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<mods:title id="8DF0C1F29C24819CFBE6E33E94DE0863">Redescription and new records of Ulomimus indicus Bates, 1873 (Coleoptera, Tenebrionidae, Tenebrioninae)</mods:title>
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<mods:namePart id="6221BD003FD2CACF8A39909FF6714647">Liu, Shanshan</mods:namePart>
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<mods:namePart id="154902755FF428D2BD4885B29F246A95">Ren, Guodong</mods:namePart>
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<mods:namePart id="AACAFFB2876D31243E80DEA5A8D56A72">Merkl, Otto</mods:namePart>
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<treatment id="BD65747AD4156E112B3CC013DB814808" ID-GBIF-Taxon="152049876" LSID="urn:lsid:plazi:treatment:BD65747AD4156E112B3CC013DB814808" httpUri="http://treatment.plazi.org/id/BD65747AD4156E112B3CC013DB814808" lastPageId="5" lastPageNumber="50" pageId="2" pageNumber="47" scope_family="Tenebrionidae" scope_order="Coleoptera">
<subSubSection id="BA826CA67D7F46863BEEF0593D5CC0C8" pageId="2" pageNumber="47" type="nomenclature">
<paragraph id="F82F4C7E3DE0F9215C2D2B6B44CA9020" pageId="2" pageNumber="47">
<taxonomicName id="A1C59ADB0A33D425C12AF72F4A353742" LSID="http://species-id.net/wiki/Ulomimus_indicus" authority="Bates, 1873" authorityName="Bates" authorityYear="1873" genus="Ulomimus" lsidName="Ulomimus indicus" pageId="2" pageNumber="47" rank="species" species="indicus">Ulomimus indicus Bates, 1873</taxonomicName>
Figs 1-13
</paragraph>
</subSubSection>
<subSubSection id="77F3842F80B472542E56A7C3C636C28A" pageId="2" pageNumber="47" type="reference_group">
<paragraph id="EE9BF96B0D52504FC12C9471A57FA4B9" pageId="2" pageNumber="47">
<taxonomicName id="311B82E6945D7B1FC325928B52A8EFEB" genus="Ulomimimus" lsidName="Ulomimimus indicus" pageId="2" pageNumber="47" rank="species" species="indicus">Ulomimimus indicus</taxonomicName>
Bates, 1873: 202.
</paragraph>
<paragraph id="EFED3423152FB16661BC278FDC9E8067" pageId="2" pageNumber="47">
<taxonomicName id="3547E8483A595433A454E71793C8E1F1" genus="Ulomimus" lsidName="Ulomimus indicus" pageId="2" pageNumber="47" rank="species" species="indicus">Ulomimus indicus</taxonomicName>
:
<bibRefCitation id="D03A9F20B0BF1D8559917DA38CF486B3" author="Gebien, H" journalOrPublisher="In: Schenkling S (Ed) Coleopterorum Catalogus Volumen XVIII. H. Gebien, Tenebrionidae, Trictenotomidae. W. Junk, Berlin" pageId="6" pageNumber="51" title="Pars 28. Tenebrionidae III. Pp. 355 - 585." year="1911">Gebien 1911</bibRefCitation>
: 399;
<bibRefCitation id="119064F8B7A1BADE58EF4CC007C3D0BF" author="Gebien, H" journalOrPublisher="Mitteilungen der Muenchener Entomologischen Gesellschaft" pageId="6" pageNumber="51" pagination="755 - 786" title="Katalog der Tenebrioniden, Teil II. [Part.]" volume="30" year="1940">Gebien 1940</bibRefCitation>
: 770 [577];
<bibRefCitation id="AF5422460EA178EF8D87957E7D81BB21" author="Kaszab, Z" journalOrPublisher="Folia entomologica hungarica" pageId="6" pageNumber="51" pagination="43 - 128" title="Faunistik der Tenebrioniden von Sri Lanka." volume="32" year="1979">Kaszab 1979</bibRefCitation>
: 88;
<bibRefCitation id="E415F030A6B66D577F076E375D752F89" author="Kaszab, Z" journalOrPublisher="Annales historico-naturales Musei Nationalis hungarici" pageId="6" pageNumber="51" pagination="169 - 221" title="Angaben zur Kenntnis der Tenebrioniden Nordvietnams (Coleoptera)." url="http://publication.nhmus.hu/pdf/annHNHM/Annals_HNHM_1980_Vol_72_169.pdf" volume="72" year="1980">Kaszab 1980</bibRefCitation>
: 175;
<bibRefCitation id="1ED9CEDBBB15CB714DE300B3BBA65584" author="Merkl, O" journalOrPublisher="Acta Zoologica Academiae Scientiarum Hungaricae" pageId="6" pageNumber="51" pagination="261 - 280" title="Tenebrionidae (Coleoptera) from Laos and Vietnam, with reclassification of Old World &quot; Doliema &quot;." url="https://www.researchgate.net/publication/257239115_Tenebrionidae_%28Coleoptera%29_from_Laos_and_Vietnam_with_reclassification_of_Old_World_Doliema?ev=prf_pub" volume="38" year="1992">Merkl 1992</bibRefCitation>
: 263.
</paragraph>
<paragraph id="D8C7DA99370E513D57B6CD98770F684F" pageId="2" pageNumber="47">
<taxonomicName id="65815CE28DE1F959F8D9F2329C4FE038" genus="Pseuduloma" lsidName="Pseuduloma cribricollis" pageId="2" pageNumber="47" rank="species" species="cribricollis">Pseuduloma cribricollis</taxonomicName>
Fairmaire, 1893: 27;
<bibRefCitation id="D3345628697E5F0E202DAD8B09C4FC98" author="Gebien, H" journalOrPublisher="In: Schenkling S (Ed) Coleopterorum Catalogus Volumen XVIII. H. Gebien, Tenebrionidae, Trictenotomidae. W. Junk, Berlin" pageId="6" pageNumber="51" title="Pars 28. Tenebrionidae III. Pp. 355 - 585." year="1911">Gebien 1911</bibRefCitation>
: 404.
</paragraph>
</subSubSection>
<subSubSection id="2F615D78D5428173AA19A1DF9A21FB5D" pageId="2" pageNumber="47" type="original description">
<paragraph id="C54EF2F6C9A4D2A1B7BD8F5B5A748B2B" pageId="2" pageNumber="47">Original description.</paragraph>
<paragraph id="2B39A2283DD3C0007758853290A04510" pageId="2" pageNumber="47">
&quot;Oblong,
<normalizedToken id="80A6E4B61D4FF0CA5CBA6F70404F20B1" originalValue="sub–parallel">sub-parallel</normalizedToken>
, moderately convex;
<normalizedToken id="D9158E28CF5891C5F18946C245081F25" originalValue="brownish–black">brownish-black</normalizedToken>
, shining, the mentum, antenna, palpi, tarsi, labrum, and margins of epistoma ferruginous, the legs
<normalizedToken id="282D79E15F81C5CD8F8B96300439FB57" originalValue="chestnut–red">chestnut-red</normalizedToken>
; head coarsely and closely
<normalizedToken id="8E1775EF2F504C11C6424C288465E14F" originalValue="reticulate–punctate">reticulate-punctate</normalizedToken>
; prothorax
<normalizedToken id="EDB718132A06F63D657D8E94D4320EAB" originalValue="punctured–sparsely">punctured-sparsely</normalizedToken>
on the
<normalizedToken id="1422CAE4458FF88B5A1260C01A3C3E1D" originalValue="disc–the">disc-the</normalizedToken>
punctures large, deep, rounded, and partly filled in with an apparent exudation of an ashy tint; scutellum smooth; elytra with nine (including the extreme marginal one) fine but deep striae, and a short scutellar one, the striae punctured (the punctures being much wider than the striae, the elytra appear
<normalizedToken id="F7D49BBE626FB3F964C8D0956E4C3B86" originalValue="crenulate–striate">crenulate-striate</normalizedToken>
), the 4th and 5th striae shortest and united at some distance from the apex; intervals convex posteriorly, very minutely and sparsely punctured;
<normalizedToken id="6493A6862F92A783A0732827C114EDA3" originalValue="pro–">pro-</normalizedToken>
and mesosterna, flanks of
<normalizedToken id="A9043C43F72175800C4AE097574D624A" originalValue="pro–">pro-</normalizedToken>
and mesothorax, and base of epipleural fold, strongly and closely punctured; metasternum, abdomen, and femora sparingly punctured, abdominal joints rugulose at the base. Long. corp. 4 lin.
</paragraph>
<paragraph id="3335911A245020A212E2C60E97513ACA" pageId="2" pageNumber="47">Hab.: East India; one example.&quot;</paragraph>
</subSubSection>
<subSubSection id="14BE594DF77436B4B9DADCC9D459E6A6" lastPageId="5" lastPageNumber="50" pageId="2" pageNumber="47" type="description">
<paragraph id="64054785BAF03109FB4A9E834E8E09EB" pageId="2" pageNumber="47">Redescription.</paragraph>
<paragraph id="ABE55F63973A59AEE4FB1708A80F46E4" lastPageId="3" lastPageNumber="48" pageId="2" pageNumber="47">
Male. Body length 8.0-9.0 mm; width 3.5-4.0 mm. Body (Fig. 1) elongate, elliptical, black or dark brown. Antennae, mouthparts and legs slightly
<pageBreakToken id="BF9CBFD973F75E999CE19C8982088372" pageId="3" pageNumber="48" start="start">paler</pageBreakToken>
. Head transverse, with small punctures on anterior half, and with sparse large punctures on posterior half; labrum transversely rectangular, densely punctate, scattered with short and yellow hairs; clypeus densely punctate, anterior margin truncate; frontoclypeal suture deeply impressed; genae feebly convex and slightly extended, temples reduced; eyes transverse, with 5-6 facets at narrowest point in lateral view; frons weakly convex, with large punctures; mentum (Fig. 4) cordate, truncate basally, with short and very dense yellow pubescence; ligula deeply emarginate anteriorly, depressed in middle; maxillary palp with narrowly trapezoidal terminal palpomere. Antennae (Fig. 3) short, not reaching half of pronotum; antennomere 1 thick, 2 very short, 3 long and narrow, 4 and 5 short, 6 to 10 gradually widening, 11 nearly globose, ratio of the length (width) of antennomeres 2-11 as follows: 4 (6): 8 (6): 5 (7): 5 (7): 5 (8): 4 (9): 5 (10): 5 (12): 5 (11): 11 (11). Antennomere 6 with one placoid sensorium on inner anterior corner, 7 with two placoid sensoria on inner corner, 8 to 9 with a few on inner and outer corners.
</paragraph>
<caption id="1929456A3652616F69BA87B08EE2613C" pageId="3" pageNumber="48">
<paragraph id="D96687468849BB555FDE4CFCD7EE586C" pageId="3" pageNumber="48">
Figures 1-2.
<taxonomicName id="3ED78C384DF1A1DA2745F47CFB7B4204" genus="Ulomimus" lsidName="Ulomimus indicus" pageId="3" pageNumber="48" rank="species" species="indicus">Ulomimus indicus</taxonomicName>
Bates, 1873. 1 Male 2 female.
</paragraph>
</caption>
<caption id="A827D4622DA719FA8FC62CDA686C5EC7" pageId="3" pageNumber="48">
<paragraph id="775C94D8D03CE629E87F76B8DC3527B2" pageId="3" pageNumber="48">
Figures 3-13.
<taxonomicName id="6AC34203BE135C7E4F7C2563A856750E" genus="Ulomimus" lsidName="Ulomimus indicus" pageId="3" pageNumber="48" rank="species" species="indicus">Ulomimus indicus</taxonomicName>
Bates, 1873. 3 Antenna, male, dorsal view 4 mentum, male, ventral view 5 pronotum, male, dorsal view 6 protibia, male, dorsal view 7 protibia, male, lateral view 8 metatarsus, male, dorsal view 9 apical aedeagus, dorsal view 10 apical aedeagus, ventral view 11 aedeagus, lateral view 12 ovipositor, dorsal view 13 ovipositor, ventral view.
</paragraph>
</caption>
<paragraph id="4088C5E3FE3151E96E8B31DB8446C914" lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48">
Pronotum (Fig. 5) transverse, about 1.35 times as wide as long, widest at anterior 1/3, with large punctures widely spaced in middle but becoming denser toward sides; anterior margin emarginate with narrow bead interrupted in middle, and with dense short hair fringes; lateral margins arcuate, strongly narrowing forward and less so from widest point to base, with narrow bead; basal margin slightly convex, with narrow bead; anterior angles nearly rectangular, posterior angles obtuse. Prosternum with sparse and large punctures, prosternal process rounded in lateral view, with small
<pageBreakToken id="BA0BFF1BDC4287855A491D2EF0D1CB23" pageId="4" pageNumber="49" start="start">subapical</pageBreakToken>
tubercle. Mesoventrite with deep triangular impression; mesepisternum, metepimeron and metepisternum coarsely and sparsely punctate.
</paragraph>
<paragraph id="53DE65D97A9246F5885B5F2471EC408D" pageId="4" pageNumber="49">
Scutellum triangular, impunctate. Elytra distinctly
<normalizedToken id="780A34682C3464599B1F90BA437F7C60" originalValue="punctato–striate">punctato-striate</normalizedToken>
, intervals weakly convex, very finely and sparsely punctate, lateral margins visible only at humeri in dorsal view. Epipleura sparsely and coarsely punctate in basal 1/3.
</paragraph>
<paragraph id="4784F38E14FD1ADE11E61E4523FD0FF8" pageId="4" pageNumber="49">Protibia (Figs 6-7) with two equal apical spurs; narrow at base, then explanate on both inner and outer edges, outer edge forming blunt subapical angulation, inner edge slightly concave at middle; outer edge without denticulation; inner edge fringed with yellow hairs becoming denser and longer toward apex; dorsal surface with low and blunt longitudinal keel and with fine and sparse punctures; ventral surface with sharp longitudinal keel and sharp tooth at middle (Fig. 7), ventral surface with a few coarse punctures and short, sparse, thick hairs. Protarsomeres 2 to 4 dilated, with long, sparse yellow dorsal hairs and dense, yellow ventral hair pads. Mesotibia and metatibia gradually dilated toward apex, outer edge with small denticles and sparse, long hairs. Length ratio of metatarsomeres 1 to 4 (Fig. 8) as follows: 10: 3: 3: 7.</paragraph>
<paragraph id="ABD8AA7B19CDD06EB7312E3071941C4E" pageId="5" pageNumber="50">
<pageBreakToken id="C9335A117D932EFA59859142EE9E0449" pageId="5" pageNumber="50" start="start">Abdominal</pageBreakToken>
ventrites finely and sparsely punctate, punctuation denser and subcontiguous toward lateral portions; last ventrite with deep apical groove.
</paragraph>
<paragraph id="F6397D27EEEB12B4E032FA989E06138D" pageId="5" pageNumber="50">
Aedeagus (Figs 9-11) with basale
<normalizedToken id="4FA35AD1BB6EEE011E9F68641DBE5576" originalValue="parallel–sided">parallel-sided</normalizedToken>
; apicale broad at base, constricted at middle, widening and truncate at apex in dorsal view, with longitudinal depression in ventral view, slightly curved in lateral view; with small oblique notch at posterior corners of widened apical part.
</paragraph>
<paragraph id="92674F02CF3ED76ABDDC0D6326CBF256" pageId="5" pageNumber="50">Female (Fig. 2). Mentum cordate, without dense pad of pubescence, but with sparse hairs and coarse wrinkles. Protibia with shape similar to or narrower than that of male, ventral surface concave, without keel and large tooth. Protarsomeres 2 to 4 not dilated and without ventral hair pads. Ovipositor (Figs 12-13) with coxites relatively smooth, bearing long sensorial hairs and a few short hairs at base.</paragraph>
</subSubSection>
<subSubSection id="F34F7431157340668AEA5689BD8EE0C2" pageId="5" pageNumber="50" type="examined materials">
<paragraph id="5983D43F7F914A566A313DA7C7908A9F" pageId="5" pageNumber="50">Examined materials.</paragraph>
<paragraph id="42418139B85B94542CD76A5C7C26C0E1" pageId="5" pageNumber="50">
1♀ (MHBU): China, Guangxi,
<normalizedToken id="96397035CC1710C68E80D904C54259BF" originalValue="Tian’e">Tian'e</normalizedToken>
County, 14 September 2002, M. Bai leg; 1♂, 1♀ (HNHM), 2♂♂, 2♀♀ (MHBU): China, Hainan, Baisha County, Nankai Town, Shenbo Village, 1 June 2007, Y. B. Ba leg; 1♀ (NMNS): Thailand, Chiang Dao Hill Resort, 10-11 November 2012, K. Masumoto &amp; K. Takahashi leg; 1♀ (HNHM): Indonesia, Sumatra, Dolok Merangir, 25 June 1970, collector unknown; 1♂ (HNHM): Indonesia, Sumatra, Palembang, date unknown, W. Knappert leg.
</paragraph>
</subSubSection>
<subSubSection id="AAC5CA0455DD4CF76A5F5AEB099EA59A" pageId="5" pageNumber="50" type="distribution">
<paragraph id="1A59D48DD5EDD07E01F62935029F1793" pageId="5" pageNumber="50">Distribution.</paragraph>
<paragraph id="88155CC5FD9F6512B9E6BAFF18F46A71" pageId="5" pageNumber="50">
&quot;East India&quot; (
<bibRefCitation id="714F757B40B2C20FF24ADA976ACFB872" author="Bates, F" journalOrPublisher="The Entomologist's monthly magazine" pageId="5" pageNumber="50" pagination="201 - 204" title="Notes on Heteromera, and descriptions of new genera and species (No. 5)." volume="9" year="1873">Bates 1873</bibRefCitation>
); Sri Lanka (
<bibRefCitation id="B5CA680315E94F729FEFF4C1E7865ABA" author="Kaszab, Z" journalOrPublisher="Folia entomologica hungarica" pageId="6" pageNumber="51" pagination="43 - 128" title="Faunistik der Tenebrioniden von Sri Lanka." volume="32" year="1979">Kaszab 1979</bibRefCitation>
); Vietnam (
<bibRefCitation id="0E8F0FB15D5F4A510468790390B7553D" author="Fairmaire, L" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="6" pageNumber="51" pagination="19 - 38" title="Contributions a la faune indo-chinoise. 11 e Memoire." volume="62" year="1893">Fairmaire 1893</bibRefCitation>
,
<bibRefCitation id="347A4E2D3DCB3B64D85A193B9CEBC331" author="Kaszab, Z" journalOrPublisher="Annales historico-naturales Musei Nationalis hungarici" pageId="6" pageNumber="51" pagination="169 - 221" title="Angaben zur Kenntnis der Tenebrioniden Nordvietnams (Coleoptera)." url="http://publication.nhmus.hu/pdf/annHNHM/Annals_HNHM_1980_Vol_72_169.pdf" volume="72" year="1980">Kaszab 1980</bibRefCitation>
,
<bibRefCitation id="0B8575941DBB6CAEDCFD6590C0AC911B" author="Merkl, O" journalOrPublisher="Acta Zoologica Academiae Scientiarum Hungaricae" pageId="6" pageNumber="51" pagination="261 - 280" title="Tenebrionidae (Coleoptera) from Laos and Vietnam, with reclassification of Old World &quot; Doliema &quot;." url="https://www.researchgate.net/publication/257239115_Tenebrionidae_%28Coleoptera%29_from_Laos_and_Vietnam_with_reclassification_of_Old_World_Doliema?ev=prf_pub" volume="38" year="1992">Merkl 1992</bibRefCitation>
); Thailand, Indonesia/Sumatra, China/Guangxi and Hainan Provinces (new records).
</paragraph>
</subSubSection>
</treatment>
</document>
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