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<document id="5B3146635B7983B8436A5C8EF231883E" ID-DOI="10.11646/phytotaxa.664.2.6" ID-ISSN="1179-3163" ID-Zenodo-Dep="14517093" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1734541779643" checkinUser="felipe" docAuthor="Dang, Minh Quan, Averyanov, Leonid V., Maisak, Tatiana V., Wook, Um Sung, Canh, Nguyen Van, Phan, Xuan Thuy, Truong, Quang Tam, Wojtas, K. Philip &amp; Truong, Ba Vuong" docDate="2024" docId="2D1387C9E942FFC22E9AFABCFD4DFA39" docLanguage="en" docName="phytotaxa.664.2.139-145.pdf" docOrigin="Phytotaxa 664 (2)" docSource="https://doi.org/10.11646/phytotaxa.664.2.6" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Chiloschista eckhardii Vuong, Aver., V. C. Nguyen, Wojtas &amp; Q. T. Truong. A. Flowering 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="142" masterDocId="D12AFFB1E943FFC12E12FF88FFAFFF9D" masterDocTitle="Chiloschista eckhardii (Orchidaceae, Aeridinae), a new miniature leafless orchid from northern Vietnam" masterLastPageNumber="145" masterPageNumber="139" pageNumber="140" updateTime="1734542172153" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E05192A87E7338E4C654C9F36C75B269">Chiloschista eckhardii (Orchidaceae, Aeridinae), a new miniature leafless orchid from northern Vietnam</mods:title>
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<mods:namePart id="4D2AE4F655522E0350399F893F975E11">Dang, Minh Quan</mods:namePart>
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<mods:namePart id="E38258130401641D5388F0C537E2AF50">Averyanov, Leonid V.</mods:namePart>
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<mods:affiliation id="F44D0D9BFF02F9CAD5D3714B398F0EAF">Komarov Botanical Institute of the Russian Academy of Sciences, Prof. Popov Street 2, 197022, St. Petersburg, Russia</mods:affiliation>
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<mods:namePart id="1B2386055169CE4A5507A339849A9C79">Wook, Um Sung</mods:namePart>
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<mods:namePart id="D1688D225697F58090E7BBEE9BFA9E61">Canh, Nguyen Van</mods:namePart>
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<mods:namePart id="9C55EC89778BD8104B4F253F32CD977B">Phan, Xuan Thuy</mods:namePart>
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<mods:namePart id="35F1EFDF3AC458FA90A36B56BE48D915">Truong, Quang Tam</mods:namePart>
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<mods:namePart id="EDB88DFBDE0C1429CFE0E1F018260CD1">Wojtas, K. Philip</mods:namePart>
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<mods:namePart id="54418F50083FF4D4B89155FEE3452064">Truong, Ba Vuong</mods:namePart>
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<emphasis id="97CEEACDE942FFC02E9AFABCFE24FAD2" bold="true" box="[136,395,1332,1359]" italics="true" pageId="1" pageNumber="140">Chiloschista eckhardii</emphasis>
Vuong, Aver., V.C.Nguyen, Wojtas &amp; Q.T.Truong
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(
<figureCitation id="3D812A5AE942FFC02E82FAD0FF42FAEF" box="[144,237,1368,1394]" captionStart="FIGURE 1" captionStartId="2.[136,229,1883,1905]" captionTargetBox="[204,1384,187,1856]" captionTargetId="figure-17@2.[204,1384,187,1856]" captionTargetPageId="2" captionText="FIGURE 1. Chiloschista eckhardii Vuong, Aver., V.C.Nguyen, Wojtas &amp; Q.T.Truong. A. Flowering plant. B. Portion of rachis with open flowers. C. Floral bract, abaxial side. D. Flower, front view. E. Median sepal, abaxial and adaxial side. F. Lateral sepals, abaxial and adaxial side. G. Petal, abaxial and adaxial side. H. Ovary, column, column foot, and lip, side view. I. Lip, view from different sides. J. Apex of median lobe and apex of lip callus. K. Lip, tangential section. L. Ovary, column, and column foot, side view. M. Column and column foot, front view. N. Column apex with stigma. O. Anther cap, views from different sides. P. Pollinaria, front and view from rear. Photos by Ba Vuong Truong from the type specimens BV 1704. Photo-correction and design by Ba Vuong Truong and L. Averyanov." figureDoi="http://doi.org/10.5281/zenodo.14517099" httpUri="https://zenodo.org/record/14517099/files/figure.png" pageId="1" pageNumber="140">Figure 1</figureCitation>
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<emphasis id="97CEEACDE942FFC02E9AFA28FF6CFA27" bold="true" box="[136,195,1440,1466]" pageId="1" pageNumber="140">Type</emphasis>
:— 
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.
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<collectingRegion id="677EF83DE942FFC02F69FA29FDFBFA27" box="[379,596,1440,1467]" country="Vietnam" name="Ha Giang" pageId="1" pageNumber="140">Ha Giang Province</collectingRegion>
, Dong Van District, 
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, elev. around 
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, 
<emphasis id="97CEEACDE942FFC02AF2FA29FCF4FA43" italics="true" pageId="1" pageNumber="140">Truong Ba Vuong, Nguyen Van Canh, Phan Xuan Thuy &amp; Um Sung Wook, BV 1704</emphasis>
(
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, 
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00067722).
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<typeStatus id="7A01887DE942FFC02EAFFA60FE96F99F" box="[189,313,1512,1538]" pageId="1" pageNumber="140" type="paratype">Paratypes</typeStatus>
:— 
<collectingCountry id="DDAD764FE942FFC02F48FA60FE4CF99F" box="[346,483,1512,1538]" name="Vietnam" pageId="1" pageNumber="140">VIETNAM</collectingCountry>
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<collectingRegion id="677EF83DE942FFC02FE3FA61FD63F99F" box="[497,716,1512,1539]" country="Vietnam" name="Ha Giang" pageId="1" pageNumber="140">Ha Giang Province</collectingRegion>
, 
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, 
<collectorName id="084F5309E942FFC02B36FA61FF6AF9BB" pageId="1" pageNumber="140">Nguyen Van Canh</collectorName>
&amp; 
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1898
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(
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</materialsCitation>
; 
<materialsCitation id="15D23C82E942FFC02C68F984FEB1F9D7" collectingDate="2020-11-18" collectionCode="VNM" collectorName="Truong Ba Vuong &amp; Ngo Quang Dang, BV" country="Vietnam" pageId="1" pageNumber="140" specimenCount="1" typeStatus="paratype">
<emphasis id="97CEEACDE942FFC02C68F984FD69F9BB" box="[634,710,1548,1574]" italics="true" pageId="1" pageNumber="140">ibidem</emphasis>
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<collectingDate id="C140E9F7E942FFC02CC7F984FC1BF9BB" box="[725,948,1548,1574]" pageId="1" pageNumber="140" value="2020-11-18">18 November 2020</collectingDate>
</date>
, 
<emphasis id="97CEEACDE942FFC02DD2F985FF6FF9D7" italics="true" pageId="1" pageNumber="140">
<collectorName id="084F5309E942FFC02DD2F985FB26F9BB" box="[960,1161,1549,1574]" pageId="1" pageNumber="140">Truong Ba Vuong</collectorName>
&amp; 
<collectorName id="084F5309E942FFC02AA2F985FA03F9BB" box="[1200,1452,1548,1574]" pageId="1" pageNumber="140">Ngo Quang Dang, BV</collectorName>
1027
</emphasis>
(
<collectionCode id="C3ABAE1AE942FFC02EDDF9B8FEB9F9D7" box="[207,278,1584,1610]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="1" pageNumber="140" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="15D23C82E942FFC02F38F9B8FB3FF9D7" box="[298,1168,1584,1611]" collectingDate="2020-05-10" collectionCode="VNM" collectorName="Truong Ba Vuong &amp; Ngo Quang Dang, BV" country="Vietnam" pageId="1" pageNumber="140" specimenCount="1" typeStatus="paratype">
<emphasis id="97CEEACDE942FFC02F38F9B8FED9F9D7" box="[298,374,1584,1610]" italics="true" pageId="1" pageNumber="140">ibidem</emphasis>
<date id="D104101FE942FFC02F6FF9B8FDBAF9D7" box="[381,533,1584,1611]" pageId="1" pageNumber="140" value="2020-05-10">
<collectingDate id="C140E9F7E942FFC02F6FF9B8FDBAF9D7" box="[381,533,1584,1611]" pageId="1" pageNumber="140" value="2020-05-10">10 May 2020</collectingDate>
</date>
, 
<emphasis id="97CEEACDE942FFC02C32F9B9FB9EF9D7" box="[544,1073,1584,1610]" italics="true" pageId="1" pageNumber="140">
<collectorName id="084F5309E942FFC02C32F9B9FD4AF9D7" box="[544,741,1585,1610]" pageId="1" pageNumber="140">Truong Ba Vuong</collectorName>
&amp; 
<collectorName id="084F5309E942FFC02D1BF9B9FBAFF9D7" box="[777,1024,1584,1610]" pageId="1" pageNumber="140">Ngo Quang Dang, BV</collectorName>
746
</emphasis>
(
<collectionCode id="C3ABAE1AE942FFC02A52F9B8FB28F9D7" box="[1088,1159,1584,1610]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="1" pageNumber="140" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EDA06554E942FFC02EAFF9DCFC98F813" pageId="1" pageNumber="140" type="diagnosis">
<paragraph id="A50536DFE942FFC02EAFF9DCFC98F813" blockId="1.[136,1452,1440,2079]" pageId="1" pageNumber="140">
<emphasis id="97CEEACDE942FFC02EAFF9DCFE9FF9F3" bold="true" box="[189,304,1620,1646]" pageId="1" pageNumber="140">Diagnosis</emphasis>
:— 
<taxonomicName id="62BA4D5CE942FFC02F46F9DCFDFDF9F3" authorityName="Vuong, Aver., V. C. Nguyen, Wojtas &amp; Q. T. Truong. A. Flowering" authorityYear="2024" box="[340,594,1620,1646]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="140" phylum="Tracheophyta" rank="species" species="eckhardii">
<emphasis id="97CEEACDE942FFC02F46F9DCFDFDF9F3" box="[340,594,1620,1646]" italics="true" pageId="1" pageNumber="140">Chiloschista eckhardii</emphasis>
</taxonomicName>
in its morphology is most similar to 
<taxonomicName id="62BA4D5CE942FFC02DEFF9DCFBD9F9F3" authorityName="Seidenfaden" authorityYear="1988" box="[1021,1142,1620,1646]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="140" phylum="Tracheophyta" rank="species" species="parishii">
<emphasis id="97CEEACDE942FFC02DEFF9DCFBD9F9F3" box="[1021,1142,1620,1646]" italics="true" pageId="1" pageNumber="140">C. parishii</emphasis>
</taxonomicName>
and 
<taxonomicName id="62BA4D5CE942FFC02AA2F9DCFACFF9F3" box="[1200,1376,1620,1646]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="140" phylum="Tracheophyta" rank="species" species="glabrisepala">
<emphasis id="97CEEACDE942FFC02AA2F9DCFACFF9F3" box="[1200,1376,1620,1646]" italics="true" pageId="1" pageNumber="140">C. glabrisepala</emphasis>
</taxonomicName>
, but it clearly differs by having a lip callus that is densely hairy in the middle (versus the lip disc and lip callus being entirely hairy in 
<taxonomicName id="62BA4D5CE942FFC02EF5F914FEF0F92B" authorityName="Seidenfaden" authorityYear="1988" box="[231,351,1692,1718]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="140" phylum="Tracheophyta" rank="species" species="parishii">
<emphasis id="97CEEACDE942FFC02EF5F914FEF0F92B" box="[231,351,1692,1718]" italics="true" pageId="1" pageNumber="140">C. parishii</emphasis>
</taxonomicName>
, and the lip callus being hairy only at the apex in 
<taxonomicName id="62BA4D5CE942FFC02D8AF914FBE7F92B" box="[920,1096,1692,1718]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="140" phylum="Tracheophyta" rank="species" species="glabrisepala">
<emphasis id="97CEEACDE942FFC02D8AF914FBE7F92B" box="[920,1096,1692,1718]" italics="true" pageId="1" pageNumber="140">C. glabrisepala</emphasis>
</taxonomicName>
). 
<taxonomicName id="62BA4D5CE942FFC02A4DF914FAF3F92B" authorityName="Vuong, Aver., V. C. Nguyen, Wojtas &amp; Q. T. Truong. A. Flowering" authorityYear="2024" box="[1119,1372,1692,1718]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="140" phylum="Tracheophyta" rank="species" species="eckhardii">
<emphasis id="97CEEACDE942FFC02A4DF914FAF3F92B" box="[1119,1372,1692,1718]" italics="true" pageId="1" pageNumber="140">Chiloschista eckhardii</emphasis>
</taxonomicName>
is also similar to 
<taxonomicName id="62BA4D5CE942FFC02EEFF948FE02F947" box="[253,429,1728,1754]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="140" phylum="Tracheophyta" rank="species" species="quangdangii">
<emphasis id="97CEEACDE942FFC02EEFF948FE02F947" box="[253,429,1728,1754]" italics="true" pageId="1" pageNumber="140">C. quangdangii</emphasis>
</taxonomicName>
but can be distinguished by its disc callus, which is slightly shorter than the median lobe (versus the disc callus being much shorter than median lobe), triangular-ovate side lobes (vs. rhomboid or oblongelliptic side lobes), and large back-wall callus (vs. an insignificant back-wall callus). In addition, 
<taxonomicName id="62BA4D5CE942FFC02AA3F880FA03F8BF" authorityName="Vuong, Aver., V. C. Nguyen, Wojtas &amp; Q. T. Truong. A. Flowering" authorityYear="2024" box="[1201,1452,1800,1826]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="140" phylum="Tracheophyta" rank="species" species="eckhardii">
<emphasis id="97CEEACDE942FFC02AA3F880FA03F8BF" box="[1201,1452,1800,1826]" italics="true" pageId="1" pageNumber="140">Chiloschista eckhardii</emphasis>
</taxonomicName>
may be misidentified as 
<taxonomicName id="62BA4D5CE942FFC02F8BF8A4FDEBF8DB" authorityName="Schlechter" authorityYear="1919" box="[409,580,1836,1862]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="140" phylum="Tracheophyta" rank="species" species="yunnanensis">
<emphasis id="97CEEACDE942FFC02F8BF8A4FDEBF8DB" box="[409,580,1836,1862]" italics="true" pageId="1" pageNumber="140">C. yunnanensis</emphasis>
</taxonomicName>
, but it can be distinguished by its tepals, which are sparsely puberulent on the outer surface and margin (vs. a puberulent surface and ciliate margin), and by its spur-like lip (vs a broadly saccate lip). A detailed morphological comparison is presented in 
<tableCitation id="E8380364E942FFC02CCCF8FCFC9EF813" box="[734,817,1908,1934]" captionStart="TABLE 1" captionStartId="3.[136,224,1492,1516]" captionText="TABLE 1. Comparison of floral morphology among Chiloschista eckhardii, C. parishii, C. yunnanensis, C. glabrisepala, and C. quangdangii." pageId="1" pageNumber="140">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EDA06554E942FFC22EAFF810FB44FD10" lastPageId="3" lastPageNumber="142" pageId="1" pageNumber="140" type="description">
<paragraph id="A50536DFE942FFC22EAFF810FB97FDF4" blockId="1.[136,1452,1440,2079]" lastBlockId="3.[136,1452,158,1445]" lastPageId="3" lastPageNumber="142" pageId="1" pageNumber="140">
<emphasis id="97CEEACDE942FFC02EAFF810FEE8F82F" bold="true" box="[189,327,1944,1970]" pageId="1" pageNumber="140">Description</emphasis>
:— 
<emphasis id="97CEEACDE942FFC02F7EF810FE05F82F" bold="true" box="[364,426,1944,1970]" pageId="1" pageNumber="140">Herb</emphasis>
leafless, miniature, monopodial, with abbreviated stem 
<emphasis id="97CEEACDE942FFC02A2EF811FBF2F82F" box="[1084,1117,1945,1970]" italics="true" pageId="1" pageNumber="140">ca.</emphasis>
<quantity id="62429B3AE942FFC02A7AF810FB01F82E" box="[1128,1198,1944,1971]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="140" unit="mm" value="3.0">3 mm</quantity>
long and many long, glabrous, flattened, green roots. 
<emphasis id="97CEEACDE942FFC02FE6F834FD3DF84B" bold="true" box="[500,658,1980,2006]" pageId="1" pageNumber="140">Inflorescence</emphasis>
green, covered by short stiff white hairs; peduncle 
<quantity id="62429B3AE942FFC02AC4F834FA8CF84A" box="[1238,1315,1980,2007]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="140" unit="cm" value="2.5">2.5 cm</quantity>
long with 3 to 4 distant bracts; rachis 
<quantity id="62429B3AE942FFC02FBCF868FD83F866" box="[430,556,2016,2043]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" metricValueMax="8.7" metricValueMin="2.5" pageId="1" pageNumber="140" unit="cm" value="5.6" valueMax="8.7" valueMin="2.5">2.5–8.7 cm</quantity>
long, with 5–9 flowers; floral bracts brown, glabrous, ovate, 
<emphasis id="97CEEACDE942FFC02AFBF869FAA4F867" box="[1257,1291,2017,2042]" italics="true" pageId="1" pageNumber="140">ca.</emphasis>
<quantity id="62429B3AE942FFC02B01F868FAC4F866" box="[1299,1387,2016,2043]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="140" unit="mm" value="2.3">2.3 mm</quantity>
long, acute, shortly mucronate; pedicel and ovary green, 
<quantity id="62429B3AE942FFC02CACF78CFC9EF782" box="[702,817,2052,2079]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="1" pageNumber="140" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity>
, with densely white stiff hairs. 
<emphasis id="97CEEACDE942FFC02A9BF78CFB47F783" bold="true" box="[1161,1256,2052,2078]" pageId="1" pageNumber="140">Flowers</emphasis>
open acropetally, ca. 
<quantity id="62429B3AE940FFC22EA3FF16FF51FF24" box="[177,254,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="142" unit="cm" value="1.6">1.6 cm</quantity>
across, tepals yellow with red-brown dots, lip white or light yellow, with many red marks on side lobes, column foot broadly trapezoidal, ca. 
<quantity id="62429B3AE940FFC22C36FF4AFDCAFF40" box="[548,613,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="142" unit="mm" value="5.0">5 mm</quantity>
long and ca. 
<quantity id="62429B3AE940FFC22CE8FF4AFC94FF40" box="[762,827,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="142" unit="mm" value="4.0">4 mm</quantity>
wide, with many brown blotches. 
<emphasis id="97CEEACDE940FFC22AAFFF4AFAFFFF41" bold="true" box="[1213,1360,194,220]" pageId="3" pageNumber="142">Dorsal sepal</emphasis>
elliptic, obovate to broadly obovate, 
<quantity id="62429B3AE940FFC22FC3FF6EFD9FFE9C" box="[465,560,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="3" pageNumber="142" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
long, 
<quantity id="62429B3AE940FFC22C69FF6EFD75FE9C" box="[635,730,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="3" pageNumber="142" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">3–7 mm</quantity>
wide, apex rounded, abaxially with minute stiff hairs. 
<emphasis id="97CEEACDE940FFC22B41FF6EFF7FFEB9" bold="true" pageId="3" pageNumber="142">Lateral sepals</emphasis>
obovate, broadly obovate, 
<quantity id="62429B3AE940FFC22FEDFE82FDF4FEB8" box="[511,603,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="3" pageNumber="142" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
long, 
<quantity id="62429B3AE940FFC22CB3FE82FD52FEB8" box="[673,765,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="3" pageNumber="142" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
wide, apex rounded, abaxially with stiff hairs. 
<emphasis id="97CEEACDE940FFC22B1AFE82FAFFFEB9" bold="true" box="[1288,1360,266,292]" pageId="3" pageNumber="142">Petals</emphasis>
oblong, abaxially with minute stiff hairs, 
<quantity id="62429B3AE940FFC22C16FEA6FDCCFED4" box="[516,611,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="3" pageNumber="142" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
long, 
<quantity id="62429B3AE940FFC22CBFFEA6FC8EFED4" box="[685,801,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="3" pageNumber="142" unit="mm" value="3.25" valueMax="3.5" valueMin="3.0">3–3.5 mm</quantity>
wide, apex truncate. 
<emphasis id="97CEEACDE940FFC22A05FEA6FBEEFED5" bold="true" box="[1047,1089,302,328]" pageId="3" pageNumber="142">Lip</emphasis>
attached to the apex of column foot, 3 lobed, spur ovoid ca. 
<quantity id="62429B3AE940FFC22FC9FEDAFDB0FEF0" box="[475,543,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="142" unit="mm" value="2.0">2 mm</quantity>
in diam.; median lobe fleshy, obscurely rectangular, ca. 
<quantity id="62429B3AE940FFC22AA0FEDAFB59FEF0" box="[1202,1270,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="142" unit="mm" value="1.0">1 mm</quantity>
long and wide, rounded, bifid; side lobes erect, not embracing column, directed forward, 
<quantity id="62429B3AE940FFC22DCBFEFEFB97FE0C" box="[985,1080,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="3" pageNumber="142" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
long, 
<quantity id="62429B3AE940FFC22A91FEFEFB4DFE0C" box="[1155,1250,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="142" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
wide, rounded at apex, adaxially with many warty streaks; central callus Y-shaped, covered with stiff and clavate hairs, apex bilobed, downcurved, shorter than median lobe, glabrous; spur backwall fleshy, covered by minute hairs; front wall callus densely hairy along median keel. 
<emphasis id="97CEEACDE940FFC22C11FE6AFDCCFE61" bold="true" box="[515,611,482,508]" pageId="3" pageNumber="142">Column</emphasis>
stout, column foot ovate-oblong, slightly incurved, ca. 
<quantity id="62429B3AE940FFC22AC5FE6AFAB7FE60" box="[1239,1304,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="142" unit="mm" value="4.0">4 mm</quantity>
long; stigma obscurely rectangular; anther cap obscurely cubic, 
<emphasis id="97CEEACDE940FFC22CDEFD8FFD42FDBD" box="[716,749,519,544]" italics="true" pageId="3" pageNumber="142">ca.</emphasis>
<quantity id="62429B3AE940FFC22CE4FD8EFC97FDBC" box="[758,824,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="142" unit="mm" value="1.0">1 mm</quantity>
long and wide, with 2 filiform setae from each side of anther cap, 
<emphasis id="97CEEACDE940FFC22F03FDA3FE9CFDD9" box="[273,307,555,580]" italics="true" pageId="3" pageNumber="142">ca.</emphasis>
<quantity id="62429B3AE940FFC22F2CFDA2FE38FDD8" box="[318,407,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="142" unit="mm" value="1.5">1.5 mm</quantity>
long; pollinarium with 2 almost globose notched pollinia, arrow-shaped stipe broadening distally, slightly contracted at the middle, and rectangular viscidium. 
<emphasis id="97CEEACDE940FFC22D9CFDC6FC63FDF5" bold="true" box="[910,972,590,616]" pageId="3" pageNumber="142">Fruit</emphasis>
not seen.
</paragraph>
<caption id="F1C56657E941FFC32E9AF8D3FA2BF7B8" ID-DOI="http://doi.org/10.5281/zenodo.14517099" ID-Zenodo-Dep="14517099" httpUri="https://zenodo.org/record/14517099/files/figure.png" pageId="2" pageNumber="141" startId="2.[136,229,1883,1905]" targetBox="[204,1384,187,1856]" targetPageId="2" targetType="figure">
<paragraph id="A50536DFE941FFC32E9AF8D3FA2BF7B8" blockId="2.[136,1453,1883,2085]" pageId="2" pageNumber="141">
<emphasis id="97CEEACDE941FFC32E9AF8D3FEAFF8ED" bold="true" box="[136,256,1883,1905]" pageId="2" pageNumber="141">FIGURE 1.</emphasis>
<taxonomicName id="62BA4D5CE941FFC32F1AF8D3FBDFF8EC" authority="Vuong, Aver., V. C. Nguyen, Wojtas &amp; Q. T. Truong. A. Flowering" authorityName="Vuong, Aver., V. C. Nguyen, Wojtas &amp; Q. T. Truong. A. Flowering" authorityYear="2024" box="[264,1136,1883,1905]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="141" phylum="Tracheophyta" rank="species" species="eckhardii">
<emphasis id="97CEEACDE941FFC32F1AF8D3FE47F8EC" bold="true" box="[264,488,1883,1905]" italics="true" pageId="2" pageNumber="141">Chiloschista eckhardii</emphasis>
Vuong, Aver., V.C.Nguyen, Wojtas &amp; Q.T.Truong. 
<emphasis id="97CEEACDE941FFC32DFFF8D3FBAAF8ED" bold="true" box="[1005,1029,1883,1904]" pageId="2" pageNumber="141">A.</emphasis>
Flowering
</taxonomicName>
plant. 
<emphasis id="97CEEACDE941FFC32AA4F8D4FB63F8EC" bold="true" box="[1206,1228,1884,1905]" pageId="2" pageNumber="141">B.</emphasis>
Portion of rachis with open flowers. 
<emphasis id="97CEEACDE941FFC32F06F8F7FE84F808" bold="true" box="[276,299,1919,1941]" pageId="2" pageNumber="141">C.</emphasis>
Floral bract, abaxial side. 
<emphasis id="97CEEACDE941FFC32C20F808FDE5F808" bold="true" box="[562,586,1920,1941]" pageId="2" pageNumber="141">D.</emphasis>
Flower, front view. 
<emphasis id="97CEEACDE941FFC32D01F808FC86F808" bold="true" box="[787,809,1920,1941]" pageId="2" pageNumber="141">E.</emphasis>
Median sepal, abaxial and adaxial side. 
<emphasis id="97CEEACDE941FFC32AAAF808FB65F808" bold="true" box="[1208,1226,1920,1941]" pageId="2" pageNumber="141">F.</emphasis>
Lateral sepals, abaxial and adaxial side. 
<emphasis id="97CEEACDE941FFC32F22F82BFEE6F824" bold="true" box="[304,329,1955,1977]" pageId="2" pageNumber="141">G.</emphasis>
Petal, abaxial and adaxial side. 
<emphasis id="97CEEACDE941FFC32C6DF82CFD37F824" bold="true" box="[639,664,1956,1977]" pageId="2" pageNumber="141">H.</emphasis>
Ovary, column, column foot, and lip, side view. 
<emphasis id="97CEEACDE941FFC32A7DF82CFBD0F824" bold="true" box="[1135,1151,1956,1977]" pageId="2" pageNumber="141">I.</emphasis>
Lip, view from different sides. 
<emphasis id="97CEEACDE941FFC32E9AF840FF35F840" bold="true" box="[136,154,1992,2013]" pageId="2" pageNumber="141">J.</emphasis>
Apex of median lobe and apex of lip callus. 
<emphasis id="97CEEACDE941FFC32C58F840FDCCF840" bold="true" box="[586,611,1992,2013]" pageId="2" pageNumber="141">K.</emphasis>
Lip, tangential section. 
<emphasis id="97CEEACDE941FFC32D59F840FCCDF840" bold="true" box="[843,866,1992,2013]" pageId="2" pageNumber="141">L.</emphasis>
Ovary, column, and column foot, side view. 
<emphasis id="97CEEACDE941FFC32B00F840FA81F840" bold="true" box="[1298,1326,1992,2013]" pageId="2" pageNumber="141">M.</emphasis>
Column and column foot, front view. 
<emphasis id="97CEEACDE941FFC32F6AF864FE3FF79C" bold="true" box="[376,400,2028,2049]" pageId="2" pageNumber="141">N.</emphasis>
Column apex with stigma. 
<emphasis id="97CEEACDE941FFC32C88F863FD1CF79C" bold="true" box="[666,691,2027,2049]" pageId="2" pageNumber="141">O.</emphasis>
Anther cap, views from different sides. 
<emphasis id="97CEEACDE941FFC32A25F864FBE6F79C" bold="true" box="[1079,1097,2028,2049]" pageId="2" pageNumber="141">P.</emphasis>
Pollinaria, front and view from rear. Photos by Ba Vuong Truong from the type specimens 
<emphasis id="97CEEACDE941FFC32C82F798FD4CF7B8" box="[656,739,2064,2085]" italics="true" pageId="2" pageNumber="141">BV 1704</emphasis>
. Photo-correction and design by Ba Vuong Truong and L. Averyanov.
</paragraph>
</caption>
<paragraph id="A50536DFE940FFC22EAFFDFAFB44FD10" blockId="3.[136,1452,158,1445]" box="[189,1259,626,653]" pageId="3" pageNumber="142">
<emphasis id="97CEEACDE940FFC22EAFFDFAFE16FD11" bold="true" box="[189,441,626,652]" pageId="3" pageNumber="142">Flowering phenology</emphasis>
: Plants in flower were observed in late November, and in March to May.
</paragraph>
</subSubSection>
<subSubSection id="EDA06554E940FFC22EAFFD1EFE22FD49" pageId="3" pageNumber="142" type="etymology">
<paragraph id="A50536DFE940FFC22EAFFD1EFE22FD49" blockId="3.[136,1452,158,1445]" pageId="3" pageNumber="142">
<emphasis id="97CEEACDE940FFC22EAFFD1EFE91FD2D" bold="true" box="[189,318,662,688]" pageId="3" pageNumber="142">Etymology</emphasis>
:—The species is named after the late Dr. Eckhard von Raab-Straube, an enthusiastic botanist from the Botanic Garden 
<collectingRegion id="677EF83DE940FFC22F52FD32FE25FD49" box="[320,394,698,724]" country="Germany" name="Berlin" pageId="3" pageNumber="142">Berlin</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="EDA06554E940FFC22EAFFD56FD4DFA39" pageId="3" pageNumber="142" type="distribution">
<paragraph id="A50536DFE940FFC22EAFFD56FA92FD65" blockId="3.[136,1452,158,1445]" box="[189,1341,734,761]" pageId="3" pageNumber="142">
<emphasis id="97CEEACDE940FFC22EAFFD56FEE0FD65" bold="true" box="[189,335,734,760]" pageId="3" pageNumber="142">Distribution</emphasis>
:—The species has so far been reported only from the 
<typeStatus id="7A01887DE940FFC22DA2FD57FC4FFD64" box="[944,992,735,761]" pageId="3" pageNumber="142">type</typeStatus>
locality in 
<collectingRegion id="677EF83DE940FFC22A4DFD57FA96FD65" box="[1119,1337,734,761]" country="Vietnam" name="Ha Giang" pageId="3" pageNumber="142">Ha Giang Province</collectingRegion>
.
</paragraph>
<paragraph id="A50536DFE940FFC22EAFFC8AFAC1FC81" blockId="3.[136,1452,158,1445]" box="[189,1390,770,797]" pageId="3" pageNumber="142">
<emphasis id="97CEEACDE940FFC22EAFFC8AFE02FC81" bold="true" box="[189,429,770,796]" pageId="3" pageNumber="142">Habitat and ecology</emphasis>
:—Branch epiphyte, in evergreen, broad-leaved forest at elevation about 
<quantity id="62429B3AE940FFC22ACCFC8AFA9CFC80" box="[1246,1331,770,797]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="142" unit="m" value="1300.0">1300 m</quantity>
a.s.l.
</paragraph>
<paragraph id="A50536DFE940FFC22EAFFCAEFA85FBFD" blockId="3.[136,1452,158,1445]" pageId="3" pageNumber="142">
<emphasis id="97CEEACDE940FFC22EAFFCAEFE5EFCDD" bold="true" box="[189,497,806,832]" pageId="3" pageNumber="142">Other Specimens studied</emphasis>
:— 
<emphasis id="97CEEACDE940FFC22C07FCAEFDCAFCDD" bold="true" box="[533,613,806,832]" pageId="3" pageNumber="142">
<collectingCountry id="DDAD764FE940FFC22C07FCAEFDCFFCDD" box="[533,608,806,832]" name="China" pageId="3" pageNumber="142">China</collectingCountry>
.
</emphasis>
<collectingRegion id="677EF83DE940FFC22C63FCAFFD65FCDC" box="[625,714,807,833]" country="China" name="Yunnan" pageId="3" pageNumber="142">Yunnan</collectingRegion>
, Szemao, 
<quantity id="62429B3AE940FFC22D57FCAEFC33FCDC" box="[837,924,806,833]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="142" unit="m" value="1500.0">1500 m</quantity>
, 
<emphasis id="97CEEACDE940FFC22DBFFCAFFBD8FCDD" box="[941,1143,806,833]" italics="true" pageId="3" pageNumber="142">A. Henry 11792A</emphasis>
(AMES00106704, https:// data.huh.harvard.edu/databases/specimen_search.php?mode=details&amp;id=38025) (AMES!- 
<typeStatus id="7A01887DE940FFC22A95FCC2FB71FCF9" box="[1159,1246,842,868]" pageId="3" pageNumber="142" type="isotype">Isotype</typeStatus>
of 
<taxonomicName id="62BA4D5CE940FFC22B0DFCC2FEC7FC15" authority="Schltr." authorityName="Schlechter" authorityYear="1919" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="yunnanensis">
<emphasis id="97CEEACDE940FFC22B0DFCC2FEBDFC15" italics="true" pageId="3" pageNumber="142">Chiloschista yunnanensis</emphasis>
Schltr.
</taxonomicName>
, image seen); 
<emphasis id="97CEEACDE940FFC22C1DFCE6FDF4FC15" box="[527,603,878,904]" italics="true" pageId="3" pageNumber="142">ibidem</emphasis>
, 
<emphasis id="97CEEACDE940FFC22C79FCE7FC9EFC15" box="[619,817,878,905]" italics="true" pageId="3" pageNumber="142">A. Henry 11792A</emphasis>
(E01313787, https://data.rbge.org.uk/search/herbarium/?cfg=fulldetails.cfg&amp;barcode=E01313787) (E!- 
<typeStatus id="7A01887DE940FFC22C85FC1AFD5EFC31" box="[663,753,914,940]" pageId="3" pageNumber="142" type="isotype">Isotype</typeStatus>
of 
<taxonomicName id="62BA4D5CE940FFC22D06FC1BFC12FC31" authorityName="Schlechter" authorityYear="1919" box="[788,957,915,941]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="yunnanensis">
<emphasis id="97CEEACDE940FFC22D06FC1BFC12FC31" box="[788,957,915,941]" italics="true" pageId="3" pageNumber="142">C. yunnanensis</emphasis>
</taxonomicName>
, image seen); 
<emphasis id="97CEEACDE940FFC22A4FFC1AFB06FC31" box="[1117,1193,914,940]" italics="true" pageId="3" pageNumber="142">ibidem</emphasis>
, 
<emphasis id="97CEEACDE940FFC22AA4FC1BFADAFC31" box="[1206,1397,914,941]" italics="true" pageId="3" pageNumber="142">A. Henry 11792A</emphasis>
(NY 4341314, https://sweetgum.nybg.org/science/vh/specimen-details/?irn=4341314) (NY!, - 
<typeStatus id="7A01887DE940FFC22A67FC3FFB66FC4C" box="[1141,1225,951,977]" pageId="3" pageNumber="142" type="isotype">Isotype</typeStatus>
of 
<taxonomicName id="62BA4D5CE940FFC22AE4FC3FFA0BFC4D" authorityName="Schlechter" authorityYear="1919" box="[1270,1444,951,977]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="yunnanensis">
<emphasis id="97CEEACDE940FFC22AE4FC3FFA0BFC4D" box="[1270,1444,951,977]" italics="true" pageId="3" pageNumber="142">C. yunnanensis</emphasis>
</taxonomicName>
, image seen); 
<collectingRegion id="677EF83DE940FFC22F09FC53FEDBFC68" box="[283,372,987,1013]" country="China" name="Yunnan" pageId="3" pageNumber="142">Yunnan</collectingRegion>
, Mengtsz, 
<emphasis id="97CEEACDE940FFC22FFEFC53FD36FC69" box="[492,665,986,1013]" italics="true" pageId="3" pageNumber="142">A. Henry 11129</emphasis>
(K001599317) (K!, 
<taxonomicName id="62BA4D5CE940FFC22D6DFC53FB86FC69" authorityName="Schlechter" authorityYear="1919" box="[895,1065,987,1013]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="yunnanensis">
<emphasis id="97CEEACDE940FFC22D6DFC53FB86FC69" box="[895,1065,987,1013]" italics="true" pageId="3" pageNumber="142">C. yunnanensis</emphasis>
</taxonomicName>
, image seen), duplicate of a single flower as spirit sample at C (C!, 
<taxonomicName id="62BA4D5CE940FFC22FFBFC77FD3DFB85" authorityName="Schlechter" authorityYear="1919" box="[489,658,1023,1049]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="yunnanensis">
<emphasis id="97CEEACDE940FFC22FFBFC77FD3DFB85" box="[489,658,1023,1049]" italics="true" pageId="3" pageNumber="142">C. yunnanensis</emphasis>
</taxonomicName>
, images seen). 
<emphasis id="97CEEACDE940FFC22D2FFC76FC7BFB85" bold="true" box="[829,980,1022,1048]" pageId="3" pageNumber="142">
<collectingCountry id="DDAD764FE940FFC22D2FFC76FC7BFB85" box="[829,980,1022,1048]" name="Myanmar" pageId="3" pageNumber="142">MYANMAR</collectingCountry>
</emphasis>
. Moulmein, 
<emphasis id="97CEEACDE940FFC22A71FC76FABCFB84" box="[1123,1299,1022,1049]" italics="true" pageId="3" pageNumber="142">C.S.P.Parish 55</emphasis>
/ 
<emphasis id="97CEEACDE940FFC22B09FC76FEC0FBA1" italics="true" pageId="3" pageNumber="142">Reichenbach Herb. Orchid. 26237</emphasis>
(
<emphasis id="97CEEACDE940FFC22F6CFBAAFE57FBA1" box="[382,504,1058,1084]" italics="true" pageId="3" pageNumber="142">W0017162</emphasis>
, https://w.jacq.org/W0017162) (W!- 
<typeStatus id="7A01887DE940FFC22D9AFBAAFC5EFBA1" box="[904,1009,1058,1084]" pageId="3" pageNumber="142" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="62BA4D5CE940FFC22A01FBAAFACCFBA1" authority="Seidenf." authorityName="Seidenfaden" authorityYear="1988" box="[1043,1379,1058,1084]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="parishii">
<emphasis id="97CEEACDE940FFC22A01FBAAFB54FBA1" box="[1043,1275,1058,1084]" italics="true" pageId="3" pageNumber="142">Chiloschista parishii</emphasis>
Seidenf.
</taxonomicName>
); icon, 
<emphasis id="97CEEACDE940FFC22E9AFBCEFDA8FBFD" box="[136,519,1094,1120]" italics="true" pageId="3" pageNumber="142">Reichenbach Herb. Orchid. 11583</emphasis>
(
<emphasis id="97CEEACDE940FFC22C0AFBCEFD3EFBFD" box="[536,657,1094,1120]" italics="true" pageId="3" pageNumber="142">W0017165</emphasis>
, https://w.jacq.org/W0017165) (W!- 
<typeStatus id="7A01887DE940FFC22A37FBCEFBD3FBFD" box="[1061,1148,1094,1120]" pageId="3" pageNumber="142" type="isotype">isotype</typeStatus>
of 
<taxonomicName id="62BA4D5CE940FFC22AB0FBCFFAB5FBFD" authorityName="Seidenfaden" authorityYear="1988" box="[1186,1306,1094,1121]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="parishii">
<emphasis id="97CEEACDE940FFC22AB0FBCFFAB5FBFD" box="[1186,1306,1094,1121]" italics="true" pageId="3" pageNumber="142">C. parishii</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="A50536DFE940FFC22EAFFBE2FAB5FA89" blockId="3.[136,1452,158,1445]" pageId="3" pageNumber="142">
<emphasis id="97CEEACDE940FFC22EAFFBE2FF5AFB19" bold="true" box="[189,245,1130,1156]" pageId="3" pageNumber="142">Note</emphasis>
:—The newly discovered plant belongs to a group of species with flowers that have yellow sepals and petals spotted with orange-brown or red-brown dots, which led to the new species being overlooked and misidentified as 
<taxonomicName id="62BA4D5CE940FFC22EB8FB3AFE37FB51" authorityName="Seidenfaden" authorityYear="1988" box="[170,408,1202,1228]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="parishii">
<emphasis id="97CEEACDE940FFC22EB8FB3AFE37FB51" box="[170,408,1202,1228]" italics="true" pageId="3" pageNumber="142">Chiloschista parishii</emphasis>
</taxonomicName>
. However, the newly described species differs from morphologically similar 
<emphasis id="97CEEACDE940FFC22B0DFB3AFE35FB6D" italics="true" pageId="3" pageNumber="142">
<taxonomicName id="62BA4D5CE940FFC22B0DFB3AFF4DFB6D" authorityName="Seidenfaden" authorityYear="1988" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="parishii">Chiloschista parishii</taxonomicName>
, 
<taxonomicName id="62BA4D5CE940FFC22EFEFB5FFE35FB6D" box="[236,410,1238,1265]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="glabrisepala">C. glabrisepala</taxonomicName>
</emphasis>
, 
<taxonomicName id="62BA4D5CE940FFC22FB5FB5FFDFAFB6D" box="[423,597,1238,1265]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="quangdangii">
<emphasis id="97CEEACDE940FFC22FB5FB5FFDFAFB6D" box="[423,597,1238,1265]" italics="true" pageId="3" pageNumber="142">C. quangdangii</emphasis>
</taxonomicName>
and 
<taxonomicName id="62BA4D5CE940FFC22C9BFB5FFC9CFB6D" authorityName="Schlechter" authorityYear="1919" box="[649,819,1239,1265]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="yunnanensis">
<emphasis id="97CEEACDE940FFC22C9BFB5FFC9CFB6D" box="[649,819,1239,1265]" italics="true" pageId="3" pageNumber="142">C. yunnanensis</emphasis>
</taxonomicName>
in many features, as detailed in 
<tableCitation id="E8380364E940FFC22A8AFB5EFB44FB6D" box="[1176,1259,1238,1264]" captionStart="TABLE 1" captionStartId="3.[136,224,1492,1516]" captionText="TABLE 1. Comparison of floral morphology among Chiloschista eckhardii, C. parishii, C. yunnanensis, C. glabrisepala, and C. quangdangii." pageId="3" pageNumber="142">Table 1</tableCitation>
. The comparison of the floral morphology among 
<taxonomicName id="62BA4D5CE940FFC22FE4FB73FD2FFA89" authorityName="Vuong, Aver., V. C. Nguyen, Wojtas &amp; Q. T. Truong. A. Flowering" authorityYear="2024" box="[502,640,1274,1301]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="eckhardii">
<emphasis id="97CEEACDE940FFC22FE4FB73FD2FFA89" box="[502,640,1274,1301]" italics="true" pageId="3" pageNumber="142">C. eckhardii</emphasis>
</taxonomicName>
, 
<taxonomicName id="62BA4D5CE940FFC22C9CFB73FC93FA89" box="[654,828,1274,1301]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="glabrisepala">
<emphasis id="97CEEACDE940FFC22C9CFB73FC93FA89" box="[654,828,1274,1301]" italics="true" pageId="3" pageNumber="142">C. glabrisepala</emphasis>
</taxonomicName>
, and 
<taxonomicName id="62BA4D5CE940FFC22D6BFB73FB87FA89" box="[889,1064,1274,1301]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="quangdangii">
<emphasis id="97CEEACDE940FFC22D6BFB73FB87FA89" box="[889,1064,1274,1301]" italics="true" pageId="3" pageNumber="142">C. quangdangii</emphasis>
</taxonomicName>
is shown in 
<figureCitation id="3D812A5AE940FFC22AA7FB72FAB9FA89" box="[1205,1302,1274,1300]" captionStart="FIGURE 2" captionStartId="4.[136,229,1940,1962]" captionTargetBox="[151,1436,985,1916]" captionTargetId="figure-252@4.[151,1436,985,1916]" captionTargetPageId="4" captionText="FIGURE 2. Morphological comparison of Chiloschista eckhardii (A), C. glabrisepala (B), and C. quangdangii (C)." figureDoi="http://doi.org/10.5281/zenodo.14517101" httpUri="https://zenodo.org/record/14517101/files/figure.png" pageId="3" pageNumber="142">Figure 2</figureCitation>
.
</paragraph>
<paragraph id="A50536DFE940FFC22EAFFA96FD4DFA39" blockId="3.[136,1452,158,1445]" pageId="3" pageNumber="142">
The morphological details of 
<taxonomicName id="62BA4D5CE940FFC22C0AFA96FCA9FAA5" authorityName="Seidenfaden" authorityYear="1988" box="[536,774,1310,1336]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="parishii">
<emphasis id="97CEEACDE940FFC22C0AFA96FCA9FAA5" box="[536,774,1310,1336]" italics="true" pageId="3" pageNumber="142">Chiloschista parishii</emphasis>
</taxonomicName>
were compiled from 
<bibRefCitation id="C12B4B2EE940FFC22A15FA96FAA4FAA5" author="Gyeltshen, C. &amp; Dalstrom, S. &amp; Gyeltshen, N. &amp; Tobgay, K." box="[1031,1291,1310,1336]" pageId="3" pageNumber="142" pagination="23 - 29" refId="ref4091" refString="Gyeltshen, C., Dalstrom, S., Gyeltshen, N. &amp; Tobgay, K. (2019) A new spotted Chiloschista (Orchidaceae: Aeridinae) from Bhutan. Lankesteriana 19 (1): 23 - 29. https: // doi. org / 10.15517 / lank. v 19 i 1.37030" type="journal article" year="2019">
Gyeltshen 
<emphasis id="97CEEACDE940FFC22A91FA97FB10FAA5" box="[1155,1215,1310,1336]" italics="true" pageId="3" pageNumber="142">et al.</emphasis>
(2019
</bibRefCitation>
&amp; 
<bibRefCitation id="C12B4B2EE940FFC22B25FA96FADEFAA5" author="Gyeltshen, N. &amp; Gyeltshen, C. &amp; Tobgay, K. &amp; Dalstrom, S. &amp; Gurung, D. B. &amp; Gyeltshen N. &amp; Ghalley, B. B." box="[1335,1393,1310,1336]" pageId="3" pageNumber="142" pagination="281 - 299" refId="ref4150" refString="Gyeltshen, N., Gyeltshen, C., Tobgay, K., Dalstrom, S., Gurung, D. B., Gyeltshen N. &amp; Ghalley, B. B. (2020) Two new spotted Chiloschista species (Orchidaceae: Aeridinae) from Bhutan. Lankesteriana 20 (3): 281 - 299. https: // doi. org / 10.15517 / lank. v 20 i 3.44149." type="journal article" year="2020">2020</bibRefCitation>
) and 
<bibRefCitation id="C12B4B2EE940FFC22E9AFACAFE54FAC1" author="Dalstrom, S. &amp; Kolanowska, M." box="[136,507,1346,1372]" pageId="3" pageNumber="142" pagination="241 - 248" refId="ref3823" refString="Dalstrom, S. &amp; Kolanowska, M. (2020) A new yellow-flowered Chiloschista (Orchidaceae: Aeridinae) from Thailand. Lankesteriana 20 (2): 241 - 248. https: // doi. org / 10.15517 / lank. v 20 i 2.43454" type="journal article" year="2020">Dalström &amp; Kolanowska (2020)</bibRefCitation>
. The morphological details of 
<taxonomicName id="62BA4D5CE940FFC22D49FACBFBA8FAC1" authorityName="Schlechter" authorityYear="1919" box="[859,1031,1347,1373]" class="Liliopsida" family="Orchidaceae" genus="Chiloschista" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="142" phylum="Tracheophyta" rank="species" species="yunnanensis">
<emphasis id="97CEEACDE940FFC22D49FACBFBA8FAC1" box="[859,1031,1347,1373]" italics="true" pageId="3" pageNumber="142">C. yunnanensis</emphasis>
</taxonomicName>
based on of Schlechter’s protologue (
<bibRefCitation id="C12B4B2EE940FFC22E9DFAEEFEE2FA1D" author="Schlechter, R." box="[143,333,1382,1408]" pageId="3" pageNumber="142" refId="ref4420" refString="Schlechter, R. (1919) Chiloschista yunnanensis. In: Orchideologiae Sino-Japonicae Prodromus. Eine kritiche Besprechung der Orchideen Ost-Asiens. Repertorium specierum novarum regni vegetabilis. Beihefte 4: 74." type="journal volume" year="1919">Schlechter 1919</bibRefCitation>
), 
<bibRefCitation id="C12B4B2EE940FFC22F70FAEEFDF2FA1D" author="Chen, X. Q. &amp; Wood, J. J." box="[354,605,1382,1408]" pageId="3" pageNumber="142" pagination="1 - 506" refId="ref3748" refString="Chen, X. Q. &amp; Wood, J. J. (2009) Chiloschista Lindley. Orchidaceae. In: Wu, Z. Y., Raven, P. H. &amp; Hong, D. Y. (Eds.) Flora of China, vol. 25. Science Press, Beijing, &amp; Missouri Botanical Garden Press, Saint Louis, pp. 1 - 506." type="book chapter" year="2009">Chen &amp; Wood (2009)</bibRefCitation>
, and the specimens 
<emphasis id="97CEEACDE940FFC22D57FAEFFBAFFA1D" box="[837,1024,1382,1409]" italics="true" pageId="3" pageNumber="142">A.Henry 11792A</emphasis>
(AMES 00106704, E 01313787, NY 4341314), and 
<emphasis id="97CEEACDE940FFC22F23FA03FE70FA39" box="[305,479,1418,1445]" italics="true" pageId="3" pageNumber="142">A. Henry 11129</emphasis>
(K 001599317 and C).
</paragraph>
</subSubSection>
</treatment>
</document>