treatments-xml/data/03/D7/FF/03D7FF1CFF1EFF38FF48FA0433F3FB1E.xml
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<emphasis id="B90A9218FF1EFF38FF48FA0430E2FE71" bold="true" box="[192,414,361,385]" italics="true" pageId="213" pageNumber="212">Tomarus ebenus</emphasis>
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<bibRefCitation id="EFEF33FBFF1EFF38FE49FAFE33F4FE55" author="De Geer, C." bookContentInfo="456 p." box="[449,648,400,421]" journalOrPublisher="Pierre Hesselberg; Stockholm" pageId="213" pageNumber="212" refId="ref198837" refString="De Geer, C. 1774. Memoires pour servir a l'histoire des insectes. Tome quatrieme. Pierre Hesselberg; Stockholm. 456 p." title="Memoires pour servir a l'histoire des insectes. Tome quatrieme" type="book" year="1774">De Geer 1774: 317</bibRefCitation>
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Distribution.
<collectingCountry id="F3690E9AFF1EFF38FEF2FAD8309BFE3E" box="[378,487,438,462]" name="Guyana" pageId="213" pageNumber="212">Guyana</collectingCountry>
:
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<bibRefCitation id="EFEF33FBFF1EFF38FE7DFAD833B7FE3E" author="Bodkin, G. E." box="[501,715,438,462]" journalOrPublisher="The Entomologist's Monthly Magazine" pageId="213" pageNumber="212" pagination="210 - 219" part="55" refId="ref194211" refString="Bodkin, G. E. 1919. Notes on the Coleoptera of British Guiana. The Entomologist's Monthly Magazine 55: 210 - 219." title="Notes on the Coleoptera of British Guiana" type="journal article" year="1919">Bodkin 1919: 215</bibRefCitation>
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<emphasis id="B90A9218FF1EFF38FD77FAD932CBFE3E" box="[767,951,438,462]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
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<bibRefCitation id="EFEF33FBFF1EFF38FC47FAD835BCFE3E" author="Endrodi, S." bookContentInfo="800 p." box="[975,1216,438,462]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="213" pageNumber="212" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">Endrödi 1985a: 254</bibRefCitation>
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<taxonomicName id="4C7E3589FF1EFF38FB7CFAD93019FE1E" baseAuthorityName="De Geer" baseAuthorityYear="1774" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus" subGenus="Euligyrus">
<emphasis id="B90A9218FF1EFF38FB7CFAD9342EFE3E" box="[1268,1362,439,462]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FAEAFAD8307FFE1E" italics="true" pageId="213" pageNumber="212">Euligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FE9BFAB83019FE1E" box="[275,357,470,494]" italics="true" pageId="213" pageNumber="212">ebenus</emphasis>
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<bibRefCitation id="EFEF33FBFF1EFF38FEF4FAB83316FE1E" author="Lachaume" box="[380,618,470,494]" firstAuthor="Lachaume" pageId="213" pageNumber="212" pagination="7 - 56" refId="ref211183" refString="Lachaume, G. 1992. Dynastidae americains Cyclocephalini - Agaocephalini - Pentodontini - Oryctini - Phileurini. Les Coleopteres du Monde 14: 7 - 56." type="journal article" year="1992">Lachaume 1992: 33</bibRefCitation>
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<emphasis id="B90A9218FF1EFF38FD14FAB9322EFE1E" box="[668,850,470,494]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
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<bibRefCitation id="EFEF33FBFF1EFF38FCE2FAB835BFFE1E" author="Escalona, H. E. &amp; L. J. Joly" box="[874,1219,470,494]" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="213" pageNumber="212" pagination="111 - 137" part="39" refId="ref201573" refString="Escalona, H. E., and L. J. Joly. 2006. El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Boletin de la Sociedad Entomologica Aragonesa 39: 111 - 137." title="El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini)" type="journal article" year="2006">Escalona and Joly 2006: 133</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FB7DFAB9301AFDFE" baseAuthorityName="De Geer" baseAuthorityYear="1774" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus" subGenus="Euligyrus">
<emphasis id="B90A9218FF1EFF38FB7DFAB9342FFE1E" box="[1269,1363,471,494]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FAEAFAB8307FFDFE" italics="true" pageId="213" pageNumber="212">Euligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FE9CFA98301AFDFE" box="[276,358,502,526]" italics="true" pageId="213" pageNumber="212">ebenus</emphasis>
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<bibRefCitation id="EFEF33FBFF1EFF38FEF7FA983390FDFE" author="Lopez-Garcia" box="[383,748,502,526]" etAl="et al." firstAuthor="Lopez-Garcia" pageId="213" pageNumber="212" pagination="491 - 509" refId="ref213049" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, R. D. Cave, and G. Amat-Garcia. 2016. An annotated checklist of the New World pentodontine scarab beetles (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Zootaxa 4170 (3): 491 - 509." type="journal article" year="2016">López-García et al. 2016: 502</bibRefCitation>
.
<emphasis id="B90A9218FF1EFF38FD70FA9832FAFDFE" bold="true" box="[760,902,502,526]" pageId="213" pageNumber="212">
<collectingCountry id="F3690E9AFF1EFF38FD70FA9832FCFDFE" box="[760,896,502,526]" name="Suriname" pageId="213" pageNumber="212">Suriname</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF1EFF38FC06FA99350FFDFE" author="De Geer, C." bookContentInfo="456 p." box="[910,1139,502,526]" journalOrPublisher="Pierre Hesselberg; Stockholm" pageId="213" pageNumber="212" refId="ref198837" refString="De Geer, C. 1774. Memoires pour servir a l'histoire des insectes. Tome quatrieme. Pierre Hesselberg; Stockholm. 456 p." title="Memoires pour servir a l'histoire des insectes. Tome quatrieme" type="book" year="1774">De Geer 1774: 317</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FB20FA9834F3FDFE" authorityName="De Geer" authorityYear="1774" box="[1192,1423,502,526]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus">
<emphasis id="B90A9218FF1EFF38FB20FA9834F3FDFE" box="[1192,1423,502,526]" italics="true" pageId="213" pageNumber="212">Scarabaeus ebenus</emphasis>
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);
<bibRefCitation id="EFEF33FBFF1EFF38FF48F97830E3FDDE" author="Endrodi, S." box="[192,415,534,558]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="213" pageNumber="212" pagination="1 - 145" part="37" refId="ref200804" refString="Endrodi, S. 1969 a. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 37 (1): 1 - 145." title="Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini" type="journal article" year="1969">Endrödi 1969a: 51</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FE58F979326BFDDE" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[464,791,534,558]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus" subGenus="Euligyrus">
<emphasis id="B90A9218FF1EFF38FE58F9793352FDDE" box="[464,558,535,558]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FDB5F97833C9FDDE" box="[573,693,534,558]" italics="true" pageId="213" pageNumber="212">Euligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FD4DF978326BFDDE" box="[709,791,534,558]" italics="true" pageId="213" pageNumber="212">ebenus</emphasis>
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<bibRefCitation id="EFEF33FBFF1EFF38FCA6F978353EFDDE" author="Dechambre, R. - P." box="[814,1090,534,558]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="213" pageNumber="212" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 168</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FBFBF9793455FDDE" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[1139,1321,534,558]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus">
<emphasis id="B90A9218FF1EFF38FBFBF9793455FDDE" box="[1139,1321,534,558]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
</taxonomicName>
); MHCN; NZCS; RMNH.
<emphasis id="B90A9218FF1EFF38FEF4F958332EFDBE" bold="true" box="[380,594,566,590]" pageId="213" pageNumber="212">
<collectingCountry id="F3690E9AFF1EFF38FEF4F9583331FDBE" box="[380,589,566,590]" name="French Guiana" pageId="213" pageNumber="212">French Guiana</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF1EFF38FDD0F9583216FDBE" author="Dechambre, R. - P." box="[600,874,566,590]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="213" pageNumber="212" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 168</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FC12F959352CFDBE" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[922,1104,566,590]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus">
<emphasis id="B90A9218FF1EFF38FC12F959352CFDBE" box="[922,1104,566,590]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
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<bibRefCitation id="EFEF33FBFF1EFF38FBEEF958340AFDBE" author="Brule, S. &amp; J. Touroult &amp; P. - H. Dalens" bookContentInfo="120 p." box="[1126,1398,566,590]" journalOrPublisher="Rapport de la Societe entomologique Antilles-Guyane, SEAG, ONF" pageId="213" pageNumber="212" refId="ref195373" refString="Brule, S., J. Touroult, and P. - H. Dalens (coord.). 2011 a. Resultats de l'inventaire entomologique du site de Saut Parare, reserve des Nouragues (Guyane), 2009 - 2010. Rapport de la Societe entomologique Antilles-Guyane, SEAG, ONF. 120 p. + annexes." title="Resultats de l'inventaire entomologique du site de Saut Parare, reserve des Nouragues (Guyane)" type="book" year="2011">Brûlé et al. 2011a: 191</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FF48F939300BFD9E" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[192,375,598,622]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus">
<emphasis id="B90A9218FF1EFF38FF48F939300BFD9E" box="[192,375,598,622]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
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<bibRefCitation id="EFEF33FBFF1EFF38FE07F9393308FD9E" author="Duranton, M." box="[399,628,598,622]" journalOrPublisher="Coleopteres, numero special" pageId="213" pageNumber="212" pagination="1 - 31" part="1" refId="ref200092" refString="Duranton, M. 2011. Chorologie et ecologie des dynastides de Guyane francaise. Coleopteres, numero special 1: 1 - 31." title="Chorologie et ecologie des dynastides de Guyane francaise" type="journal article" year="2011">Duranton 2011: 20</bibRefCitation>
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<emphasis id="B90A9218FF1EFF38FD2FF9393222FD9E" box="[679,862,598,622]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
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<bibRefCitation id="EFEF33FBFF1EFF38FCFEF9383539FD9E" author="Ponchel" box="[886,1093,598,622]" firstAuthor="Ponchel" pageId="213" pageNumber="212" pagination="60 - 61" refId="ref220051" refString="Ponchel, Y. 2011. Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 4: 60 - 61. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." type="journal article" year="2011">Ponchel 2011: 61</bibRefCitation>
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<emphasis id="B90A9218FF1EFF38FBF0F9393453FD9E" box="[1144,1327,598,622]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
</taxonomicName>
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<bibRefCitation id="EFEF33FBFF1EFF38FACFF938303AFD7E" author="Dupuis, F." journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="213" pageNumber="212" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis 2016b: 110</bibRefCitation>
; MHCN; NZCS.
<emphasis id="B90A9218FF1EFF38FD83F91833E2FD7E" bold="true" box="[523,670,630,654]" pageId="213" pageNumber="212">“Guianas”:</emphasis>
<bibRefCitation id="EFEF33FBFF1EFF38FD2CF9193216FD7E" author="Arrow, G. J." bookContentInfo="124 p." box="[676,874,630,655]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="213" pageNumber="212" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 37</bibRefCitation>
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<emphasis id="B90A9218FF1EFF38FC11F9193532FD7E" box="[921,1102,630,654]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
</taxonomicName>
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<bibRefCitation id="EFEF33FBFF1EFF38FBEDF918340AFD7E" author="Blackwelder, R. E." box="[1125,1398,630,654]" journalOrPublisher="United States National Museum Bulletin" pageId="213" pageNumber="212" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 254</bibRefCitation>
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<taxonomicName id="4C7E3589FF1EFF38FF48F9F9300BFD5E" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[192,375,662,686]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus">
<emphasis id="B90A9218FF1EFF38FF48F9F9300BFD5E" box="[192,375,662,686]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FE07F9F83313FD5E" author="Endrodi, S." box="[399,623,662,686]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="213" pageNumber="212" pagination="1 - 145" part="37" refId="ref200804" refString="Endrodi, S. 1969 a. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 37 (1): 1 - 145." title="Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini" type="journal article" year="1969">Endrödi 1969a: 51</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FD2AF9F93296FD5E" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[674,1002,662,686]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus" subGenus="Euligyrus">
<emphasis id="B90A9218FF1EFF38FD2AF9F9327CFD5E" box="[674,768,663,686]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FC98F9F832F4FD5E" box="[784,904,662,686]" italics="true" pageId="213" pageNumber="212">Euligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FC10F9F83296FD5E" box="[920,1002,662,686]" italics="true" pageId="213" pageNumber="212">ebenus</emphasis>
</taxonomicName>
).
<emphasis id="B90A9218FF1EFF38FB8AF9F835D8FD5E" bold="true" box="[1026,1188,662,686]" pageId="213" pageNumber="212">
<collectingCountry id="F3690E9AFF1EFF38FB8AF9F8352AFD5E" box="[1026,1110,662,686]" name="Brazil" pageId="213" pageNumber="212">Brazil</collectingCountry>
(NA):
</emphasis>
<bibRefCitation id="EFEF33FBFF1EFF38FB23F9F93408FD5E" author="Arrow, G. J." bookContentInfo="124 p." box="[1195,1396,662,687]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="213" pageNumber="212" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 37</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FF48F9D9300AFD3E" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[192,374,694,718]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus">
<emphasis id="B90A9218FF1EFF38FF48F9D9300AFD3E" box="[192,374,694,718]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FE06F9D833DEFD3E" author="Blackwelder, R. E." box="[398,674,694,718]" journalOrPublisher="United States National Museum Bulletin" pageId="213" pageNumber="212" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 254</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FD5BF9D932F6FD3E" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[723,906,694,718]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus">
<emphasis id="B90A9218FF1EFF38FD5BF9D932F6FD3E" box="[723,906,694,718]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FC29F9D835FCFD3E" author="Endrodi, S." box="[929,1152,694,718]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="213" pageNumber="212" pagination="1 - 145" part="37" refId="ref200804" refString="Endrodi, S. 1969 a. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 37 (1): 1 - 145." title="Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini" type="journal article" year="1969">Endrödi 1969a: 51</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FB3AF9D9306EFD1E" baseAuthorityName="De Geer" baseAuthorityYear="1774" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus" subGenus="Euligyrus">
<emphasis id="B90A9218FF1EFF38FB3AF9D9346CFD3E" box="[1202,1296,695,718]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FA97F9D834EBFD3E" box="[1311,1431,694,718]" italics="true" pageId="213" pageNumber="212">Euligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FF48F9B8306EFD1E" box="[192,274,726,750]" italics="true" pageId="213" pageNumber="212">ebenus</emphasis>
</taxonomicName>
) (NA), 1985a: 254 (as
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<emphasis id="B90A9218FF1EFF38FDA0F9B933FAFD1E" box="[552,646,727,750]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FD1FF9B83273FD1E" box="[663,783,726,750]" italics="true" pageId="213" pageNumber="212">Euligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FCA8F9B8320EFD1E" box="[800,882,726,750]" italics="true" pageId="213" pageNumber="212">ebenus</emphasis>
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<bibRefCitation id="EFEF33FBFF1EFF38FC03F9B835DEFD1E" author="Dechambre, R. - P." box="[907,1186,726,750]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="213" pageNumber="212" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 168</bibRefCitation>
(as
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<emphasis id="B90A9218FF1EFF38FB5FF9B934F3FD1E" box="[1239,1423,726,750]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FF48F99830CFFCFE" author="Lachaume" box="[192,435,758,782]" firstAuthor="Lachaume" pageId="213" pageNumber="212" pagination="7 - 56" refId="ref211183" refString="Lachaume, G. 1992. Dynastidae americains Cyclocephalini - Agaocephalini - Pentodontini - Oryctini - Phileurini. Les Coleopteres du Monde 14: 7 - 56." type="journal article" year="1992">Lachaume 1992: 33</bibRefCitation>
(as
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<emphasis id="B90A9218FF1EFF38FE64F99933D9FCFE" box="[492,677,758,782]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
</taxonomicName>
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<bibRefCitation id="EFEF33FBFF1EFF38FD48F9983525FCFE" author="Andreazze, R. &amp; C. R. V. da Fonseca" box="[704,1113,758,782]" journalOrPublisher="Acta Amazonica" pageId="213" pageNumber="212" pagination="59 - 66" part="28" refId="ref190842" refString="Andreazze, R., and C. R. V. da Fonseca. 1998. Dinastineos (Coleoptera, Scarabaeoidea, Melolonthidae) em uma area de terra firme na Amazonia central, Brasil. Acta Amazonica 28 (1): 59 - 66." title="Dinastineos (Coleoptera, Scarabaeoidea, Melolonthidae) em uma area de terra firme na Amazonia central, Brasil" type="journal article" year="1998">Andreazze and Fonseca 1998: 62</bibRefCitation>
(as
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<emphasis id="B90A9218FF1EFF38FB19F9993437FCFE" box="[1169,1355,758,782]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
</taxonomicName>
) (NA);
<bibRefCitation id="EFEF33FBFF1EFF38FF48F87830D7FCDE" author="Ratcliffe" box="[192,427,790,814]" firstAuthor="Ratcliffe" pageId="213" pageNumber="212" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe 2003: 274</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF1EFF38FE30F8783266FCDE" author="Escalona, H. E. &amp; L. J. Joly" box="[440,794,790,814]" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="213" pageNumber="212" pagination="111 - 137" part="39" refId="ref201573" refString="Escalona, H. E., and L. J. Joly. 2006. El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Boletin de la Sociedad Entomologica Aragonesa 39: 111 - 137." title="El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini)" type="journal article" year="2006">Escalona and Joly 2006: 133</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FCD8F87935E7FCDE" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[848,1179,790,814]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus" subGenus="Euligyrus">
<emphasis id="B90A9218FF1EFF38FCD8F87932D2FCDE" box="[848,942,791,814]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FC37F878354BFCDE" box="[959,1079,790,814]" italics="true" pageId="213" pageNumber="212">Euligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FBC1F87835E7FCDE" box="[1097,1179,790,814]" italics="true" pageId="213" pageNumber="212">ebenus</emphasis>
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<bibRefCitation id="EFEF33FBFF1EFF38FB3DF8783044FCBE" author="Lopez-Garcia" etAl="et al." firstAuthor="Lopez-Garcia" pageId="213" pageNumber="212" pagination="451 - 492" refId="ref212996" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, and G. Amat-Garcia. 2015. The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution. Zootaxa 4048 (4): 451 - 492." type="journal article" year="2015">López-García et al. 2015: 481</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF1EFF38FECCF85830C1FCBE" author="Lopez-Garcia" box="[324,445,822,846]" firstAuthor="Lopez-Garcia" pageId="213" pageNumber="212" pagination="491 - 509" refId="ref213049" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, R. D. Cave, and G. Amat-Garcia. 2016. An annotated checklist of the New World pentodontine scarab beetles (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Zootaxa 4170 (3): 491 - 509." type="journal article" year="2016">2016: 502</bibRefCitation>
.
<emphasis id="B90A9218FF1EFF38FE40F85833D8FCBE" bold="true" box="[456,676,822,846]" pageId="213" pageNumber="212">
<collectingCountry id="F3690E9AFF1EFF38FE40F858332AFCBE" box="[456,598,822,846]" name="Venezuela" pageId="213" pageNumber="212">Venezuela</collectingCountry>
(EO):
</emphasis>
<bibRefCitation id="EFEF33FBFF1EFF38FD23F85832F6FCBE" author="Endrodi, S." box="[683,906,822,846]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="213" pageNumber="212" pagination="1 - 145" part="37" refId="ref200804" refString="Endrodi, S. 1969 a. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 37 (1): 1 - 145." title="Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini" type="journal article" year="1969">Endrödi 1969a: 51</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FC34F8593478FCBE" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[956,1284,822,846]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus" subGenus="Euligyrus">
<emphasis id="B90A9218FF1EFF38FC34F8593566FCBE" box="[956,1050,823,846]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FBA2F85835DEFCBE" box="[1066,1186,822,846]" italics="true" pageId="213" pageNumber="212">Euligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FB3AF8583478FCBE" box="[1202,1284,822,846]" italics="true" pageId="213" pageNumber="212">ebenus</emphasis>
</taxonomicName>
), 1985a: 254 (as
<taxonomicName id="4C7E3589FF1EFF38FF66F8393340FC9E" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[238,572,854,878]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus" subGenus="Euligyrus">
<emphasis id="B90A9218FF1EFF38FF66F8393030FC9E" box="[238,332,855,878]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FED7F83830ABFC9E" box="[351,471,854,878]" italics="true" pageId="213" pageNumber="212">Euligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FE62F8383340FC9E" box="[490,572,854,878]" italics="true" pageId="213" pageNumber="212">ebenus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FDDEF838320DFC9E" author="Dechambre, R. - P." box="[598,881,854,878]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="213" pageNumber="212" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 168</bibRefCitation>
(as
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<emphasis id="B90A9218FF1EFF38FC21F839351FFC9E" box="[937,1123,854,878]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FBF6F838340DFC9E" author="Lachaume" box="[1150,1393,854,878]" firstAuthor="Lachaume" pageId="213" pageNumber="212" pagination="7 - 56" refId="ref211183" refString="Lachaume, G. 1992. Dynastidae americains Cyclocephalini - Agaocephalini - Pentodontini - Oryctini - Phileurini. Les Coleopteres du Monde 14: 7 - 56." type="journal article" year="1992">Lachaume 1992: 33</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FF48F8193006FC7E" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[192,378,886,910]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus">
<emphasis id="B90A9218FF1EFF38FF48F8193006FC7E" box="[192,378,886,910]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FE1CF8183385FC7E" author="Escalona, H. E. &amp; L. J. Joly" box="[404,761,886,910]" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="213" pageNumber="212" pagination="111 - 137" part="39" refId="ref201573" refString="Escalona, H. E., and L. J. Joly. 2006. El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Boletin de la Sociedad Entomologica Aragonesa 39: 111 - 137." title="El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini)" type="journal article" year="2006">Escalona and Joly 2006: 133</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FCB9F8193502FC7E" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[817,1150,886,910]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus" subGenus="Euligyrus">
<emphasis id="B90A9218FF1EFF38FCB9F81932F3FC7E" box="[817,911,887,910]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FC29F8183565FC7E" box="[929,1049,886,910]" italics="true" pageId="213" pageNumber="212">Euligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FBA4F8183502FC7E" box="[1068,1150,886,910]" italics="true" pageId="213" pageNumber="212">ebenus</emphasis>
</taxonomicName>
) (EO);
<bibRefCitation id="EFEF33FBFF1EFF38FB54F818301DFC5E" author="Lopez-Garcia" etAl="et al." firstAuthor="Lopez-Garcia" pageId="213" pageNumber="212" pagination="451 - 492" refId="ref212996" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, and G. Amat-Garcia. 2015. The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution. Zootaxa 4048 (4): 451 - 492." type="journal article" year="2015">López-García et al. 2015: 481</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF1EFF38FEE6F8F83094FC5E" author="Lopez-Garcia" box="[366,488,918,942]" firstAuthor="Lopez-Garcia" pageId="213" pageNumber="212" pagination="491 - 509" refId="ref213049" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, R. D. Cave, and G. Amat-Garcia. 2016. An annotated checklist of the New World pentodontine scarab beetles (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Zootaxa 4170 (3): 491 - 509." type="journal article" year="2016">2016: 502</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF1EFF38FE7DF8F8320AFC5E" author="Kirmse and Ratcliffe" box="[501,886,918,942]" firstAuthor="Kirmse" pageId="213" pageNumber="212" pagination="149 - 167" refId="ref210433" refString="Kirmse, S., and B. C. Ratcliffe. 2019. Composition and host-use patterns of a scarab beetle (Coleoptera: Scarabaeidae) community inhabiting the canopy of a lowland tropical rainforest in southern Venezuela. The Coleopterists Bulletin 73 (1): 149 - 167." type="journal article" year="2019">Kirmse and Ratcliffe 2019: 153</bibRefCitation>
(EO).
<emphasis id="B90A9218FF1EFF38FC40F8F8355CFC5E" bold="true" box="[968,1056,918,942]" pageId="213" pageNumber="212">Other:</emphasis>
<bibRefCitation id="EFEF33FBFF1EFF38FBA0F8F93588FC5E" author="Arrow, G. J." bookContentInfo="124 p." box="[1064,1268,918,943]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="213" pageNumber="212" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 37</bibRefCitation>
(Guadeloupe, Mexico, Peru - as
<taxonomicName id="4C7E3589FF1EFF38FE07F8D9333EFC3E" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[399,578,950,974]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus">
<emphasis id="B90A9218FF1EFF38FE07F8D9333EFC3E" box="[399,578,950,974]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FDDFF8D8321AFC3E" author="Blackwelder, R. E." box="[599,870,950,974]" journalOrPublisher="United States National Museum Bulletin" pageId="213" pageNumber="212" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 254</bibRefCitation>
(Mexico, Peru, West Indies - as
<taxonomicName id="4C7E3589FF1EFF38FB55F8D934ECFC3E" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[1245,1424,950,974]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus">
<emphasis id="B90A9218FF1EFF38FB55F8D934ECFC3E" box="[1245,1424,950,974]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FF48F8B830D4FC1E" author="Endrodi, S." box="[192,424,982,1006]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="213" pageNumber="212" pagination="1 - 145" part="37" refId="ref200804" refString="Endrodi, S. 1969 a. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 37 (1): 1 - 145." title="Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini" type="journal article" year="1969">Endrödi 1969a: 51</bibRefCitation>
(Bolivia, Colombia, Mexico, Peru, Trinidad and Tobago - as
<taxonomicName id="4C7E3589FF1EFF38FB25F8B9306EFBFE" baseAuthorityName="De Geer" baseAuthorityYear="1774" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus" subGenus="Euligyrus">
<emphasis id="B90A9218FF1EFF38FB25F8B93477FC1E" box="[1197,1291,983,1006]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FA97F8B834EBFC1E" box="[1311,1431,982,1006]" italics="true" pageId="213" pageNumber="212">Euligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FF48F898306EFBFE" box="[192,274,1014,1038]" italics="true" pageId="213" pageNumber="212">ebenus</emphasis>
</taxonomicName>
), 1985a: 254 (Bolivia, Colombia, Mexico, Peru, Trinidad and Tobago - as
<taxonomicName id="4C7E3589FF1EFF38FB26F899306EFBDE" baseAuthorityName="De Geer" baseAuthorityYear="1774" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus" subGenus="Euligyrus">
<emphasis id="B90A9218FF1EFF38FB26F8993470FBFE" box="[1198,1292,1015,1038]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FA97F89834EBFBFE" box="[1311,1431,1014,1038]" italics="true" pageId="213" pageNumber="212">Euligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FF48FF78306EFBDE" box="[192,274,1046,1070]" italics="true" pageId="213" pageNumber="212">ebenus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FEA5FF78333BFBDE" author="Dechambre, R. - P." box="[301,583,1046,1070]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="213" pageNumber="212" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre 1979c: 168</bibRefCitation>
(Bolivia, Colombia, Mexico, Peru, Trinidad and Tobago - as
<taxonomicName id="4C7E3589FF1EFF38FACAFF79306EFBBE" baseAuthorityName="De Geer" baseAuthorityYear="1774" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus">
<emphasis id="B90A9218FF1EFF38FACAFF79306EFBBE" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FEA0FF58336DFBBE" author="Lachaume" box="[296,529,1078,1102]" firstAuthor="Lachaume" pageId="213" pageNumber="212" pagination="7 - 56" refId="ref211183" refString="Lachaume, G. 1992. Dynastidae americains Cyclocephalini - Agaocephalini - Pentodontini - Oryctini - Phileurini. Les Coleopteres du Monde 14: 7 - 56." type="journal article" year="1992">Lachaume 1992: 33</bibRefCitation>
(Bolivia, Colombia, Mexico, Peru, Trinidad and Tobago - as
<taxonomicName id="4C7E3589FF1EFF38FB53FF5934ECFBBE" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[1243,1424,1078,1102]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus">
<emphasis id="B90A9218FF1EFF38FB53FF5934ECFBBE" box="[1243,1424,1078,1102]" italics="true" pageId="213" pageNumber="212">Ligyrus ebenus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FF48FF3830DDFB9E" author="Ratcliffe" box="[192,417,1110,1134]" firstAuthor="Ratcliffe" pageId="213" pageNumber="212" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe 2003: 274</bibRefCitation>
(Brazil/BoliviaMexico);
<bibRefCitation id="EFEF33FBFF1EFF38FD5AFF38355AFB9E" author="Escalona, H. E. &amp; L. J. Joly" box="[722,1062,1110,1134]" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="213" pageNumber="212" pagination="111 - 137" part="39" refId="ref201573" refString="Escalona, H. E., and L. J. Joly. 2006. El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Boletin de la Sociedad Entomologica Aragonesa 39: 111 - 137." title="El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini)" type="journal article" year="2006">Escalona and Joly 2006: 133</bibRefCitation>
(Bolivia, Colombia, Costa Rica, Mexico, Panama, Peru, Trinidad and Tobago - as
<taxonomicName id="4C7E3589FF1EFF38FC9DFF193527FB7E" baseAuthorityName="De Geer" baseAuthorityYear="1774" box="[789,1115,1142,1166]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="ebenus" subGenus="Euligyrus">
<emphasis id="B90A9218FF1EFF38FC9DFF19320FFB7E" box="[789,883,1143,1166]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FC09FF183285FB7E" box="[897,1017,1142,1166]" italics="true" pageId="213" pageNumber="212">Euligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FB81FF183527FB7E" box="[1033,1115,1142,1166]" italics="true" pageId="213" pageNumber="212">ebenus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FBFAFF183190FB5E" author="Lopez-Garcia" etAl="et al." firstAuthor="Lopez-Garcia" pageId="213" pageNumber="212" pagination="451 - 492" refId="ref212996" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, and G. Amat-Garcia. 2015. The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution. Zootaxa 4048 (4): 451 - 492." type="journal article" year="2015">López-García et al. 2015: 481</bibRefCitation>
(Belize, Bolivia, Colombia, Guatemala, Honduras, Lesser Antilles, Mexico, Nicaragua, Peru), 2016: 502 (Belize, Bolivia, Colombia, Costa Rica, Guatemala, Honduras, Lesser Antilles, Mexico, Nicaragua, Panama, Peru, Trinidad and Tobago).
</paragraph>
</subSubSection>
</treatment>
</document>