589 lines
75 KiB
XML
589 lines
75 KiB
XML
<document id="2D0229D080A810D29C3C6F8C15194F61" ID-CLB-Dataset="92619" ID-DOI="10.5852/ejt.2022.815.1749" ID-GBIF-Dataset="9a798cae-f10e-44c2-a65c-cdadbff28cb4" ID-ISSN="2118-9773" ID-Zenodo-Dep="6484153" ID-ZooBank="6AD0082E-7349-48DE-AFCA-1EE0BFBB3887" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1650913068508" checkinUser="felipe" docAuthor="Vranken, Nathan, Steenberge, Maarten Van, Heylen, Annelies, Decru, Eva & Snoeks, Jos" docDate="2022" docId="038F87A6613EDA65FDB756882794FA9A" docLanguage="en" docName="ejt-815_1-94.pdf" docOrigin="European Journal of Taxonomy 815" docSource="http://dx.doi.org/10.5852/ejt.2022.815.1749" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Haplochromis pardus Vranken & Steenberge & Heylen & Decru & Snoeks 2022, sp. nov." docType="treatment" docUuid="8DDACAC6-C993-4750-BE1B-1BF4C0312DB7" docUuidSource="ZooBank" docVersion="10" lastPageNumber="62" masterDocId="FFB6FFDE6107DA58FFD257322629FC0A" masterDocTitle="From a pair to a dozen: the piscivorous species of Haplochromis (Cichlidae) from the Lake Edward system" masterLastPageNumber="94" masterPageNumber="1" pageNumber="58" updateTime="1699329164302" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F8DA79FBD06E280D17D24C520C817857">From a pair to a dozen: the piscivorous species of Haplochromis (Cichlidae) from the Lake Edward system</mods:title>
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<mods:namePart id="6C131CA4E4A7CA887927614FC04A8907">Vranken, Nathan</mods:namePart>
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<mods:affiliation id="85884D0FA59797DE45915857640DD86C">Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium. & Royal Museum for Central Africa, Biology department, Section Vertebrates, Leuvensesteenweg 13, 3080 Tervuren, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="D7A1184B1D936BBCEB6D9F76DB199CDF" type="email">nathan.vranken@kuleuven.be</mods:nameIdentifier>
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<mods:namePart id="60AE596E943052ABB2EB7E5BE8FE7510">Steenberge, Maarten Van</mods:namePart>
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<mods:affiliation id="A1ACA71DD4A51A75FDD63161DF6EED84">Royal Belgian Institute of Natural Sciences, Operational Directorate Taxonomy and Phylogeny, Vautierstraat 29, 1000 Brussels, Belgium. & Masaryk University, Department of Botany and Zoology, Kotlářská 2, 611 37 Brno, Czech Republic.</mods:affiliation>
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<mods:nameIdentifier id="5327D755DDEFBDDAFE0E44509CAA60F1" type="email">mvansteenberge@naturalsciences.be</mods:nameIdentifier>
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<mods:namePart id="C76E1716D8C685E12CD3AD0F5F1425CB">Heylen, Annelies</mods:namePart>
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<mods:affiliation id="50C3A9577170FFCF8129DE42ACAD14F6">KU Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="A5F9C444227C0C8B292429F8750815AD" type="email">annelies.heylen@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="887F20757059155103FB686E9CB6A603">Decru, Eva</mods:namePart>
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<mods:affiliation id="71A1D4CBC5E7B5086F7DFA32081E28FF">KU Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="32CDCF90B5A7B75B15C5E5FD8049D89D" type="email">eva.decru.icht@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FA14CA78A7FCFACD493F69E97EBF969D">Snoeks, Jos</mods:namePart>
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<mods:affiliation id="D34F39A473F61E7CBEEB46D562048E53">Royal Museum for Central Africa, Biology department, Section Vertebrates, Leuvensesteenweg 13, 3080 Tervuren, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="DD2FB73E4295F7DCAC145406F04BBB3B" type="email">jos.snoeks@africamuseum.be</mods:nameIdentifier>
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<mods:title id="87E6E07070C461CC3976A35F303505A6">European Journal of Taxonomy</mods:title>
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<mods:date id="76578A402DAF66579BF1917130C57A90">2022</mods:date>
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<treatment id="038F87A6613EDA65FDB756882794FA9A" ID-DOI="http://doi.org/10.5281/zenodo.6502617" ID-GBIF-Taxon="197197500" ID-Zenodo-Dep="6502617" LSID="urn:lsid:zoobank.org:act:8DDACAC6-C993-4750-BE1B-1BF4C0312DB7" httpUri="http://treatment.plazi.org/id/038F87A6613EDA65FDB756882794FA9A" lastPageId="61" lastPageNumber="62" pageId="57" pageNumber="58">
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<heading id="D0D181DC613EDA61FDB7568825E6FDDF" box="[613,975,442,469]" centered="true" fontSize="11" level="2" pageId="57" pageNumber="58" reason="2">
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<taxonomicName id="4C264D33613EDA61FDB756882545FDDF" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[613,876,442,469]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="pardus" status="sp. nov.">
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<emphasis id="B952EAA2613EDA61FDB756882545FDDF" bold="true" box="[613,876,442,469]" italics="true" pageId="57" pageNumber="58">Haplochromis pardus</emphasis>
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<taxonomicNameLabel id="A26157D9613EDA61FCA6568925E6FDDF" box="[884,975,443,469]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C33C653B613EDA61FEB056EC22F8FDF3" box="[354,1233,478,505]" pageId="57" pageNumber="58" type="multiple">
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<paragraph id="8B9936B0613EDA61FEB056EC22F8FDF3" blockId="57.[354,1233,442,541]" box="[354,1233,478,505]" pageId="57" pageNumber="58">
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</paragraph>
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<subSubSection id="C33C653B613EDA61FD5A55302585FE17" box="[648,940,514,541]" pageId="57" pageNumber="58" type="description">
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<paragraph id="8B9936B0613EDA61FD5A55302585FE17" blockId="57.[354,1233,442,541]" box="[648,940,514,541]" pageId="57" pageNumber="58">
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<figureCitation id="131D2A35613EDA61FD5A553024D9FE17" box="[648,752,514,541]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="5.[190,234,1822,1848]" captionStartId-1="6.[189,231,1892,1918]" captionTargetBox-0="[193,1398,734,1726]" captionTargetBox-1="[208,1335,272,1830]" captionTargetId-0="figure-217@5.[189,1399,730,1726]" captionTargetId-1="figure-18@6.[208,1335,272,1830]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Fig. 1. The Lake Edward system, consisting of Lakes Edward and George that are connected through the Kazinga Channel. The catch localities of all examined specimens are indicated: Haplochromis sp. ‘latifrons’ (), H. mentatus Regan, 1925 (), H. sp. ‘rex’ (), H. sp. ‘simba’ (▼), H. sp. ‘glaucus’ (), H. sp. ‘aquila’ (), H. sp. ‘kimondo’ (●), H. sp. ‘falcatus’ (), H. sp. ‘curvidens’ (), H. sp. ‘pardus’ (), H. sp. ‘quasimodo’ (▲), and H. squamipinnis Regan, 1921 (█). The type localities of H. mentatus and H. squamipinnis are not shown as the exact locations are unknown." captionText-1="Fig. 2. a. Differences in total number of outer teeth and premaxillary dentition between the macro- and microdontic groups of piscivores from the Lake Edward system.b. Schematic presentation of the dominant male colour patterns in life, in the various groups studied. For H. sp. ‘latifrons’ the female and for H. sp. ‘curvidens’ the non-dominant colour patterns are shown, as those of dominant males are unknown." figureDoi-0="http://doi.org/10.5281/zenodo.6484155" figureDoi-1="http://doi.org/10.5281/zenodo.6484157" httpUri-0="https://zenodo.org/record/6484155/files/figure.png" httpUri-1="https://zenodo.org/record/6484157/files/figure.png" pageId="57" pageNumber="58">Figs 1–2</figureCitation>
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,
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<figureCitation id="131D2A35613EDA61FD2E55312560FE17" box="[764,841,515,541]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="58.[189,232,1963,1989]" captionStartId-1="59.[189,232,1892,1918]" captionStartId-2="60.[189,232,1963,1989]" captionTargetBox-0="[190,1398,1197,1855]" captionTargetBox-1="[242,1338,284,1857]" captionTargetBox-2="[189,1394,1035,1887]" captionTargetId-0="figure-363@58.[189,1399,1195,1855]" captionTargetId-1="figure-12@59.[242,1338,265,1857]" captionTargetId-2="figure-283@60.[189,1399,1035,1889]" captionTargetPageId-0="58" captionTargetPageId-1="59" captionTargetPageId-2="60" captionText-0="Fig. 32. Haplochromis pardus sp. nov., holotype, ♂, 89.2 mm SL (RMCA 2016.035.P.0202). Drawn by N. Vranken." captionText-1="Fig. 33. Haplochromis pardus sp. nov. a. Photograph of preserved holotype (RMCA 2016.035.P.0202; 89.2 mm SL). b. X-ray image of holotype. c–d. Photographs of freshly caught specimens. c. Dominant male (RMCA 2017.006.P (HP1463); 81.4 mm SL). d. Female (RMCA 2017.006.P.0342; 75.9 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." captionText-2="Fig. 34. Haplochromis pardus sp. nov. (RMCA 2016.035.P.0207; 96.1 mm SL). a. Dorsal view of the lower pharyngeal jaw. b. Lateral view of the lower pharyngeal jaw." figureDoi-0="http://doi.org/10.5281/zenodo.6484234" figureDoi-1="http://doi.org/10.5281/zenodo.6484236" figureDoi-2="http://doi.org/10.5281/zenodo.6484238" httpUri-0="https://zenodo.org/record/6484234/files/figure.png" httpUri-1="https://zenodo.org/record/6484236/files/figure.png" httpUri-2="https://zenodo.org/record/6484238/files/figure.png" pageId="57" pageNumber="58">32–34</figureCitation>
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;
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<tableCitation id="C6A4030B613EDA61FC8755302585FE17" box="[853,940,514,541]" captionStart="Table 1" captionStartId="9.[182,252,548,576]" captionTargetPageId="9" captionText="Table 1. (continued on the next three pages). Measurements and counts of the piscivorous species of Haplochromis from the Lake Edward system." httpUri="http://table.plazi.org/id/DF596638610EDA51FF6455162158FE4A" pageId="57" pageNumber="58" tableUuid="DF596638610EDA51FF6455162158FE4A">Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C653B613EDA61FF6F5576279BFA1B" pageId="57" pageNumber="58" type="diagnosis">
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<paragraph id="8B9936B0613EDA61FF6F557627E0FE54" blockId="57.[189,457,580,606]" box="[189,457,580,606]" pageId="57" pageNumber="58">
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<heading id="D0D181DC613EDA61FF6F557627E0FE54" bold="true" box="[189,457,580,606]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
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<emphasis id="B952EAA2613EDA61FF6F557627E0FE54" bold="true" box="[189,457,580,606]" pageId="57" pageNumber="58">Differential diagnosis</emphasis>
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<paragraph id="8B9936B0613EDA61FF6F554622FCFEBB" blockId="57.[189,1398,628,690]" pageId="57" pageNumber="58">
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Species with a piscivorous morphology; adult size small (max. known size
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<quantity id="4CDE9B55613EDA61FBE2554622ABFE84" box="[1072,1154,628,654]" metricMagnitude="-2" metricUnit="m" metricValue="9.6" pageId="57" pageNumber="58" unit="mm" value="96.0">96 mm</quantity>
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SL); outer oral teeth many and small [UOT 39–56 (median 58)]; dominant males speckled to uniformly black.
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</paragraph>
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<paragraph id="8B9936B0613EDA61FF6F55E825BFFF31" blockId="57.[188,1399,730,828]" pageId="57" pageNumber="58">
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Amongst piscivorous species from the Lake Edward system,
|
||
<taxonomicName id="4C264D33613EDA61FC4355E8222EFEFE" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[913,1031,730,756]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="pardus" status="sp. nov.">
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<emphasis id="B952EAA2613EDA61FC4355E8258FFEF9" box="[913,934,730,755]" italics="true" pageId="57" pageNumber="58">H</emphasis>
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.
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<emphasis id="B952EAA2613EDA61FC6455E8222EFEFE" box="[950,1031,730,756]" italics="true" pageId="57" pageNumber="58">pardus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26157D9613EDA61FBDD55E82242FEFE" box="[1039,1131,730,756]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
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differs from all by the combination of a smaller adult size (max. 96 vs
|
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<quantity id="4CDE9B55613EDA61FD3955CC25ACFF12" box="[747,901,766,792]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" metricValueMax="2.11" metricValueMin="1.09" pageId="57" pageNumber="58" unit="mm" value="160.0" valueMax="211.0" valueMin="109.0">109–211 mm</quantity>
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SL) and colour pattern of small specimens (<
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SL) speckled to uniformly black vs light coloured.
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</paragraph>
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<paragraph id="8B9936B0613EDA61FF6F54562378FFCC" blockId="57.[189,1399,867,966]" pageId="57" pageNumber="58">
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It further differs from
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<taxonomicName id="4C264D33613EDA61FE1654562460FF77" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[452,585,867,894]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="latifrons" status="sp. nov.">
|
||
<emphasis id="B952EAA2613EDA61FE16545627F0FF77" box="[452,473,868,893]" italics="true" pageId="57" pageNumber="58">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA2613EDA61FE3854512460FF77" box="[490,585,867,893]" italics="true" pageId="57" pageNumber="58">latifrons</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9613EDA61FD8154562499FF74" box="[595,688,868,894]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C264D33613EDA61FD6D54562562FF77" authorityName="Regan" authorityYear="1925" box="[703,843,868,894]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="mentatus">
|
||
<emphasis id="B952EAA2613EDA61FD6D545624FDFF77" box="[703,724,868,893]" italics="true" pageId="57" pageNumber="58">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA2613EDA61FD3654562562FF77" box="[740,843,868,893]" italics="true" pageId="57" pageNumber="58">mentatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C264D33613EDA61FC8E545625F2FF77" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[860,987,867,894]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="glaucus" status="sp. nov.">
|
||
<emphasis id="B952EAA2613EDA61FC8E54562558FF77" box="[860,881,868,893]" italics="true" pageId="57" pageNumber="58">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA2613EDA61FC50545125F2FF77" box="[898,987,867,893]" italics="true" pageId="57" pageNumber="58">glaucus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9613EDA61FC365456226BFF74" box="[996,1090,868,894]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C264D33613EDA61FB82545622F1FF77" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[1104,1240,867,894]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="kimondo" status="sp. nov.">
|
||
<emphasis id="B952EAA2613EDA61FB825456224CFF77" box="[1104,1125,868,893]" italics="true" pageId="57" pageNumber="58">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA2613EDA61FBA4545122F1FF77" box="[1142,1240,867,893]" italics="true" pageId="57" pageNumber="58">kimondo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9613EDA61FB3054562316FF74" box="[1250,1343,868,894]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C264D33613EDA61FF6F54BA275EFFAB" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[189,375,904,930]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="squamipinnis">
|
||
<emphasis id="B952EAA2613EDA61FF6F54BA26FBFFAB" box="[189,210,904,929]" italics="true" pageId="57" pageNumber="58">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA2613EDA61FF0D54BA275EFFAB" box="[223,375,904,929]" italics="true" pageId="57" pageNumber="58">squamipinnis</emphasis>
|
||
</taxonomicName>
|
||
by the combination of a shallower cheek [ChD 20.8–24.4 (mean 22.5) vs 23.8–36.0 (26.4– 30.9) % HL] and a narrower interorbital area [IOW 39.3–48.4 (44.6) vs 48.6–63.3 (51.9–60.0) % HW].
|
||
</paragraph>
|
||
<paragraph id="8B9936B0613EDA61FF6F54DC2217F845" blockId="57.[188,1398,1005,1104]" pageId="57" pageNumber="58">
|
||
It further differs from
|
||
<taxonomicName id="4C264D33613EDA61FE1354DC2423F80D" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[449,522,1006,1032]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="rex" status="sp. nov.">
|
||
<emphasis id="B952EAA2613EDA61FE1354DC27FFF80D" box="[449,470,1006,1031]" italics="true" pageId="57" pageNumber="58">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA2613EDA61FE3754DC2423F80D" box="[485,522,1006,1031]" italics="true" pageId="57" pageNumber="58">rex</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9613EDA61FDC354DC2445F802" box="[529,620,1006,1032]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C264D33613EDA61FDA854DC24CCF80D" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[634,741,1005,1032]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="simba" status="sp. nov.">
|
||
<emphasis id="B952EAA2613EDA61FDA854DC24A6F80D" box="[634,655,1006,1031]" italics="true" pageId="57" pageNumber="58">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA2613EDA61FD4C54DF24CCF80D" box="[670,741,1005,1031]" italics="true" pageId="57" pageNumber="58">simba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9613EDA61FD3F54DC2563F802" box="[749,842,1006,1032]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C264D33613EDA61FC5554DC25DEF80D" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[903,1015,1005,1032]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="aquila" status="sp. nov.">
|
||
<emphasis id="B952EAA2613EDA61FC5554DC25B5F80D" box="[903,924,1006,1031]" italics="true" pageId="57" pageNumber="58">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA2613EDA61FC7954DF25DEF80D" box="[939,1015,1005,1031]" italics="true" pageId="57" pageNumber="58">aquila</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9613EDA61FC2D54DC2270F802" box="[1023,1113,1006,1032]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the combination of a shallower cheek [ChD 20.8–24.4 (22.5) vs 26.8–33.5 (28.3–31.1) % HL], smaller outer oral teeth, and a larger number of outer upper jaw teeth [UOT 39–51 (45) vs 22–37 (27–31)].
|
||
</paragraph>
|
||
<paragraph id="8B9936B0613EDA61FF6F534A2574F8F7" blockId="57.[188,1399,1143,1278]" pageId="57" pageNumber="58">
|
||
It further differs from
|
||
<taxonomicName id="4C264D33613EDA61FE07534A2475F89B" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[469,604,1143,1170]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="falcatus" status="sp. nov.">
|
||
<emphasis id="B952EAA2613EDA61FE07534A27C3F89B" box="[469,490,1144,1169]" italics="true" pageId="57" pageNumber="58">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA2613EDA61FE2C53452475F89B" box="[510,604,1143,1169]" italics="true" pageId="57" pageNumber="58">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9613EDA61FDBB534A24E0F898" box="[617,713,1144,1170]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the combination of by weakly recurved vs strongly recurved outer jaw teeth, a shallower cheek [ChD 20.8–24.4 (22.5) vs 25.1–28.0 (exceptionally
|
||
<quantity id="4CDE9B55613EDA61FAF753A9235EF8BF" box="[1317,1399,1179,1205]" metricMagnitude="-1" metricUnit="m" metricValue="5.9182" pageId="57" pageNumber="58" unit="in" value="23.3">23.3 in</quantity>
|
||
<specimenCount id="9D20FD39613EDA61FF6F538D2776F8D0" box="[189,351,1215,1242]" pageId="57" pageNumber="58" type="generic">one specimen</specimenCount>
|
||
) (mean 26.0) % HL], and a shorter pre-dorsal distance [PrD 34.1–37.8 (36.0) vs 38.2–41.1 (exceptionally
|
||
<quantity id="4CDE9B55613EDA61FEB953D12794F8F7" box="[363,445,1251,1277]" metricMagnitude="-1" metricUnit="m" metricValue="9.3726" pageId="57" pageNumber="58" unit="in" value="36.9">36.9 in</quantity>
|
||
<specimenCount id="9D20FD39613EDA61FE1653D1244FF8F4" box="[452,614,1251,1278]" pageId="57" pageNumber="58" type="generic">one specimen</specimenCount>
|
||
) (mean 39.5) % SL].
|
||
</paragraph>
|
||
<paragraph id="8B9936B0613EDA61FF6F521727C5F98D" blockId="57.[189,1398,1317,1415]" pageId="57" pageNumber="58">
|
||
It further differs from
|
||
<taxonomicName id="4C264D33613EDA61FE6852142478F935" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[442,593,1317,1343]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="curvidens" status="sp. nov.">
|
||
<emphasis id="B952EAA2613EDA61FE68521427E6F935" box="[442,463,1318,1343]" italics="true" pageId="57" pageNumber="58">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA2613EDA61FE0C52172478F935" box="[478,593,1317,1343]" italics="true" pageId="57" pageNumber="58">curvidens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9613EDA61FD8A52172498F935" box="[600,689,1317,1343]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the combination of a longer anal-fin base [AFB 19.2–22.2 (20.5) vs 17.9–18.6 (18.3) % SL] and a slightly narrower interorbital area [IOW 39.3–48.4 (44.6) vs 46.4–52.5 (49.1) % HW].
|
||
</paragraph>
|
||
<paragraph id="8B9936B0613EDA61FF6F529D279BFA1B" blockId="57.[189,1398,1455,1553]" pageId="57" pageNumber="58">
|
||
It further differs from
|
||
<taxonomicName id="4C264D33613EDA61FE11529D2440F9C3" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[451,617,1455,1481]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="quasimodo" status="sp. nov.">
|
||
<emphasis id="B952EAA2613EDA61FE11529D27F1F9C2" box="[451,472,1455,1480]" italics="true" pageId="57" pageNumber="58">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA2613EDA61FE35529D2440F9C3" box="[487,617,1455,1481]" italics="true" pageId="57" pageNumber="58">quasimodo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9613EDA61FDA3529D24E5F9C3" box="[625,716,1455,1481]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the combination of a shallower cheek [ChD 20.8–24.4 (22.5) vs 23.7–32.9 (27.5) % HL] and a smaller number of caudal peduncle scales (CPS 16, rarely 17 vs 17–20, rarely 16).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C653B613EDA61FF6F510B2275FAAF" pageId="57" pageNumber="58" type="etymology">
|
||
<paragraph id="8B9936B0613EDA61FF6F510B276DFA59" blockId="57.[189,324,1593,1619]" box="[189,324,1593,1619]" pageId="57" pageNumber="58">
|
||
<heading id="D0D181DC613EDA61FF6F510B276DFA59" bold="true" box="[189,324,1593,1619]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
|
||
<emphasis id="B952EAA2613EDA61FF6F510B276DFA59" bold="true" box="[189,324,1593,1619]" pageId="57" pageNumber="58">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B0613EDA61FF6F515A2275FAAF" blockId="57.[189,1399,1640,1702]" pageId="57" pageNumber="58">
|
||
Specific name from the Latin ‘
|
||
<taxonomicName id="4C264D33613EDA61FDFF515A2457FA88" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[557,638,1640,1666]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="pardus">
|
||
<emphasis id="B952EAA2613EDA61FDFF515A2457FA88" box="[557,638,1640,1666]" italics="true" pageId="57" pageNumber="58">pardus</emphasis>
|
||
</taxonomicName>
|
||
’ for ‘leopard’; referring to nearly uniform black to yellow-pink flanks with clear black blotches, i.e., interrupted horizontal and vertical stripes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C653B613EDA62FF6F51FF22F2FE6A" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="materials_examined">
|
||
<paragraph id="8B9936B0613EDA61FF6F51FF2783FAED" blockId="57.[189,426,1741,1767]" box="[189,426,1741,1767]" pageId="57" pageNumber="58">
|
||
<heading id="D0D181DC613EDA61FF6F51FF2783FAED" bold="true" box="[189,426,1741,1767]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
|
||
<emphasis id="B952EAA2613EDA61FF6F51FF2783FAED" bold="true" box="[189,426,1741,1767]" pageId="57" pageNumber="58">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B0613EDA61FF3851CF2200FB55" blockId="57.[189,1399,1789,1887]" pageId="57" pageNumber="58">
|
||
<materialsCitation id="3B4E3CED613EDA61FF3851CF2200FB55" ID-GBIF-Occurrence="3861203371" collectingDate="2016-10-23" collectionCode="RMCA" collectorName="HIPE 1 exped." country="Uganda" latitude="-0.192" location="Mweya" longLatPrecision="1" longitude="29.874" municipality="Mukutu Kihinga" pageId="57" pageNumber="58" specimenCode="RMCA 2016.035.P.0202" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="holotype">
|
||
<emphasis id="B952EAA2613EDA61FF3851CF2775FB1D" bold="true" box="[234,348,1789,1815]" pageId="57" pageNumber="58">
|
||
<typeStatus id="549D8812613EDA61FF3851CF2775FB1D" box="[234,348,1789,1815]" pageId="57" pageNumber="58" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F3317620613EDA61FF6F50132713FB31" box="[189,314,1825,1851]" name="Uganda" pageId="57" pageNumber="58">UGANDA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D20FD39613EDA61FE8750132744FB31" box="[341,365,1825,1851]" pageId="57" pageNumber="58" type="male">♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B55613EDA61FEA8501327CAFB31" box="[378,483,1825,1851]" metricMagnitude="-2" metricUnit="m" metricValue="8.92" pageId="57" pageNumber="58" unit="mm" value="89.2">89.2 mm</quantity>
|
||
SL;
|
||
<collectingRegion id="49E2F852613EDA61FDCC50132497FB31" box="[542,702,1825,1851]" pageId="57" pageNumber="58">Lake Edward</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BFDACCA613EDA61FD19501325BBFB31" box="[715,914,1825,1851]" pageId="57" pageNumber="58">Mukutu Kihinga</collectingMunicipality>
|
||
, rocky offshore of
|
||
<location id="8EF9606B613EDA61FBA0501322E7FB31" LSID="urn:lsid:plazi:treatment:038F87A6613EDA65FDB756882794FA9A:8EF9606B613EDA61FBA0501322E7FB31" box="[1138,1230,1825,1851]" country="Uganda" latitude="-0.192" longLatPrecision="1" longitude="29.874" municipality="Mukutu Kihinga" name="Mweya" pageId="57" pageNumber="58" stateProvince="Lake Edward">Mweya</location>
|
||
;
|
||
<geoCoordinate id="EE125077613EDA61FB08502D2358FB31" box="[1242,1393,1823,1851]" degrees="0" direction="south" minutes="11" orientation="latitude" pageId="57" pageNumber="58" precision="1" seconds="31.2" value="-0.192">0°11′31.2″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077613EDA61FF6F5071274FFB55" box="[189,358,1859,1887]" degrees="29" direction="east" minutes="52" orientation="longitude" pageId="57" pageNumber="58" precision="1" seconds="26.4" value="29.874">29°52′26.4″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070613EDA61FEA050772427FB54" box="[370,526,1860,1887]" pageId="57" pageNumber="58" value="2016-10-23">
|
||
<collectingDate id="EFDCE998613EDA61FEA050772427FB54" box="[370,526,1860,1887]" pageId="57" pageNumber="58" value="2016-10-23">23 Oct. 2016</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366613EDA61FDC850772496FB54" box="[538,703,1860,1887]" pageId="57" pageNumber="58">HIPE1 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB613EDA61FCD35077220DFB55" box="[769,1060,1860,1887]" pageId="57" pageNumber="58">RMCA 2016.035.P.0202</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9936B0613EDA62FF3850B522F2FE6A" blockId="57.[189,1399,1927,2025]" lastBlockId="58.[189,1399,263,608]" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
|
||
<materialsCitation id="3B4E3CED613EDA61FF3850B52231FBE3" ID-GBIF-Occurrence="3861203353" collectingDate="2016-10-23" collectionCode="HIPE, RMCA" country="Uganda" lastPageId="57" lastPageNumber="59" latitude="-0.192" location="Mweya" longLatPrecision="1" longitude="29.874" municipality="Mukutu Kihinga" pageId="57" pageNumber="58" specimenCode="HIPE1, RMCA 2016.035.P.0203" specimenCount="1" specimenCount-female="1" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<emphasis id="B952EAA2613EDA61FF3850B5274EFBAB" bold="true" box="[234,359,1927,1953]" pageId="57" pageNumber="58">
|
||
<typeStatus id="549D8812613EDA61FF3850B5274EFBAB" box="[234,359,1927,1953]" pageId="57" pageNumber="58" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F3317620613EDA61FF6F50992713FBCF" box="[189,314,1963,1989]" name="Uganda" pageId="57" pageNumber="58">UGANDA</collectingCountry>
|
||
–
|
||
<emphasis id="B952EAA2613EDA61FE82509927D2FBCF" bold="true" box="[336,507,1963,1989]" pageId="57" pageNumber="58">
|
||
<collectingRegion id="49E2F852613EDA61FE82509927D2FBCF" box="[336,507,1963,1989]" pageId="57" pageNumber="58">Lake Edward</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D20FD39613EDA61FDC050992412FBCF" box="[530,571,1963,1989]" pageId="57" pageNumber="58" type="female">1 ♀</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B55613EDA61FD9550992487FBCF" box="[583,686,1963,1989]" metricMagnitude="-2" metricUnit="m" metricValue="8.49" pageId="57" pageNumber="58" unit="mm" value="84.9">84.9 mm</quantity>
|
||
SL;
|
||
<collectingMunicipality id="6BFDACCA613EDA61FD335098258CFBCE" box="[737,933,1962,1988]" pageId="57" pageNumber="58">Mukutu Kihinga</collectingMunicipality>
|
||
, rocky offshore of
|
||
<location id="8EF9606B613EDA61FBAA509922FDFBCF" LSID="urn:lsid:plazi:treatment:038F87A6613EDA65FDB756882794FA9A:8EF9606B613EDA61FBAA509922FDFBCF" box="[1144,1236,1963,1989]" country="Uganda" latitude="-0.192" longLatPrecision="1" longitude="29.874" municipality="Mukutu Kihinga" name="Mweya" pageId="57" pageNumber="58" stateProvince="Lake Edward">Mweya</location>
|
||
;
|
||
<geoCoordinate id="EE125077613EDA61FB0F509B2358FBCF" box="[1245,1393,1961,1989]" degrees="0" direction="south" minutes="11" orientation="latitude" pageId="57" pageNumber="58" precision="1" seconds="31.2" value="-0.192">0°11′31.2″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077613EDA61FF6F50FF274CFBE3" box="[189,357,1997,2025]" degrees="29" direction="east" minutes="52" orientation="longitude" pageId="57" pageNumber="58" precision="1" seconds="26.4" value="29.874">29°52′26.4″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070613EDA61FEA250FC2423FBE2" box="[368,522,1998,2024]" pageId="57" pageNumber="58" value="2016-10-23">
|
||
<collectingDate id="EFDCE998613EDA61FEA250FC2423FBE2" box="[368,522,1998,2024]" pageId="57" pageNumber="58" value="2016-10-23">23 Oct. 2016</collectingDate>
|
||
</date>
|
||
;
|
||
<specimenCode id="DB809ECB613EDA61FDC750FC244CFBE2" box="[533,613,1998,2024]" pageId="57" pageNumber="58">HIPE1</specimenCode>
|
||
exped. leg.;
|
||
<specimenCode id="DB809ECB613EDA61FD2B50FC2231FBE3" box="[761,1048,1998,2025]" pageId="57" pageNumber="58">RMCA 2016.035.P.0203</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED613EDA62FBE250FC27C4FD4C" ID-GBIF-Occurrence="3861203408" collectingDate="2016-10-24" collectionCode="RMCA" collectorName="HIPE 1 exped." country="Uganda" lastPageId="58" lastPageNumber="59" latitude="-0.209" location="Kazinga Channel" longLatPrecision="1" longitude="29.885" pageId="57" pageNumber="58" specimenCode="RMCA 2016.035.P.0204, RMCA 2016.035.P.0205, RMCA 2016.035.P.0206, RMCA 2016.035.P.0207" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<specimenCount id="9D20FD39613EDA61FBE250FC225AFBE3" box="[1072,1139,1998,2025]" pageId="57" pageNumber="58" type="male">2 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D20FD39613EDA61FBAC50FC22E9FBE3" box="[1150,1216,1998,2025]" pageId="57" pageNumber="58" type="female">2 ♀♀</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B55613EDA61FB1F50FC235EFBE3" box="[1229,1399,1998,2025]" metricMagnitude="-2" metricUnit="m" metricValue="8.985" metricValueMax="9.61" metricValueMin="8.36" pageId="57" pageNumber="58" unit="mm" value="89.85" valueMax="96.1" valueMin="83.6">83.6–96.1 mm</quantity>
|
||
SL; mouth of
|
||
<location id="8EF9606B613DDA62FEB3563B2419FD29" LSID="urn:lsid:plazi:treatment:038F87A6613EDA65FDB756882794FA9A:8EF9606B613DDA62FEB3563B2419FD29" box="[353,560,265,291]" country="Uganda" latitude="-0.209" longLatPrecision="1" longitude="29.885" name="Kazinga Channel" pageId="58" pageNumber="59" stateProvince="Lake Edward">Kazinga Channel</location>
|
||
;
|
||
<geoCoordinate id="EE125077613DDA62FDE9563524FBFD29" box="[571,722,263,291]" degrees="0" direction="south" minutes="12" orientation="latitude" pageId="58" pageNumber="59" precision="1" seconds="32.4" value="-0.209">0°12′32.4″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077613DDA62FD0D563525AEFD29" box="[735,903,263,291]" degrees="29" direction="east" minutes="53" orientation="longitude" pageId="58" pageNumber="59" precision="1" seconds="06.0" value="29.885">29°53′06.0″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070613DDA62FC46563B2206FD29" box="[916,1071,265,291]" pageId="58" pageNumber="59" value="2016-10-24">
|
||
<collectingDate id="EFDCE998613DDA62FC46563B2206FD29" box="[916,1071,265,291]" pageId="58" pageNumber="59" value="2016-10-24">24 Oct. 2016</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366613DDA62FBE9563B22C9FD29" box="[1083,1248,265,291]" pageId="58" pageNumber="59">HIPE1 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB613DDA62FAF0563B27C4FD4C" pageId="58" pageNumber="59">RMCA 2016.035.P.0204 to 0207</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED613DDA62FDC2561E2290FD63" ID-GBIF-Occurrence="3861203354" collectingDate="2017-03-26" collectionCode="RMCA" collectorName="HIPE 2 exped." country="Uganda" lastPageId="58" lastPageNumber="59" latitude="-0.40158334" location="Rwenshama" longLatPrecision="1" longitude="29.776417" pageId="58" pageNumber="58" specimenCode="RMCA 2017.006.P.0342, RMCA 2017.006.P.0343" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<specimenCount id="9D20FD39613DDA62FDC2561E246BFD4D" box="[528,578,300,327]" pageId="58" pageNumber="59" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D20FD39613DDA62FD84561E24A0FD4D" box="[598,649,300,327]" pageId="58" pageNumber="59" type="male">1 ♂</specimenCount>
|
||
, 75.9,
|
||
<quantity id="4CDE9B55613DDA62FD31561E2578FD4D" box="[739,849,300,327]" metricMagnitude="-2" metricUnit="m" metricValue="8.47" pageId="58" pageNumber="59" unit="mm" value="84.7">84.7 mm</quantity>
|
||
SL;
|
||
<location id="8EF9606B613DDA62FC46561E220AFD4C" LSID="urn:lsid:plazi:treatment:038F87A6613EDA65FDB756882794FA9A:8EF9606B613DDA62FC46561E220AFD4C" box="[916,1059,300,326]" country="Uganda" latitude="-0.40158334" longLatPrecision="1" longitude="29.776417" name="Rwenshama" pageId="58" pageNumber="59" stateProvince="Lake Edward">Rwenshama</location>
|
||
rocky shore;
|
||
<geoCoordinate id="EE125077613DDA62FB0656192358FD4C" box="[1236,1393,299,327]" degrees="0" direction="south" minutes="24" orientation="latitude" pageId="58" pageNumber="59" precision="1" seconds="05.7" value="-0.40158334">0°24′05.7″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077613DDA62FF6F567C2743FD60" box="[189,362,334,362]" degrees="29" direction="east" minutes="46" orientation="longitude" pageId="58" pageNumber="59" precision="1" seconds="35.1" value="29.776417">29°46′35.1″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070613DDA62FEA9567D240CFD63" box="[379,549,335,362]" pageId="58" pageNumber="59" value="2017-03-26">
|
||
<collectingDate id="EFDCE998613DDA62FEA9567D240CFD63" box="[379,549,335,362]" pageId="58" pageNumber="59" value="2017-03-26">26 Mar. 2017</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366613DDA62FDE4567D24F6FD63" box="[566,735,335,361]" pageId="58" pageNumber="59">HIPE2 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB613DDA62FCF8567D2290FD63" box="[810,1209,335,362]" pageId="58" pageNumber="59">RMCA 2017.006.P.0342 to 0343</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED613DDA62FB09567D27C8FDBA" ID-GBIF-Occurrence="3861203379" collectingDate="2017-03-30" collectionCode="RMCA" collectorName="HIPE 2 exped." country="Uganda" lastPageId="58" lastPageNumber="59" latitude="-0.093" location="Kayanja" longLatPrecision="1" longitude="29.758" pageId="58" pageNumber="58" specimenCode="RMCA 2017.006.P.0346, RMCA 2017.006.P.0347, RMCA 2017.006.P.0348" specimenCount="3" specimenCount-female="3" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<specimenCount id="9D20FD39613DDA62FB09567D230BFD60" box="[1243,1314,335,362]" pageId="58" pageNumber="59" type="female">3 ♀♀</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B55613DDA62FAE6567D270CFD87" metricMagnitude="-2" metricUnit="m" metricValue="6.895" metricValueMax="7.05" metricValueMin="6.74" pageId="58" pageNumber="59" unit="mm" value="68.95" valueMax="70.5" valueMin="67.4">67.4– 70.5 mm</quantity>
|
||
SL;
|
||
<location id="8EF9606B613DDA62FE8C564027E9FD86" LSID="urn:lsid:plazi:treatment:038F87A6613EDA65FDB756882794FA9A:8EF9606B613DDA62FE8C564027E9FD86" box="[350,448,370,396]" country="Uganda" latitude="-0.093" longLatPrecision="1" longitude="29.758" name="Kayanja" pageId="58" pageNumber="59" stateProvince="Lake Edward">Kayanja</location>
|
||
offshore;
|
||
<geoCoordinate id="EE125077613DDA62FDE5564324E7FD87" box="[567,718,369,397]" degrees="0" direction="south" minutes="05" orientation="latitude" pageId="58" pageNumber="59" precision="1" seconds="34.8" value="-0.093">0°05′34.8″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077613DDA62FD09564325AAFD87" box="[731,899,369,397]" degrees="29" direction="east" minutes="45" orientation="longitude" pageId="58" pageNumber="59" precision="1" seconds="28.8" value="29.758">29°45′28.8″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070613DDA62FC4256412219FD87" box="[912,1072,371,397]" pageId="58" pageNumber="59" value="2017-03-30">
|
||
<collectingDate id="EFDCE998613DDA62FC4256412219FD87" box="[912,1072,371,397]" pageId="58" pageNumber="59" value="2017-03-30">30 Mar. 2017</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366613DDA62FBEE564122C8FD86" box="[1084,1249,370,397]" pageId="58" pageNumber="59">HIPE2 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB613DDA62FAF0564127C8FDBA" pageId="58" pageNumber="59">RMCA 2017.006.P.0346 to 0348</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED613DDA62FE2B56A425CCFDDE" ID-GBIF-Occurrence="3861203384" collectingDate="2017-03-30" collectionCode="RMCA" collectorName="HIPE 2 exped." country="Uganda" lastPageId="58" lastPageNumber="59" latitude="-0.093" location="Kayanja" longLatPrecision="1" longitude="29.758" pageId="58" pageNumber="58" specimenCode="RMCA 2017.006.P.0344, RMCA 2017.006.P.0345" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<specimenCount id="9D20FD39613DDA62FE2B56A4240DFDBA" box="[505,548,406,432]" pageId="58" pageNumber="59" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D20FD39613DDA62FDE056A42449FDBA" box="[562,608,406,432]" pageId="58" pageNumber="59" type="male">1 ♂</specimenCount>
|
||
, 67.7,
|
||
<quantity id="4CDE9B55613DDA62FD7F56A4253FFDBA" box="[685,790,406,432]" metricMagnitude="-2" metricUnit="m" metricValue="7.81" pageId="58" pageNumber="59" unit="mm" value="78.1">78.1 mm</quantity>
|
||
SL;
|
||
<location id="8EF9606B613DDA62FC9F56A42586FDBA" LSID="urn:lsid:plazi:treatment:038F87A6613EDA65FDB756882794FA9A:8EF9606B613DDA62FC9F56A42586FDBA" box="[845,943,406,432]" country="Uganda" latitude="-0.093" longLatPrecision="1" longitude="29.758" name="Kayanja" pageId="58" pageNumber="59" stateProvince="Lake Edward">Kayanja</location>
|
||
offshore;
|
||
<geoCoordinate id="EE125077613DDA62FBF456A62294FDBA" box="[1062,1213,404,432]" degrees="0" direction="south" minutes="05" orientation="latitude" pageId="58" pageNumber="59" precision="1" seconds="34.8" value="-0.093">0°05′34.8″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077613DDA62FB1B56A62358FDBA" box="[1225,1393,404,432]" degrees="29" direction="east" minutes="45" orientation="longitude" pageId="58" pageNumber="59" precision="1" seconds="28.8" value="29.758">29°45′28.8″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070613DDA62FF6F568B274BFDD9" box="[189,354,441,467]" pageId="58" pageNumber="59" value="2017-03-30">
|
||
<collectingDate id="EFDCE998613DDA62FF6F568B274BFDD9" box="[189,354,441,467]" pageId="58" pageNumber="59" value="2017-03-30">30 Mar. 2017</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366613DDA62FEA2568B2431FDD9" box="[368,536,441,467]" pageId="58" pageNumber="59">HIPE2 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB613DDA62FD8F568B25CCFDDE" box="[605,997,441,468]" pageId="58" pageNumber="59">RMCA 2017.006.P.0344 to 0345</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED613DDA62FBD0568B230EFDFC" ID-GBIF-Occurrence="3861203392" collectingDate="2018-01-19" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" lastPageId="58" lastPageNumber="59" latitude="-0.16802779" location="Katwe" longLatPrecision="1" longitude="29.874277" pageId="58" pageNumber="58" specimenCode="RMCA 2018.008.P.0331" specimenCount="1" specimenCount-female="1" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<specimenCount id="9D20FD39613DDA62FBD0568B2219FDD9" box="[1026,1072,441,467]" pageId="58" pageNumber="59" type="female">1 ♀</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B55613DDA62FB93568B2285FDD9" box="[1089,1196,441,467]" metricMagnitude="-2" metricUnit="m" metricValue="9.24" pageId="58" pageNumber="59" unit="mm" value="92.4">92.4 mm</quantity>
|
||
SL; islands near
|
||
<location id="8EF9606B613DDA62FF6F56EF2724FDFD" LSID="urn:lsid:plazi:treatment:038F87A6613EDA65FDB756882794FA9A:8EF9606B613DDA62FF6F56EF2724FDFD" box="[189,269,477,503]" country="Uganda" latitude="-0.16802779" longLatPrecision="1" longitude="29.874277" name="Katwe" pageId="58" pageNumber="59" stateProvince="Lake Edward">Katwe</location>
|
||
;
|
||
<geoCoordinate id="EE125077613DDA62FEC856E9279BFDFC" box="[282,434,475,503]" degrees="0" direction="south" minutes="10" orientation="latitude" pageId="58" pageNumber="59" precision="1" seconds="04.9" value="-0.16802779">0°10′04.9″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077613DDA62FE6D56E92440FDFD" box="[447,617,475,503]" degrees="29" direction="east" minutes="52" orientation="longitude" pageId="58" pageNumber="59" precision="1" seconds="27.4" value="29.874277">29°52′27.4″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070613DDA62FDA456EE2539FDFC" box="[630,784,476,503]" pageId="58" pageNumber="59" value="2018-01-19">
|
||
<collectingDate id="EFDCE998613DDA62FDA456EE2539FDFC" box="[630,784,476,503]" pageId="58" pageNumber="59" value="2018-01-19">19 Jan. 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366613DDA62FCCF56EE25EAFDFC" box="[797,963,476,502]" pageId="58" pageNumber="59">HIPE3 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB613DDA62FBD456EE230EFDFC" box="[1030,1319,476,502]" pageId="58" pageNumber="59">RMCA 2018.008.P.0331</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED613DDA62FA9056EE24F7FE37" ID-GBIF-Occurrence="3861203380" collectingDate="2018-01-24" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" lastPageId="58" lastPageNumber="59" latitude="-0.40158334" location="Rwenshama" longLatPrecision="1" longitude="29.776417" pageId="58" pageNumber="58" specimenCode="RMCA 2018.008.P.0332, RMCA 2018.008.P.0333" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<specimenCount id="9D20FD39613DDA62FA9056EE2346FDFD" box="[1346,1391,476,503]" pageId="58" pageNumber="59" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D20FD39613DDA62FF6F56CD26C4FE10" box="[189,237,511,538]" pageId="58" pageNumber="59" type="male">1 ♂</specimenCount>
|
||
, 72.3,
|
||
<quantity id="4CDE9B55613DDA62FEED56CD2783FE10" box="[319,426,511,538]" metricMagnitude="-2" metricUnit="m" metricValue="8.17" pageId="58" pageNumber="59" unit="mm" value="81.7">81.7 mm</quantity>
|
||
SL;
|
||
<location id="8EF9606B613DDA62FE3456CD245CFE13" LSID="urn:lsid:plazi:treatment:038F87A6613EDA65FDB756882794FA9A:8EF9606B613DDA62FE3456CD245CFE13" box="[486,629,511,537]" country="Uganda" latitude="-0.40158334" longLatPrecision="1" longitude="29.776417" name="Rwenshama" pageId="58" pageNumber="59" stateProvince="Lake Edward">Rwenshama</location>
|
||
rocky shore;
|
||
<geoCoordinate id="EE125077613DDA62FCCE56CC259CFE13" box="[796,949,510,538]" degrees="0" direction="south" minutes="24" orientation="latitude" pageId="58" pageNumber="59" precision="1" seconds="05.7" value="-0.40158334">0°24′05.7″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077613DDA62FC1656CC2247FE10" box="[964,1134,510,538]" degrees="29" direction="east" minutes="46" orientation="longitude" pageId="58" pageNumber="59" precision="1" seconds="35.1" value="29.776417">29°46′35.1″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070613DDA62FBAE56CD2330FE13" box="[1148,1305,511,538]" pageId="58" pageNumber="59" value="2018-01-24">
|
||
<collectingDate id="EFDCE998613DDA62FBAE56CD2330FE13" box="[1148,1305,511,538]" pageId="58" pageNumber="59" value="2018-01-24">24 Jan. 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366613DDA62FAF556CD2722FE36" pageId="58" pageNumber="59">HIPE3 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB613DDA62FE81551124F7FE37" box="[339,734,547,573]" pageId="58" pageNumber="59">RMCA 2018.008.P.0332 to 0333</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED613DDA62FD2F551122F1FE6A" ID-GBIF-Occurrence="3861203347" collectingDate="2017-03-26" collectionCode="IRSNB" collectorName="HIPE 2 exped." country="Uganda" lastPageId="58" lastPageNumber="59" latitude="-0.40158334" location="Rwenshama" longLatPrecision="1" longitude="29.776417" pageId="58" pageNumber="58" specimenCode="IRSNB 920, IRSNB 921, IRSNB 922" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<specimenCount id="9D20FD39613DDA62FD2F5511256DFE37" box="[765,836,547,573]" pageId="58" pageNumber="59" type="female">2 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D20FD39613DDA62FC87551125B5FE37" box="[853,924,547,573]" pageId="58" pageNumber="59" type="male">2 ♂♂</specimenCount>
|
||
, 71.0–
|
||
<quantity id="4CDE9B55613DDA62FC3D55112273FE37" box="[1007,1114,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="8.49" pageId="58" pageNumber="59" unit="mm" value="84.9">84.9 mm</quantity>
|
||
SL;
|
||
<location id="8EF9606B613DDA62FB4855102300FE36" LSID="urn:lsid:plazi:treatment:038F87A6613EDA65FDB756882794FA9A:8EF9606B613DDA62FB4855102300FE36" box="[1178,1321,546,572]" country="Uganda" latitude="-0.40158334" longLatPrecision="1" longitude="29.776417" name="Rwenshama" pageId="58" pageNumber="59" stateProvince="Lake Edward">Rwenshama</location>
|
||
rocky shore;
|
||
<geoCoordinate id="EE125077613DDA62FEDE5576278AFE6A" box="[268,419,580,608]" degrees="0" direction="south" minutes="24" orientation="latitude" pageId="58" pageNumber="59" precision="1" seconds="05.7" value="-0.40158334">0°24′05.7″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE125077613DDA62FE6255762470FE6A" box="[432,601,580,608]" degrees="29" direction="east" minutes="46" orientation="longitude" pageId="58" pageNumber="59" precision="1" seconds="35.1" value="29.776417">29°46′35.1″ E</geoCoordinate>
|
||
;
|
||
<date id="FF981070613DDA62FDB75574252FFE6A" box="[613,774,582,608]" pageId="58" pageNumber="59" value="2017-03-26">
|
||
<collectingDate id="EFDCE998613DDA62FDB75574252FFE6A" box="[613,774,582,608]" pageId="58" pageNumber="59" value="2017-03-26">26 Mar. 2017</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D35366613DDA62FCC05574259EFE6A" box="[786,951,582,608]" pageId="58" pageNumber="59">HIPE2 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB613DDA62FC2A557422F1FE6A" box="[1016,1240,582,608]" pageId="58" pageNumber="59">IRSNB 920 to 922</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C653B613DDA65FF6F55B527F7F9FC" lastPageId="61" lastPageNumber="62" pageId="58" pageNumber="59" type="description">
|
||
<paragraph id="8B9936B0613DDA62FF6F55B52767FEAB" blockId="58.[189,334,647,673]" box="[189,334,647,673]" pageId="58" pageNumber="59">
|
||
<heading id="D0D181DC613DDA62FF6F55B52767FEAB" bold="true" box="[189,334,647,673]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
|
||
<emphasis id="B952EAA2613DDA62FF6F55B52767FEAB" bold="true" box="[189,334,647,673]" pageId="58" pageNumber="59">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B0613DDA62FF6F55842497F85E" blockId="58.[187,1400,694,1108]" pageId="58" pageNumber="59">
|
||
Based on
|
||
<specimenCount id="9D20FD39613DDA62FEEA558427F3FEDA" box="[312,474,694,720]" pageId="58" pageNumber="59" type="generic">20 specimens</specimenCount>
|
||
(
|
||
<quantity id="4CDE9B55613DDA62FE3C558424B7FEDA" box="[494,670,694,720]" metricMagnitude="-2" metricUnit="m" metricValue="8.175" metricValueMax="9.61" metricValueMin="6.74" pageId="58" pageNumber="59" unit="mm" value="81.75" valueMax="96.1" valueMin="67.4">67.4–96.1 mm</quantity>
|
||
SL); body shallow (
|
||
<tableCitation id="C6A4030B613DDA62FC49558425DFFEDA" box="[923,1014,694,720]" captionStart="Table 1" captionStartId="9.[182,252,548,576]" captionTargetPageId="9" captionText="Table 1. (continued on the next three pages). Measurements and counts of the piscivorous species of Haplochromis from the Lake Edward system." httpUri="http://table.plazi.org/id/DF596638610EDA51FF6455162158FE4A" pageId="58" pageNumber="59" tableUuid="DF596638610EDA51FF6455162158FE4A">Table 1</tableCitation>
|
||
) and oval to slightly rhomboid (
|
||
<figureCitation id="131D2A35613DDA62FF1455EB2708FEF9" box="[198,289,729,755]" captionStart="Fig" captionStartId="58.[189,232,1963,1989]" captionTargetBox="[190,1398,1197,1855]" captionTargetId="figure-363@58.[189,1399,1195,1855]" captionTargetPageId="58" captionText="Fig. 32. Haplochromis pardus sp. nov., holotype, ♂, 89.2 mm SL (RMCA 2016.035.P.0202). Drawn by N. Vranken." figureDoi="http://doi.org/10.5281/zenodo.6484234" httpUri="https://zenodo.org/record/6484234/files/figure.png" pageId="58" pageNumber="59">Fig. 32</figureCitation>
|
||
). Head narrow, shallow, and with a straight dorsal outline with a concavity above eye; eye average in size in comparison to generalised
|
||
<taxonomicName id="4C264D33613DDA62FD1155CF2568FF1C" authorityName="Trewavas" authorityYear="1933" box="[707,833,764,791]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="elegans">
|
||
<emphasis id="B952EAA2613DDA62FD1155CF24F1FF1C" box="[707,728,765,790]" italics="true" pageId="58" pageNumber="59">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA2613DDA62FD3555CE2568FF1C" box="[743,833,764,790]" italics="true" pageId="58" pageNumber="59">elegans</emphasis>
|
||
</taxonomicName>
|
||
(but large for a piscivorous species); interorbital area very narrow; cheek shallow; lacrimal average in depth. Snout average in length, very acute, and slopes gently at 30–40°; premaxillary pedicel long and prominent. Jaws iso- to slightly prognathous, average in length, narrow, and rounded in dorsal view; gape large and slopes gently at 15–25°; maxilla extends to between verticals through anterior margins of orbit and pupil. Lower jaw shallow and with straight ventral outline in lateral view, mental prominence absent, and lower jaw side nearly flat with an inclination of 15–30° to horizontal in anterior view. Upper jaw weakly expanded anteriorly and ventrally. Lips and oral mucosa large. Neurocranium average in depth, ethmo-vomerine block decurved, preorbital region very shallow (18–22% NL), orbital region average in depth (30–33% NL), and supraoccipital crest shallow and wedge-shaped (
|
||
<figureCitation id="131D2A35613DDA62FD98530B249BF85E" box="[586,690,1081,1108]" captionStart="Fig" captionStartId="59.[189,232,1892,1918]" captionTargetBox="[242,1338,284,1857]" captionTargetId="figure-12@59.[242,1338,265,1857]" captionTargetPageId="59" captionText="Fig. 33. Haplochromis pardus sp. nov. a. Photograph of preserved holotype (RMCA 2016.035.P.0202; 89.2 mm SL). b. X-ray image of holotype. c–d. Photographs of freshly caught specimens. c. Dominant male (RMCA 2017.006.P (HP1463); 81.4 mm SL). d. Female (RMCA 2017.006.P.0342; 75.9 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484236" httpUri="https://zenodo.org/record/6484236/files/figure.png" pageId="58" pageNumber="59">Fig. 33b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF596638613DDA62FF6F50992762FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484234" ID-Zenodo-Dep="6484234" httpUri="https://zenodo.org/record/6484234/files/figure.png" pageId="58" pageNumber="59" startId="58.[189,232,1963,1989]" targetBox="[190,1398,1197,1855]" targetPageId="58">
|
||
<paragraph id="8B9936B0613DDA62FF6F50992762FBE2" blockId="58.[189,1398,1963,2025]" pageId="58" pageNumber="59">
|
||
<emphasis id="B952EAA2613DDA62FF6F50992733FBCF" bold="true" box="[189,282,1963,1989]" pageId="58" pageNumber="59">Fig. 32.</emphasis>
|
||
<taxonomicName id="4C264D33613DDA62FEF350992408FBCF" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[289,545,1963,1989]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="pardus" status="sp. nov.">
|
||
<emphasis id="B952EAA2613DDA62FEF350992408FBCF" box="[289,545,1963,1989]" italics="true" pageId="58" pageNumber="59">Haplochromis pardus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9613DDA62FDFA509E24ACFBCC" box="[552,645,1964,1990]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂, 89.2 mm SL (RMCA 2016.035.P.0202). Drawn by N. Vranken.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF596638613CDA63FF6F50562223FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484236" ID-Zenodo-Dep="6484236" httpUri="https://zenodo.org/record/6484236/files/figure.png" pageId="59" pageNumber="60" startId="59.[189,232,1892,1918]" targetBox="[242,1338,284,1857]" targetPageId="59">
|
||
<paragraph id="8B9936B0613CDA63FF6F50562223FBE2" blockId="59.[189,1398,1892,2025]" pageId="59" pageNumber="60">
|
||
<emphasis id="B952EAA2613CDA63FF6F50562732FB75" bold="true" box="[189,283,1892,1919]" pageId="59" pageNumber="60">Fig. 33.</emphasis>
|
||
<taxonomicName id="4C264D33613CDA63FEF05057240DFB75" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[290,548,1893,1919]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="59" pageNumber="60" phylum="Chordata" rank="species" species="pardus" status="sp. nov.">
|
||
<emphasis id="B952EAA2613CDA63FEF05057240DFB75" box="[290,548,1893,1919]" italics="true" pageId="59" pageNumber="60">Haplochromis pardus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9613CDA63FDFF505724A1FB75" box="[557,648,1893,1919]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B952EAA2613CDA63FD5C505724B4FB75" bold="true" box="[654,669,1893,1919]" pageId="59" pageNumber="60">a</emphasis>
|
||
. Photograph of preserved holotype (RMCA 2016.035.P.0202; 89.2 mm SL).
|
||
<emphasis id="B952EAA2613CDA63FEBA50BA2751FBA8" bold="true" box="[360,376,1928,1954]" pageId="59" pageNumber="60">b</emphasis>
|
||
. X-ray image of holotype.
|
||
<emphasis id="B952EAA2613CDA63FD6A50BA24CDFBA8" bold="true" box="[696,740,1928,1954]" pageId="59" pageNumber="60">c–d</emphasis>
|
||
. Photographs of freshly caught specimens.
|
||
<emphasis id="B952EAA2613CDA63FB3750BA22DBFBA8" bold="true" box="[1253,1266,1928,1954]" pageId="59" pageNumber="60">c</emphasis>
|
||
. Dominant male (RMCA 2017.006.P (HP1463); 81.4 mm SL).
|
||
<emphasis id="B952EAA2613CDA63FCF350992518FBCF" bold="true" box="[801,817,1963,1989]" pageId="59" pageNumber="60">d</emphasis>
|
||
. Female (RMCA 2017.006.P.0342; 75.9 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9936B0613BDA64FF6F563B25F1FE89" blockId="60.[188,1400,265,643]" pageId="60" pageNumber="61">
|
||
Outer oral teeth numerous, small, and weakly embedded in oral mucosa. Necks stout, cylindrical, and straight; crowns weakly recurved, unicuspid in large specimens (<
|
||
<quantity id="4CDE9B55613BDA64FC32561E2210FD4D" box="[992,1081,300,327]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="60" pageNumber="61" unit="mm" value="80.0">80 mm</quantity>
|
||
), bicuspid with posteriorly some weakly tricuspid teeth in small specimens (>
|
||
<quantity id="4CDE9B55613BDA64FCDD567D254DFD60" box="[783,868,335,362]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="60" pageNumber="61" unit="mm" value="70.0">70 mm</quantity>
|
||
), and acutely pointed in all specimens. Dental arcades rounded and with anterior half weakly expanded laterally. Outer teeth closely and regularly set with neck-distances of ½ neck-width; lateral outer teeth implanted slightly labially. In upper jaw, 1–3 posteriormost teeth enlarged. Inner teeth small, recurved, unicuspid in large specimens (>
|
||
<quantity id="4CDE9B55613BDA64FADD568B2343FDD9" box="[1295,1386,441,467]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="60" pageNumber="61" unit="mm" value="80.0">80 mm</quantity>
|
||
), tricuspid in small specimens (<
|
||
<quantity id="4CDE9B55613BDA64FDE356EE24AEFDFD" box="[561,647,476,503]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="60" pageNumber="61" unit="mm" value="70.0">70 mm</quantity>
|
||
), and acutely pointed in all specimens. Tooth bands very slender crescent-shaped with 2 (rarely 3) rows of inner teeth, and narrow posteriorly until only outer row remains past ⅔ length of tooth band. Inner rows closely and regularly set on 1 neck-width from outer row in lower jaw, on 1–2 neck-widths from outer row in upper jaw; implantation erect in first row and recumbent in subsequent rows; size uniform throughout tooth band.
|
||
</paragraph>
|
||
<paragraph id="8B9936B0613BDA64FF6F55992731FF7F" blockId="60.[187,1399,682,885]" pageId="60" pageNumber="61">
|
||
Lower pharyngeal bone average in length, narrow, slim, and shallow over entire length (
|
||
<figureCitation id="131D2A35613BDA64FADE55982340FECF" box="[1292,1385,682,709]" captionStart="Fig" captionStartId="60.[189,232,1963,1989]" captionTargetBox="[189,1394,1035,1887]" captionTargetId="figure-283@60.[189,1399,1035,1889]" captionTargetPageId="60" captionText="Fig. 34. Haplochromis pardus sp. nov. (RMCA 2016.035.P.0207; 96.1 mm SL). a. Dorsal view of the lower pharyngeal jaw. b. Lateral view of the lower pharyngeal jaw." figureDoi="http://doi.org/10.5281/zenodo.6484238" httpUri="https://zenodo.org/record/6484238/files/figure.png" pageId="60" pageNumber="61">Fig. 34</figureCitation>
|
||
). Pharyngeal teeth small and slender; major cusps acutely pointed; cusp gaps nearly straight; minor cusps and cusp protuberances small. Teeth in two median longitudinal rows equal in size and form to lateral teeth,
|
||
<quantity id="4CDE9B55613BDA64FED454262747FF24" box="[262,366,788,814]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" metricValueMax="3.048" metricValueMin="2.794" pageId="60" pageNumber="61" unit="in" value="11.5" valueMax="12.0" valueMin="11.0">11–12 in</quantity>
|
||
each row. Posterior transverse row with 22 teeth, implanted erectly with a slight lateral inclination; major cusps weakly recurved, bluntly pointed, and laterally compressed; minor cusps mostly present.
|
||
</paragraph>
|
||
<paragraph id="8B9936B0613BDA65FF6F54AE25D4FD63" blockId="60.[189,1399,924,950]" lastBlockId="61.[189,1399,265,362]" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">Chest scales small; transition to larger flank scales gradual. Minute scales on proximal half of caudal fin. Caudal fin emarginate; dorsal and anal fins reach to between verticals through one scale anterior and one scale posterior to caudal-fin base. Pectoral and pelvic fins reach to genital opening; pelvic fin reaches to first anal-fin spine in males; first branched pelvic-fin ray not elongated.</paragraph>
|
||
<caption id="DF596638613BDA64FF6F509925F8FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484238" ID-Zenodo-Dep="6484238" httpUri="https://zenodo.org/record/6484238/files/figure.png" pageId="60" pageNumber="61" startId="60.[189,232,1963,1989]" targetBox="[189,1394,1035,1887]" targetPageId="60">
|
||
<paragraph id="8B9936B0613BDA64FF6F509925F8FBE2" blockId="60.[189,1399,1963,2025]" pageId="60" pageNumber="61">
|
||
<emphasis id="B952EAA2613BDA64FF6F50992735FBCF" bold="true" box="[189,284,1963,1989]" pageId="60" pageNumber="61">Fig. 34.</emphasis>
|
||
<taxonomicName id="4C264D33613BDA64FEF65099240FFBCF" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[292,550,1963,1989]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="pardus" status="sp. nov.">
|
||
<emphasis id="B952EAA2613BDA64FEF65099240FFBCF" box="[292,550,1963,1989]" italics="true" pageId="60" pageNumber="61">Haplochromis pardus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9613BDA64FDE2509E24A2FBCC" box="[560,651,1964,1990]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(RMCA 2016.035.P.0207; 96.1 mm SL).
|
||
<emphasis id="B952EAA2613BDA64FBAE509922A3FBCF" bold="true" box="[1148,1162,1963,1989]" pageId="60" pageNumber="61">a</emphasis>
|
||
. Dorsal view of the lower pharyngeal jaw.
|
||
<emphasis id="B952EAA2613BDA64FE1A50FC27F1FBE2" bold="true" box="[456,472,1998,2024]" pageId="60" pageNumber="61">b</emphasis>
|
||
. Lateral view of the lower pharyngeal jaw.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9936B0613ADA65FF6F56A32200FDC4" blockId="61.[189,1399,401,462]" pageId="61" pageNumber="62">Ceratobranchial gill rakers in outer row of first gill arch short, stout, and simple; posteriormost rakers mostly weakly bifid. Epibranchial gill rakers relatively slender and simple.</paragraph>
|
||
<paragraph id="8B9936B0613ADA65FF6F56C7278AFE05" blockId="61.[189,419,501,527]" box="[189,419,501,527]" pageId="61" pageNumber="62">
|
||
<heading id="D0D181DC613ADA65FF6F56C7278AFE05" bold="true" box="[189,419,501,527]" fontSize="11" level="3" pageId="61" pageNumber="62" reason="3">
|
||
<emphasis id="B952EAA2613ADA65FF6F56C7278AFE05" bold="true" box="[189,419,501,527]" pageId="61" pageNumber="62">Colouration in life</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B0613ADA65FF6F551626D7FF71" blockId="61.[188,1399,548,891]" pageId="61" pageNumber="62">
|
||
Dominant males: body speckled to uniformly black with a blue sheen; dorsum and dorsal part of head pink to speckled black; chest white; belly, caudal peduncle, operculum, lower jaw, and lips black; cheek, lacrimal, and snout speckled to uniformly black, cheek with yellow sheen; eye with grey to dark outer ring and silver to dark inner ring (
|
||
<figureCitation id="131D2A35613ADA65FD8E55BF24EDFEA2" box="[604,708,653,680]" captionStart="Fig" captionStartId="59.[189,232,1892,1918]" captionTargetBox="[242,1338,284,1857]" captionTargetId="figure-12@59.[242,1338,265,1857]" captionTargetPageId="59" captionText="Fig. 33. Haplochromis pardus sp. nov. a. Photograph of preserved holotype (RMCA 2016.035.P.0202; 89.2 mm SL). b. X-ray image of holotype. c–d. Photographs of freshly caught specimens. c. Dominant male (RMCA 2017.006.P (HP1463); 81.4 mm SL). d. Female (RMCA 2017.006.P.0342; 75.9 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484236" httpUri="https://zenodo.org/record/6484236/files/figure.png" pageId="61" pageNumber="62">Fig. 33c</figureCitation>
|
||
). Flank with dorsal-lateral and mid-lateral bands and 5–6 vertical stripes, all black, broad, well-defined, but interrupted, hereby body seemingly blotched. Nostril, interorbital, supraorbital, lacrimal, vertical preopercular stripes, nape band, mental blotch, and black posterior margin of operculum well-defined. Pectoral, dorsal, and anal fins dusky; lappets, bases, and posterior parts of anal and dorsal fins black; anal fin with faint crimson flush and 1–3 small orange eggspots with dusky rings. Pelvic and caudal fins black; caudal fin with dusky distal part and a faint crimson flush.
|
||
</paragraph>
|
||
<paragraph id="8B9936B0613ADA65FF6F549022E8F89A" blockId="61.[188,1400,930,1168]" pageId="61" pageNumber="62">
|
||
Females and juveniles: body yellowish with a dusky sheen; dorsum and dorsal part of head pink; belly, chest, operculum, and cheek, white; lacrimal speckled black; snout dusky to black; eye with grey to dark outer ring and silver to dark inner ring (
|
||
<figureCitation id="131D2A35613ADA65FD4A54DB24D6F80E" box="[664,767,1001,1028]" captionStart="Fig" captionStartId="59.[189,232,1892,1918]" captionTargetBox="[242,1338,284,1857]" captionTargetId="figure-12@59.[242,1338,265,1857]" captionTargetPageId="59" captionText="Fig. 33. Haplochromis pardus sp. nov. a. Photograph of preserved holotype (RMCA 2016.035.P.0202; 89.2 mm SL). b. X-ray image of holotype. c–d. Photographs of freshly caught specimens. c. Dominant male (RMCA 2017.006.P (HP1463); 81.4 mm SL). d. Female (RMCA 2017.006.P.0342; 75.9 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484236" httpUri="https://zenodo.org/record/6484236/files/figure.png" pageId="61" pageNumber="62">Fig. 33d</figureCitation>
|
||
). Flank with dorsal-lateral and mid-lateral bands and 5–6 vertical stripes, all faint and interrupted. Nostril and interorbital stripes faint; supraorbital, lacrimal, vertical preopercular stripes, mental blotch, and nape band well-defined. Pectoral, pelvic, and dorsal fins dusky; anal and caudal fins dusky with yellow sheen; dorsal and anal fins with black lappets and posterodistal part; caudal fin with black base and 1–2 small spots resembling egg-spots.
|
||
</paragraph>
|
||
<paragraph id="8B9936B0613ADA65FF6F538527F9F8DB" blockId="61.[189,464,1207,1233]" box="[189,464,1207,1233]" pageId="61" pageNumber="62">
|
||
<heading id="D0D181DC613ADA65FF6F538527F9F8DB" bold="true" box="[189,464,1207,1233]" fontSize="11" level="3" pageId="61" pageNumber="62" reason="3">
|
||
<emphasis id="B952EAA2613ADA65FF6F538527F9F8DB" bold="true" box="[189,464,1207,1233]" pageId="61" pageNumber="62">Preserved colouration</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B0613ADA65FF6F53D427F7F9FC" blockId="61.[187,1400,1253,1526]" pageId="61" pageNumber="62">
|
||
In all specimens, dorsal part of body dark brown to speckled black; ventral part of body dark brown to uniformly black; belly black; chest whitish (
|
||
<figureCitation id="131D2A35613ADA65FD26523A2575F929" box="[756,860,1288,1315]" captionStart="Fig" captionStartId="59.[189,232,1892,1918]" captionTargetBox="[242,1338,284,1857]" captionTargetId="figure-12@59.[242,1338,265,1857]" captionTargetPageId="59" captionText="Fig. 33. Haplochromis pardus sp. nov. a. Photograph of preserved holotype (RMCA 2016.035.P.0202; 89.2 mm SL). b. X-ray image of holotype. c–d. Photographs of freshly caught specimens. c. Dominant male (RMCA 2017.006.P (HP1463); 81.4 mm SL). d. Female (RMCA 2017.006.P.0342; 75.9 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484236" httpUri="https://zenodo.org/record/6484236/files/figure.png" pageId="61" pageNumber="62">Fig. 33a</figureCitation>
|
||
). Flank with faint to well-defined, broad, but interrupted mid-lateral, dorsal-lateral, and dorsal-medial bands and 5–6 vertical stripes. Snout dusky; lips dusky with well-defined black spots; cheek yellowish to black; operculum black. Nostril, interorbital, supraorbital, lacrimal, vertical preopercular stripes, nape band, mental blotch, and black posterior margin of operculum well-defined. Pectoral, dorsal, and anal fins dusky; lappets, bases, and posterior parts of anal and dorsal fins black; anal fin with 1–3 egg-spots. Pelvic and caudal fins dusky to black; caudal fin with blackish distal part.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C653B613ADA65FF6F512F2794FA9A" pageId="61" pageNumber="62" type="distribution">
|
||
<paragraph id="8B9936B0613ADA65FF6F512F27D8FA3D" blockId="61.[189,497,1565,1591]" box="[189,497,1565,1591]" pageId="61" pageNumber="62">
|
||
<heading id="D0D181DC613ADA65FF6F512F27D8FA3D" bold="true" box="[189,497,1565,1591]" fontSize="11" level="3" pageId="61" pageNumber="62" reason="3">
|
||
<emphasis id="B952EAA2613ADA65FF6F512F27D8FA3D" bold="true" box="[189,497,1565,1591]" pageId="61" pageNumber="62">Distribution and ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B0613ADA65FF6F517D2794FA9A" blockId="61.[189,1398,1614,1680]" pageId="61" pageNumber="62">Only known from Lake Edward; found in inshore areas. Based on its morphology, most probably a piscivorous species.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |