treatments-xml/data/4D/2C/C2/4D2CC257FF8A7F2019C612404473FA0E.xml

<document id="2C6C8A8C8642DF92480FD8E395DF108E" ID-CLB-Dataset="25354" ID-DOI="10.11646/zootaxa.4679.1.4" ID-GBIF-Dataset="96b40299-afd6-442d-ad37-898a3728b2d9" ID-ISSN="1175-5326" ID-Zenodo-Dep="3470070" ID-ZooBank="C6356637-AFA5-4553-A5F8-F6821BE73122" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1569910593809" checkinUser="plazi" docAuthor="Starkevich, Pavel, Men, Qiu-Lei &amp; Saldaitis, Aidas" docDate="2019" docId="4D2CC257FF8A7F2019C612404473FA0E" docLanguage="en" docName="zootaxa.4679.1.4.pdf" docOrigin="Zootaxa 4679 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tipula (Vestiplex) maoxianensis Starkevich &amp; Men &amp; Saldaitis 2019, sp. nov." docType="treatment" docUuid="3747C906-EBE5-4A24-8CE7-FC40D2E6C789" docUuidSource="ZooBank" docVersion="10" lastPageNumber="77" masterDocId="B115BA2FFF8F7F28195114754078FF91" masterDocTitle="Four new crane fly species of subgenus Tipula (Vestiplex) (Diptera, Tipulidae) from China" masterLastPageNumber="86" masterPageNumber="69" pageNumber="74" updateTime="1698756632134" updateUser="ExternalLinkService">
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<mods:title id="F65A072D77753FB975DFDA9AEBF41133">Four new crane fly species of subgenus Tipula (Vestiplex) (Diptera, Tipulidae) from China</mods:title>
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<mods:namePart id="66214CEDF7225653842AFF4D76A80E95">Men, Qiu-Lei</mods:namePart>
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<treatment id="4D2CC257FF8A7F2019C612404473FA0E" ID-DOI="http://doi.org/10.5281/zenodo.5935864" ID-GBIF-Taxon="159609610" ID-Zenodo-Dep="5935864" LSID="urn:lsid:zoobank.org:act:3747C906-EBE5-4A24-8CE7-FC40D2E6C789" httpUri="http://treatment.plazi.org/id/4D2CC257FF8A7F2019C612404473FA0E" lastPageId="8" lastPageNumber="77" pageId="5" pageNumber="74">
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<emphasis id="F7F1AF53FF8A7F2D19C612404211F9C1" bold="true" box="[151,617,1589,1616]" pageId="5" pageNumber="74">
<taxonomicName id="028508C2FF8A7F2D19C612404271F9C1" authority="Starkevich &amp; Men &amp; Saldaitis, 2019" authorityName="Starkevich &amp; Men &amp; Saldaitis" authorityYear="2019" box="[151,521,1589,1616]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="maoxianensis" status="sp. nov." subGenus="Vestiplex">
<emphasis id="F7F1AF53FF8A7F2D19C61240409AF9C1" bold="true" box="[151,226,1589,1616]" italics="true" pageId="5" pageNumber="74">Tipula</emphasis>
(
<emphasis id="F7F1AF53FF8A7F2D19A212404122F9C1" bold="true" box="[243,346,1589,1616]" italics="true" pageId="5" pageNumber="74">Vestiplex</emphasis>
) 
<emphasis id="F7F1AF53FF8A7F2D183B12434271F9C1" bold="true" box="[362,521,1590,1616]" italics="true" pageId="5" pageNumber="74">maoxianensis</emphasis>
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</emphasis>
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<subSubSection id="8D9F20CAFF8A7F2D19C6122F4161F9E5" box="[151,281,1626,1652]" pageId="5" pageNumber="74" type="description">
<paragraph id="C53A7341FF8A7F2D19C6122F4161F9E5" blockId="5.[151,617,1589,1652]" box="[151,281,1626,1652]" pageId="5" pageNumber="74">
(
<figureCitation id="5DBE6FC4FF8A7F2D19CE122F4169F9E5" box="[159,273,1626,1652]" captionStart-0="FIGURES 9–10" captionStart-1="FIGURES 11–17" captionStartId-0="6.[151,264,1166,1191]" captionStartId-1="7.[151,264,1837,1862]" captionTargetBox-0="[151,1424,200,1132]" captionTargetBox-1="[156,1420,422,1802]" captionTargetId-0="figure@6.[151,1435,181,1141]" captionTargetId-1="figure@7.[151,1435,417,1812]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 9–10. Tipula (Vestiplex) maoxianensis sp. nov. 9: adult male, general view of paratype; 10: wing, paratype." captionText-1="FIGURES 11–17. Hypopygium of male Tipula (Vestiplex) maoxianensis sp. nov. 11: hypopygium, lateral view; 12: hypopygium, dorsal view; 13: hypopygium, ventral view; 14: gonocoxite, lateral view; 15: left outer gonostylus; 16: left inner gonostylus, lateral view; 17: adminiculum, lateral view. Abbreviations: adm, adminiculum; admr, adminicular rod; gcx, gonocoxite; gph, gonapophysis; ig, inner gonostylus; og, outer gonostylus; st9, sternite 9; t9, tergite 9." figureDoi-0="http://doi.org/10.5281/zenodo.3470076" figureDoi-1="http://doi.org/10.5281/zenodo.3470078" httpUri-0="https://zenodo.org/record/3470076/files/figure.png" httpUri-1="https://zenodo.org/record/3470078/files/figure.png" pageId="5" pageNumber="74">Figs 9–17</figureCitation>
)
</paragraph>
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<paragraph id="C53A7341FF8A7F2D19C612E842A9F94A" blockId="5.[151,1437,1692,2042]" pageId="5" pageNumber="74">
<materialsCitation id="75ED791CFF8A7F2D19C612E842A9F94A" ID-GBIF-Occurrence="2423311343" collectingDate="2017-05-20" collectionCode="NRC" collectorName="N. Maoxian &amp; A. Saldaitis &amp; No. PS" country="China" elevation="1820" latitude="31.771833" location="Male" longLatPrecision="1" longitude="103.714966" pageId="5" pageNumber="74" specimenCount="1" stateProvince="Sichuan" typeStatus="holotype">
<emphasis id="F7F1AF53FF8A7F2D19C612E84175F926" bold="true" box="[151,269,1693,1719]" pageId="5" pageNumber="74">
<typeStatus id="1A3ECDE3FF8A7F2D19C612E84170F926" box="[151,264,1693,1719]" pageId="5" pageNumber="74" type="holotype">Holotype</typeStatus>
:
</emphasis>
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, 
<collectingCountry id="BD9233D1FF8A7F2D180C12E941DBF927" box="[349,419,1692,1718]" name="China" pageId="5" pageNumber="74">China</collectingCountry>
, N. 
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, 
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<collectorName id="68701697FF8A7F2D1BD912E84368F927" box="[648,784,1692,1719]" pageId="5" pageNumber="74">N. Maoxian</collectorName>
, elev. 
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, 
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, 
<geoCoordinate id="A0B11586FF8A7F2D1D0A12E84482F926" box="[1115,1274,1693,1719]" degrees="103" direction="east" minutes="42.898" orientation="longitude" pageId="5" pageNumber="74" precision="1" value="103.714966">E103°42.898’</geoCoordinate>
, 
<date id="B13B5581FF8A7F2D1C5712E8450FF926" box="[1286,1399,1693,1719]" pageId="5" pageNumber="74" value="2017-05-20">
<collectingDate id="A17FAC69FF8A7F2D1C5712E8450FF926" box="[1286,1399,1693,1719]" pageId="5" pageNumber="74" value="2017-05-20">20.v.2017</collectingDate>
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, 
<collectorName id="68701697FF8A7F2D1CD012E8408FF94B" pageId="5" pageNumber="74">A. Saldaitis</collectorName>
leg., genitalia slide 
<collectorName id="68701697FF8A7F2D188812B44250F94A" box="[473,552,1729,1755]" pageId="5" pageNumber="74">No. PS</collectorName>
0413m (
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).
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</paragraph>
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<materialsCitation id="75ED791CFF8A7F2D1996129043E2F96E" ID-GBIF-Occurrence="2423311331" box="[199,922,1764,1791]" collectionCode="NRC" collectorName="No. PS" pageId="5" pageNumber="74" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<emphasis id="F7F1AF53FF8A7F2D199612904130F96E" bold="true" box="[199,328,1765,1791]" pageId="5" pageNumber="74">
<typeStatus id="1A3ECDE3FF8A7F2D19961290413BF96E" box="[199,323,1765,1791]" pageId="5" pageNumber="74" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="D383B8C8FF8A7F2D181E129041D0F96F" box="[335,424,1764,1791]" pageId="5" pageNumber="74" type="male">3 males</specimenCount>
, topotypic, wing slide 
<collectorName id="68701697FF8A7F2D1BF71290428CF96E" box="[678,756,1765,1791]" pageId="5" pageNumber="74">No. PS</collectorName>
0414m (
<collectionCode id="A394EB84FF8A7F2D1A03129043EAF96E" box="[850,914,1765,1791]" country="Canada" httpUri="http://grbio.org/cool/shqc-84rv" name="Division of Biological Sciences, National Research Council of Canada" pageId="5" pageNumber="74">NRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8D9F20CAFF8A7F2D1996137D44A2F86B" pageId="5" pageNumber="74" type="diagnosis">
<paragraph id="C53A7341FF8A7F2D1996137D44A2F86B" blockId="5.[151,1437,1692,2042]" pageId="5" pageNumber="74">
<emphasis id="F7F1AF53FF8A7F2D1996137D413AF8B2" bold="true" box="[199,322,1800,1827]" pageId="5" pageNumber="74">Diagnosis.</emphasis>
A medium-sized crane fly with brownish-gray thorax, yellowish-brown abdomen and yellowishbrown wing, spotted within translucent areas. Antenna, reaching the fifth abdominal segment if bent backward. Tergite 
<quantity id="027DDEA4FF8A7F2D19BD1324416FF8FB" box="[236,279,1872,1899]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="5" pageNumber="74" unit="in" value="9.0">9 in</quantity>
the shape of a narrowed concave saucer, with main body light brown, medially with pale area, posterior margin of tergal saucer deeply emarginated, medially strongly sclerotized, anterior portion of tergal saucer raised into sclerotised border, medially interrupted by pale area. Gonocoxite in the shape of a large triangular plate, apex with black tooth. Inner gonostylus in the shape of a large plate, middorsal edge with blackened plate, upper beak sclerotised and rounded, lower beak sclerotised and swollen, both beaks separated by deep notch.
</paragraph>
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<paragraph id="C53A7341FF897F2E19C610FB4533FB39" blockId="6.[151,1355,1166,1192]" box="[151,1355,1166,1192]" pageId="6" pageNumber="75">
<emphasis id="F7F1AF53FF897F2E19C610FB413FFB36" bold="true" box="[151,327,1166,1191]" pageId="6" pageNumber="75">FIGURES 9–10.</emphasis>
<taxonomicName id="028508C2FF897F2E181C10FA42E8FB36" authority="Starkevich &amp; Men &amp; Saldaitis, 2019" authorityName="Starkevich &amp; Men &amp; Saldaitis" authorityYear="2019" box="[333,656,1167,1191]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="75" phylum="Arthropoda" rank="species" species="maoxianensis" status="sp. nov." subGenus="Vestiplex">
<emphasis id="F7F1AF53FF897F2E181C10FA41F6FB36" box="[333,398,1167,1191]" italics="true" pageId="6" pageNumber="75">Tipula</emphasis>
(
<emphasis id="F7F1AF53FF897F2E18CC10FA418FFB36" box="[413,503,1167,1191]" italics="true" pageId="6" pageNumber="75">Vestiplex</emphasis>
) 
<emphasis id="F7F1AF53FF897F2E1B5710E542E8FB36" box="[518,656,1168,1191]" italics="true" pageId="6" pageNumber="75">maoxianensis</emphasis>
</taxonomicName>
<emphasis id="F7F1AF53FF897F2E1BC710FA429FFB36" bold="true" box="[662,743,1167,1191]" pageId="6" pageNumber="75">
<taxonomicNameLabel id="ECC21228FF897F2E1BC710FA429FFB36" box="[662,743,1167,1191]" pageId="6" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
9: adult male, general view of paratype; 10: wing, paratype.
</paragraph>
</caption>
<subSubSection id="8D9F20CAFF897F20199610A042D5FDF2" lastPageId="8" lastPageNumber="77" pageId="6" pageNumber="75" type="description">
<paragraph id="C53A7341FF897F2E199610A041B5FA85" blockId="6.[151,1437,1237,2020]" pageId="6" pageNumber="75">
<emphasis id="F7F1AF53FF897F2E199610A04121FB61" bold="true" box="[199,345,1237,1264]" pageId="6" pageNumber="75">Description.</emphasis>
Adult (
<figureCitation id="5DBE6FC4FF897F2E18FF10A0418BFB61" box="[430,499,1237,1264]" captionStart="FIGURES 9–10" captionStartId="6.[151,264,1166,1191]" captionTargetBox="[151,1424,200,1132]" captionTargetId="figure@6.[151,1435,181,1141]" captionTargetPageId="6" captionText="FIGURES 9–10. Tipula (Vestiplex) maoxianensis sp. nov. 9: adult male, general view of paratype; 10: wing, paratype." figureDoi="http://doi.org/10.5281/zenodo.3470076" httpUri="https://zenodo.org/record/3470076/files/figure.png" pageId="6" pageNumber="75">Fig. 9</figureCitation>
), male (N=4). General body coloration brownish-gray. Body length 
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, wing length 
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.
</paragraph>
<paragraph id="C53A7341FF897F2E19961168454CFA7A" blockId="6.[151,1437,1237,2020]" pageId="6" pageNumber="75">
<emphasis id="F7F1AF53FF897F2E19961168417CFAA6" box="[199,260,1309,1335]" italics="true" pageId="6" pageNumber="75">Head</emphasis>
. Brownish-gray, vertex and occiput gray pruinose, with dark brown median line. Rostrum brownishgray, nasus gray, very short. Antenna 12-segmented, reaching the fifth abdominal segment if bent backward. Scape brown, more than three times as long as pedicel, expanded apically. Pedicel brown, very short. Flagellum with first flagellomere cylindrical, entirely brown, subequal in length to second flagellomere, with subsequent flagellomeres changing to brownish-black, lighter at apex and enlarged basally, gradually shorter towards apex, basal enlargement with abundant black verticils, distinctly shorter than length of corresponding flagellomeres. Palpus black.
</paragraph>
<paragraph id="C53A7341FF897F2E199611804236F8C2" blockId="6.[151,1437,1237,2020]" pageId="6" pageNumber="75">
<emphasis id="F7F1AF53FF897F2E199611804160F99E" box="[199,280,1525,1551]" italics="true" pageId="6" pageNumber="75">Thorax</emphasis>
. Brownish-gray. Pronotum dark brown, gray pruinose. Mesonotal prescutum brown, gray pruinose with four darker indistinct longitudinal stripes. Median pair narrowly divided by brown line. Thoracic dorsum nearly bare, with very scarce short and light setae only between median and lateral stripes. Scutum dark brown, gray pruinose with two darkened spots. Scutellum and mediotergite brown, gray pruinose, with latter bearing black line medially. Pleuron dark brown, thinly dusted with gray. Leg slender, coxa brown, thinly dusted with gray, trochanter, femur and tibia brown with dark brown tips, tarsal segments dark brown. Tarsal claw simple without tooth. Wing (
<figureCitation id="5DBE6FC4FF897F2E19F112B8408AF979" box="[160,242,1741,1768]" captionStart="FIGURES 9–10" captionStartId="6.[151,264,1166,1191]" captionTargetBox="[151,1424,200,1132]" captionTargetId="figure@6.[151,1435,181,1141]" captionTargetPageId="6" captionText="FIGURES 9–10. Tipula (Vestiplex) maoxianensis sp. nov. 9: adult male, general view of paratype; 10: wing, paratype." figureDoi="http://doi.org/10.5281/zenodo.3470076" httpUri="https://zenodo.org/record/3470076/files/figure.png" pageId="6" pageNumber="75">Fig. 10</figureCitation>
) yellowish-brown, wing cells marbled with darker spots: faint brown marks at origin of 
<emphasis id="F7F1AF53FF897F2E1D8B12BB448EF976" box="[1242,1270,1742,1767]" italics="true" pageId="6" pageNumber="75">Rs</emphasis>
and at stigmal area, with diffuse brown patches at distal and middle area of 
<emphasis id="F7F1AF53FF897F2E1A101284431BF89A" box="[833,867,1777,1803]" italics="true" pageId="6" pageNumber="75">bm</emphasis>
, with latter extended along 
<emphasis id="F7F1AF53FF897F2E1DCA128744C4F89D" box="[1179,1212,1778,1804]" italics="true" pageId="6" pageNumber="75">Cu</emphasis>
. Discal cell narrow, elongated, petiole of cell 
<emphasis id="F7F1AF53FF897F2E18FF136341BAF8BE" box="[430,450,1814,1839]" italics="true" pageId="6" pageNumber="75">m</emphasis>
<subScript id="59017104FF897F2E1892135241B3F8A4" attach="left" box="[451,459,1831,1845]" fontSize="6" pageId="6" pageNumber="75">1</subScript>
distinctly shorter than discal cell, cell 
<emphasis id="F7F1AF53FF897F2E1A22136343F7F8A4" box="[883,911,1814,1845]" italics="true" pageId="6" pageNumber="75">
m 
<subScript id="59017104FF897F2E1AD6135243F7F8A4" attach="left" box="[903,911,1831,1845]" fontSize="6" pageId="6" pageNumber="75">1</subScript>
</emphasis>
more than three times as long as petiole. Halter with stem yellowish, knob dark brown.
</paragraph>
<paragraph id="C53A7341FF897F2E1996132843D7F80D" blockId="6.[151,1437,1237,2020]" pageId="6" pageNumber="75">
<emphasis id="F7F1AF53FF897F2E199613284149F8E6" box="[199,305,1885,1911]" italics="true" pageId="6" pageNumber="75">Abdomen</emphasis>
. Abdominal segments 1–3 yellowish-brown, gradually darker starting from segment 4. Dorsal median stripe interrupted at caudal margins of segments. Lateral stripe narrow.
</paragraph>
<paragraph id="C53A7341FF897F2F19C613D343ABFE1A" blockId="6.[151,1437,1237,2020]" lastBlockId="7.[151,1437,153,395]" lastPageId="7" lastPageNumber="76" pageId="6" pageNumber="75">
<emphasis id="F7F1AF53FF897F2E19C613D3415EF82E" box="[151,294,1958,1983]" italics="true" pageId="6" pageNumber="75">Hypopygium</emphasis>
. Yellowish-brown. Tergite 
<quantity id="027DDEA4FF897F2E1B3113D342F6F82E" box="[608,654,1957,1984]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="6" pageNumber="75" unit="in" value="9.0">9 in</quantity>
the shape of narrowed concave saucer (
<figureCitation id="5DBE6FC4FF897F2E1D3113D044CEF851" box="[1120,1206,1957,1984]" captionStart="FIGURES 11–17" captionStartId="7.[151,264,1837,1862]" captionTargetBox="[156,1420,422,1802]" captionTargetId="figure@7.[151,1435,417,1812]" captionTargetPageId="7" captionText="FIGURES 11–17. Hypopygium of male Tipula (Vestiplex) maoxianensis sp. nov. 11: hypopygium, lateral view; 12: hypopygium, dorsal view; 13: hypopygium, ventral view; 14: gonocoxite, lateral view; 15: left outer gonostylus; 16: left inner gonostylus, lateral view; 17: adminiculum, lateral view. Abbreviations: adm, adminiculum; admr, adminicular rod; gcx, gonocoxite; gph, gonapophysis; ig, inner gonostylus; og, outer gonostylus; st9, sternite 9; t9, tergite 9." figureDoi="http://doi.org/10.5281/zenodo.3470078" httpUri="https://zenodo.org/record/3470078/files/figure.png" pageId="6" pageNumber="75">Fig. 12</figureCitation>
). The main body of tergal saucer light brown, medially with pale area, posterior margin of tergal saucer deeply emarginated, medially strongly sclerotised; lateral angles of tergal saucer obtuse; anterior portion of tergal saucer raised into sclerotised border medially broadly interrupted by pale area. Gonocoxite in the shape of a large triangular plate, apex with black tooth (
<figureCitation id="5DBE6FC4FF887F2F19B01494411CFF6A" box="[225,356,225,251]" captionStart="FIGURES 11–17" captionStartId="7.[151,264,1837,1862]" captionTargetBox="[156,1420,422,1802]" captionTargetId="figure@7.[151,1435,417,1812]" captionTargetPageId="7" captionText="FIGURES 11–17. Hypopygium of male Tipula (Vestiplex) maoxianensis sp. nov. 11: hypopygium, lateral view; 12: hypopygium, dorsal view; 13: hypopygium, ventral view; 14: gonocoxite, lateral view; 15: left outer gonostylus; 16: left inner gonostylus, lateral view; 17: adminiculum, lateral view. Abbreviations: adm, adminiculum; admr, adminicular rod; gcx, gonocoxite; gph, gonapophysis; ig, inner gonostylus; og, outer gonostylus; st9, sternite 9; t9, tergite 9." figureDoi="http://doi.org/10.5281/zenodo.3470078" httpUri="https://zenodo.org/record/3470078/files/figure.png" pageId="7" pageNumber="76">Figs 11, 14</figureCitation>
). Outer gonostylus flattened, widest at the middle with apex obtuse (
<figureCitation id="5DBE6FC4FF887F2F1D24149444B2FF6A" box="[1141,1226,224,251]" captionStart="FIGURES 11–17" captionStartId="7.[151,264,1837,1862]" captionTargetBox="[156,1420,422,1802]" captionTargetId="figure@7.[151,1435,417,1812]" captionTargetPageId="7" captionText="FIGURES 11–17. Hypopygium of male Tipula (Vestiplex) maoxianensis sp. nov. 11: hypopygium, lateral view; 12: hypopygium, dorsal view; 13: hypopygium, ventral view; 14: gonocoxite, lateral view; 15: left outer gonostylus; 16: left inner gonostylus, lateral view; 17: adminiculum, lateral view. Abbreviations: adm, adminiculum; admr, adminicular rod; gcx, gonocoxite; gph, gonapophysis; ig, inner gonostylus; og, outer gonostylus; st9, sternite 9; t9, tergite 9." figureDoi="http://doi.org/10.5281/zenodo.3470078" httpUri="https://zenodo.org/record/3470078/files/figure.png" pageId="7" pageNumber="76">Fig. 15</figureCitation>
). Inner gonostylus in the shape of large plate, middorsal edge with blackened plate, upper beak sclerotised and rounded, lower beak sclerotised and swollen, both beaks separated by deep notch (
<figureCitation id="5DBE6FC4FF887F2F1A19155C43E4FED5" box="[840,924,297,324]" captionStart="FIGURES 11–17" captionStartId="7.[151,264,1837,1862]" captionTargetBox="[156,1420,422,1802]" captionTargetId="figure@7.[151,1435,417,1812]" captionTargetPageId="7" captionText="FIGURES 11–17. Hypopygium of male Tipula (Vestiplex) maoxianensis sp. nov. 11: hypopygium, lateral view; 12: hypopygium, dorsal view; 13: hypopygium, ventral view; 14: gonocoxite, lateral view; 15: left outer gonostylus; 16: left inner gonostylus, lateral view; 17: adminiculum, lateral view. Abbreviations: adm, adminiculum; admr, adminicular rod; gcx, gonocoxite; gph, gonapophysis; ig, inner gonostylus; og, outer gonostylus; st9, sternite 9; t9, tergite 9." figureDoi="http://doi.org/10.5281/zenodo.3470078" httpUri="https://zenodo.org/record/3470078/files/figure.png" pageId="7" pageNumber="76">Fig. 16</figureCitation>
). Adminiculum in shape of triangular groove, broad at base and gradually narrowed to apex in dorsal view, gonapophyses short and wrinkled, adminicular rods long (
<figureCitation id="5DBE6FC4FF887F2F198915044122FE1A" box="[216,346,369,395]" captionStart="FIGURES 11–17" captionStartId="7.[151,264,1837,1862]" captionTargetBox="[156,1420,422,1802]" captionTargetId="figure@7.[151,1435,417,1812]" captionTargetPageId="7" captionText="FIGURES 11–17. Hypopygium of male Tipula (Vestiplex) maoxianensis sp. nov. 11: hypopygium, lateral view; 12: hypopygium, dorsal view; 13: hypopygium, ventral view; 14: gonocoxite, lateral view; 15: left outer gonostylus; 16: left inner gonostylus, lateral view; 17: adminiculum, lateral view. Abbreviations: adm, adminiculum; admr, adminicular rod; gcx, gonocoxite; gph, gonapophysis; ig, inner gonostylus; og, outer gonostylus; st9, sternite 9; t9, tergite 9." figureDoi="http://doi.org/10.5281/zenodo.3470078" httpUri="https://zenodo.org/record/3470078/files/figure.png" pageId="7" pageNumber="76">Figs 13, 17</figureCitation>
). Sternite 9 with two small blackened sclerites (
<figureCitation id="5DBE6FC4FF887F2F1A23150443BDFE1A" box="[882,965,369,395]" captionStart="FIGURES 11–17" captionStartId="7.[151,264,1837,1862]" captionTargetBox="[156,1420,422,1802]" captionTargetId="figure@7.[151,1435,417,1812]" captionTargetPageId="7" captionText="FIGURES 11–17. Hypopygium of male Tipula (Vestiplex) maoxianensis sp. nov. 11: hypopygium, lateral view; 12: hypopygium, dorsal view; 13: hypopygium, ventral view; 14: gonocoxite, lateral view; 15: left outer gonostylus; 16: left inner gonostylus, lateral view; 17: adminiculum, lateral view. Abbreviations: adm, adminiculum; admr, adminicular rod; gcx, gonocoxite; gph, gonapophysis; ig, inner gonostylus; og, outer gonostylus; st9, sternite 9; t9, tergite 9." figureDoi="http://doi.org/10.5281/zenodo.3470078" httpUri="https://zenodo.org/record/3470078/files/figure.png" pageId="7" pageNumber="76">Fig. 13</figureCitation>
).
</paragraph>
<caption id="91FA23C9FF887F2F19C61358445BF823" ID-DOI="http://doi.org/10.5281/zenodo.3470078" ID-Zenodo-Dep="3470078" httpUri="https://zenodo.org/record/3470078/files/figure.png" pageId="7" pageNumber="76" startId="7.[151,264,1837,1862]" targetBox="[156,1420,422,1802]" targetPageId="7">
<paragraph id="C53A7341FF887F2F19C61358445BF823" blockId="7.[151,1436,1837,1970]" pageId="7" pageNumber="76">
<emphasis id="F7F1AF53FF887F2F19C61358412CF8D7" bold="true" box="[151,340,1837,1862]" pageId="7" pageNumber="76">FIGURES 11–17.</emphasis>
Hypopygium of male 
<taxonomicName id="028508C2FF887F2F1B6F135B43FBF8D4" authority="Starkevich &amp; Men &amp; Saldaitis, 2019" authorityName="Starkevich &amp; Men &amp; Saldaitis" authorityYear="2019" box="[574,899,1838,1862]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="76" phylum="Arthropoda" rank="species" species="maoxianensis" status="sp. nov." subGenus="Vestiplex">
<emphasis id="F7F1AF53FF887F2F1B6F135B4207F8D7" box="[574,639,1838,1862]" italics="true" pageId="7" pageNumber="76">Tipula</emphasis>
(
<emphasis id="F7F1AF53FF887F2F1BDE135B4291F8D7" box="[655,745,1838,1862]" italics="true" pageId="7" pageNumber="76">Vestiplex</emphasis>
) 
<emphasis id="F7F1AF53FF887F2F1BA8135B43FBF8D4" box="[761,899,1838,1861]" italics="true" pageId="7" pageNumber="76">maoxianensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECC21228FF887F2F1ADB135B43A4F8D7" box="[906,988,1838,1862]" pageId="7" pageNumber="76" rank="species">
<emphasis id="F7F1AF53FF887F2F1ADB135B43A4F8D7" bold="true" box="[906,988,1838,1862]" pageId="7" pageNumber="76">sp. nov.</emphasis>
</taxonomicNameLabel>
11: hypopygium, lateral view; 12: hypopygium, dorsal view; 13: hypopygium, ventral view; 14: gonocoxite, lateral view; 15: left outer gonostylus; 16: left inner gonostylus, lateral view; 17: adminiculum, lateral view. Abbreviations: adm, adminiculum; admr, adminicular rod; gcx, gonocoxite; gph, gonapophysis; ig, inner gonostylus; og, outer gonostylus; st9, sternite 9; t9, tergite 9.
</paragraph>
</caption>
<paragraph id="C53A7341FF877F20199614ED42D5FDF2" blockId="8.[151,1437,152,935]" pageId="8" pageNumber="77">
<emphasis id="F7F1AF53FF877F20199614ED422FFF22" bold="true" box="[199,599,152,179]" pageId="8" pageNumber="77">Comparative material examined:</emphasis>
<taxonomicName id="028508C2FF877F201B0E14EC4283FF23" authorityName="Savchenko" authorityYear="1960" box="[607,763,153,179]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="aestiva" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F201B0E14EC420CFF23" box="[607,628,153,178]" italics="true" pageId="8" pageNumber="77">T.</emphasis>
(
<emphasis id="F7F1AF53FF877F201BD714EC42E3FF23" box="[646,667,153,178]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F201BFD14EC4283FF23" box="[684,763,153,178]" italics="true" pageId="8" pageNumber="77">aestiva</emphasis>
</taxonomicName>
: 
<materialsCitation id="75ED791CFF877F201A5A14ED443EFE1B" ID-GBIF-Occurrence="2423311329" collectingDate="1900-07" collectingDateMax="2011-07-23" collectingDateMin="1900-07" collectionCode="ZIN, HNHM, ANSP" collectorName="SW Rand &amp; Exp. Dr. Z. &amp; Tsenkhermandal Soum &amp; Tsenkher Gol" country="China" county="Kuku-nor lake" determinerName="Mannheims" elevation="1650" latitude="47.92564" location="Yangtze river" longLatPrecision="1" longitude="109.16114" municipality="Sogon-Gomba" pageId="8" pageNumber="77" specimenCount="19" specimenCount-female="3" specimenCount-male="16" stateProvince="Hovsgol" typeStatus="holotype">
<typeStatus id="1A3ECDE3FF877F201A5A14ED430EFF23" box="[779,886,152,178]" pageId="8" pageNumber="77" type="holotype">Holotype</typeStatus>
, male, 
<collectingCountry id="BD9233D1FF877F201A9814ED4477FF23" box="[969,1039,152,178]" name="China" pageId="8" pageNumber="77">China</collectingCountry>
, Qinghai, south shore of 
<collectingCounty id="2C5B0BCDFF877F201C7E14ED40B0FF47" pageId="8" pageNumber="77">Kuku-nor lake</collectingCounty>
, 
<date id="B13B5581FF877F20198714C8413DFF46" box="[214,325,189,215]" pageId="8" pageNumber="77" value="1901-08">
<collectingDate id="A17FAC69FF877F20198714C8413DFF46" box="[214,325,189,215]" pageId="8" pageNumber="77" value="1901-08">VIII.1901</collectingDate>
</date>
(Kozlov); 
<typeStatus id="1A3ECDE3FF877F20189514C8425FFF46" box="[452,551,189,215]" pageId="8" pageNumber="77" type="paratype">paratype</typeStatus>
, female, topotypic; 
<typeStatus id="1A3ECDE3FF877F201A5D14C84317FF46" box="[780,879,189,215]" pageId="8" pageNumber="77" type="paratype">paratype</typeStatus>
, male, 
<collectingMunicipality id="255EE93BFF877F201A9214C94411FF47" box="[963,1129,188,214]" pageId="8" pageNumber="77">Sogon-Gomba</collectingMunicipality>
, I-chu river, up to 
<location id="C05A259AFF877F201C1314C840B6FF6B" LSID="urn:lsid:plazi:treatment:4D2CC257FF8A7F2019C612404473FA0E:C05A259AFF877F201C1314C840B6FF6B" country="China" county="Kuku-nor lake" latitude="47.92564" longLatPrecision="1" longitude="109.16114" municipality="Sogon-Gomba" name="Yangtze river" pageId="8" pageNumber="77" stateProvince="Hovsgol">Yangtze river</location>
, 
<date id="B13B5581FF877F20198B14944147FF6A" box="[218,319,225,251]" pageId="8" pageNumber="77" value="1900-07">
<collectingDate id="A17FAC69FF877F20198B14944147FF6A" box="[218,319,225,251]" pageId="8" pageNumber="77" value="1900-07">VII.1900</collectingDate>
</date>
(Kozlov); 
<typeStatus id="1A3ECDE3FF877F2018EF14944259FF6A" box="[446,545,225,251]" pageId="8" pageNumber="77" type="paratype">paratype</typeStatus>
, male, shore of 
<location id="C05A259AFF877F201B881495430DFF6B" LSID="urn:lsid:plazi:treatment:4D2CC257FF8A7F2019C612404473FA0E:C05A259AFF877F201B881495430DFF6B" box="[729,885,224,250]" country="China" county="Kuku-nor lake" latitude="47.92564" longLatPrecision="1" longitude="109.16114" municipality="Sogon-Gomba" name="Orin-nor lake" pageId="8" pageNumber="77" stateProvince="Hovsgol">Orin-nor lake</location>
, 
<location id="C05A259AFF877F201AD314954444FF6B" LSID="urn:lsid:plazi:treatment:4D2CC257FF8A7F2019C612404473FA0E:C05A259AFF877F201AD314954444FF6B" box="[898,1084,224,250]" country="China" county="Kuku-nor lake" latitude="47.92564" longLatPrecision="1" longitude="109.16114" municipality="Sogon-Gomba" name="Huang-he Basin" pageId="8" pageNumber="77" stateProvince="Hovsgol">Huang-he Basin</location>
, V-VI.1901 (Kozlov) (
<collectionCode id="A394EB84FF877F201C6F1494450AFF6A" box="[1342,1394,225,251]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="77" type="Museum">ZIN</collectionCode>
); 
<taxonomicName id="028508C2FF877F201CD91494411BFE8F" authorityName="Nielsen" authorityYear="1929" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="longitudinalis" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F201CD9149445E5FF6B" box="[1416,1437,225,250]" italics="true" pageId="8" pageNumber="77">T.</emphasis>
(
<emphasis id="F7F1AF53FF877F2019F0157040CEFE8F" box="[161,182,261,286]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F2019961571411BFE8F" box="[199,355,260,286]" italics="true" pageId="8" pageNumber="77">longitudinalis</emphasis>
</taxonomicName>
: 
<specimenCount id="D383B8C8FF877F201822157041BBFE8F" box="[371,451,260,287]" pageId="8" pageNumber="77" type="male">1 male</specimenCount>
, 
<specimenCount id="D383B8C8FF877F20188115704239FE8F" box="[464,577,260,287]" pageId="8" pageNumber="77" type="female">2 females</specimenCount>
, 
<collectingCountry id="BD9233D1FF877F201B1F157142C6FE8F" box="[590,702,260,286]" name="Mongolia" pageId="8" pageNumber="77">Mongolia</collectingCountry>
, 
<collectingRegion id="0741BDA3FF877F201B9B157143FDFE8F" box="[714,901,260,286]" country="Mongolia" name="Hovsgol" pageId="8" pageNumber="77">Chövsgöl aimak</collectingRegion>
, 
<locationDeviation id="D4886A63FF877F201AC3157044E9FE8F" box="[914,1169,260,287]" location="Somon Chatgal" pageId="8" pageNumber="77">N von Somon Chatgal</locationDeviation>
am 
<collectorName id="68701697FF877F201D941570454AFE8F" box="[1221,1330,260,287]" pageId="8" pageNumber="77">
SW 
<location id="C05A259AFF877F201DA61571454AFE8F" LSID="urn:lsid:plazi:treatment:4D2CC257FF8A7F2019C612404473FA0E:C05A259AFF877F201DA61571454AFE8F" box="[1271,1330,260,286]" country="China" county="Kuku-nor lake" latitude="47.92564" longLatPrecision="1" longitude="109.16114" municipality="Sogon-Gomba" name="Rand des Sees" pageId="8" pageNumber="77" stateProvince="Hovsgol">Rand</location>
</collectorName>
des Sees 
<collectingRegion id="0741BDA3FF877F2019C6155D417AFED3" box="[151,258,296,322]" country="Mongolia" name="Hovsgol" pageId="8" pageNumber="77">Chövsgöl</collectingRegion>
nuur, 
<quantity id="027DDEA4FF877F201803155D41D4FED2" box="[338,428,296,323]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="8" pageNumber="77" unit="m" value="1650.0">
<elevation id="4EA89472FF877F201803155D41D4FED2" box="[338,428,296,323]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="8" pageNumber="77" unit="m" value="1650.0">1650 m</elevation>
</quantity>
, 
<collectorName id="68701697FF877F2018ED155C4239FED2" box="[444,577,297,323]" pageId="8" pageNumber="77">Exp. Dr. Z.</collectorName>
KASZAB, 
<determinerName id="30E69F85FF877F201B9F155D432AFED3" box="[718,850,296,322]" pageId="8" pageNumber="77">Mannheims</determinerName>
det. 1969, 
<date id="B13B5581FF877F201A8A155D441CFED3" box="[987,1124,296,322]" pageId="8" pageNumber="77" value="1968-07-18">
<collectingDate id="A17FAC69FF877F201A8A155D441CFED3" box="[987,1124,296,322]" pageId="8" pageNumber="77" value="1968-07-18">18.VII.1968</collectingDate>
</date>
(
<collectionCode id="A394EB84FF877F201D26155C44ABFED2" box="[1143,1235,297,323]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="8" pageNumber="77">HNHM</collectionCode>
); 
<specimenCount id="D383B8C8FF877F201DBB155D452DFED3" box="[1258,1365,296,323]" pageId="8" pageNumber="77" type="male">15 males</specimenCount>
, 
<specimenCount id="D383B8C8FF877F201C35155C40A8FEF7" pageId="8" pageNumber="77" type="female">1 female</specimenCount>
, 
<collectingRegion id="0741BDA3FF877F20198C153941FFFEF7" box="[221,391,332,358]" country="Mongolia" name="Hentiy" pageId="8" pageNumber="77">Khentii Aimag</collectingRegion>
, 
<collectorName id="68701697FF877F2018C2153942E1FEF6" box="[403,665,332,359]" pageId="8" pageNumber="77">Tsenkhermandal Soum</collectorName>
, spring headwaters 
<collectorName id="68701697FF877F201A2B1539446EFEF7" box="[890,1046,332,358]" pageId="8" pageNumber="77">Tsenkher Gol</collectorName>
, 
<locationDeviation id="D4886A63FF877F201D73153845E1FEF7" box="[1058,1433,332,359]" location="North of Tsenkhermandal" pageId="8" pageNumber="77">
<quantity id="027DDEA4FF877F201D7315384413FEF7" box="[1058,1131,332,359]" metricMagnitude="4" metricUnit="m" metricValue="2.9" pageId="8" pageNumber="77" unit="km" value="29.0">29 km</quantity>
North of Tsenkhermandal
</locationDeviation>
, 
<geoCoordinate id="A0B11586FF877F2019C61505416FFE1A" box="[151,279,368,395]" degrees="47.92564" direction="north" orientation="latitude" pageId="8" pageNumber="77" precision="1" value="47.92564">N47.92564</geoCoordinate>
, 
<geoCoordinate id="A0B11586FF877F201872150541D5FE1B" box="[291,429,368,394]" degrees="109.16114" direction="east" orientation="longitude" pageId="8" pageNumber="77" precision="1" value="109.16114">E109.16114</geoCoordinate>
, elev. 
<quantity id="027DDEA4FF877F2018A215054232FE1A" box="[499,586,368,395]" metricMagnitude="3" metricUnit="m" metricValue="1.589" pageId="8" pageNumber="77" unit="m" value="1589.0">
<elevation id="4EA89472FF877F2018A215054232FE1A" box="[499,586,368,395]" metricMagnitude="3" metricUnit="m" metricValue="1.589" pageId="8" pageNumber="77" unit="m" value="1589.0">1589 m</elevation>
</quantity>
, 
<date id="B13B5581FF877F201B07150442A6FE1A" box="[598,734,369,395]" pageId="8" pageNumber="77" value="2011-07-23">
<collectingDate id="A17FAC69FF877F201B07150442A6FE1A" box="[598,734,369,395]" pageId="8" pageNumber="77" value="2011-07-23">23.VII.2011</collectingDate>
</date>
(
<collectionCode id="A394EB84FF877F201BBC15044343FE1A" box="[749,827,369,395]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="8" pageNumber="77">ANSP</collectionCode>
); 
<taxonomicName id="028508C2FF877F201A1F1504443EFE1B" authorityName="Savchenko" authorityYear="1955" box="[846,1094,368,395]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="opilionimorpha" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F201A1F1504431BFE1B" box="[846,867,369,394]" italics="true" pageId="8" pageNumber="77">T.</emphasis>
(
<emphasis id="F7F1AF53FF877F201A22150443F0FE1B" box="[883,904,369,394]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F201AC91505443EFE1B" box="[920,1094,368,394]" italics="true" pageId="8" pageNumber="77">opilionimorpha</emphasis>
</taxonomicName>
</materialsCitation>
: 
<materialsCitation id="75ED791CFF877F201D04150442D1FDF2" ID-GBIF-Occurrence="2423311346" collectingDate="1900-07-01" collectingDateMax="2011-07-04" collectingDateMin="1900-07-01" collectionCode="ZIN, HNHM, ANSP" collectorName="Exp. Dr. Z. &amp; Changaj Gebirge &amp; Ulaan &amp; Pass Egijn &amp; Erdenesogt Soum &amp; Shargaljuut Gol &amp; Ugalzyn Gol" country="China" county="river Dzagyn-gol" elevation="2080" latitude="46.35902" location="Changaj Gebirge" longLatPrecision="1" longitude="101.27559" municipality="Ubruchangaj" pageId="8" pageNumber="77" specimenCount="23" specimenCount-female="6" specimenCount-male="17" stateProvince="Bayanhongor" typeStatus="lectotype">
<typeStatus id="1A3ECDE3FF877F201D04150444B0FE1A" box="[1109,1224,369,395]" pageId="8" pageNumber="77" type="lectotype">Lectotype</typeStatus>
, male, 
<collectingCountry id="BD9233D1FF877F201C4915054526FE1B" box="[1304,1374,368,394]" name="China" pageId="8" pageNumber="77">China</collectingCountry>
, Qinghai, valley of 
<collectingCounty id="2C5B0BCDFF877F20186E15E1427AFE3F" box="[319,514,404,430]" pageId="8" pageNumber="77">river Dzagyn-gol</collectingCounty>
, 
<date id="B13B5581FF877F201B5F15E142DDFE3F" box="[526,677,404,430]" pageId="8" pageNumber="77" value="1900-07-01" valueMax="1900-07-06" valueMin="1900-07-01">
<collectingDate id="A17FAC69FF877F201B5F15E142DDFE3F" box="[526,677,404,430]" pageId="8" pageNumber="77" value="1900-07-01" valueMax="1900-07-06" valueMin="1900-07-01">1–6.VII.1900</collectingDate>
</date>
(Kozlov); 
<typeStatus id="1A3ECDE3FF877F201A7115E143BDFE3F" box="[800,965,404,430]" pageId="8" pageNumber="77" type="paralectotype">paralectotypes</typeStatus>
, 
<specimenCount id="D383B8C8FF877F201A9E15E04466FE3F" box="[975,1054,404,431]" pageId="8" pageNumber="77" type="male">1 male</specimenCount>
, 
<specimenCount id="D383B8C8FF877F201D7815E044F5FE3F" box="[1065,1165,404,431]" pageId="8" pageNumber="77" type="female">1 female</specimenCount>
, topotypic (
<collectionCode id="A394EB84FF877F201C4015E0453DFE3E" box="[1297,1349,405,431]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="77" type="Museum">ZIN</collectionCode>
); 
<taxonomicName id="028508C2FF877F201C0815E04085FE43" authorityName="Riedel" authorityYear="1913" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="virgatula" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F201C0815E04517FE3F" box="[1369,1391,405,430]" italics="true" pageId="8" pageNumber="77">T.</emphasis>
(
<emphasis id="F7F1AF53FF877F201C2F15E045ECFE3F" box="[1406,1428,405,430]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F2019C615CD4085FE43" box="[151,253,440,466]" italics="true" pageId="8" pageNumber="77">virgatula</emphasis>
</taxonomicName>
: 
<specimenCount id="D383B8C8FF877F20185A15CC411CFE43" box="[267,356,440,467]" pageId="8" pageNumber="77" type="male">3 males</specimenCount>
, 
<collectingCountry id="BD9233D1FF877F20183E15CD41A7FE43" box="[367,479,440,466]" name="Mongolia" pageId="8" pageNumber="77">Mongolia</collectingCountry>
, 
<collectingMunicipality id="255EE93BFF877F2018B815CD4202FE43" box="[489,634,440,466]" pageId="8" pageNumber="77">Ubruchangaj</collectingMunicipality>
aimak, 
<location id="C05A259AFF877F201B8215CD43ECFE43" LSID="urn:lsid:plazi:treatment:4D2CC257FF8A7F2019C612404473FA0E:C05A259AFF877F201B8215CD43ECFE43" box="[723,916,440,466]" country="China" county="river Dzagyn-gol" latitude="46.35902" longLatPrecision="1" longitude="101.27559" municipality="Ubruchangaj" name="Changaj Gebirge" pageId="8" pageNumber="77" stateProvince="Bayanhongor">Changaj Gebirge</location>
, 
<quantity id="027DDEA4FF877F201ACF15CC439DFE43" box="[926,997,440,467]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="77" unit="km" value="21.0">21 km</quantity>
O von somon 
<location id="C05A259AFF877F201DD915CD4483FE43" LSID="urn:lsid:plazi:treatment:4D2CC257FF8A7F2019C612404473FA0E:C05A259AFF877F201DD915CD4483FE43" box="[1160,1275,440,466]" country="China" county="river Dzagyn-gol" latitude="46.35902" longLatPrecision="1" longitude="101.27559" municipality="Ubruchangaj" name="Narijnteel" pageId="8" pageNumber="77" stateProvince="Bayanhongor">Narijnteel</location>
, 
<quantity id="027DDEA4FF877F201C5715CC4525FE42" box="[1286,1373,441,467]" metricMagnitude="3" metricUnit="m" metricValue="2.08" pageId="8" pageNumber="77" unit="m" value="2080.0">
<elevation id="4EA89472FF877F201C5715CC4525FE42" box="[1286,1373,441,467]" metricMagnitude="3" metricUnit="m" metricValue="2.08" pageId="8" pageNumber="77" unit="m" value="2080.0">2080 m</elevation>
</quantity>
, 
<collectorName id="68701697FF877F201C3915CC40A2FE66" pageId="8" pageNumber="77">Exp. Dr. Z.</collectorName>
KASZAB, 1964, 
<date id="B13B5581FF877F2018F915A9425FFE67" box="[424,551,476,502]" pageId="8" pageNumber="77" value="1964-06-27">
<collectingDate id="A17FAC69FF877F2018F915A9425FFE67" box="[424,551,476,502]" pageId="8" pageNumber="77" value="1964-06-27">27.VI.1964</collectingDate>
</date>
(
<collectionCode id="A394EB84FF877F201B6615A842EBFE66" box="[567,659,477,503]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="8" pageNumber="77">HNHM</collectionCode>
); 
<specimenCount id="D383B8C8FF877F201BF615A9436FFE67" box="[679,791,476,503]" pageId="8" pageNumber="77" type="female">5 females</specimenCount>
, 
<collectingRegion id="0741BDA3FF877F201A7215A94477FE67" box="[803,1039,476,502]" country="Mongolia" name="Bayanhongor" pageId="8" pageNumber="77">Bajanchongor aimak</collectingRegion>
, 
<collectorName id="68701697FF877F201D4A15A944A5FE67" box="[1051,1245,476,502]" pageId="8" pageNumber="77">Changaj Gebirge</collectorName>
, 
<collectorName id="68701697FF877F201DB915A94553FE67" box="[1256,1323,476,502]" pageId="8" pageNumber="77">Ulaan</collectorName>
colon, 
<quantity id="027DDEA4FF877F201CD115A840C3FD8B" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="8" pageNumber="77" unit="km" value="18.0">18 km</quantity>
S vom 
<collectorName id="68701697FF877F20185F167441FBFD8B" box="[270,387,512,539]" pageId="8" pageNumber="77">Pass Egijn</collectorName>
davaa, 
<quantity id="027DDEA4FF877F20188716744254FD8A" box="[470,556,513,539]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="77" unit="m" value="2300.0">
<elevation id="4EA89472FF877F20188716744254FD8A" box="[470,556,513,539]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="77" unit="m" value="2300.0">2300 m</elevation>
</quantity>
, 
<collectorName id="68701697FF877F201B66167442CAFD8A" box="[567,690,513,539]" pageId="8" pageNumber="77">Exp. Dr. Z.</collectorName>
KASZAB, 
<date id="B13B5581FF877F201A65167543CCFD8B" box="[820,948,512,538]" pageId="8" pageNumber="77" value="1966-06-21">
<collectingDate id="A17FAC69FF877F201A65167543CCFD8B" box="[820,948,512,538]" pageId="8" pageNumber="77" value="1966-06-21">21.VI.1966</collectingDate>
</date>
(
<collectionCode id="A394EB84FF877F201A9316744466FD8A" box="[962,1054,513,539]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="8" pageNumber="77">HNHM</collectionCode>
); 
<specimenCount id="D383B8C8FF877F201D61167444EEFD8B" box="[1072,1174,512,539]" pageId="8" pageNumber="77" type="male">13 males</specimenCount>
, 
<collectingRegion id="0741BDA3FF877F201DF1167545E0FD8B" box="[1184,1432,512,538]" country="Mongolia" name="Bayanhongor" pageId="8" pageNumber="77">Bayankhongor Aimag</collectingRegion>
, 
<collectorName id="68701697FF877F2019C61651411AFDAE" box="[151,354,548,575]" pageId="8" pageNumber="77">Erdenesogt Soum</collectorName>
, 
<collectorName id="68701697FF877F20183E1651425EFDAF" box="[367,550,548,574]" pageId="8" pageNumber="77">
<location id="C05A259AFF877F20183E16514188FDAF" LSID="urn:lsid:plazi:treatment:4D2CC257FF8A7F2019C612404473FA0E:C05A259AFF877F20183E16514188FDAF" box="[367,496,548,574]" country="China" county="river Dzagyn-gol" latitude="46.35902" longLatPrecision="1" longitude="101.27559" municipality="Ubruchangaj" name="Shargaljuut" pageId="8" pageNumber="77" stateProvince="Bayanhongor">Shargaljuut</location>
Gol
</collectorName>
, 
<locationDeviation id="D4886A63FF877F201B621650443CFDAF" box="[563,1092,548,575]" location="Shargaljuut Rashaan Hot Spring" pageId="8" pageNumber="77">
<quantity id="027DDEA4FF877F201B6216504204FDAF" box="[563,636,548,575]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="77" unit="km" value="10.0">10 km</quantity>
NE of Shargaljuut Rashaan Hot Spring
</locationDeviation>
, jct 
<collectorName id="68701697FF877F201D2716514573FDAF" box="[1142,1291,548,574]" pageId="8" pageNumber="77">Ugalzyn Gol</collectorName>
, 
<geoCoordinate id="A0B11586FF877F201C49165145E0FDAE" box="[1304,1432,548,575]" degrees="46.35902" direction="north" orientation="latitude" pageId="8" pageNumber="77" precision="1" value="46.35902">N46.35902</geoCoordinate>
, 
<geoCoordinate id="A0B11586FF877F2019C6163D415BFDF2" box="[151,291,584,611]" degrees="101.27559" direction="east" orientation="longitude" pageId="8" pageNumber="77" precision="1" value="101.27559">E101.27559</geoCoordinate>
, elev. 
<quantity id="027DDEA4FF877F201839163D41C7FDF2" box="[360,447,584,611]" metricMagnitude="3" metricUnit="m" metricValue="2.186" pageId="8" pageNumber="77" unit="m" value="2186.0">
<elevation id="4EA89472FF877F201839163D41C7FDF2" box="[360,447,584,611]" metricMagnitude="3" metricUnit="m" metricValue="2.186" pageId="8" pageNumber="77" unit="m" value="2186.0">2186 m</elevation>
</quantity>
, 
<date id="B13B5581FF877F20189A163C423DFDF2" box="[459,581,585,611]" pageId="8" pageNumber="77" value="2011-07-04">
<collectingDate id="A17FAC69FF877F20189A163C423DFDF2" box="[459,581,585,611]" pageId="8" pageNumber="77" value="2011-07-04">4.VII.2011</collectingDate>
</date>
(
<collectionCode id="A394EB84FF877F201B05163C42DAFDF2" box="[596,674,585,611]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="8" pageNumber="77">ANSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8D9F20CAFF877F201996161944AFFD5E" pageId="8" pageNumber="77" type="biology_ecology">
<paragraph id="C53A7341FF877F201996161944AFFD5E" blockId="8.[151,1437,152,935]" pageId="8" pageNumber="77">
<emphasis id="F7F1AF53FF877F2019961619418AFD16" bold="true" box="[199,498,620,647]" pageId="8" pageNumber="77">Biology and distribution.</emphasis>
Four males were collected during a single rainy night on 
<date id="B13B5581FF877F201D2A161844ABFD16" box="[1147,1235,621,647]" pageId="8" pageNumber="77" value="1920-05">20 May</date>
, 
<quantity id="027DDEA4FF877F201D8F1618454CFD17" box="[1246,1332,620,647]" metricMagnitude="1" metricUnit="m" metricValue="5.12318" pageId="8" pageNumber="77" unit="in" value="2017.0">2017 in</quantity>
a remote part of west 
<collectingCountry id="BD9233D1FF877F20184E16E5411DFD3B" box="[287,357,656,682]" name="China" pageId="8" pageNumber="77">China</collectingCountry>
, 
<collectingRegion id="0741BDA3FF877F20183E16E5424CFD3B" box="[367,564,656,682]" country="China" name="Sichuan" pageId="8" pageNumber="77">Sichuan Province</collectingRegion>
near the Maoxian. The new species was collected at an altitude of 1800 meters in a valley sparsely covered by mixed forests dominated by deciduous trees and bushes (
<figureCitation id="5DBE6FC4FF877F201D2716C144B3FD5E" box="[1142,1227,692,719]" captionStart="FIGURE 37" captionStartId="15.[151,250,1029,1054]" captionTargetBox="[242,1345,179,1007]" captionTargetId="figure@15.[242,1345,179,1007]" captionTargetPageId="15" captionText="FIGURE 37. Type locality of Tipula (Vestiplex) borthi sp. nov. and T. (Vestiplex) maoxianensis sp. nov. China, N. Sichuan, 20 km. N. Maoxian, N31°46.310’, E103°42.898’, altitude 1820 m." figureDoi="http://doi.org/10.5281/zenodo.3470088" httpUri="https://zenodo.org/record/3470088/files/figure.png" pageId="8" pageNumber="77">Fig. 37</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8D9F20CAFF877F20199616AD4477FCCE" pageId="8" pageNumber="77" type="discussion">
<paragraph id="C53A7341FF877F20199616AD4477FCCE" blockId="8.[151,1437,152,935]" pageId="8" pageNumber="77">
<emphasis id="F7F1AF53FF877F20199616AD4134FD62" bold="true" box="[199,332,728,755]" pageId="8" pageNumber="77">Discussion.</emphasis>
The new species is most similar to 
<taxonomicName id="028508C2FF877F201BA016AC43E9FD63" authorityName="Savchenko" authorityYear="1960" box="[753,913,729,755]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="aestiva" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F201BA016AC4379FD63" box="[753,769,729,754]" italics="true" pageId="8" pageNumber="77">T</emphasis>
. (
<emphasis id="F7F1AF53FF877F201A4A16AC4348FD63" box="[795,816,729,754]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F201A1316AC43E9FD63" box="[834,913,729,754]" italics="true" pageId="8" pageNumber="77">aestiva</emphasis>
</taxonomicName>
but differs by details of tergite 9. 
<taxonomicName id="028508C2FF877F201C7516AD4157FC87" authority="Starkevich &amp; Men &amp; Saldaitis, 2019" authorityName="Starkevich &amp; Men &amp; Saldaitis" authorityYear="2019" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="maoxianensis" status="sp. nov." subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F201C7516AD4514FD63" box="[1316,1388,728,754]" italics="true" pageId="8" pageNumber="77">Tipula</emphasis>
(
<emphasis id="F7F1AF53FF877F201C2F16AC45ECFD63" box="[1406,1428,729,754]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F2019C616884157FC87" box="[151,303,765,790]" italics="true" pageId="8" pageNumber="77">maoxianensis</emphasis>
</taxonomicName>
<emphasis id="F7F1AF53FF877F201868168841EDFC86" bold="true" box="[313,405,765,791]" pageId="8" pageNumber="77">
<taxonomicNameLabel id="ECC21228FF877F201868168841EDFC86" box="[313,405,765,791]" pageId="8" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has tergite 9 with lateral corners rounded, while 
<taxonomicName id="028508C2FF877F201A9F16884415FC87" authorityName="Savchenko" authorityYear="1960" box="[974,1133,765,791]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="aestiva" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F201A9F168843A6FC87" box="[974,990,765,790]" italics="true" pageId="8" pageNumber="77">T</emphasis>
. (
<emphasis id="F7F1AF53FF877F201AA616884474FC87" box="[1015,1036,765,790]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F201D4F16884415FC87" box="[1054,1133,765,790]" italics="true" pageId="8" pageNumber="77">aestiva</emphasis>
</taxonomicName>
lateral corners obtuse and distinctly blackened. They can also be separated by antenna reaching the fifth abdominal segment in 
<taxonomicName id="028508C2FF877F201C5517544093FCCF" authority="Starkevich &amp; Men &amp; Saldaitis, 2019" authorityName="Starkevich &amp; Men &amp; Saldaitis" authorityYear="2019" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="maoxianensis" status="sp. nov." subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F201C5517544562FCAB" box="[1284,1306,801,826]" italics="true" pageId="8" pageNumber="77">T.</emphasis>
(
<emphasis id="F7F1AF53FF877F201C7817544547FCAB" box="[1321,1343,801,826]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F201C1F17544093FCCF" italics="true" pageId="8" pageNumber="77">maoxianensis</emphasis>
</taxonomicName>
<emphasis id="F7F1AF53FF877F2019A317304134FCCE" bold="true" box="[242,332,837,863]" pageId="8" pageNumber="77">
<taxonomicNameLabel id="ECC21228FF877F2019A317304134FCCE" box="[242,332,837,863]" pageId="8" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
while antenna reach the base of the abdomen in 
<taxonomicName id="028508C2FF877F201A3C17304470FCCF" authorityName="Savchenko" authorityYear="1960" box="[877,1032,837,863]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="aestiva" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F201A3C17304305FCCF" box="[877,893,837,862]" italics="true" pageId="8" pageNumber="77">T</emphasis>
. (
<emphasis id="F7F1AF53FF877F201AC5173043D2FCCF" box="[916,938,837,862]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F201AE817304470FCCF" box="[953,1032,837,862]" italics="true" pageId="8" pageNumber="77">aestiva</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="8D9F20CAFF877F201996171C4473FA0E" pageId="8" pageNumber="77" type="etymology">
<paragraph id="C53A7341FF877F201996171C4137FC37" blockId="8.[151,1437,152,935]" pageId="8" pageNumber="77">
<emphasis id="F7F1AF53FF877F201996171C4136FC12" bold="true" box="[199,334,873,899]" pageId="8" pageNumber="77">Etymology.</emphasis>
The specific epithet is a noun ‘ 
<emphasis id="F7F1AF53FF877F201BE2171C436BFC13" box="[691,787,873,898]" italics="true" pageId="8" pageNumber="77">maoxian</emphasis>
’ with Latin suffix ‘ 
<emphasis id="F7F1AF53FF877F201AA1171C4450FC13" box="[1008,1064,873,898]" italics="true" pageId="8" pageNumber="77">ensis</emphasis>
’, referring to the 
<typeStatus id="1A3ECDE3FF877F201DA0171C4559FC12" box="[1265,1313,873,899]" pageId="8" pageNumber="77">type</typeStatus>
locality of the new species.
</paragraph>
<paragraph id="C53A7341FF877F2019C6178C42CDFB82" blockId="8.[151,693,1016,1043]" box="[151,693,1016,1043]" pageId="8" pageNumber="77">
<taxonomicName id="028508C2FF877F2019C6178C427FFB82" authorityName="Alexander" authorityYear="1932" box="[151,519,1016,1043]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="divisotergata" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F2019C6178C409AFB82" box="[151,226,1017,1043]" italics="true" pageId="8" pageNumber="77">Tipula</emphasis>
<emphasis id="F7F1AF53FF877F2019BB178D408CFB83" bold="true" box="[234,244,1016,1042]" pageId="8" pageNumber="77">(</emphasis>
<emphasis id="F7F1AF53FF877F2019A2178C4123FB82" box="[243,347,1017,1043]" italics="true" pageId="8" pageNumber="77">Vestiplex</emphasis>
<emphasis id="F7F1AF53FF877F20180D178D411EFB83" bold="true" box="[348,358,1016,1042]" pageId="8" pageNumber="77">)</emphasis>
<emphasis id="F7F1AF53FF877F20183C178C427FFB82" box="[365,519,1017,1043]" italics="true" pageId="8" pageNumber="77">divisotergata</emphasis>
</taxonomicName>
<emphasis id="F7F1AF53FF877F201B5F178D42CDFB82" bold="true" box="[526,693,1016,1043]" pageId="8" pageNumber="77">species group</emphasis>
</paragraph>
<paragraph id="C53A7341FF877F2019C610354473FA0E" blockId="8.[151,1437,1088,1439]" pageId="8" pageNumber="77">
The 
<taxonomicName id="028508C2FF877F20199810354124FBCB" authorityName="Alexander" authorityYear="1932" box="[201,348,1088,1114]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="divisotergata">
<emphasis id="F7F1AF53FF877F20199810354124FBCB" box="[201,348,1088,1114]" italics="true" pageId="8" pageNumber="77">divisotergata</emphasis>
</taxonomicName>
species group was proposed by 
<bibRefCitation id="A1140EB0FF877F201B92103543F7FBCA" author="Savchenko, E. N." box="[707,911,1088,1115]" pageId="8" pageNumber="77" pagination="1 - 503" refId="ref11220" refString="Savchenko, E. N. (1964) Crane-flies (Diptera, Tipulidae), Subfam. Tipulinae, Genus Tipula L., 2. Fauna USSR, New Series, 89 (Insecta Diptera), 4, 1 - 503. [in Russian]" type="journal article" year="1964">Savchenko (1964)</bibRefCitation>
based on the shape of the male gonocoxite and tergite 9. A list of species belonging to the 
<taxonomicName id="028508C2FF877F201B2A10114376FBEF" authorityName="Alexander" authorityYear="1932" box="[635,782,1124,1150]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="divisotergata">
<emphasis id="F7F1AF53FF877F201B2A10114376FBEF" box="[635,782,1124,1150]" italics="true" pageId="8" pageNumber="77">divisotergata</emphasis>
</taxonomicName>
group is revised and the following species are recognized in 
<collectingCountry id="BD9233D1FF877F2019E210FD4082FB33" box="[179,250,1160,1186]" name="China" pageId="8" pageNumber="77">China</collectingCountry>
: 
<taxonomicName id="028508C2FF877F20185510FC42F2FB33" authority="Alexander, 1963" authorityName="Alexander" authorityYear="1963" box="[260,650,1160,1187]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="adungensis" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F20185510FC4161FB33" box="[260,281,1161,1186]" italics="true" pageId="8" pageNumber="77">T.</emphasis>
(
<emphasis id="F7F1AF53FF877F20187910FC4145FB33" box="[296,317,1161,1186]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F20181D10FD41B2FB33" box="[332,458,1160,1186]" italics="true" pageId="8" pageNumber="77">adungensis</emphasis>
<bibRefCitation id="A1140EB0FF877F20189F10FD42F2FB33" author="Alexander, C. P." box="[462,650,1160,1186]" pageId="8" pageNumber="77" pagination="319 - 340" refId="ref10593" refString="Alexander, C. P. (1963) Some Tipulidae from Tibet and upper Burma in the British museum (Natural History) (Diptera). Bulletin of the British Museum (Natural History), Entomology, 14, 319 - 340. https: // doi. org / 10.5962 / bhl. part. 8785" type="journal article" year="1963">Alexander, 1963</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="028508C2FF877F201BC510FC4457FB32" authority="Alexander, 1932" authorityName="Alexander" authorityYear="1932" box="[660,1071,1160,1187]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="divisotergata" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F201BC510FC42D1FB33" box="[660,681,1161,1186]" italics="true" pageId="8" pageNumber="77">T.</emphasis>
(
<emphasis id="F7F1AF53FF877F201BE910FC42B5FB33" box="[696,717,1161,1186]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F201B8D10FD4317FB33" box="[732,879,1160,1186]" italics="true" pageId="8" pageNumber="77">divisotergata</emphasis>
<bibRefCitation id="A1140EB0FF877F201A2210FD4457FB32" author="Alexander, C. P." box="[883,1071,1160,1187]" pageId="8" pageNumber="77" pagination="105 - 136" refId="ref10389" refString="Alexander, C. P. (1932) New or little-known Tipulidae from eastern Asia (Diptera). X. Philippine Journal of Science, 49, 105 - 136." type="journal article" year="1932">Alexander, 1932</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="028508C2FF877F201D6810FC45E0FB32" authority="Alexander, 1928" authorityName="Alexander" authorityYear="1928" box="[1081,1432,1160,1187]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="nokonis" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F201D6810FC4436FB33" box="[1081,1102,1161,1186]" italics="true" pageId="8" pageNumber="77">T.</emphasis>
(
<emphasis id="F7F1AF53FF877F201D0C10FC440AFB33" box="[1117,1138,1161,1186]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F201DD010FD44A0FB33" box="[1153,1240,1160,1186]" italics="true" pageId="8" pageNumber="77">nokonis</emphasis>
Alexander, 1928
</taxonomicName>
, 
<taxonomicName id="028508C2FF877F2019C610D8425EFB57" authority="Alexander, 1963" authorityName="Alexander" authorityYear="1963" box="[151,550,1196,1223]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="rongtoensis" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F2019C610D840D4FB57" box="[151,172,1197,1222]" italics="true" pageId="8" pageNumber="77">T.</emphasis>
(
<emphasis id="F7F1AF53FF877F2019ED10D840AAFB57" box="[188,210,1197,1222]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F2019B310D8411CFB57" box="[226,356,1197,1222]" italics="true" pageId="8" pageNumber="77">rongtoensis</emphasis>
<bibRefCitation id="A1140EB0FF877F20183810D9425EFB57" author="Alexander, C. P." box="[361,550,1196,1222]" pageId="8" pageNumber="77" pagination="319 - 340" refId="ref10593" refString="Alexander, C. P. (1963) Some Tipulidae from Tibet and upper Burma in the British museum (Natural History) (Diptera). Bulletin of the British Museum (Natural History), Entomology, 14, 319 - 340. https: // doi. org / 10.5962 / bhl. part. 8785" type="journal article" year="1963">Alexander, 1963</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="028508C2FF877F201B6310D8439BFB56" authority="Alexander, 1920" authorityName="Alexander" authorityYear="1920" box="[562,995,1196,1223]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="subapterogyne" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F201B6310D8423FFB57" box="[562,583,1197,1222]" italics="true" pageId="8" pageNumber="77">T.</emphasis>
(
<emphasis id="F7F1AF53FF877F201B0610D84214FB57" box="[599,620,1197,1222]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F201B2D10D94358FB57" box="[636,800,1196,1222]" italics="true" pageId="8" pageNumber="77">subapterogyne</emphasis>
Alexander, 1920
</taxonomicName>
, 
<taxonomicName id="028508C2FF877F201ABF10D84514FB56" authority="Alexander, 1924" authorityName="Alexander" authorityYear="1924" box="[1006,1388,1196,1223]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="verecunda" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F201ABF10D8447CFB57" box="[1006,1028,1197,1222]" italics="true" pageId="8" pageNumber="77">T.</emphasis>
(
<emphasis id="F7F1AF53FF877F201D4510D84451FB57" box="[1044,1065,1197,1222]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F201D6810D944D5FB57" box="[1081,1197,1196,1222]" italics="true" pageId="8" pageNumber="77">verecunda</emphasis>
Alexander, 1924
</taxonomicName>
and 
<taxonomicName id="028508C2FF877F2019C610A4426BFB7B" authority="Alexander, 1963" authorityName="Alexander" authorityYear="1963" box="[151,531,1232,1259]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="zayulensis" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F2019C610A440D4FB7B" box="[151,172,1233,1258]" italics="true" pageId="8" pageNumber="77">T.</emphasis>
(
<emphasis id="F7F1AF53FF877F2019EA10A440A9FB7B" box="[187,209,1233,1258]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F20198E10A5412AFB7B" box="[223,338,1232,1258]" italics="true" pageId="8" pageNumber="77">zayulensis</emphasis>
<bibRefCitation id="A1140EB0FF877F20180610A5426BFB7B" author="Alexander, C. P." box="[343,531,1232,1258]" pageId="8" pageNumber="77" pagination="319 - 340" refId="ref10593" refString="Alexander, C. P. (1963) Some Tipulidae from Tibet and upper Burma in the British museum (Natural History) (Diptera). Bulletin of the British Museum (Natural History), Entomology, 14, 319 - 340. https: // doi. org / 10.5962 / bhl. part. 8785" type="journal article" year="1963">Alexander, 1963</bibRefCitation>
</taxonomicName>
. The 
<taxonomicName id="028508C2FF877F201B1F10A54299FB7B" authorityName="Alexander" authorityYear="1932" box="[590,737,1232,1258]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="divisotergata">
<emphasis id="F7F1AF53FF877F201B1F10A54299FB7B" box="[590,737,1232,1258]" italics="true" pageId="8" pageNumber="77">divisotergata</emphasis>
</taxonomicName>
species group belongs to the part of 
<emphasis id="F7F1AF53FF877F201D2710A444F3FB7B" box="[1142,1163,1233,1258]" italics="true" pageId="8" pageNumber="77">T.</emphasis>
(
<taxonomicName id="028508C2FF877F201DCB10A54486FB7B" authorityName="Bezzi" authorityYear="1924" box="[1178,1278,1232,1258]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="86" phylum="Arthropoda" rank="subGenus" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F201DCB10A54486FB7B" box="[1178,1278,1232,1258]" italics="true" pageId="8" pageNumber="77">Vestiplex</emphasis>
</taxonomicName>
) whose males are characterized by tergite 9 divided by a pale membrane. In this part males of the 
<taxonomicName id="028508C2FF877F201D6D108144B7FA9F" authorityName="Alexander" authorityYear="1932" box="[1084,1231,1268,1294]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="divisotergata">
<emphasis id="F7F1AF53FF877F201D6D108144B7FA9F" box="[1084,1231,1268,1294]" italics="true" pageId="8" pageNumber="77">divisotergata</emphasis>
</taxonomicName>
species group can be easily recognized by simple unarmed gonocoxite, inner gonostylus has lower beak with roundly swollen area and usually well-developed dorsal lobe of appendage of sternite 9. Ventral lobes of tergite 9 appear as small narrow sclerotised plates in 
<taxonomicName id="028508C2FF877F20182C11144244FAEB" authorityName="Alexander" authorityYear="1924" box="[381,572,1376,1403]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="verecunda" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F20182C111441EAFAEB" box="[381,402,1377,1402]" italics="true" pageId="8" pageNumber="77">T.</emphasis>
(
<emphasis id="F7F1AF53FF877F2018F2111441C0FAEB" box="[419,440,1377,1402]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F20189911154244FAEB" box="[456,572,1376,1402]" italics="true" pageId="8" pageNumber="77">verecunda</emphasis>
</taxonomicName>
, 
<taxonomicName id="028508C2FF877F201B1B1114436CFAEB" authorityName="Alexander" authorityYear="1963" box="[586,788,1376,1403]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="adungensis" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F201B1B11144218FAEB" box="[586,608,1377,1402]" italics="true" pageId="8" pageNumber="77">T.</emphasis>
(
<emphasis id="F7F1AF53FF877F201B21111442FDFAEB" box="[624,645,1377,1402]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F201BC71115436CFAEB" box="[662,788,1376,1402]" italics="true" pageId="8" pageNumber="77">adungensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="028508C2FF877F201A7311144398FAEB" authorityName="Alexander" authorityYear="1963" box="[802,992,1376,1403]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="zayulensis" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F201A731114434FFAEB" box="[802,823,1377,1402]" italics="true" pageId="8" pageNumber="77">T.</emphasis>
(
<emphasis id="F7F1AF53FF877F201A1911144325FAEB" box="[840,861,1377,1402]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F201A3C11154398FAEB" box="[877,992,1376,1402]" italics="true" pageId="8" pageNumber="77">zayulensis</emphasis>
</taxonomicName>
, as bilobed foot-shaped structures in 
<taxonomicName id="028508C2FF877F201CD911144120FA0F" authorityName="Alexander" authorityYear="1932" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="divisotergata" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F201CD9111445E5FAEB" box="[1416,1437,1377,1402]" italics="true" pageId="8" pageNumber="77">T.</emphasis>
(
<emphasis id="F7F1AF53FF877F2019F011F040CEFA0F" box="[161,182,1413,1438]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F20199411F14120FA0F" box="[197,344,1412,1438]" italics="true" pageId="8" pageNumber="77">divisotergata</emphasis>
</taxonomicName>
or as pale rods in 
<taxonomicName id="028508C2FF877F201B7411F04289FA0F" authorityName="Alexander" authorityYear="1963" box="[549,753,1413,1439]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="77" phylum="Arthropoda" rank="species" species="rongtoensis" subGenus="Vestiplex">
<emphasis id="F7F1AF53FF877F201B7411F04243FA0F" box="[549,571,1413,1438]" italics="true" pageId="8" pageNumber="77">T.</emphasis>
(
<emphasis id="F7F1AF53FF877F201B1B11F04218FA0F" box="[586,608,1413,1438]" italics="true" pageId="8" pageNumber="77">V.</emphasis>
) 
<emphasis id="F7F1AF53FF877F201B3E11F04289FA0F" box="[623,753,1413,1438]" italics="true" pageId="8" pageNumber="77">rongtoensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="A1140EB0FF877F201BAE11F143C0FA0E" author="Alexander, C. P." box="[767,952,1412,1439]" pageId="8" pageNumber="77" pagination="105 - 136" refId="ref10389" refString="Alexander, C. P. (1932) New or little-known Tipulidae from eastern Asia (Diptera). X. Philippine Journal of Science, 49, 105 - 136." type="journal article" year="1932">Alexander 1932</bibRefCitation>
, 
<bibRefCitation id="A1140EB0FF877F201A9211F14387FA0E" author="Alexander, C. P." box="[963,1023,1412,1439]" pageId="8" pageNumber="77" pagination="319 - 340" refId="ref10593" refString="Alexander, C. P. (1963) Some Tipulidae from Tibet and upper Burma in the British museum (Natural History) (Diptera). Bulletin of the British Museum (Natural History), Entomology, 14, 319 - 340. https: // doi. org / 10.5962 / bhl. part. 8785" type="journal article" year="1963">1963</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>