarahae n. sp., holotype, female, 6.6 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view. Scale bars: A, B = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8036047" httpUri="https://zenodo.org/record/8036047/files/figure.png" pageId="5" pageNumber="466">Figs. 2A</figureCitation>
, 
<figureCitation id="8DF66FCDF65CEC339CFF9CC21CBFFCFA" box="[953,988,763,789]" captionStart="FIGURE 3" captionStartId="4.[151,250,1575,1599]" captionTargetBox="[178,1414,185,1549]" captionTargetId="figure-26@4.[170,1416,181,1551]" captionTargetPageId="4" captionText="FIGURE 3. Ione sarahae n. sp., paratype, female, 4.3 mm TL (MZUCR 3791-01). A, right antennula; B, right antenna; C, barbula; D, right maxilliped; E, right pereopod 1; F, right pereopod 7; G, right oostegite 1, external view; H, right oostegite 1, internal view. Scale bars: A, B, E, F = 100 µm; C = 500 µm; G, H = 1 mm; D = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.8036049" httpUri="https://zenodo.org/record/8036049/files/figure.png" pageId="5" pageNumber="466">3B</figureCitation>
, 
<figureCitation id="8DF66FCDF65CEC339CAE9CC21B48FCFA" box="[1000,1067,763,789]" captionStart="FIGURE 4" captionStartId="5.[151,250,1977,2001]" captionTargetBox="[169,1421,828,1951]" captionTargetId="figure-356@5.[164,1423,813,1954]" captionTargetPageId="5" captionText="FIGURE 4. Ione sarahae n. sp., allotype, male, 3.5 mm TL (MZUCR 3789-01). A, habitus, dorsal view; B, habitus, ventral view; C, left antennula; D, left antenna; E, left pereopod 1. Scale bars: A, B = 250 µm; C, D = 50 µm; E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.8036051" httpUri="https://zenodo.org/record/8036051/files/figure.png" pageId="5" pageNumber="466">4A, B</figureCitation>
herein).
</paragraph>
</subSubSection>
</treatment>
</document>