treatments-xml/data/22/8C/96/228C962509E1470F58AE17EC00D94D1F.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.768.24423" ID-PMC="PMC6019434" ID-Pensoft-Pub="1313-2970-768-113" ID-Pensoft-UUID="CB01CA30FFB4F96DFFA2FF86FFAEFFB6" ID-PubMed="29955215" ID-Zenodo-Dep="1298776" ID-ZooBank="399BCC3E9D6F4231870E05C79B9FD4B0" ModsDocID="1313-2970-768-113" checkinTime="1529468775230" checkinUser="pensoft" docAuthor="Giron, Jennifer C. &amp; Short, Andrew Edward Z." docDate="2018" docId="228C962509E1470F58AE17EC00D94D1F" docLanguage="en" docName="ZooKeys 768: 113-158" docOrigin="ZooKeys 768" docPubDate="2018-06-19" docSource="http://dx.doi.org/10.3897/zookeys.768.24423" docTitle="Chasmogenus cremnobates Girón &amp; Short 2018, comb. n." docType="treatment" docVersion="6" id="CB01CA30FFB4F96DFFA2FF86FFAEFFB6" lastPageNumber="148" masterDocId="CB01CA30FFB4F96DFFA2FF86FFAEFFB6" masterDocTitle="Three new genera of acidocerine water scavenger beetles from tropical South America (Coleoptera, Hydrophilidae, Acidocerinae)" masterLastPageNumber="158" masterPageNumber="113" pageNumber="147" updateTime="1668165917053" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Three new genera of acidocerine water scavenger beetles from tropical South America (Coleoptera, Hydrophilidae, Acidocerinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Giron, Jennifer C.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0851-6883</mods:nameIdentifier>
<mods:affiliation>Department of Ecology &amp; Evolutionary Biology, and Division of Entomology, Biodiversity Institute, University of Kansas, Lawrence, KS 66045, USA</mods:affiliation>
<mods:nameIdentifier type="email">entiminae@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Short, Andrew Edward Z.</mods:namePart>
<mods:affiliation>Department of Ecology &amp; Evolutionary Biology, and Division of Entomology, Biodiversity Institute, University of Kansas, Lawrence, KS 66045, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-06-19</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>768</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>113</mods:start>
<mods:end>158</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.768.24423</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.768.24423</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-768-113</mods:identifier>
<mods:identifier type="ZooBank">399BCC3E9D6F4231870E05C79B9FD4B0</mods:identifier>
<mods:identifier type="Pensoft-UUID">CB01CA30FFB4F96DFFA2FF86FFAEFFB6</mods:identifier>
<mods:identifier type="Zenodo-Dep">1298776</mods:identifier>
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<subSection lastPageId="35" lastPageNumber="148" pageId="34" pageNumber="147" type="taxonomy">
<treatment ID-GBIF-Taxon="146002390" LSID="urn:lsid:plazi:treatment:228C962509E1470F58AE17EC00D94D1F" httpUri="http://treatment.plazi.org/id/228C962509E1470F58AE17EC00D94D1F" lastPageId="35" lastPageNumber="148" pageId="34" pageNumber="147">
<subSubSection pageId="34" pageNumber="147" type="nomenclature">
<paragraph pageId="34" pageNumber="147">
<taxonomicName LSID="228C9625-09E1-470F-58AE-17EC00D94D1F" authority="(Spangler, 1979)" authorityName="Girón &amp; Short" authorityYear="2018" baseAuthorityName="Spangler" baseAuthorityYear="1979" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chasmogenus cremnobates" order="Coleoptera" pageId="34" pageNumber="147" phylum="Arthropoda" rank="species" species="cremnobates" status="comb. n.">Chasmogenus cremnobates (Spangler, 1979)</taxonomicName>
<taxonomicNameLabel pageId="34" pageNumber="147">comb. n.</taxonomicNameLabel>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Habitus of Chasmogenus cremnobates: A dorsal view B lateral view C ventral view D head E aedeagus. Scale bar 1 mm." figureDoi="10.3897/zookeys.768.24423.figure21" httpUri="https://binary.pensoft.net/fig/211234" pageId="34" pageNumber="147">Fig. 21</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="148" type="reference_group">
<paragraph pageId="35" pageNumber="148">
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Dieroxenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dieroxenus cremnobates" order="Coleoptera" pageId="35" pageNumber="148" phylum="Arthropoda" rank="species" species="cremnobates" status="comb. n.">
<pageBreakToken pageId="35" pageNumber="148" start="start">Dieroxenus</pageBreakToken>
cremnobates
</taxonomicName>
Spangler, 1979: 754
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="148" type="type material">
<paragraph pageId="35" pageNumber="148">Type material examined.</paragraph>
<paragraph pageId="35" pageNumber="148">
<emphasis bold="true" pageId="35" pageNumber="148">Holotype</emphasis>
: &quot;ECUADOR: Napo, Baeza (72 km E), 16 May 1975, Spangler, Langley, and Cohen&quot; (USNM). 
<emphasis bold="true" pageId="35" pageNumber="148">Paratypes: ECUADOR: Napo</emphasis>
: same data as holotype (44 males; 42 females); 
<emphasis bold="true" pageId="35" pageNumber="148">Tungurahua</emphasis>
: 
<normalizedToken originalValue="“Baños">&quot;Banos</normalizedToken>
(35 km E), 29 May 1975, Langley, Cohen, and Monnig (8 males; 6 females); 
<normalizedToken originalValue="“Baños">&quot;Banos</normalizedToken>
(18 km E), 25 Jan 1976, Spangler, et al. (15 males; 18 females); 
<normalizedToken originalValue="“Baños">&quot;Banos</normalizedToken>
(20 km E), 28 Jan 1976, Spangler, et al. (7 males; 6 females); 
<normalizedToken originalValue="“Baños">&quot;Banos</normalizedToken>
(39 km E), 28 Jan 1976, Spangler, et al. (2 females).
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="148" type="materials_examined">
<paragraph pageId="35" pageNumber="148">Additional material.</paragraph>
<paragraph pageId="35" pageNumber="148">
<materialsCitation collectingDate="1978-01-17" collectorName="J. Anderson" country="ECUADOR" elevation="1610" location="Borja" municipality="San Francisco" specimenCount="12" specimenCount-female="6" specimenCount-male="6" stateProvince="Napo">
<emphasis bold="true" pageId="35" pageNumber="148">
<collectingCountry name="Ecuador">ECUADOR</collectingCountry>
: 
<collectingRegion country="Ecuador" name="Napo">Napo</collectingRegion>
</emphasis>
: &quot; 
<collectingMunicipality>San Francisco</collectingMunicipality>
de / 
<location LSID="urn:lsid:plazi:treatment:228C962509E1470F58AE17EC00D94D1F:7A197600DDA3D7327DB3A85C4CE03639" country="ECUADOR" municipality="San Francisco" name="Borja" stateProvince="Napo">Borja</location>
<collectingDate value="1978-01-17">17 Jan 1978</collectingDate>
/ 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.61" unit="m" value="1610.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.61" unit="m" value="1610.0">1610 m</elevation>
</quantity>
PJ Spangler/ &amp; 
<collectorName>J. Anderson</collectorName>
// seep on stream bank&quot; (
<specimenCount type="male">6 males</specimenCount>
, 
<specimenCount type="female">6 females</specimenCount>
; USNM)
</materialsCitation>
.
</paragraph>
<caption ID-Zenodo-Dep="1298838" doi="10.3897/zookeys.768.24423.figure21" httpUri="https://binary.pensoft.net/fig/211234" pageId="35" pageNumber="148" start="Figure 21" startId="F21">
<paragraph pageId="35" pageNumber="148">
<emphasis bold="true" pageId="35" pageNumber="148">Figure 21.</emphasis>
Habitus of 
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" baseAuthorityName="Spangler" baseAuthorityYear="1979" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chasmogenus cremnobates" order="Coleoptera" pageId="35" pageNumber="148" phylum="Arthropoda" rank="species" species="cremnobates">
<emphasis italics="true" pageId="35" pageNumber="148">Chasmogenus cremnobates</emphasis>
</taxonomicName>
: 
<emphasis bold="true" pageId="35" pageNumber="148">A</emphasis>
dorsal view 
<emphasis bold="true" pageId="35" pageNumber="148">B</emphasis>
lateral view 
<emphasis bold="true" pageId="35" pageNumber="148">C</emphasis>
ventral view 
<emphasis bold="true" pageId="35" pageNumber="148">D</emphasis>
head 
<emphasis bold="true" pageId="35" pageNumber="148">E</emphasis>
aedeagus. Scale bar 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="35" pageNumber="148" type="key">
<paragraph pageId="35" pageNumber="148">
Key to the genera of New World 
<taxonomicName baseAuthorityName="Zaitzev" baseAuthorityYear="1908" lsidName="" pageId="35" pageNumber="148" rank="subFamily" subFamily="Acidocerinae">Acidocerinae</taxonomicName>
</paragraph>
<paragraph pageId="35" pageNumber="148">
<table inLine="true" pageId="35" pageNumber="148">
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">1</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Eyes absent. Known only from a cave in Ecuador</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<taxonomicName authorityName="Spangler" authorityYear="1981" class="Insecta" family="Hydrophilidae" genus="Troglochares" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Troglochares" order="Coleoptera" pageId="35" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis bold="true" italics="true" pageId="35" pageNumber="148">Troglochares</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">-</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Eyes present</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<emphasis bold="true" pageId="35" pageNumber="148">2</emphasis>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">2</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Eyes completely divided into dorsal and ventral sections by a lateral projection of frons. Size small (&lt;3 mm)</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<taxonomicName authorityName="Hansen" authorityYear="1999" class="Insecta" family="Hydrophilidae" genus="Quadriops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quadriops" order="Coleoptera" pageId="35" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis bold="true" italics="true" pageId="35" pageNumber="148">Quadriops</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">-</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Eyes not divided into dorsal and ventral sections by frons. Size variable</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<emphasis bold="true" pageId="35" pageNumber="148">3</emphasis>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">3</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Labrum concealed by clypeus, elytral margins broadly explanate. Body extremely dorsoventrally compressed</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<taxonomicName authorityName="Bergroth" authorityYear="1888" class="Insecta" family="Hydrophilidae" genus="Helobata" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Helobata" order="Coleoptera" pageId="35" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis bold="true" italics="true" pageId="35" pageNumber="148">Helobata</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">-</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Labrum not concealed by clypeus elytral margins not or at most weakly explanate. Body form variable but rarely dorsoventrally compressed</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<emphasis bold="true" pageId="35" pageNumber="148">4</emphasis>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">4</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Elytra with distinctly impressed sutural striae. Only Neotropical region</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chasmogenus" order="Coleoptera" pageId="35" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis bold="true" italics="true" pageId="35" pageNumber="148">Chasmogenus</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">-</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Elytra without sutural striae. Both Neotropical and Nearctic</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<emphasis bold="true" pageId="35" pageNumber="148">5</emphasis>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">5</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Prosternum with strongly elevated median carina</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Crucisternum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crucisternum" order="Coleoptera" pageId="35" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis bold="true" italics="true" pageId="35" pageNumber="148">Crucisternum</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">-</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Prosternum without median carina; at most tectiform medially</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<emphasis bold="true" pageId="35" pageNumber="148">6</emphasis>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">6</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Metafemora entire glabrous. Size small (&lt;3 mm)</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<taxonomicName authorityName="Short &amp; Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="35" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis bold="true" italics="true" pageId="35" pageNumber="148">Tobochares</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">-</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Metafemora pubescent at least on basal half or anterior third. Size variable</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<emphasis bold="true" pageId="35" pageNumber="148">7</emphasis>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">7</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Fifth ventrite entire, without apical emargination or truncation. Antennae with nine antennomeres. Maxillary palps shorter than the width of the head</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Radicitus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Radicitus" order="Coleoptera" pageId="35" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis bold="true" italics="true" pageId="35" pageNumber="148">Radicitus</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">-</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Fifth ventrite with apical emargination or truncation. Antennae with eight or nine antennomeres. Maxillary palps variable in length</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<emphasis bold="true" pageId="35" pageNumber="148">8</emphasis>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">8</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Elytral systematic punctures very distinct, distinctly larger than surrounding ground punctation. Antennae with nine antennomeres</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Katasophistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Katasophistes" order="Coleoptera" pageId="35" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis bold="true" italics="true" pageId="35" pageNumber="148">Katasophistes</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">-</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Elytral systematic punctures indistinct, usually blending with surrounding ground punctation. Antennae with eight or nine antennomeres</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<emphasis bold="true" pageId="35" pageNumber="148">9</emphasis>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">9</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Antennae with nine antennomeres. Size variable, but rarely less than 4 mm. Extremely common and widespread throughout the New World</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<taxonomicName class="Hexapoda" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Helochares" order="Coleoptera" pageId="35" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis bold="true" italics="true" pageId="35" pageNumber="148">Helochares</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">-</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Antennae with eight antennomeres. Rare and only known from the Guiana Shield region of South America. Size very small (&lt;3 mm)</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<emphasis bold="true" pageId="35" pageNumber="148">10</emphasis>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">10</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Body form circular, rounded. Size very small (1.9-2.3 mm)</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<taxonomicName authorityName="Garcia" authorityYear="2001" class="Insecta" family="Hydrophilidae" genus="Globulosis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Globulosis" order="Coleoptera" pageId="35" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis bold="true" italics="true" pageId="35" pageNumber="148">Globulosis</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="35" pageNumber="148">
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">-</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">Body form ovoid, parallel sided. Size exceedingly small (1.1-1.5 mm)</td>
<td colspan="1" pageId="35" pageNumber="148" rowspan="1">
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Nanosaphes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanosaphes" order="Coleoptera" pageId="35" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis bold="true" italics="true" pageId="35" pageNumber="148">Nanosaphes</emphasis>
</taxonomicName>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>