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<mods:title id="232B548E291808E56BAFCC9D55F9C5B9">An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas</mods:title>
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<treatment id="03D7FF1CFF72FF54FF48F93D30BCFA6E" ID-DOI="http://doi.org/10.5281/zenodo.3683294" ID-GBIF-Taxon="162246696" ID-Zenodo-Dep="3683294" LSID="urn:lsid:plazi:treatment:03D7FF1CFF72FF54FF48F93D30BCFA6E" httpUri="http://treatment.plazi.org/id/03D7FF1CFF72FF54FF48F93D30BCFA6E" lastPageNumber="184" pageId="185" pageNumber="184">
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<heading id="D089F966FF72FF54FF48F93D3271FD9B" bold="true" box="[192,781,594,619]" fontSize="10" level="2" pageId="185" pageNumber="184" reason="3">
<emphasis id="B90A9218FF72FF54FF48F93D3271FD9B" bold="true" box="[192,781,594,619]" pageId="185" pageNumber="184">
§
<taxonomicName id="4C7E3589FF72FF54FF5DF93C3271FD9B" authority="(Fabricius, 1801)" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[213,781,594,619]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="B90A9218FF72FF54FF5DF93C335CFD9A" bold="true" box="[213,544,594,618]" italics="true" pageId="185" pageNumber="184">Cyclocephala tridentata</emphasis>
(
<bibRefCitation id="EFEF33FBFF72FF54FDB9F93D3279FD9B" author="Fabricius, J. C." bookContentInfo="506 p." box="[561,773,595,619]" journalOrPublisher="Tomus I. Impensis Bibliopolii Academici Novi; Kiel, Germany" pageId="185" pageNumber="184" refId="ref201994" refString="Fabricius, J. C. 1801 a. Systema eleutheratorum secundum ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus. Tomus I. Impensis Bibliopolii Academici Novi; Kiel, Germany. 506 p." title="Systema eleutheratorum secundum ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus" type="book" year="1801">Fabricius, 1801</bibRefCitation>
)
</taxonomicName>
</emphasis>
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</paragraph>
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<subSubSection id="C3641D81FF72FF54FF60F91734E6FD15" pageId="185" pageNumber="184" type="reference_group">
<paragraph id="8BC14E0AFF72FF54FF60F9173221FD7F" blockId="185.[232,1440,633,742]" box="[232,861,633,655]" pageId="185" pageNumber="184">
<treatmentCitationGroup id="AB6E6924FF72FF54FF60F9173221FD7F" box="[232,861,633,655]" pageId="185" pageNumber="184">
<taxonomicName id="4C7E3589FF72FF54FF60F9173221FD7F" authority="Fabricius 1801 b: 170" authorityName="Fabricius" authorityPageNumber="170" authorityYear="1801" box="[232,861,633,655]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="species" species="tridentata">
<treatmentCitation id="0ADF681BFF72FF54FF60F91733BAFD7F" author="Fabricius, J. C." box="[232,710,633,655]" page="170" pageId="185" pageNumber="184" year="1801">
<emphasis id="B90A9218FF72FF54FF60F91730A0FD7E" box="[232,476,633,654]" italics="true" pageId="185" pageNumber="184">Melolontha tridentata</emphasis>
<bibRefCitation id="EFEF33FBFF72FF54FE6BF91733BAFD7F" author="Fabricius, J. C." bookContentInfo="687 p." box="[483,710,633,655]" journalOrPublisher="Tomus II. Impensis Bibliopolii Academici Novi; Kiel, Germany" pageId="185" pageNumber="184" refId="ref202035" refString="Fabricius, J. C. 1801 b. Systema eleutheratorum secundum ordines, genera, species: adjectis synonymis, locis, observationibus, descriptionibus. Tomus II. Impensis Bibliopolii Academici Novi; Kiel, Germany. 687 p." title="Systema eleutheratorum secundum ordines, genera, species: adjectis synonymis, locis, observationibus, descriptionibus" type="book" year="1801">Fabricius 1801b: 170</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F3690E9AFF72FF54FD5CF917322BFD7F" box="[724,855,633,655]" name="Guadeloupe" pageId="185" pageNumber="184">Guadeloupe</collectingCountry>
)
</taxonomicName>
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</paragraph>
<paragraph id="8BC14E0AFF72FF54FF60F9F930BAFD39" blockId="185.[232,1440,633,742]" pageId="185" pageNumber="184">
<treatmentCitationGroup id="AB6E6924FF72FF54FF60F9F930BAFD39" pageId="185" pageNumber="184">
=
<taxonomicName id="4C7E3589FF72FF54FF75F9F835C2FD5C" authority="Cartwright and Chalumeau 1978: 25" authorityName="Cartwright and Chalumeau" authorityPageNumber="25" authorityYear="1978" box="[253,1214,662,684]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="subSpecies" species="tridentata" subSpecies="dominicensis">
<treatmentCitation id="0ADF681BFF72FF54FF75F9F83540FD5C" author="Cartwright, O. L. &amp; F. E. Chalumeau" box="[253,1084,662,684]" page="25" pageId="185" pageNumber="184" year="1978">
<emphasis id="B90A9218FF72FF54FF75F9F833E0FD5B" box="[253,668,662,683]" italics="true" pageId="185" pageNumber="184">Cyclocephala tridentata dominicensis</emphasis>
<bibRefCitation id="EFEF33FBFF72FF54FD2BF9F83540FD5C" author="Cartwright, O. L. &amp; F. E. Chalumeau" box="[675,1084,662,684]" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="185" pageNumber="184" pagination="1 - 32" part="279" refId="ref196093" refString="Cartwright, O. L., and F. E. Chalumeau. 1978. Bredin-Archbold-Smithsonian biological survey of Dominica: The superfamily Scarabaeoidea (Coleoptera). Smithsonian Contributions to Zoology 279: 1 - 32." title="Bredin-Archbold-Smithsonian biological survey of Dominica: The superfamily Scarabaeoidea (Coleoptera)" type="journal article" year="1978">Cartwright and Chalumeau 1978: 25</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F3690E9AFF72FF54FBC2F9F835CBFD5C" box="[1098,1207,662,684]" name="Dominica" pageId="185" pageNumber="184">Dominica</collectingCountry>
)
</taxonomicName>
-
<treatmentCitation id="0ADF681BFF72FF54FB5BF9F8302DFD38" author="Ratcliffe and Cave" page="113" pageId="185" pageNumber="184" year="2015">
<bibRefCitation id="EFEF33FBFF72FF54FB5BF9F8302DFD38" author="Ratcliffe and Cave" firstAuthor="Ratcliffe" pageId="185" pageNumber="184" pagination="1 - 343" refId="ref222299" refString="Ratcliffe, B. C., and R. D. Cave. 2015. The dynastine scarab beetles of the West Indies (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 28: 1 - 343." type="journal article" year="2015">Ratcliffe and Cave 2015: 113</bibRefCitation>
</treatmentCitation>
(syn. nov.)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC14E0AFF72FF54FF60F9BE34E6FD15" blockId="185.[232,1440,633,742]" box="[232,1434,720,742]" pageId="185" pageNumber="184">
<treatmentCitationGroup id="AB6E6924FF72FF54FF60F9BE34E6FD15" box="[232,1434,720,742]" pageId="185" pageNumber="184">
=
<taxonomicName id="4C7E3589FF72FF54FF75F9BE32AFFD16" authority="Dutrillaux et al. 2013: 64" authorityName="Dutrillaux" authorityPageNumber="64" authorityYear="2013" box="[253,979,720,742]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="species" species="annamariae">
<treatmentCitation id="0ADF681BFF72FF54FF75F9BE3240FD16" author="Dutrillaux, B. &amp; F. Chalumeau &amp; A. - M. Dutrillaux &amp; T. Giannoulis &amp; Z. Mamuris" box="[253,828,720,742]" page="64" pageId="185" pageNumber="184" year="2013">
<emphasis id="B90A9218FF72FF54FF75F9BE3361FD15" box="[253,541,720,741]" italics="true" pageId="185" pageNumber="184">Cyclocephala annamariae</emphasis>
<bibRefCitation id="EFEF33FBFF72FF54FDACF9BE3240FD16" author="Dutrillaux, B. &amp; F. Chalumeau &amp; A. - M. Dutrillaux &amp; T. Giannoulis &amp; Z. Mamuris" box="[548,828,720,742]" journalOrPublisher="Annales de la Societe entomologique de France (N. S.)" pageId="185" pageNumber="184" pagination="61 - 67" part="49" refId="ref200117" refString="Dutrillaux, B., F. Chalumeau, A. - M. Dutrillaux, T. Giannoulis, and Z. Mamuris. 2013. Separation taxonomique en trois especes au sein des populations de Cyclocephala tridentata Fabricius (Coleoptera: Scarabaeidae: Dynastinae), sur la base de criteres genetiques, chromosomiques et geographiques. Annales de la Societe entomologique de France (N. S.) 49 (1): 61 - 67." title="Separation taxonomique en trois especes au sein des populations de Cyclocephala tridentata Fabricius (Coleoptera: Scarabaeidae: Dynastinae), sur la base de criteres genetiques, chromosomiques et geographiques" type="journal article" year="2013">Dutrillaux et al. 2013: 64</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F3690E9AFF72FF54FCC3F9BE32B0FD16" box="[843,972,720,742]" name="Martinique" pageId="185" pageNumber="184">Martinique</collectingCountry>
)
</taxonomicName>
-
<treatmentCitation id="0ADF681BFF72FF54FC60F9BE3459FD15" author="Ratcliffe and Cave" box="[1000,1317,720,742]" page="113" pageId="185" pageNumber="184" year="2015">
<bibRefCitation id="EFEF33FBFF72FF54FC60F9BE3459FD15" author="Ratcliffe and Cave" box="[1000,1317,720,742]" firstAuthor="Ratcliffe" pageId="185" pageNumber="184" pagination="1 - 343" refId="ref222299" refString="Ratcliffe, B. C., and R. D. Cave. 2015. The dynastine scarab beetles of the West Indies (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 28: 1 - 343." type="journal article" year="2015">Ratcliffe and Cave 2015: 113</bibRefCitation>
</treatmentCitation>
(syn. nov.)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3641D81FF72FF54FF48F99830BCFA6E" pageId="185" pageNumber="184" type="distribution">
<paragraph id="8BC14E0AFF72FF54FF48F99833E2FC1E" blockId="185.[192,1440,758,1006]" pageId="185" pageNumber="184">
<emphasis id="B90A9218FF72FF54FF48F9983374FCFE" bold="true" box="[192,520,758,782]" pageId="185" pageNumber="184">
Distribution.
<collectingCountry id="F3690E9AFF72FF54FEF1F998337DFCFE" box="[377,513,758,782]" name="Suriname" pageId="185" pageNumber="184">Suriname</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF72FF54FD87F998327CFCFE" author="Endrodi, S." box="[527,768,758,782]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="185" pageNumber="184" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 321</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF72FF54FC84F99832F8FCFE" author="Endrodi, S." bookContentInfo="800 p." box="[780,900,758,782]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="185" pageNumber="184" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">1985a: 99</bibRefCitation>
, 138;
<bibRefCitation id="EFEF33FBFF72FF54FC42F9993596FCFE" author="Moore" box="[970,1258,758,782]" etAl="et al." firstAuthor="Moore" pageId="185" pageNumber="184" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 255</bibRefCitation>
.
<emphasis id="B90A9218FF72FF54FB7DF9983431FCFE" bold="true" box="[1269,1357,758,782]" pageId="185" pageNumber="184">Other:</emphasis>
<bibRefCitation id="EFEF33FBFF72FF54FADCF9983002FCDE" author="Fabricius, J. C." bookContentInfo="687 p." journalOrPublisher="Tomus II. Impensis Bibliopolii Academici Novi; Kiel, Germany" pageId="185" pageNumber="184" refId="ref202035" refString="Fabricius, J. C. 1801 b. Systema eleutheratorum secundum ordines, genera, species: adjectis synonymis, locis, observationibus, descriptionibus. Tomus II. Impensis Bibliopolii Academici Novi; Kiel, Germany. 687 p." title="Systema eleutheratorum secundum ordines, genera, species: adjectis synonymis, locis, observationibus, descriptionibus" type="book" year="1801">Fabricius 1801b: 170</bibRefCitation>
(Guadeloupe - as
<taxonomicName id="4C7E3589FF72FF54FDD7F8783211FCDE" authorityName="Fabricius" authorityYear="1801" box="[607,877,790,814]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="B90A9218FF72FF54FDD7F8783211FCDE" box="[607,877,790,814]" italics="true" pageId="185" pageNumber="184">Melolontha tridentata</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF72FF54FC0DF879352DFCDE" author="Arrow, G. J." bookContentInfo="124 p." box="[901,1105,790,815]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="185" pageNumber="184" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 16</bibRefCitation>
(Guadeloupe, Martinique);
<bibRefCitation id="EFEF33FBFF72FF54FF48F85830AFFCBE" author="Blackwelder, R. E." box="[192,467,822,846]" journalOrPublisher="United States National Museum Bulletin" pageId="185" pageNumber="184" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 253</bibRefCitation>
(Dominica, Guadeloupe, Martinique);
<bibRefCitation id="EFEF33FBFF72FF54FC2FF85835E8FCBE" author="Endrodi, S." box="[935,1172,822,846]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="185" pageNumber="184" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 321</bibRefCitation>
(Barbados, Colombia, Dominican Republic [&gt; Dominica!], Guadeloupe, Martinique), 1985a: 99, 138 (Barbados, Colombia, Dominica, Guadeloupe, Martinique);
<bibRefCitation id="EFEF33FBFF72FF54FD0DF818353AFC7E" author="Cartwright, O. L. &amp; F. E. Chalumeau" box="[645,1094,886,910]" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="185" pageNumber="184" pagination="1 - 32" part="279" refId="ref196093" refString="Cartwright, O. L., and F. E. Chalumeau. 1978. Bredin-Archbold-Smithsonian biological survey of Dominica: The superfamily Scarabaeoidea (Coleoptera). Smithsonian Contributions to Zoology 279: 1 - 32." title="Bredin-Archbold-Smithsonian biological survey of Dominica: The superfamily Scarabaeoidea (Coleoptera)" type="journal article" year="1978">Cartwright and Chalumeau 1978: 25</bibRefCitation>
(Dominica - as
<taxonomicName id="4C7E3589FF72FF54FA8BF8183023FC5E" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="subSpecies" species="tridentata" subSpecies="dominicensis">
<emphasis id="B90A9218FF72FF54FA8BF8183023FC5E" italics="true" pageId="185" pageNumber="184">C. tridentata dominicensis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF72FF54FEFDF8F8336AFC5E" author="Ponchel" box="[373,534,918,942]" firstAuthor="Ponchel" pageId="185" pageNumber="184" pagination="60 - 61" refId="ref220051" refString="Ponchel, Y. 2011. Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 4: 60 - 61. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." type="journal article" year="2011">Ponchel 2011</bibRefCitation>
(not mentioned);
<bibRefCitation id="EFEF33FBFF72FF54FD61F8F83569FC5E" author="Dutrillaux, B. &amp; F. Chalumeau &amp; A. - M. Dutrillaux &amp; T. Giannoulis &amp; Z. Mamuris" box="[745,1045,918,942]" journalOrPublisher="Annales de la Societe entomologique de France (N. S.)" pageId="185" pageNumber="184" pagination="61 - 67" part="49" refId="ref200117" refString="Dutrillaux, B., F. Chalumeau, A. - M. Dutrillaux, T. Giannoulis, and Z. Mamuris. 2013. Separation taxonomique en trois especes au sein des populations de Cyclocephala tridentata Fabricius (Coleoptera: Scarabaeidae: Dynastinae), sur la base de criteres genetiques, chromosomiques et geographiques. Annales de la Societe entomologique de France (N. S.) 49 (1): 61 - 67." title="Separation taxonomique en trois especes au sein des populations de Cyclocephala tridentata Fabricius (Coleoptera: Scarabaeidae: Dynastinae), sur la base de criteres genetiques, chromosomiques et geographiques" type="journal article" year="2013">Dutrillaux et al. 2013: 64</bibRefCitation>
(Martinique - as
<taxonomicName id="4C7E3589FF72FF54FB6BF8F834E4FC5E" box="[1251,1432,918,942]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="species" species="annamariae">
<emphasis id="B90A9218FF72FF54FB6BF8F834E4FC5E" box="[1251,1432,918,942]" italics="true" pageId="185" pageNumber="184">C. annamariae</emphasis>
</taxonomicName>
; Dominica - as
<taxonomicName id="4C7E3589FF72FF54FEE4F8D83350FC3E" box="[364,556,950,974]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="species" species="dominicensis">
<emphasis id="B90A9218FF72FF54FEE4F8D83350FC3E" box="[364,556,950,974]" italics="true" pageId="185" pageNumber="184">C. dominicensis</emphasis>
</taxonomicName>
; Guadeloupe - as
<taxonomicName id="4C7E3589FF72FF54FC8CF8D832DCFC3E" authorityName="Fabricius" authorityYear="1801" box="[772,928,950,974]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="B90A9218FF72FF54FC8CF8D832DCFC3E" box="[772,928,950,974]" italics="true" pageId="185" pageNumber="184">C. tridentata</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF72FF54FC3EF8D83523FC3E" author="Dupuis, F." box="[950,1119,950,974]" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="185" pageNumber="184" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis 2016b</bibRefCitation>
(not mentioned);
<bibRefCitation id="EFEF33FBFF72FF54FABDF8D93016FC1E" author="Moore" etAl="et al." firstAuthor="Moore" pageId="185" pageNumber="184" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 255</bibRefCitation>
(Colombia, West Indies).
</paragraph>
<paragraph id="8BC14E0AFF72FF54FF48F89032D5FB26" blockId="185.[192,1440,1022,1238]" pageId="185" pageNumber="184">
<emphasis id="B90A9218FF72FF54FF48F890305EFBE6" bold="true" box="[192,290,1022,1046]" pageId="185" pageNumber="184">Note 1.</emphasis>
<bibRefCitation id="EFEF33FBFF72FF54FEA4F8903332FBE6" author="Dutrillaux, B. &amp; F. Chalumeau &amp; A. - M. Dutrillaux &amp; T. Giannoulis &amp; Z. Mamuris" box="[300,590,1022,1046]" journalOrPublisher="Annales de la Societe entomologique de France (N. S.)" pageId="185" pageNumber="184" pagination="61 - 67" part="49" refId="ref200117" refString="Dutrillaux, B., F. Chalumeau, A. - M. Dutrillaux, T. Giannoulis, and Z. Mamuris. 2013. Separation taxonomique en trois especes au sein des populations de Cyclocephala tridentata Fabricius (Coleoptera: Scarabaeidae: Dynastinae), sur la base de criteres genetiques, chromosomiques et geographiques. Annales de la Societe entomologique de France (N. S.) 49 (1): 61 - 67." title="Separation taxonomique en trois especes au sein des populations de Cyclocephala tridentata Fabricius (Coleoptera: Scarabaeidae: Dynastinae), sur la base de criteres genetiques, chromosomiques et geographiques" type="journal article" year="2013">Dutrillaux et al. (2013)</bibRefCitation>
split the populations of
<taxonomicName id="4C7E3589FF72FF54FC0BF89035D5FBE6" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[899,1193,1022,1046]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="B90A9218FF72FF54FC0BF89035D5FBE6" box="[899,1193,1022,1046]" italics="true" pageId="185" pageNumber="184">Cyclocephala tridentata</emphasis>
</taxonomicName>
from the islands of
<collectingCountry id="F3690E9AFF72FF54FF48FF703030FBC6" box="[192,332,1054,1078]" name="Martinique" pageId="185" pageNumber="184">Martinique</collectingCountry>
,
<collectingCountry id="F3690E9AFF72FF54FEDEFF7030B6FBC6" box="[342,458,1054,1078]" name="Dominica" pageId="185" pageNumber="184">Dominica</collectingCountry>
and
<collectingCountry id="F3690E9AFF72FF54FD8AFF7033EFFBC6" box="[514,659,1054,1078]" name="Guadeloupe" pageId="185" pageNumber="184">Guadeloupe</collectingCountry>
(Lesser Antilles) into
<taxonomicName id="4C7E3589FF72FF54FC1FFF7035A6FBC6" authority="(Guadeloupe)" baseAuthorityName="Guadeloupe" baseAuthorityYear="1801" box="[919,1242,1054,1078]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="B90A9218FF72FF54FC1FFF70354CFBC6" box="[919,1072,1054,1078]" italics="true" pageId="185" pageNumber="184">C. tridentata</emphasis>
(
<collectingCountry id="F3690E9AFF72FF54FBB5FF7035AEFBC6" box="[1085,1234,1054,1078]" name="Guadeloupe" pageId="185" pageNumber="184">Guadeloupe</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="4C7E3589FF72FF54FB6BFF7034DCFBC6" box="[1251,1440,1054,1078]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="species" species="dominicensis">
<emphasis id="B90A9218FF72FF54FB6BFF7034DCFBC6" box="[1251,1440,1054,1078]" italics="true" pageId="185" pageNumber="184">C. dominicensis</emphasis>
</taxonomicName>
(was
<taxonomicName id="4C7E3589FF72FF54FF75FF5032AEFBA6" authority="Cartwright and Chalumeau, 1978" authorityName="Cartwright and Chalumeau" authorityYear="1978" box="[253,978,1086,1110]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="subSpecies" species="tridentata" subSpecies="dominicensis">
<emphasis id="B90A9218FF72FF54FF75FF503344FBA6" box="[253,568,1086,1110]" italics="true" pageId="185" pageNumber="184">C. tridentata dominicensis</emphasis>
<bibRefCitation id="EFEF33FBFF72FF54FDB6FF5032AEFBA6" author="Cartwright, O. L. &amp; F. E. Chalumeau" box="[574,978,1086,1110]" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="185" pageNumber="184" pagination="1 - 32" part="279" refId="ref196093" refString="Cartwright, O. L., and F. E. Chalumeau. 1978. Bredin-Archbold-Smithsonian biological survey of Dominica: The superfamily Scarabaeoidea (Coleoptera). Smithsonian Contributions to Zoology 279: 1 - 32." title="Bredin-Archbold-Smithsonian biological survey of Dominica: The superfamily Scarabaeoidea (Coleoptera)" type="journal article" year="1978">Cartwright and Chalumeau, 1978</bibRefCitation>
</taxonomicName>
(p. 25)) (
<collectingCountry id="F3690E9AFF72FF54FBBFFF5035D0FBA6" box="[1079,1196,1086,1110]" name="Dominica" pageId="185" pageNumber="184">Dominica</collectingCountry>
) and
<taxonomicName id="4C7E3589FF72FF54FB65FF503020FB86" authority="(Martinique)" baseAuthorityName="Martinique" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="species" species="annamariae">
<emphasis id="B90A9218FF72FF54FB65FF5034DCFBA6" box="[1261,1440,1086,1110]" italics="true" pageId="185" pageNumber="184">C. annamariae</emphasis>
(
<collectingCountry id="F3690E9AFF72FF54FF40FF303028FB86" box="[200,340,1118,1142]" name="Martinique" pageId="185" pageNumber="184">Martinique</collectingCountry>
)
</taxonomicName>
based on differences in their mitochondrial gene CO1. The publication does not include information regarding mainland specimens.
<bibRefCitation id="EFEF33FBFF72FF54FD7AFF10350FFB66" author="Ratcliffe and Cave" box="[754,1139,1150,1174]" firstAuthor="Ratcliffe" pageId="185" pageNumber="184" pagination="1 - 343" refId="ref222299" refString="Ratcliffe, B. C., and R. D. Cave. 2015. The dynastine scarab beetles of the West Indies (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 28: 1 - 343." type="journal article" year="2015">Ratcliffe and Cave (2015: 113)</bibRefCitation>
reduce
<taxonomicName id="4C7E3589FF72FF54FB53FF1034DCFB66" box="[1243,1440,1150,1174]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="species" species="dominicensis">
<emphasis id="B90A9218FF72FF54FB53FF1034DCFB66" box="[1243,1440,1150,1174]" italics="true" pageId="185" pageNumber="184">C. dominicensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7E3589FF72FF54FF7EFFF030D2FB46" box="[246,430,1182,1206]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="species" species="annamariae">
<emphasis id="B90A9218FF72FF54FF7EFFF030D2FB46" box="[246,430,1182,1206]" italics="true" pageId="185" pageNumber="184">C. annamariae</emphasis>
</taxonomicName>
to synonyms of
<taxonomicName id="4C7E3589FF72FF54FDFEFFF03268FB46" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[630,788,1182,1206]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="B90A9218FF72FF54FDFEFFF03268FB46" box="[630,788,1182,1206]" italics="true" pageId="185" pageNumber="184">C. tridentata</emphasis>
</taxonomicName>
. The authors and year of description of
<taxonomicName id="4C7E3589FF72FF54FA8AFFF03023FB26" authorityName="Cartwright and Chalumeau" authorityYear="1978" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="subSpecies" species="tridentata" subSpecies="dominicensis">
<emphasis id="B90A9218FF72FF54FA8AFFF03023FB26" italics="true" pageId="185" pageNumber="184">C. tridentata dominicensis</emphasis>
</taxonomicName>
are discussed in
<bibRefCitation id="EFEF33FBFF72FF54FDB9FFD032D9FB26" author="Ratcliffe and Cave" box="[561,933,1214,1238]" firstAuthor="Ratcliffe" pageId="185" pageNumber="184" pagination="1 - 343" refId="ref222299" refString="Ratcliffe, B. C., and R. D. Cave. 2015. The dynastine scarab beetles of the West Indies (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 28: 1 - 343." type="journal article" year="2015">Ratcliffe and Cave (2015: 116)</bibRefCitation>
.
</paragraph>
<paragraph id="8BC14E0AFF72FF54FF48FF8830BCFA6E" blockId="185.[192,1440,1254,1438]" pageId="185" pageNumber="184">
<emphasis id="B90A9218FF72FF54FF48FF883059FB0E" bold="true" box="[192,293,1254,1278]" pageId="185" pageNumber="184">Note 2.</emphasis>
<bibRefCitation id="EFEF33FBFF72FF54FEB9FF883345FB0E" author="Endrodi, S." box="[305,569,1254,1278]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="185" pageNumber="184" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi (1966a: 321</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF72FF54FDC2FF8833B5FB0E" author="Endrodi, S." bookContentInfo="800 p." box="[586,713,1254,1278]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="185" pageNumber="184" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">1985a: 99</bibRefCitation>
, 138) records
<taxonomicName id="4C7E3589FF72FF54FC0AFF8835CDFB0E" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[898,1201,1254,1278]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="184" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="B90A9218FF72FF54FC0AFF8835CDFB0E" box="[898,1201,1254,1278]" italics="true" pageId="185" pageNumber="184">Cyclocephala tridentata</emphasis>
</taxonomicName>
for
<collectingCountry id="F3690E9AFF72FF54FB62FF88341AFB0E" box="[1258,1382,1254,1278]" name="Suriname" pageId="185" pageNumber="184">Suriname</collectingCountry>
and
<collectingCountry id="F3690E9AFF72FF54FF48FE683044FAEE" box="[192,312,1286,1310]" name="Colombia" pageId="185" pageNumber="184">Colombia</collectingCountry>
, and these records are repeated in
<bibRefCitation id="EFEF33FBFF72FF54FD61FE693563FAEE" author="Moore" box="[745,1055,1286,1310]" etAl="et al." firstAuthor="Moore" pageId="185" pageNumber="184" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. (2018b: 255)</bibRefCitation>
.
<bibRefCitation id="EFEF33FBFF72FF54FBA3FE6834E3FAEE" author="Ratcliffe and Cave" box="[1067,1439,1286,1310]" firstAuthor="Ratcliffe" pageId="185" pageNumber="184" pagination="1 - 343" refId="ref222299" refString="Ratcliffe, B. C., and R. D. Cave. 2015. The dynastine scarab beetles of the West Indies (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 28: 1 - 343." type="journal article" year="2015">Ratcliffe and Cave (2015: 114)</bibRefCitation>
mention they have not seen specimens from
<collectingCountry id="F3690E9AFF72FF54FD67FE483215FACE" box="[751,873,1318,1342]" name="Suriname" pageId="185" pageNumber="184">Suriname</collectingCountry>
or
<collectingCountry id="F3690E9AFF72FF54FC1CFE483571FACE" box="[916,1037,1318,1342]" name="Colombia" pageId="185" pageNumber="184">Colombia</collectingCountry>
. There are also no other records for the mainland of South America that we are aware of, and it may well be that Endrödi’s record for
<collectingCountry id="F3690E9AFF72FF54FF48FE083046FA8E" box="[192,314,1382,1406]" name="Suriname" pageId="185" pageNumber="184">Suriname</collectingCountry>
is based on a mislabeled or misidentified specimen. We regard this species as not occurring in the research area.
</paragraph>
</subSubSection>
</treatment>
</document>
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