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<document id="A4CCBC9B81F5D3B561C7A07F7FD2A29D" ID-CLB-Dataset="20950" ID-DOI="10.5852/ejt.2020.717.1095" ID-GBIF-Dataset="b9fb54f7-1883-4a6d-99b4-76f855a4d1cd" ID-Zenodo-Dep="4043697" ID-ZooBank="1358E5B2-056C-4DF2-9C4E-0189E1086712" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1600783222892" checkinUser="valdenar" docAuthor="Ševčík, Jan, Burdíková, Nikola, Kaspřák, David &amp; Kurina, Olavi" docDate="2020" docId="5012270FFFADFD14BC5EEFA40307F9DF" docLanguage="en" docName="ejt-717_sevcik_burdikova_kasprak.pdf.imf" docOrigin="European Journal of Taxonomy 717" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Docosia anatolica Sevcik 2020, sp. nov." docType="treatment" docUuid="7E7CA4EE-F77C-4B6F-87FA-A42B0F0B5A35" docUuidSource="ZooBank" docVersion="10" lastPageNumber="10" masterDocId="AC2B5F77FFA8FD13BE0FEC6D023EFFE6" masterDocTitle="Five new Palaearctic species of Docosia (Diptera: Mycetophilidae), with updated molecular phylogeny of the genus" masterLastPageNumber="26" masterPageNumber="3" pageNumber="8" updateTime="1698845880826" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C838D232CCC3A87D1E0CD7EC2A3356A4">Five new Palaearctic species of Docosia (Diptera: Mycetophilidae), with updated molecular phylogeny of the genus</mods:title>
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<mods:namePart id="E7BC633FDDF0D4804170257DF83D0136">Ševčík, Jan</mods:namePart>
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<mods:affiliation id="06E44DFF4B864A17EA6ED319A8BACEE8">University of Ostrava, Faculty of Science, Department of Biology and Ecology, Chittussiho 10, CZ- 710 00 Ostrava, Czech Republic. Silesian Museum, Nádražní okruh 31, CZ- 746 01 Opava, Czech Republic. Institute of Agricultural and Environmental Sciences, Estonian University of Life Sciences, Kreutzwaldi st 5 D, 51006 Tartu, Estonia.</mods:affiliation>
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<mods:namePart id="3C7FF2E0709E7EAA96325511AC122F0E">Burdíková, Nikola</mods:namePart>
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<mods:affiliation id="9765D720736A1EC066BBDDBC7C51A287">University of Ostrava, Faculty of Science, Department of Biology and Ecology, Chittussiho 10, CZ- 710 00 Ostrava, Czech Republic. Silesian Museum, Nádražní okruh 31, CZ- 746 01 Opava, Czech Republic. Institute of Agricultural and Environmental Sciences, Estonian University of Life Sciences, Kreutzwaldi st 5 D, 51006 Tartu, Estonia.</mods:affiliation>
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<mods:namePart id="7E2042068F88098886F7E9F42D9E26D5">Kaspřák, David</mods:namePart>
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<mods:affiliation id="6FF426D2D09B3070A465FB7BB1944F63">University of Ostrava, Faculty of Science, Department of Biology and Ecology, Chittussiho 10, CZ- 710 00 Ostrava, Czech Republic. Silesian Museum, Nádražní okruh 31, CZ- 746 01 Opava, Czech Republic. Institute of Agricultural and Environmental Sciences, Estonian University of Life Sciences, Kreutzwaldi st 5 D, 51006 Tartu, Estonia.</mods:affiliation>
<mods:nameIdentifier id="81C6304C1FC6961533D3A2E3E0D37C36" type="email">davidkasprak@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B9CECC07CF5D86917E2CC9237812BF09">Kurina, Olavi</mods:namePart>
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<mods:affiliation id="87BCB23BE9AE9235BBEB40F46C0EE299">Institute of Agricultural and Environmental Sciences, Estonian University of Life Sciences, Kreutzwaldi st 5 D, 51006 Tartu, Estonia.</mods:affiliation>
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<mods:date id="1CEF5CE6334263CEAF27976606997234">2020</mods:date>
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<treatment id="5012270FFFADFD14BC5EEFA40307F9DF" ID-DOI="http://doi.org/10.5281/zenodo.4331807" ID-GBIF-Taxon="167778118" ID-Zenodo-Dep="4331807" LSID="urn:lsid:zoobank.org:act:7E7CA4EE-F77C-4B6F-87FA-A42B0F0B5A35" httpUri="http://treatment.plazi.org/id/5012270FFFADFD14BC5EEFA40307F9DF" lastPageId="7" lastPageNumber="10" pageId="5" pageNumber="8">
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<paragraph id="D8049619FFADFD16BC5EEFA401DDFC05" blockId="5.[360,1227,968,1070]" box="[593,995,968,997]" pageId="5" pageNumber="8">
<heading id="834C2175FFADFD16BC5EEFA401DDFC05" box="[593,995,968,997]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="8" reason="2">
<taxonomicName id="1FBBED9AFFADFD16BC5EEFA401BFFC03" ID-CoL="9HHZL" authority="Sevcik" authorityName="Sevcik" authorityYear="2020" box="[593,897,968,997]" class="Insecta" family="Mycetophilidae" genus="Docosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="8" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="EACF4A0BFFADFD16BC5EEFA40114FC05" bold="true" box="[593,810,968,995]" italics="true" pageId="5" pageNumber="8">Docosia anatolica</emphasis>
Ševčík
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<taxonomicNameLabel id="F1FCF770FFADFD16BD86EFA401DDFC05" box="[905,995,969,995]" pageId="5" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<paragraph id="D8049619FFADFD16BCD2E8790166FBC8" blockId="5.[360,1227,968,1070]" box="[733,856,1044,1070]" pageId="5" pageNumber="8">
<figureCitation id="40808A9CFFADFD16BCD2E8790116FBC8" box="[733,808,1044,1070]" captionStart="Fig" captionStartId="6.[189,232,1726,1752]" captionTargetBox="[202,1360,266,1681]" captionTargetId="figure-16@6.[189,1398,265,1688]" captionTargetPageId="6" captionText="Fig. 2. Docosia anatolica Ševčík sp. nov., ♂, terminalia. A. Lateral view. B. Posterior view. C. Ventral view of gonostylus. D. Dorsal view of cerci. E. Dorsal view of tergite 9. F. Ventral view of aedeagal complex. G. Lateral view of aedeagal complex. Abbreviations: aed = aedeagus; ej ap = ejaculatory apodeme; gc = gonocoxite; gc pvm = posteroventral margin of gonocoxite; gst vl = ventral lobe of gonostylus; gst dl = dorsal lobe of gonostylus; par = paramere; par ap = parameral apodeme; ret = combs of retinacula; tg = tergite. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4043701" httpUri="https://zenodo.org/record/4043701/files/figure.png" pageId="5" pageNumber="8">Figs 2</figureCitation>
,
<figureCitation id="40808A9CFFADFD16BD3BE8790166FBC8" box="[820,856,1044,1070]" captionStart="Fig" captionStartId="17.[189,232,1878,1904]" captionTargetBox="[189,1398,474,1835]" captionTargetId="figure-65@17.[189,1398,474,1840]" captionTargetPageId="17" captionText="Fig. 7. Male terminalia, ventroposterior view. A. Docosia anatolica Ševčík sp. nov. B. D. japonica Kurina sp. nov. C. D. polyspina Kurina sp. nov. D. D. peloponnensis Ševčík sp. nov. E. D. svanetica Kurina sp. nov. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4043711" httpUri="https://zenodo.org/record/4043711/files/figure.png" pageId="5" pageNumber="8">7A</figureCitation>
</paragraph>
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<paragraph id="D8049619FFADFD16BEB2E83A03F8FB97" blockId="5.[189,454,1111,1137]" box="[189,454,1111,1137]" pageId="5" pageNumber="8">
<heading id="834C2175FFADFD16BEB2E83A03F8FB97" bold="true" box="[189,454,1111,1137]" fontSize="11" level="3" pageId="5" pageNumber="8" reason="3">
<emphasis id="EACF4A0BFFADFD16BEB2E83A03F8FB97" bold="true" box="[189,454,1111,1137]" pageId="5" pageNumber="8">Differential diagnosis</emphasis>
</heading>
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<paragraph id="D8049619FFADFD16BEB2E8E40341FA13" blockId="5.[189,1399,1161,1525]" pageId="5" pageNumber="8">
By structure of the male terminalia,
<taxonomicName id="1FBBED9AFFADFD16BC7FE8E4013BFB45" authority="Ševčík &amp; Burdíková &amp; Kaspřák &amp; Kurina, 2020" authorityName="Ševčík &amp; Burdíková &amp; Kaspřák &amp; Kurina" authorityYear="2020" box="[624,773,1161,1187]" class="Insecta" family="Mycetophilidae" genus="Docosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="8" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="EACF4A0BFFADFD16BC7FE8E4013BFB45" box="[624,773,1161,1187]" italics="true" pageId="5" pageNumber="8">D. anatolica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1FCF770FFADFD16BD02E8E40154FB45" box="[781,874,1161,1187]" pageId="5" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="1FBBED9AFFADFD16BDFEE8E40749FB45" authority="Chandler, 2004" authorityName="Chandler" authorityYear="2004" box="[1009,1399,1161,1187]" class="Insecta" family="Mycetophilidae" genus="Docosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="8" phylum="Arthropoda" rank="species" species="juxtamontana">
<emphasis id="EACF4A0BFFADFD16BDFEE8E40686FB44" box="[1009,1208,1161,1186]" italics="true" pageId="5" pageNumber="8">D. juxtamontana</emphasis>
Chandler, 2004
</taxonomicName>
and
<taxonomicName id="1FBBED9AFFADFD16BEFDE8C20046FB2E" authority="Chandler, 2004" authorityName="Chandler" authorityYear="2004" box="[242,632,1198,1224]" class="Insecta" family="Mycetophilidae" genus="Docosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="8" phylum="Arthropoda" rank="species" species="incolamontis">
<emphasis id="EACF4A0BFFADFD16BEFDE8C2038CFB2E" box="[242,434,1198,1224]" italics="true" pageId="5" pageNumber="8">D. incolamontis</emphasis>
Chandler, 2004
</taxonomicName>
. All species have a bifid gonostylus and posteroventral margin of gonocoxites with a prominent medial process surrounded by two small lobes bearing setae. These setae are apically ramified in
<taxonomicName id="1FBBED9AFFADFD16BC29E8970082FAF5" authority="Ševčík &amp; Burdíková &amp; Kaspřák &amp; Kurina, 2020" authorityName="Ševčík &amp; Burdíková &amp; Kaspřák &amp; Kurina" authorityYear="2020" box="[550,700,1273,1299]" class="Insecta" family="Mycetophilidae" genus="Docosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="8" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="EACF4A0BFFADFD16BC29E8970082FAF5" box="[550,700,1273,1299]" italics="true" pageId="5" pageNumber="8">D. anatolica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1FCF770FFADFD16BCC9E8970119FAF2" box="[710,807,1274,1300]" pageId="5" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
, but with an unknown structure in the other two species. Apically modified setae at the posteroventral margin of gonocoxites are frequent in Central- Asian species (cf.
<bibRefCitation id="BC2AEBE8FFADFD16BF91E929008CFAB8" author="Kurina &amp; Sevcik" box="[414,690,1348,1377]" firstAuthor="Kurina" pageId="5" pageNumber="8" pagination="25 - 40" refId="ref9730" refString="Kurina O. &amp; Sevcik J. 2012. Notes on Docosia Winnertz (Diptera: Mycetophilidae), with description of six new species from Central Asia and the first generic record from the Afrotropical region. Zootaxa 3570: 25 - 40. https: // doi. org / 10.11646 / zootaxa. 3570.1.2" type="journal article" year="2012">Kurina &amp; Ševčík 2012</bibRefCitation>
) but they are also observable in other species, e.g., in the European
<taxonomicName id="1FBBED9AFFADFD16BF38E906012BFA62" authority="Lastovka &amp; Sevcik, 2006" authorityName="Lastovka &amp; Sevcik" authorityYear="2006" box="[311,789,1386,1414]" class="Insecta" family="Mycetophilidae" genus="Docosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="8" phylum="Arthropoda" rank="species" species="pannonica">
<emphasis id="EACF4A0BFFADFD16BF38E90603E6FA62" box="[311,472,1387,1412]" italics="true" pageId="5" pageNumber="8">D. pannonica</emphasis>
<bibRefCitation id="BC2AEBE8FFADFD16BFEEE907012BFA62" author="Lastovka &amp; Sevcik" box="[481,789,1386,1414]" firstAuthor="Lastovka" pageId="5" pageNumber="8" pagination="1 - 37" refId="ref9787" refString="Lastovka P. &amp; Sevcik J. 2006. A review of the Czech and Slovak species of Docosia Winnertz (Diptera: Mycetophilidae), with atlas of the male and female terminalia. Casopis Slezskeho zemskeho muzea Opava (A) 55: 1 - 37." type="journal article" year="2006">Laštovka &amp; Ševčík, 2006</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="1FBBED9AFFADFD16BD2DE90601C2FA62" authority="Ševčík &amp; Burdíková &amp; Kaspřák &amp; Kurina, 2020" authorityName="Ševčík &amp; Burdíková &amp; Kaspřák &amp; Kurina" authorityYear="2020" box="[802,1020,1386,1412]" class="Insecta" family="Mycetophilidae" genus="Docosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="8" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="EACF4A0BFFADFD16BD2DE90601C2FA62" box="[802,1020,1386,1412]" italics="true" pageId="5" pageNumber="8">Docosia anatolica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1FCF770FFADFD16BA0AE907065FFA62" box="[1029,1121,1386,1412]" pageId="5" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
has the ventral lobe of the gonostylus obtriangular, with spines at both posterior corners, while it is slender and furcated, with medial and apical spines in
<taxonomicName id="1FBBED9AFFADFD16BC0DE9DB0094FA29" box="[514,682,1462,1487]" class="Insecta" family="Mycetophilidae" genus="Docosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="8" phylum="Arthropoda" rank="species" species="juxtamonta">
<emphasis id="EACF4A0BFFADFD16BC0DE9DB0094FA29" box="[514,682,1462,1487]" italics="true" pageId="5" pageNumber="8">D. juxtamonta</emphasis>
</taxonomicName>
and irregularly crescent-shaped with two subapical spines in
<taxonomicName id="1FBBED9AFFADFD16BEB2E9B60347FA12" authorityName="Chandler" authorityYear="2004" box="[189,377,1498,1524]" class="Insecta" family="Mycetophilidae" genus="Docosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="8" phylum="Arthropoda" rank="species" species="incolamontis">
<emphasis id="EACF4A0BFFADFD16BEB2E9B60347FA12" box="[189,377,1498,1524]" italics="true" pageId="5" pageNumber="8">D. incolamontis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="90A1C592FFADFD16BEB2EA7301D2F98C" box="[189,1004,1566,1642]" pageId="5" pageNumber="8" type="etymology">
<paragraph id="D8049619FFADFD16BEB2EA73037AF9DE" blockId="5.[189,324,1566,1592]" box="[189,324,1566,1592]" pageId="5" pageNumber="8">
<heading id="834C2175FFADFD16BEB2EA73037AF9DE" bold="true" box="[189,324,1566,1592]" fontSize="11" level="3" pageId="5" pageNumber="8" reason="3">
<emphasis id="EACF4A0BFFADFD16BEB2EA73037AF9DE" bold="true" box="[189,324,1566,1592]" pageId="5" pageNumber="8">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="D8049619FFADFD16BEB2EA2201D2F98C" blockId="5.[189,1004,1615,1642]" box="[189,1004,1615,1642]" pageId="5" pageNumber="8">
The species name refers to the
<typeStatus id="070028BBFFADFD16BC27EA220065F98F" box="[552,603,1615,1641]" pageId="5" pageNumber="8">type</typeStatus>
locality in
<collectingCountry id="A0ACD689FFADFD16BCD1EA22017DF98F" box="[734,835,1615,1641]" name="Turkey" pageId="5" pageNumber="8">Anatolia</collectingCountry>
(
<collectingCountry id="A0ACD689FFADFD16BD5DEA2201DEF98C" box="[850,992,1615,1642]" name="Turkey" pageId="5" pageNumber="8">Asia Minor</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="90A1C592FFADFD16BEB2EAFE01EFF8A8" pageId="5" pageNumber="8" type="materials_examined">
<paragraph id="D8049619FFADFD16BEB2EAFE0352F94A" blockId="5.[189,364,1682,1709]" box="[189,364,1682,1709]" pageId="5" pageNumber="8">
<heading id="834C2175FFADFD16BEB2EAFE0352F94A" bold="true" box="[189,364,1682,1709]" fontSize="11" level="3" pageId="5" pageNumber="8" reason="3">
<emphasis id="EACF4A0BFFADFD16BEB2EAFE0352F94A" bold="true" box="[189,364,1682,1709]" pageId="5" pageNumber="8">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="68D39C44FFADFD16BEE5EAA901EFF8A8" ID-GBIF-Occurrence="2863261317" collectingDate="2016-04-22" collectingMethod="Malaise trap" collectionCode="MT" collectorName="M. Bartak &amp; S. Kubik" country="Turkey" elevation="1100" location="University Campus" pageId="5" pageNumber="8" specimenCount="1" stateProvince="Mugla" typeStatus="holotype">
<paragraph id="D8049619FFADFD16BEE5EAA90362F938" blockId="5.[189,1398,1732,1871]" box="[234,348,1732,1758]" pageId="5" pageNumber="8">
<emphasis id="EACF4A0BFFADFD16BEE5EAA90362F938" bold="true" box="[234,348,1732,1758]" pageId="5" pageNumber="8">
<typeStatus id="070028BBFFADFD16BEE5EAA90362F938" box="[234,348,1732,1758]" pageId="5" pageNumber="8" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="D8049619FFADFD16BEB2EA8701EFF8A8" blockId="5.[189,1398,1732,1871]" pageId="5" pageNumber="8">
<collectingCountry id="A0ACD689FFADFD16BEB2EA87030DF8E2" box="[189,307,1770,1796]" name="Turkey" pageId="5" pageNumber="8">TURKEY</collectingCountry>
<specimenCount id="CEBD5D90FFADFD16BF40EA8A0358F8E5" box="[335,358,1767,1795]" pageId="5" pageNumber="8" type="male"></specimenCount>
;
<collectingRegion id="1A7F58FBFFADFD16BF7AEA8403FBF8E5" box="[373,453,1769,1795]" country="Turkey" name="Mugla" pageId="5" pageNumber="8">Mugla</collectingRegion>
,
<location id="DD64C0C2FFADFD16BFDDEA840085F8E5" LSID="urn:lsid:plazi:treatment:5012270FFFADFD14BC5EEFA40307F9DF:DD64C0C2FFADFD16BFDDEA840085F8E5" box="[466,699,1769,1795]" country="Turkey" name="University Campus" pageId="5" pageNumber="8" stateProvince="Mugla">University Campus</location>
;
<elevation id="5396712AFFADFD16BCC8EA84015CF8E5" box="[711,866,1769,1796]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="8" unit="m" value="1100.0">
<quantity id="1F433BFCFFADFD16BCC8EA84011FF8E2" box="[711,801,1769,1796]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="8" unit="m" value="1100.0">1100 m</quantity>
a.s.l.
</elevation>
;
<date id="AC05B0D9FFADFD16BD7EEA840631F8E5" box="[881,1039,1769,1795]" pageId="5" pageNumber="8" value="2016-04-22">
<collectingDate id="BC414931FFADFD16BD7EEA840631F8E5" box="[881,1039,1769,1795]" pageId="5" pageNumber="8" value="2016-04-22">22 Apr. 2016</collectingDate>
</date>
;
<collectorName id="754EF3CFFFADFD16BA12EA8706ABF8E5" box="[1053,1173,1769,1796]" pageId="5" pageNumber="8">M. Barták</collectorName>
and
<collectorName id="754EF3CFFFADFD16BADEEA870704F8E5" box="[1233,1338,1769,1798]" pageId="5" pageNumber="8">Š. Kubík</collectorName>
leg.;
<collectingMethod id="01FAEE0EFFADFD16BEB2EB62036EF8CF" box="[189,336,1807,1833]" pageId="5" pageNumber="8">Malaise trap</collectingMethod>
; in a pinned microvial with glycerol; specimen after DNA extraction; GenBank:
<accessionNumber id="C7E80BFAFFADFD16BAE4EB62074CF8CF" box="[1259,1394,1807,1833]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT293019" pageId="5" pageNumber="8">MT293019</accessionNumber>
,
<accessionNumber id="C7E80BFAFFADFD16BEB2EB59037AF8A9" box="[189,324,1844,1871]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT293050" pageId="5" pageNumber="8">MT293050</accessionNumber>
,
<accessionNumber id="C7E80BFAFFADFD16BF5FEB5903E9F8A8" box="[336,471,1844,1870]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT293033" pageId="5" pageNumber="8">MT293033</accessionNumber>
,
<accessionNumber id="C7E80BFAFFADFD16BFEDEB590057F8A8" box="[482,617,1844,1870]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT293067" pageId="5" pageNumber="8">MT293067</accessionNumber>
,
<accessionNumber id="C7E80BFAFFADFD16BC7AEB5900C3F8A8" box="[629,765,1844,1870]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT293001" pageId="5" pageNumber="8">MT293001</accessionNumber>
; NMPC-JSDO26.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="90A1C592FFADFD14BEB2EB150307FA21" lastPageId="7" lastPageNumber="10" pageId="5" pageNumber="8" type="description">
<paragraph id="D8049619FFADFD16BEB2EB150370F874" blockId="5.[189,334,1912,1938]" box="[189,334,1912,1938]" pageId="5" pageNumber="8">
<heading id="834C2175FFADFD16BEB2EB150370F874" bold="true" box="[189,334,1912,1938]" fontSize="11" level="3" pageId="5" pageNumber="8" reason="3">
<emphasis id="EACF4A0BFFADFD16BEB2EB150370F874" bold="true" box="[189,334,1912,1938]" pageId="5" pageNumber="8">Description</emphasis>
</heading>
</paragraph>
<paragraph id="D8049619FFADFD16BEE5EBC403BCF825" blockId="5.[189,673,1961,2025]" box="[234,386,1961,1987]" pageId="5" pageNumber="8">
<emphasis id="EACF4A0BFFADFD16BEE5EBC40314F825" bold="true" box="[234,298,1961,1987]" pageId="5" pageNumber="8">Male</emphasis>
(n = 1)
</paragraph>
<paragraph id="D8049619FFADFD16BEB2EBA3009FF80F" blockId="5.[189,673,1961,2025]" box="[189,673,1998,2025]" pageId="5" pageNumber="8">
MEASUREMENTS. Length of wing
<quantity id="1F433BFCFFADFD16BC4FEBA300A3F80F" box="[576,669,1998,2025]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="5" pageNumber="8" unit="mm" value="4.2">4.2 mm</quantity>
.
</paragraph>
<caption id="8CC4C691FFAEFD15BEB2EAD300D9F86F" ID-DOI="http://doi.org/10.5281/zenodo.4043701" ID-Zenodo-Dep="4043701" httpUri="https://zenodo.org/record/4043701/files/figure.png" pageId="6" pageNumber="9" startId="6.[189,232,1726,1752]" targetBox="[202,1360,266,1681]" targetPageId="6">
<paragraph id="D8049619FFAEFD15BEB2EAD300D9F86F" blockId="6.[189,1399,1724,1929]" pageId="6" pageNumber="9">
<emphasis id="EACF4A0BFFAEFD15BEB2EAD30335F93E" bold="true" box="[189,267,1726,1752]" pageId="6" pageNumber="9">Fig. 2.</emphasis>
<taxonomicName id="1FBBED9AFFAEFD15BF1DEAD2007CF93D" authority="Sevcik" authorityName="Sevcik" authorityYear="2020" box="[274,578,1726,1755]" class="Insecta" family="Mycetophilidae" genus="Docosia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="9" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="EACF4A0BFFAEFD15BF1DEAD203D5F93E" box="[274,491,1726,1752]" italics="true" pageId="6" pageNumber="9">Docosia anatolica</emphasis>
Ševčík
</taxonomicName>
<taxonomicNameLabel id="F1FCF770FFAEFD15BC45EAD20099F93F" box="[586,679,1727,1753]" pageId="6" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, terminalia.
<emphasis id="EACF4A0BFFAEFD15BD50EAD3014AF93E" bold="true" box="[863,884,1726,1752]" pageId="6" pageNumber="9">A</emphasis>
. Lateral view.
<emphasis id="EACF4A0BFFAEFD15BA2BEAD20606F93F" bold="true" box="[1060,1080,1727,1753]" pageId="6" pageNumber="9">B</emphasis>
. Posterior view.
<emphasis id="EACF4A0BFFAEFD15BAF1EAD3072DF93E" bold="true" box="[1278,1299,1726,1752]" pageId="6" pageNumber="9">C</emphasis>
. Ventral view of gonostylus.
<emphasis id="EACF4A0BFFAEFD15BFBFEA8F03FBF91A" bold="true" box="[432,453,1762,1788]" pageId="6" pageNumber="9">D</emphasis>
. Dorsal view of cerci.
<emphasis id="EACF4A0BFFAEFD15BCD4EA8F00D1F91A" bold="true" box="[731,751,1762,1788]" pageId="6" pageNumber="9">E</emphasis>
. Dorsal view of tergite 9.
<emphasis id="EACF4A0BFFAEFD15BA23EA8F0600F91A" bold="true" box="[1068,1086,1762,1788]" pageId="6" pageNumber="9">F</emphasis>
. Ventral view of aedeagal complex.
<emphasis id="EACF4A0BFFAEFD15BF3BEB680375F8F9" bold="true" box="[308,331,1797,1823]" pageId="6" pageNumber="9">G</emphasis>
. Lateral view of aedeagal complex. Abbreviations: aed = aedeagus; ej ap = ejaculatory apodeme; gc = gonocoxite; gc pvm = posteroventral margin of gonocoxite; gst vl = ventral lobe of gonostylus; gst dl = dorsal lobe of gonostylus; par = paramere; par ap = parameral apodeme; ret = combs of retinacula; tg = tergite. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="D8049619FFAFFD14BEB2ED640197FE55" blockId="7.[189,1399,265,435]" pageId="7" pageNumber="10">HEAD. Blackish brown with numerous pale setae. Three ocelli, lateral ones almost touching compound eyes, separated from eye margins by less than half of their own diameter. Mouthparts light brownish. Palpus brownish yellow, basal segment and apical part of last palpomere darker. Scape, pedicel and all flagellomeres dark brown. Flagellomeres cylindrical, flagellomeres 1 and 2 about twice as long as broad, other flagellomeres (3 to 14) about three times as long as broad.</paragraph>
<paragraph id="D8049619FFAFFD14BEB2EDB6073CFDDB" blockId="7.[189,1399,475,573]" pageId="7" pageNumber="10">THORAX.All parts of thorax blackish brown, with light setae. Scutellum with numerous setae and several submarginal pale bristles about twice as long as scutellum. Antepronotum and proepisternum with pale bristles and short darker setae. Laterotergite and other pleural parts bare. Haltere pale yellow.</paragraph>
<paragraph id="D8049619FFAFFD14BEB2EE0803DFFCD5" blockId="7.[189,1399,613,819]" pageId="7" pageNumber="10">LEGS. All coxae yellow, with basal third darkened. Femora mostly yellow, fore and mid femur dark along ventral margin, more extensively in proximal third; similarly hind femur, which is brownish also around tip. Trochanters blackish brown. Fore tibia brownish yellow, apicomedially with simple semicircular tibial organ (anteroapical depressed area), without strong setae, only densely covered with fine setulae. Mid tibia darkened dorsally. Hind tibia mostly brownish yellow, with small black dot approximately in middle of dorsal surface.</paragraph>
<paragraph id="D8049619FFAFFD14BEB2EF360600FC04" blockId="7.[189,1399,859,994]" pageId="7" pageNumber="10">
WINGS. Hyaline, unmarked. Radial veins and r-m dark brown, other veins paler, basal parts of m faint, almost not traceable. Sc, Rs and basal third of cu-stem asetose, other veins setose. Costa reaches to about one third of distance between R
<subScript id="443F945CFFAFFD14BC37EFDB007FFC22" attach="left" box="[568,577,950,964]" fontSize="6" pageId="7" pageNumber="10">5</subScript>
and M
<subScript id="443F945CFFAFFD14BC99EFDB00A1FC22" attach="left" box="[662,671,950,964]" fontSize="6" pageId="7" pageNumber="10">1</subScript>
. Sc ends in R before level of beginning of m-stem. Posterior fork begins before anterior fork, approximately at level of basal third of r-m.
</paragraph>
<paragraph id="D8049619FFAFFD14BEB2E86403C5FBC5" blockId="7.[189,507,1033,1059]" box="[189,507,1033,1059]" pageId="7" pageNumber="10">ABDOMEN. All dark brown.</paragraph>
<paragraph id="D8049619FFAFFD14BEB2E821073EFA87" blockId="7.[189,1399,1099,1378]" pageId="7" pageNumber="10">
TERMINALIA (
<figureCitation id="40808A9CFFAFFD14BF55E826039FFB80" box="[346,417,1099,1126]" captionStart="Fig" captionStartId="6.[189,232,1726,1752]" captionTargetBox="[202,1360,266,1681]" captionTargetId="figure-16@6.[189,1398,265,1688]" captionTargetPageId="6" captionText="Fig. 2. Docosia anatolica Ševčík sp. nov., ♂, terminalia. A. Lateral view. B. Posterior view. C. Ventral view of gonostylus. D. Dorsal view of cerci. E. Dorsal view of tergite 9. F. Ventral view of aedeagal complex. G. Lateral view of aedeagal complex. Abbreviations: aed = aedeagus; ej ap = ejaculatory apodeme; gc = gonocoxite; gc pvm = posteroventral margin of gonocoxite; gst vl = ventral lobe of gonostylus; gst dl = dorsal lobe of gonostylus; par = paramere; par ap = parameral apodeme; ret = combs of retinacula; tg = tergite. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4043701" httpUri="https://zenodo.org/record/4043701/files/figure.png" pageId="7" pageNumber="10">Figs 2</figureCitation>
AE, 7A). Dark brown except lighter gonostyli and posterior margin of gonocoxites. Tergite 9 subrectangular, about 1.5 times as long as broad, slightly broadening posteriorly, posterior margin with medial incision. Posteroventral margin of gonocoxites with blunt lateral projections and prominent rounded process medioventrally, surrounded by two shorter processes bearing apically ramified thick setae. Gonostylus bifid. Ventral lobe of gonostylus obtriangular in ventral view, with dark tooth at both posterior corners. Dorsal lobe of gonostylus larger than ventral lobe, bent inwards and rounded apically, with large aggregation of setulae along ventral surface anteriorly, prominent subapical seta posteriorly and with about 10 setae posterolaterally. Cercus with 16 combs of retinacula.
</paragraph>
<paragraph id="D8049619FFAFFD14BEE5E9E7037BFA42" blockId="7.[189,325,1418,1479]" box="[234,325,1418,1444]" pageId="7" pageNumber="10">
<emphasis id="EACF4A0BFFAFFD14BEE5E9E7037BFA42" bold="true" box="[234,325,1418,1444]" pageId="7" pageNumber="10">Female</emphasis>
</paragraph>
<paragraph id="D8049619FFAFFD14BEB2E9C00307FA21" blockId="7.[189,325,1418,1479]" box="[189,313,1453,1479]" pageId="7" pageNumber="10">Unknown.</paragraph>
</subSubSection>
<subSubSection id="90A1C592FFAFFD14BEB2E9820307F9DF" box="[189,313,1519,1593]" pageId="7" pageNumber="10" type="biology_ecology">
<paragraph id="D8049619FFAFFD14BEB2E9820322F9EF" blockId="7.[189,284,1519,1545]" box="[189,284,1519,1545]" pageId="7" pageNumber="10">
<heading id="834C2175FFAFFD14BEB2E9820322F9EF" bold="true" box="[189,284,1519,1545]" fontSize="11" level="3" pageId="7" pageNumber="10" reason="3">
<emphasis id="EACF4A0BFFAFFD14BEB2E9820322F9EF" bold="true" box="[189,284,1519,1545]" pageId="7" pageNumber="10">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="D8049619FFAFFD14BEB2EA720307F9DF" blockId="7.[189,313,1567,1593]" box="[189,313,1567,1593]" pageId="7" pageNumber="10">Unknown.</paragraph>
</subSubSection>
</treatment>
</document>