treatments-xml/data/AC/88/8F/AC888F6EC3F6588F9EEAB351452E904E.xml
2024-06-21 12:47:31 +02:00

226 lines
24 KiB
XML
Raw Blame History

This file contains invisible Unicode characters

This file contains invisible Unicode characters that are indistinguishable to humans but may be processed differently by a computer. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1192.111275" ID-Pensoft-Pub="1313-2970-1192-111" ID-Pensoft-UUID="436A6870AC145BE79A56D786FB1C9716" ID-ZooBank="94FB491FB5A54514A1EF062B6A216D11" ModsDocID="1313-2970-1192-111" checkinTime="1708593701100" checkinUser="pensoft" docAuthor="Doan, Lydia M., Miller, James S., Brown, John W., Forister, Matthew L. &amp; Dyer, Lee A." docDate="2024" docId="AC888F6EC3F6588F9EEAB351452E904E" docLanguage="en" docName="ZooKeys 1192: 111-140" docOrigin="ZooKeys 1192" docPubDate="2024-02-21" docSource="http://dx.doi.org/10.3897/zookeys.1192.111275" docTitle="Eois parumsimii Doan 2024, sp. nov." docType="treatment" docUuid="4A65A58B-A430-4E22-A1F0-79D75A7267F8" docUuidSource="ZooBank" docVersion="1" id="436A6870AC145BE79A56D786FB1C9716" lastPageNumber="111" masterDocId="436A6870AC145BE79A56D786FB1C9716" masterDocTitle="Two new species of the hyperdiverse geometrid moth genus Eois (Lepidoptera, Geometridae, Larentiinae) from Ecuador, with descriptions of early stages" masterLastPageNumber="140" masterPageNumber="111" pageNumber="111" updateTime="1708593701100" updateUser="pensoft">
<mods:mods id="DB8FD4AFD62F31BD8480F55ECDD1FE6A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CDE408017E0971FB5FCA1742E561A668">
<mods:title id="00E141DBFA2A5E1DFE6C3D8B2030B736">Two new species of the hyperdiverse geometrid moth genus Eois (Lepidoptera, Geometridae, Larentiinae) from Ecuador, with descriptions of early stages</mods:title>
</mods:titleInfo>
<mods:name id="B5F34C2073236FAFDF9FF3264F34B922" type="personal">
<mods:role id="B5E03C57420332C054D51832BA09FC0F">
<mods:roleTerm id="9110365A5221BE0D1B4E26282E1041ED">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7C4846183C53A7E45ADC37BD54D374B5">Doan, Lydia M.</mods:namePart>
<mods:nameIdentifier id="8DDE912494659515CBF8090945B0B1DD" type="ORCID">https://orcid.org/0000-0002-7039-8814</mods:nameIdentifier>
<mods:affiliation id="DDA8A23E69E5029E298579AD12FD6C9B">Department of Biology, Ecology, Evolution and Conservation of Biology, University of Nevada, Reno, NV 89557, USA</mods:affiliation>
</mods:name>
<mods:name id="501FFA2F4D9296087CECD7E7270A877C" type="personal">
<mods:role id="A383628945041388456560D5A1BF0D18">
<mods:roleTerm id="ABFFB82BF65B146EF5A3EF648BC39EF4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8A0904A8FE0DBB63C52DC89FA060C8CE">Miller, James S.</mods:namePart>
<mods:affiliation id="0C88C6DCD11BEF915D1FD158A39625F1">Entomology Department, American Museum of Natural History, New York, NY, 10024, USA</mods:affiliation>
</mods:name>
<mods:name id="5E7C3C5AC64B1AAC7A1F3E6DB60BAD44" type="personal">
<mods:role id="96315683E130016F37B5FE17DCD08834">
<mods:roleTerm id="BA34C762C21411225B66AE08FF268FB4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="569A3BA4AE196818A031EF4EF0C1E9DF">Brown, John W.</mods:namePart>
<mods:nameIdentifier id="CB4B6851FB25F24CB5514E2BBB284A10" type="ORCID">https://orcid.org/0000-0001-5610-9855</mods:nameIdentifier>
<mods:affiliation id="12660C104B5B2E8D5D66C1743D3B046C">Entomology Department, National Museum of Natural History, Smithsonian Institution, Washington, DC, 20560, USA</mods:affiliation>
</mods:name>
<mods:name id="DBEA6520EC4D93E0DD6FFA1563FA7D34" type="personal">
<mods:role id="823BE6525E221A02B51F58CCB4117B60">
<mods:roleTerm id="893926A67F501B2F6C95653E97D58D53">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0FE9206619B1A0A51D710EFDE99AFDB0">Forister, Matthew L.</mods:namePart>
<mods:nameIdentifier id="DAFBD5D696162E7C3FC6D6167EABCD96" type="ORCID">https://orcid.org/0000-0003-2765-4779</mods:nameIdentifier>
<mods:affiliation id="64AC5ACC6B8F4ECE2D7936400B39C8D8">Department of Biology, Ecology, Evolution and Conservation of Biology, University of Nevada, Reno, NV 89557, USA</mods:affiliation>
</mods:name>
<mods:name id="ACC013DD781C6F4BC0FB54D7940E0FEC" type="personal">
<mods:role id="599FB4E6D0771FA86E75FF1DC6E662A8">
<mods:roleTerm id="19CB2A0FBD27629E38D37B79FEA10E11">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1B471FB28B138E49EE468DBF6CC59EC4">Dyer, Lee A.</mods:namePart>
<mods:nameIdentifier id="DC904EA48E3F2C302A2853A3DE0460CA" type="ORCID">https://orcid.org/0000-0002-0867-8874</mods:nameIdentifier>
<mods:affiliation id="914B0BD435736445B4584BDFD3AF44D9">Department of Biology, Ecology, Evolution and Conservation of Biology, University of Nevada, Reno, NV 89557, USA</mods:affiliation>
<mods:nameIdentifier id="D2D7DB4E55E8C19AF883EFC6384FFA52" type="email">ldyer@unr.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="5B47EFF5E64C8AAE79E6C43BC9D2EC83">text</mods:typeOfResource>
<mods:relatedItem id="178D92A438A67A745A7AAC42700858FA" type="host">
<mods:titleInfo id="F3CE647662FD628F46AA94AC1A4AE3ED">
<mods:title id="632E515FEEC0226158ED45996BA2E706">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="CA86D174175369D5D2F043C270BF361E">
<mods:date id="A3D6C866D848F21BDC2B30C7D7813CAF">2024</mods:date>
<mods:detail id="666CEC27CC1531AFD0070703A4092600" type="pubDate">
<mods:number id="FDFB1CB96468146607F5E29A2B793967">2024-02-21</mods:number>
</mods:detail>
<mods:detail id="E13E80905552EB5E781BC6B330DBB315" type="volume">
<mods:number id="A6C48A2424F49C02E76FF32E5CDF2949">1192</mods:number>
</mods:detail>
<mods:extent id="4D084479B17210840675EC35ED68FCE7" unit="page">
<mods:start id="490665D2273D0F8929EB4A7666116BC2">111</mods:start>
<mods:end id="D4D0B0CF5210EBD7F919C7FC3F1B9A26">140</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="E5EB1DA86B29F048F19CC2D70C895066">
<mods:url id="A96E1E4A879207573CCD9C195B42387C">http://dx.doi.org/10.3897/zookeys.1192.111275</mods:url>
</mods:location>
<mods:classification id="6232117D9191E104F52FEEDD8E771F04">journal article</mods:classification>
<mods:identifier id="0B81A26D4889C4EAA47F98515BCB1A19" type="DOI">http://dx.doi.org/10.3897/zookeys.1192.111275</mods:identifier>
<mods:identifier id="311E63241AE351BA7545EEFF20D47FE5" type="Pensoft-Pub">1313-2970-1192-111</mods:identifier>
<mods:identifier id="038E69DCCDD2A67C4D1654CE30FF45E8" type="ZooBank">94FB491FB5A54514A1EF062B6A216D11</mods:identifier>
<mods:identifier id="CE011A44CF01D550C6D384796701B943" type="Pensoft-UUID">436A6870AC145BE79A56D786FB1C9716</mods:identifier>
</mods:mods>
<subSection id="8DC4DEEB733F24FD7570A0A5347D3D50" pageId="0" pageNumber="111" type="results">
<treatment id="AC888F6EC3F6588F9EEAB351452E904E" LSID="urn:lsid:zoobank.org:act:4A65A58B-A430-4E22-A1F0-79D75A7267F8" httpUri="http://treatment.plazi.org/id/AC888F6EC3F6588F9EEAB351452E904E" lastPageNumber="111" pageId="0" pageNumber="111">
<subSubSection id="A5C15DA9E507B702635B76DF533D2B83" pageId="0" pageNumber="111" type="nomenclature">
<paragraph id="A4AE78F5AB50698F3802BC47956B69E5" pageId="0" pageNumber="111">
<taxonomicName id="C5D6C6DD2F7D6C10DB57FD5442B7A4CE" LSID="https://zoobank.org/4A65A58B-A430-4E22-A1F0-79D75A7267F8" authority="Doan" authorityName="Doan" authorityYear="2024" class="Insecta" family="Geometridae" genus="Eois" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eois parumsimii" order="Lepidoptera" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="parumsimii" status="sp. nov.">Eois parumsimii Doan</taxonomicName>
<taxonomicNameLabel id="9DEAAA375CA8D3667D857AAABC6E6389" pageId="0" pageNumber="111">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="16C7E57D685BF3EDBF1BE460F26B5E0A" pageId="0" pageNumber="111" type="description">
<paragraph id="033B2C985AC6D5FE52693B3319FE2052" pageId="0" pageNumber="111">
<figureCitation id="B92DA373CBDCC2CC3CA2A24061160E3C" captionStart="Figures 2023" captionStartId="F7" captionText="Figures 20 - 23. Eois parumsimii 20 holotype upperside (UNR) 21 holotype underside (UNR) 22 male genitalia, slide 75819 (UNR) 23 female genitalia, slide 75961 (UNR)." figureDoi="10.3897/zookeys.1192.111275.figures20-23" httpUri="https://binary.pensoft.net/fig/990480" pageId="0" pageNumber="111">Figs 20-23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8AEC73D9B38BAF29A9C526A3548D6D8C" pageId="0" pageNumber="111" type="type material">
<paragraph id="990590451C6536FB464EC1A6162E4D7B" pageId="0" pageNumber="111">Type material.</paragraph>
<paragraph id="118040BF89E0336F618A856824267CBD" pageId="0" pageNumber="111">
<emphasis id="CDE0C2E32E16F2BF7E359C76EF1F2328" bold="true" italics="true" pageId="0" pageNumber="111">Holotype</emphasis>
♂, Ecuador, Napo, Yanayacu Biological Station, 2113.9 m,
<geoCoordinate id="6D88D1A8861C26542CF5FA6CAF696AF8" degrees="00" direction="south" minutes="35" orientation="latitude" precision="1" seconds="48.998" value="-0.59694386">00°35'48.998&quot;S</geoCoordinate>
,
<geoCoordinate id="152656D5FAB78F7246CCF2DF35037D18" degrees="77" direction="west" minutes="53" orientation="longitude" precision="1" seconds="17.998" value="-77.88833">77°53'17.998&quot;W</geoCoordinate>
, Nov 2012, r.f. 71361, Earthwatch (UNR).
<emphasis id="7F9C466DD7DE30F5A6DB8CA21EC627AA" bold="true" italics="true" pageId="0" pageNumber="111">Paratypes</emphasis>
(5♂, 2♀). Ecuador: Napo: Yanayacu Biological Station, 2066.8 m,
<geoCoordinate id="A144EEF5B3732222A476666D9DDB7652" degrees="00" direction="south" minutes="34" orientation="latitude" precision="1" seconds="0.001" value="-0.56666696">00°34'0.001&quot;S</geoCoordinate>
,
<geoCoordinate id="659B1A505440D1FC2E00D2FECFEDCC85" degrees="77" direction="west" minutes="52" orientation="longitude" precision="1" seconds="0.001" value="-77.86667">77°52'0.001&quot;W</geoCoordinate>
, Feb 2005, r.f. 1673, 1674 (2♂), Sept 2010, r.f. 51792 (1♂), Earthwatch (UNR); 2188.4 m,
<geoCoordinate id="AB2C833FD0EA0D954EE9C3A000EA97C5" degrees="00" direction="south" minutes="35" orientation="latitude" precision="15" seconds="54" value="-0.5983333">00°35'54&quot;S</geoCoordinate>
,
<geoCoordinate id="8851556D5C7D1E8F4CD04C3AB3BA506C" degrees="77" direction="west" minutes="53" orientation="longitude" precision="1" seconds="44.34" value="-77.895645">77°53'44.34&quot;W</geoCoordinate>
, Nov 2005, r.f. 9442 (1♂), Earthwatch (UNR); 1240.7 m,
<geoCoordinate id="2E0C02EADBBE92C72F2E9109CBD14504" degrees="00" direction="south" minutes="43" orientation="latitude" precision="1" seconds="38.798" value="-0.7274439">00°43'38.798&quot;S</geoCoordinate>
,
<geoCoordinate id="2E4675D0D2BE138704F43B56EC68609D" degrees="77" direction="west" minutes="46" orientation="longitude" precision="15" seconds="22" value="-77.77278">77°46'22&quot;W</geoCoordinate>
, Jun 2014, r.f. 84256 (1♀), Earthwatch (UNR); 1871.9 m,
<geoCoordinate id="42EAFC8F03C310C75D1AC0063C19AF35" degrees="00" direction="south" minutes="31" orientation="latitude" precision="1" seconds="31.2" value="-0.52533334">00°31'31.2&quot;S</geoCoordinate>
,
<geoCoordinate id="6A21361CBC088FDD977787ED8F5C4B36" degrees="77" direction="west" minutes="52" orientation="longitude" precision="1" seconds="35.399" value="-77.8765">77°52'35.399&quot;W</geoCoordinate>
, Aug 2014, r.f. 85960, 85963 (1♂, 1♀), Earthwatch (UNR).
</paragraph>
</subSubSection>
<subSubSection id="CCEF8AAB215B17745781125D8AA85418" pageId="0" pageNumber="111" type="diagnosis">
<paragraph id="F5854BE4C5F81AED4597BD2D684956C0" pageId="0" pageNumber="111">Diagnosis.</paragraph>
<paragraph id="31E7FE8031D6EC6C48E5E96C6D27CF90" pageId="0" pageNumber="111">
This species is described from specimens reared from larvae collected at Yanayacu Biological Station in Ecuador. Externally,
<taxonomicName id="4B4A9E060481B2374DEE5C10CC7B5F18" lsidName="E. parumsimii" pageId="0" pageNumber="111" rank="species" species="parumsimii">
<emphasis id="2FAD0C75F333B2820C0728E2BCFC53D7" italics="true" pageId="0" pageNumber="111">E. parumsimii</emphasis>
</taxonomicName>
is distinct from all of species in the genus, with a much broader, yellow, postmedial line on a pale pinkish gray ground color (Fig.
<figureCitation id="8BEAAD43D59F83B2E7F1084791EE19FB" captionStart="Figures 2023" captionStartId="F7" captionText="Figures 20 - 23. Eois parumsimii 20 holotype upperside (UNR) 21 holotype underside (UNR) 22 male genitalia, slide 75819 (UNR) 23 female genitalia, slide 75961 (UNR)." figureDoi="10.3897/zookeys.1192.111275.figures20-23" httpUri="https://binary.pensoft.net/fig/990480" pageId="0" pageNumber="111">20</figureCitation>
). The male genitalia of
<taxonomicName id="9164652FAB6BFB1101A94B19BAD25B6E" lsidName="E. parumsimii" pageId="0" pageNumber="111" rank="species" species="parumsimii">
<emphasis id="3C3B5DBFF4A1E36FB3B366A70187FE30" italics="true" pageId="0" pageNumber="111">E. parumsimii</emphasis>
</taxonomicName>
can be distinguished from those of other members of the complex by the following combination of character states: the ventral margin of the saccus forming a blunt conical pocket (vs a small, transverse-ovoid pocket in most other species); and the membrane surrounding the phallus with a large dorsal field of short spines arranged in a series of longitudinal rows (vs arranged in two longitudinal rows most other in species). The female genitalia of
<taxonomicName id="9CFEF30681E43DAA22AAE42100E45D44" lsidName="E. parumsimii" pageId="0" pageNumber="111" rank="species" species="parumsimii">
<emphasis id="84FDEAFF023A6FE2E003D97B46A7EC90" italics="true" pageId="0" pageNumber="111">E. parumsimii</emphasis>
</taxonomicName>
have short, trapezoidal papillae anales, whereas the papillae anales are slenderer in most other species; and the signum is located laterally on the left side of the corpus bursae, whereas it located ventrally in many other species of the complex.
</paragraph>
</subSubSection>
<subSubSection id="3376ED4D0D107E6D8D10035CE8AD98E5" pageId="0" pageNumber="111" type="description">
<paragraph id="BDA79C027B83D7D41A7ADB945355D22F" pageId="0" pageNumber="111">Description.</paragraph>
<paragraph id="DCD8BD3B43A3467C328EB8FD9B0B5CE8" pageId="0" pageNumber="111">
<emphasis id="3FB87E04306613896E28FB18CBB6401F" bold="true" pageId="0" pageNumber="111">
Male.
<emphasis id="8A67AFDBB8B67F5F35624274DF0EE855" italics="true" pageId="0" pageNumber="111">Head</emphasis>
</emphasis>
: Essentially as described for species complex.
<emphasis id="C80B6E89B58234071BF89EC913C55B6F" bold="true" italics="true" pageId="0" pageNumber="111">Thorax</emphasis>
: Essentially as described for species complex, except forewing length 8.0-9.0 mm (
<emphasis id="F70E09CAF83AC5FFB14ED1348B8C302B" italics="true" pageId="0" pageNumber="111">n</emphasis>
= 8); ground color manzanilla olive, anterior portion of antemedial line very faint, posterior portion prominent, pale yellow; postmedial line well defined, wavy, pale yellow; discal spot well defined, red brown; basal 0.66 of costal margin concolorous with ground color, distal portion with clay-brown markings; termen with slender clay-brown line; fringe two-toned, mostly pale yellow, with red-brown incursion between veins M2 and M3. Forewing underside ground color pale yellow, suffused with red-brown, inverse to dorsal pattern; antemedial line absent; postmedial line prominent, width variable, but broader near costa and posterior margin; discal spot round, faint, clay-brown; termen clay brown. Fringe pale yellow. Hindwing upperside ground color clay-brown; antemedial line faint; postmedial line wavy, pale yellow; discal spot round, well defined, orange-brown; termen clay-brown. Fringe pale yellow. Hindwing with pattern similar to forewing, with prominent pale yellow antemedial and postmedial lines and faint discal spot.
<emphasis id="0E9DABD1CA0BC8D38227A9373F79FE73" bold="true" italics="true" pageId="0" pageNumber="111">Abdomen</emphasis>
: Genitalia (Fig.
<figureCitation id="8E8B467006FAE9DA67E03771ACB0C6DD" captionStart="Figures 2023" captionStartId="F7" captionText="Figures 20 - 23. Eois parumsimii 20 holotype upperside (UNR) 21 holotype underside (UNR) 22 male genitalia, slide 75819 (UNR) 23 female genitalia, slide 75961 (UNR)." figureDoi="10.3897/zookeys.1192.111275.figures20-23" httpUri="https://binary.pensoft.net/fig/990480" pageId="0" pageNumber="111">22</figureCitation>
) with tegumen arms forming rounded triangular dorsal arch, curving slightly posterad; ventral margin of saccus forming a blunt conical pocket; transtilla slender, V-shaped; juxta wide basally, abruptly narrowed in distal 0.33 with acute dorsal part; dorsal margin of juxta narrow, truncate, with a small down-curved lip; valva subrectangular, weakly constricted on ventral margin near distal end of sacculus; a brush of bristle-like setae near apex, contrasting with remaining setae; setae at apex of sacculus longer than width of valva; ventral margin of sacculus bowed outward; base of phallus narrow, horn-shaped, membrane surrounding phallus with large dorsal field of short spines arranged in longitudinal rows; phallus ca as long as valva; vesica bifurcated distally with two appendices, one with distal group of spine-like cornuti, the other at base of vesica with a single, large scobinate plate.
</paragraph>
<paragraph id="C693874BFB0B7CFF60AFEF6115FDFD13" pageId="0" pageNumber="111">
<emphasis id="1527FF956DBC68084A84604A2A828A41" bold="true" pageId="0" pageNumber="111">
Female.
<emphasis id="45BBB72B44A403D3627A252C29563739" italics="true" pageId="0" pageNumber="111">Head</emphasis>
</emphasis>
and
<emphasis id="DDB9B475668B16CEFB7416836C028ADB" bold="true" italics="true" pageId="0" pageNumber="111">Thorax</emphasis>
: Essentially as described for male except lacking rami on antenna.
<emphasis id="52855314E7CBA7C0A3E8456FB1411823" bold="true" pageId="0" pageNumber="111">Abdomen</emphasis>
: Genitalia (Fig.
<figureCitation id="03B64805C0AC70951FF166E07B920F3E" captionStart="Figures 2023" captionStartId="F7" captionText="Figures 20 - 23. Eois parumsimii 20 holotype upperside (UNR) 21 holotype underside (UNR) 22 male genitalia, slide 75819 (UNR) 23 female genitalia, slide 75961 (UNR)." figureDoi="10.3897/zookeys.1192.111275.figures20-23" httpUri="https://binary.pensoft.net/fig/990480" pageId="0" pageNumber="111">23</figureCitation>
) with papillae anales short, trapezoidal. Dorsal membrane between Tg8 and papillae anales bearing a large dorsal sac. Ductus bursae wide; ductus seminalis arising from elongate, triangular appendix at base of corpus bursae; Corpus bursae oblong, without mesal constriction, distal appendix absent; signum wing-shaped with serrate lateral margins, located laterally on left side of corpus bursae, internal part horn-like.
</paragraph>
<caption id="460AF140C72127ECA32871F8F3D9FEB6" doi="10.3897/zookeys.1192.111275.figures20-23" httpUri="https://binary.pensoft.net/fig/990480" pageId="0" pageNumber="111" start="Figures 2023" startId="F7">
<paragraph id="084C7E64EC46EBAE87627C48ED1ED6A2" pageId="0" pageNumber="111">
<emphasis id="EADBA90E2C0ACB928360476C33F2E8DD" bold="true" pageId="0" pageNumber="111">Figures 20-23.</emphasis>
<taxonomicName id="E39BBBF308E9965125549C9274216652" authorityName="Doan" authorityYear="2024" class="Insecta" family="Geometridae" genus="Eois" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eois parumsimii" order="Lepidoptera" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="parumsimii">
<emphasis id="E8AC8BFF8F5CD5E79A2072B73EC5DA02" italics="true" pageId="0" pageNumber="111">Eois parumsimii</emphasis>
</taxonomicName>
<emphasis id="624D8BAF07ED8F185D085DDE28B4F61B" bold="true" pageId="0" pageNumber="111">20</emphasis>
holotype upperside (UNR)
<emphasis id="76CF2DF92EA871D1A86940767D205E42" bold="true" pageId="0" pageNumber="111">21</emphasis>
holotype underside (UNR)
<emphasis id="31888B77ECC2E6FDEBF5632F5458FC79" bold="true" pageId="0" pageNumber="111">22</emphasis>
male genitalia, slide 75819 (UNR)
<emphasis id="D35D904F4393814BC5B004A0C0A2889C" bold="true" pageId="0" pageNumber="111">23</emphasis>
female genitalia, slide 75961 (UNR).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="790006EAD3772AA0C865735EA4A0F37D" pageId="0" pageNumber="111" type="biology_ecology">
<paragraph id="7D8ED038602FB65F59086ABAF94ADE78" pageId="0" pageNumber="111">Biology and distribution.</paragraph>
<paragraph id="4BC85DA24EE48BD5199663D828818F93" pageId="0" pageNumber="111">
This species is known only from Napo Province, Ecuador. Adults were reared from larvae (
<emphasis id="39C63ECD8836F2CD0A8D83BB088A33CD" italics="true" pageId="0" pageNumber="111">n</emphasis>
= 25) discovered in the field on
<taxonomicName id="2BB4182768A51E2501B5793AC50ADB25" class="Hexapoda" family="Piperaceae" genus="Piper" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Piper baezanum" order="Lepidoptera" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="baezanum">
<emphasis id="D75D0C546EB5F5BFDC417A456F23FEAA" italics="true" pageId="0" pageNumber="111">Piper baezanum</emphasis>
</taxonomicName>
, which is a threatened species endemic to Ecuador (
<bibRefCitation id="729B1DE1B4C45D18B68958D3CB1870F4" DOI="https://doi.org/10.2305/IUCN.UK.2004.RLTS.T45817A11017241.en" author="Santiana, J" journalOrPublisher="Springer Nature, Berlin" pageId="0" pageNumber="111" refId="B27" refString="Santiana, J, Pitman, N, 2004. Piper baezanum. IUCN Red List of Threatened Species 2004: e.T45817A11017241. https://doi.org/10.2305/IUCN.UK.2004.RLTS.T45817A11017241.en" title="Piper baezanum. IUCN Red List of Threatened Species 2004: e. T 45817 A 11017241." url="https://doi.org/10.2305/IUCN.UK.2004.RLTS.T45817A11017241.en" year="2004">Santiana and Pitman 2004</bibRefCitation>
).
</paragraph>
<paragraph id="B20160BE3CAEFF18CCE14356D72CBA1C" pageId="0" pageNumber="111">
The eggs, larvae, and pupae of
<taxonomicName id="7B591E73F5C3D60BB97195E626DDFCDB" lsidName="E. parumsimii" pageId="0" pageNumber="111" rank="species" species="parumsimii">
<emphasis id="452E20B70A20C802B17E48266ECEB4D5" italics="true" pageId="0" pageNumber="111">E. parumsimii</emphasis>
</taxonomicName>
have all the same general characteristics described above for
<taxonomicName id="8CDEE990D9B2AFFAE3CBDDA87627DE7A" authorityName="Hubner" authorityYear="1818" class="Insecta" family="Geometridae" genus="Eois" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eois" order="Lepidoptera" pageId="0" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="BF8324EE22F14B301665A44B869F7A3F" italics="true" pageId="0" pageNumber="111">Eois</emphasis>
</taxonomicName>
with no modifications (Figs
<figureCitation id="5DAE9CF84E7D5D823E0368EE5766024B" captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. Early stages of Eois pseudolivacea from Ecuador 1 egg 2 second instar larva 3 fifth instar larva 4 pupa." figureDoi="10.3897/zookeys.1192.111275.figures1-4" httpUri="https://binary.pensoft.net/fig/990474" pageId="0" pageNumber="111">1-4</figureCitation>
) and are difficult to distinguish from those of
<taxonomicName id="4EA6CFB89A5E728A9D83D9195A5126B2" lsidName="E. pseudolivacea" pageId="0" pageNumber="111" rank="species" species="pseudolivacea">
<emphasis id="F5F55E5CB0A390DCF7EE49991D7339EF" italics="true" pageId="0" pageNumber="111">E. pseudolivacea</emphasis>
</taxonomicName>
. All larval instars (until the prepupal stage) are similarly colored, with a translucent beige head capsule, pale green thorax and abdomen, translucent pinacula, and paired subdorsal yellow spots on all segments form T2-A6. Abdominal segments are separated by slight constrictions. The prothoracic legs and abdominal prolegs are tan. The prepupa is translucent.
</paragraph>
</subSubSection>
<subSubSection id="F8DC6D104A598E6307E4D61518628EFF" pageId="0" pageNumber="111" type="etymology">
<paragraph id="20FD30A6547E58CA800A8415B38C4C64" pageId="0" pageNumber="111">Etymology.</paragraph>
<paragraph id="49970B87CD9AE0FFD7DCB55242500C6D" pageId="0" pageNumber="111">
The species epithet
<taxonomicName id="4996E1B96D62BF17A164EC0E7E195C3B" authorityName="Doan" authorityYear="2024" class="Insecta" family="Geometridae" genus="Eois" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eois parumsimii" order="Lepidoptera" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="parumsimii">
<emphasis id="F9DF5D92F5BB4ED0DAA1DB74C5BCA18F" italics="true" pageId="0" pageNumber="111">Eois parumsimii</emphasis>
</taxonomicName>
is a patronym for Michael Lumibao, who is the partner and long-time supporter of the first author; the name is derived from his Chinese zodiac animal.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>