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<document id="806FC5B8265E8E83D45C9BC6BED1ECE8" ID-DOI="10.11646/phytotaxa.295.3.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13688365" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725496115713" checkinUser="felipe" docAuthor="Steyn, Hester M. &amp; Van Wyk, Abraham E." docDate="2017" docId="03D7F475622EFFC0FF79FCE4FBF5F8A3" docLanguage="en" docName="phytotaxa.295.3.1.pdf" docOrigin="Phytotaxa 295 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.295.3.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Acanthopsis tetragona H. M. Steyn 2017, sp. nov." docType="treatment" docVersion="2" lastPageNumber="212" masterDocId="FFEE8C0D6225FFCBFFF1FFE0FFEEFFF6" masterDocTitle="Taxonomic notes on Acanthopsis Harv. (Acanthaceae, tribe Acantheae): the group with trifid bracts" masterLastPageNumber="217" masterPageNumber="201" pageNumber="212" updateTime="1725496529253" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="39317C67A81FA09AB3CB8CC9B3CC3AAC">Taxonomic notes on Acanthopsis Harv. (Acanthaceae, tribe Acantheae): the group with trifid bracts</mods:title>
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<mods:namePart id="3CEBE35565BCA37DCA9E26972C2E7442">Steyn, Hester M.</mods:namePart>
<mods:affiliation id="55D838488556935B5B9F4E29365B0B9A">National Herbarium, South African National Biodiversity Institute, Private Bag X 101, Pretoria 0001, South Africa.</mods:affiliation>
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<mods:title id="91A3A141D6D58B383B3F5BC4214A86D0">Phytotaxa</mods:title>
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<mods:date id="01F850C3040A6898EE4BA40BB984A678">2017</mods:date>
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<mods:identifier id="A459C350AB54526898D98843BF69A410" type="DOI">10.11646/phytotaxa.295.3.1</mods:identifier>
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<treatment id="03D7F475622EFFC0FF79FCE4FBF5F8A3" ID-DOI="http://doi.org/10.5281/zenodo.13688375" ID-Zenodo-Dep="13688375" LSID="urn:lsid:plazi:treatment:03D7F475622EFFC0FF79FCE4FBF5F8A3" httpUri="http://treatment.plazi.org/id/03D7F475622EFFC0FF79FCE4FBF5F8A3" lastPageNumber="212" pageId="11" pageNumber="212">
<subSubSection id="C36416E8622EFFC0FF79FCE4FD65FCEB" box="[136,651,770,797]" pageId="11" pageNumber="212" type="nomenclature">
<paragraph id="8BC14563622EFFC0FF79FCE4FD65FCEB" blockId="11.[136,651,770,797]" box="[136,651,770,797]" pageId="11" pageNumber="212">
<heading id="D089F20F622EFFC0FF79FCE4FD65FCEB" box="[136,651,770,797]" fontSize="11" level="2" pageId="11" pageNumber="212" reason="4">
<emphasis id="B90A9971622EFFC0FF79FCE4FF73FCEB" bold="true" box="[136,157,771,797]" pageId="11" pageNumber="212">3.</emphasis>
<taxonomicName id="4C7E3EE0622EFFC0FF52FCE2FDC4FCEB" ID-CoL="8ZLL" authority="H.M.Steyn" authorityName="H. M. Steyn" authorityYear="2017" box="[163,554,770,797]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="212" phylum="Tracheophyta" rank="species" species="tetragona" status="sp. nov.">
<emphasis id="B90A9971622EFFC0FF52FCE2FE48FCEB" bold="true" box="[163,422,770,797]" italics="true" pageId="11" pageNumber="212">Acanthopsis tetragona</emphasis>
H.M.Steyn
</taxonomicName>
,
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<taxonomicNameLabel id="A239240A622EFFC0FDC7FCE4FD65FCEB" box="[566,651,772,797]" pageId="11" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C36416E8622EFFC0FF79FCAEFCE6FC71" pageId="11" pageNumber="212" type="materials_examined">
<paragraph id="8BC14563622EFFC0FF79FCAEFCE6FC71" blockId="11.[136,1451,846,903]" pageId="11" pageNumber="212">
<emphasis id="B90A9971622EFFC0FF79FCAEFF2CFC95" bold="true" box="[136,194,846,867]" pageId="11" pageNumber="212">Type:</emphasis>
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<collectingCountry id="F36905F3622EFFC0FF2AFCAEFE65FC95" box="[219,395,846,867]" name="South Africa" pageId="11" pageNumber="212">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="49BA8B81622EFFC0FE64FCAEFDC6FC95" box="[405,552,846,867]" country="South Africa" name="Northern Cape" pageId="11" pageNumber="212">Northern Cape</collectingRegion>
:
<collectingCounty id="62A03DEF622EFFC0FDC2FCAEFCE4FC95" box="[563,778,846,867]" pageId="11" pageNumber="212">Namaqualand District</collectingCounty>
,
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,
<collectionCode id="ED6FDDA6622EFFC0FC6AFCAEFC46FC95" box="[923,936,846,867]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="11" pageNumber="212" type="Herbarium">S</collectionCode>
of
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camp site,
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, (2817
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),
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<collectingDate id="EF849A4B622EFFC0FAA5FCAEFF00FC71" pageId="11" pageNumber="212" value="2011-08-08">8 August 2011</collectingDate>
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,
<emphasis id="B90A9971622EFFC0FF08FC92FE8DFC71" box="[249,355,882,903]" italics="true" pageId="11" pageNumber="212">Steyn 1848</emphasis>
(
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<specimenCode id="DBD8ED18622EFFC0FE3BFC92FDBAFC71" box="[458,596,882,903]" collectionCode="PRE" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="11" pageNumber="212" type="Herbarium">PRE0861471</specimenCode>
-0!;
<typeStatus id="54C5FBC1622EFFC0FD82FC92FD56FC71" box="[627,696,882,903]" pageId="11" pageNumber="212" type="isotype">isotype</typeStatus>
<collectionCode id="ED6FDDA6622EFFC0FD4FFC92FD17FC71" box="[702,761,882,903]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15565" lsid="urn:lsid:biocol.org:col:15565" name="South African National Biodiversity Institute" pageId="11" pageNumber="212" type="Herbarium">NBG</collectionCode>
!)
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.
</paragraph>
</subSubSection>
<subSubSection id="C36416E8622EFFC0FF79FC58FD48F8FB" pageId="11" pageNumber="212" type="diagnosis">
<paragraph id="8BC14563622EFFC0FF79FC58FF08FBCB" blockId="11.[136,1458,952,1877]" pageId="11" pageNumber="212">
<emphasis id="B90A9971622EFFC0FF79FC58FEEBFC27" bold="true" box="[136,261,952,977]" pageId="11" pageNumber="212">Diagnosis:</emphasis>
<taxonomicName id="4C7E3EE0622EFFC0FED0FC58FDFEFC27" box="[289,528,952,977]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="212" phylum="Tracheophyta" rank="species" species="tetragon">
<emphasis id="B90A9971622EFFC0FED0FC58FDFEFC27" box="[289,528,952,977]" italics="true" pageId="11" pageNumber="212">Acanthopsis tetragon</emphasis>
</taxonomicName>
a differs from
<taxonomicName id="4C7E3EE0622EFFC0FD40FC58FCF3FC27" authorityName="Schinz" authorityYear="1890" baseAuthorityName="Nees" box="[689,797,952,977]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="212" phylum="Tracheophyta" rank="species" species="glauca">
<emphasis id="B90A9971622EFFC0FD40FC58FCF3FC27" box="[689,797,952,977]" italics="true" pageId="11" pageNumber="212">A. glauca</emphasis>
</taxonomicName>
in bracts being short-spined (not with excurrent spines up to
<quantity id="4C86E886622EFFC0FF55FC3CFF1DFC03" box="[164,243,988,1013]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="212" unit="mm" value="20.0">20 mm</quantity>
long), long, viscid glandular hairs (not short, eglandular hairs), inflorescences thicker, (12)1416(20) mm (not
<quantity id="4C86E886622EFFC0FF4FFBE0FEC5FBEF" box="[190,299,1024,1049]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="11" pageNumber="212" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">810 mm</quantity>
) in diameter and from
<taxonomicName id="4C7E3EE0622EFFC0FDC2FBE0FD53FBEF" box="[563,701,1024,1049]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="212" phylum="Tracheophyta" rank="species" species="dregeana">
<emphasis id="B90A9971622EFFC0FDC2FBE0FD53FBEF" box="[563,701,1024,1049]" italics="true" pageId="11" pageNumber="212">A. dregeana</emphasis>
</taxonomicName>
in being a compact subshrub
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tall (not a shrublet
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tall).
</paragraph>
<paragraph id="8BC14563622EFFC0FF4CFBA8FD48F8FB" blockId="11.[136,1458,952,1877]" pageId="11" pageNumber="212">
Perennial subshrub, (8)
<quantity id="4C86E886622EFFC0FE28FBA8FDBFFB97" box="[473,593,1096,1121]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" metricValueMax="2.0" metricValueMin="1.2" pageId="11" pageNumber="212" unit="cm" value="16.0" valueMax="20.0" valueMin="12.0">1220 cm</quantity>
tall. Leaves sessile; lamina oblanceolate, 4070(75) × (8)
<quantity id="4C86E886622EFFC0FAD4FBA8FA49FB97" box="[1317,1447,1096,1121]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="11" pageNumber="212" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">1015 mm</quantity>
, with appressed to strigose short, white hairs and subsessile glands; base attenuate, often decurrent; margin undulate, dentate-spinose to coarsely dentate-spinose, spines rigid (2)46(8) mm long, yellowish or maroonish brown; apex cuspidate. Inflorescences subsessile to shortly pedunculate, cylindrical to turbinate,
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long (excluding peduncle), (12)1416(20) mm in diameter. Bracts ovate to obovate, (22)2328(30) mm long, lamina (12)
<quantity id="4C86E886622EFFC0FA99FB38FF5AFAE3" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="11" pageNumber="212" unit="mm" value="15.5" valueMax="18.0" valueMin="13.0">1318 mm</quantity>
long; lower bracts ending in 5(7) primary spines, central primary spine nearly always simple; middle to upper bracts with 3(5) primary spines, usually ending in 3 broadly triangular to ovate, mucronate lobes with 3 main central veins usually converging towards tip of central primary spine, central primary spine usually to nearly always simple or with 2 pairs of very short, marginal secondary spines, primary spine tips usually recurved in fruit; lower bracts hirsute with deflexed to spreading short, white hairs; middle to upper bracts with spreading short, white hairs together with long, viscid glandular hairs, lamina often with long, spreading silky hairs, usually also with glandular hairs adaxially. Bracteoles narrowly lanceolate to linear, (5)
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long, silky-hairy, also with glandular hairs. Calyx with dorsal sepal ovate, acuminate to cuspidate, (11)
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long, silky-hairy, also with long, glandular hairs abaxially, especially towards tip, 57-veined from base; ventral sepal ovate, (11)
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long, silky-hairy, also with long, glandular hairs abaxially, especially towards tip, 57-veined from base; lateral sepals lanceolate, ending in spinous tip, broader at base,
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long, densely silky-hairy, also with glandular hairs. Corolla blue-purple or lilac with lemon-yellow throat, (25)2627(30) ×
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, tube (8)
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long; central lobe broader than long or equal, constricted at base, truncate to emarginate. Filaments 57(8) mm long, glandular; anthers beige,
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long. Style with patch of glandular hairs at base. Capsules ovate, flattened, glabrous, shiny, 7(9) × 3(4) mm. Seeds ovate, 6 ×
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, covered with long white hygroscopic hairs.
</paragraph>
</subSubSection>
<subSubSection id="C36416E8622EFFC0FF4CF8F8FBF5F8A3" pageId="11" pageNumber="212" type="etymology">
<paragraph id="8BC14563622EFFC0FF4CF8F8FBF5F8A3" blockId="11.[136,1458,952,1877]" pageId="11" pageNumber="212">
<emphasis id="B90A9971622EFFC0FF4CF8F8FEA6F8C7" bold="true" box="[189,328,1816,1841]" pageId="11" pageNumber="212">Etymology:</emphasis>
—The specific epithet
<taxonomicName id="4C7E3EE0622EFFC0FDB4F8F8FD50F8C7" authorityName="H. M. Steyn" authorityYear="2017" box="[581,702,1816,1841]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="212" phylum="Tracheophyta" rank="species" species="tetragona">
<emphasis id="B90A9971622EFFC0FDA0F8F8FD50F8C7" box="[593,702,1816,1841]" italics="true" pageId="11" pageNumber="212">tetragona</emphasis>
</taxonomicName>
” refers to the regular decussate arrangement of the bracts that gives the inflorescence a distinctly “four-angled” appearance when viewed from above.
</paragraph>
</subSubSection>
</treatment>
</document>