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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.956.51592" ID-GBIF-Dataset="b5cb4b83-ae17-48b3-8b26-f0e09c01d152" ID-PMC="PMC7426283" ID-Pensoft-Pub="1313-2970-956-49" ID-Pensoft-UUID="DE73B9FFE81C556285DDAB038D9FF8CB" ID-PubMed="32855600" ID-ZooBank="9B9E6A85D8C94794AC84B4D1965C2015" ModsDocID="1313-2970-956-49" checkinTime="1596777994909" checkinUser="pensoft" docAuthor="Espeland, Marianne, Sjoeberg, Tin &amp; Johanson, Kjell Arne" docDate="2020" docId="8DF29A394D9C517F8D8BCC67E8B1A4E5" docLanguage="en" docName="ZooKeys 956: 49-162" docOrigin="ZooKeys 956" docSource="http://dx.doi.org/10.3897/zookeys.956.51592" docTitle="Agmina cornuta Espeland &amp; Sjöberg &amp; Johanson 2020, sp. nov." docType="treatment" docUuid="F11DF372-A11F-4D40-AC2A-2625787E0DF3" docUuidSource="ZooBank" docVersion="4" id="DE73B9FFE81C556285DDAB038D9FF8CB" lastPageNumber="49" masterDocId="DE73B9FFE81C556285DDAB038D9FF8CB" masterDocTitle="Description of 47 new species of the New Caledonian endemic caddisfly genus Agmina Ward &amp; Schefter (Trichoptera, Ecnomidae)" masterLastPageNumber="162" masterPageNumber="49" pageNumber="49" updateTime="1668169213298" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Description of 47 new species of the New Caledonian endemic caddisfly genus Agmina Ward &amp; Schefter (Trichoptera, Ecnomidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Espeland, Marianne</mods:namePart>
<mods:affiliation>Arthropoda Department, Zoological Research Museum Alexander Koenig, Bonn, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sjoeberg, Tin</mods:namePart>
<mods:affiliation>Zoology Department, Swedish Museum of Natural History, Box 50007, 10405 Stockholm, Sweden</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Johanson, Kjell Arne</mods:namePart>
<mods:affiliation>Zoology Department, Swedish Museum of Natural History, Box 50007, 10405 Stockholm, Sweden</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-1893-3429</mods:nameIdentifier>
<mods:nameIdentifier type="email">kjell.arne.johanson@nrm.se</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>956</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>49</mods:start>
<mods:end>162</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.956.51592</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.956.51592</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-956-49</mods:identifier>
<mods:identifier type="ZooBank">9B9E6A85D8C94794AC84B4D1965C2015</mods:identifier>
<mods:identifier type="Pensoft-UUID">DE73B9FFE81C556285DDAB038D9FF8CB</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="166097650" LSID="urn:lsid:zoobank.org:act:F11DF372-A11F-4D40-AC2A-2625787E0DF3" httpUri="http://treatment.plazi.org/id/8DF29A394D9C517F8D8BCC67E8B1A4E5" lastPageNumber="49" pageId="0" pageNumber="49">
<subSubSection pageId="0" pageNumber="49" type="nomenclature">
<paragraph pageId="0" pageNumber="49">
<taxonomicName LSID="8DF29A39-4D9C-517F-8D8B-CC67E8B1A4E5" authority="Espeland &amp; Sjöberg &amp; Johanson, 2020" authorityName="Espeland &amp; Sjöberg &amp; Johanson" authorityYear="2020" class="Insecta" family="Ecnomidae" genus="Agmina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Agmina cornuta" order="Trichoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="cornuta" status="sp. nov.">Agmina cornuta</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="49">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 2530" captionStartId="F6" captionText="Figures 25 - 30. Agmina cornuta sp. nov. male holotype 25 genitalia, left lateral view 26 genitalia, dorsal view 27 end of superior appendages, showing mesal spine, dorsal view 28 genitalia, ventral view 29 phallus, lateral view 30 phallus, ventral view." figureDoi="10.3897/zookeys.956.51592.figures25-30" httpUri="https://binary.pensoft.net/fig/439924" pageId="0" pageNumber="49">Figs 25-30</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="diagnosis">
<paragraph pageId="0" pageNumber="49">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="49">
<taxonomicName authorityName="Espeland &amp; Sjöberg &amp; Johanson" authorityYear="2020" class="Insecta" family="Ecnomidae" genus="Agmina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Agmina cornuta" order="Trichoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis italics="true" pageId="0" pageNumber="49">Agmina cornuta</emphasis>
</taxonomicName>
sp. nov. is distinguished from the other
<taxonomicName authorityName="JB Ward &amp; PW Schefter" authorityYear="2000" class="Insecta" family="Ecnomidae" genus="Agmina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Agmina" order="Trichoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="49">Agmina</emphasis>
</taxonomicName>
species by the presence of a pair of almost uniformly oval superior appendages in lateral view, and a pair of branches of the inferior appendages which appear beak-like in lateral view and are approx. as long as the superior appendages. It resembles
<taxonomicName lsidName="A. multidentata" pageId="0" pageNumber="49" rank="species" species="multidentata">
<emphasis italics="true" pageId="0" pageNumber="49">A. multidentata</emphasis>
</taxonomicName>
sp. nov. in the shape of tergum X, but in
<taxonomicName lsidName="A. cornuta" pageId="0" pageNumber="49" rank="species" species="cornuta">
<emphasis italics="true" pageId="0" pageNumber="49">A. cornuta</emphasis>
</taxonomicName>
sp. nov. the inferior appendages are lower in lateral view and the superior appendages are rounder posteriorly compared to those in
<taxonomicName lsidName="A. multidentata" pageId="0" pageNumber="49" rank="species" species="multidentata">
<emphasis italics="true" pageId="0" pageNumber="49">A. multidentata</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
<caption doi="10.3897/zookeys.956.51592.figures25-30" httpUri="https://binary.pensoft.net/fig/439924" pageId="0" pageNumber="49" start="Figures 2530" startId="F6">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Figures 25-30.</emphasis>
<taxonomicName authorityName="Espeland &amp; Sjöberg &amp; Johanson" authorityYear="2020" class="Insecta" family="Ecnomidae" genus="Agmina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Agmina cornuta" order="Trichoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis italics="true" pageId="0" pageNumber="49">Agmina cornuta</emphasis>
</taxonomicName>
sp. nov. male holotype
<emphasis bold="true" pageId="0" pageNumber="49">25</emphasis>
genitalia, left lateral view
<emphasis bold="true" pageId="0" pageNumber="49">26</emphasis>
genitalia, dorsal view
<emphasis bold="true" pageId="0" pageNumber="49">27</emphasis>
end of superior appendages, showing mesal spine, dorsal view
<emphasis bold="true" pageId="0" pageNumber="49">28</emphasis>
genitalia, ventral view
<emphasis bold="true" pageId="0" pageNumber="49">29</emphasis>
phallus, lateral view
<emphasis bold="true" pageId="0" pageNumber="49">30</emphasis>
phallus, ventral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="etymology">
<paragraph pageId="0" pageNumber="49">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="49">
<emphasis italics="true" pageId="0" pageNumber="49">Cornuta</emphasis>
, from Latin, meaning horned. Named for the shape of the inferior appendage in lateral view, which has two long, horn-shaped, dorsal branches.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="materials_examined">
<paragraph pageId="0" pageNumber="49">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" italics="true" pageId="0" pageNumber="49">Holotype</emphasis>
: New Caledonia -
<emphasis bold="true" pageId="0" pageNumber="49">Province Sud</emphasis>
• ♂; Plateau de Dogny, northern part;
<geoCoordinate degrees="21" direction="south" minutes="36.853" orientation="latitude" precision="1" value="-21.614216">21°36.853'S</geoCoordinate>
,
<geoCoordinate degrees="165" direction="east" minutes="52.548" orientation="longitude" precision="1" value="165.8758">165°52.548'E</geoCoordinate>
; loc#159 (32-2001); Malaise trap; 2-5.xii-2001; leg. KA Johanson, T Pape &amp; B Viklund. MNHN.
</paragraph>
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" italics="true" pageId="0" pageNumber="49">Paratypes</emphasis>
: New Caledonia -
<emphasis bold="true" pageId="0" pageNumber="49">Province Sud</emphasis>
• ♂;
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="spéciale">speciale</normalizedToken>
de faune de la haute
<normalizedToken originalValue="Yaté">Yate</normalizedToken>
, along road on southern part of Marais de la
<normalizedToken originalValue="Rivière">Riviere</normalizedToken>
Blanche, stream draining to Marais de la
<normalizedToken originalValue="Rivière">Riviere</normalizedToken>
Blanche, 2.25 km SW Pont
<normalizedToken originalValue="Pérignon">Perignon</normalizedToken>
; 180 m; 6-16.xi.2003; Malaise trap; loc#010a; leg. KA Johanson; NHRS; • ♂; Mt. Dzumac, source stream of Ouinne River, near crossing point to mountain track;
<geoCoordinate degrees="22" direction="south" minutes="02.073" orientation="latitude" precision="1" value="-22.03455">22°02.073'S</geoCoordinate>
,
<geoCoordinate degrees="166" direction="east" minutes="28.460" orientation="longitude" precision="1" value="166.47433">166°28.460'E</geoCoordinate>
; 810 m; 18.xi-4.xii.2003; Malaise trap; loc#030; leg. KA Johanson; NHRS; • ♂; Monts des Koghis, ca 300 m S Koghi Restaurant;
<geoCoordinate degrees="22.18288" direction="south" orientation="latitude" precision="1" value="-22.18288">22.18288S</geoCoordinate>
,
<geoCoordinate degrees="166.50167" direction="east" orientation="longitude" precision="1" value="166.50166">166.50167E</geoCoordinate>
; 417 m; 2-16.xi.2003; Malaise trap; loc#004; leg. KA Johanson; NHRS.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="description">
<paragraph pageId="0" pageNumber="49">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="49">
Fore wing length 3.2-4.1 mm (
<emphasis italics="true" pageId="0" pageNumber="49">N</emphasis>
= 4). Total length of genitalia: 0.5 mm.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="description">
<paragraph pageId="0" pageNumber="49">Description.</paragraph>
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" italics="true" pageId="0" pageNumber="49">Genitalia</emphasis>
: In lateral view, segment IX semi-triangular anteriorly, apex located dorsally; in ventral view anteriorly widely U-shaped. Sternal processes, lateral view, with each apex, not exceeding posterior apex of tergum X, very short, triangular; in ventral view, short, slender, not reaching mid-length of inferior appendage, each with apex slightly curving mesad. Tergum X small, in lateral view approx. half the length of superior appendage, outer margin irregularly convex, slightly converging towards posterior with straight inner margin; in dorsal view, mesally connected, axe-shaped with inner margins forming a right triangle. Parameres starting at tergum X, in lateral view semi-circular, each with narrow protrusion orientated posteroventrad with blunt apex, approx. half the length of the main parameres; in dorsal view, separated, slightly diverging towards apices, outer margins straight for first 2/3, then abruptly curving inwards at obtuse angle, inner margin irregularly lobed. Superior appendages, in lateral view, approx. twice as long as tergum X, semi-ovoid with truncated, irregularly dentate apex, mesal process absent; in dorsal view almost parallel, each gently widening to mid-length, then inner margin abruptly widening, with apical half irregularly dentate. Mesal processes absent. Inferior appendages, in lateral view, with posterad orientated long, knife-shaped dorsal branch; ventral branch present as short, obtuse process at posteroventral margin; in ventral view convex at base, gradually widening for first 1/3, then abruptly narrowing towards apex, long, narrow, slightly tapering dorsal branches orientated posteriorly, slightly curving inwards, apices pointed, ventral branch indiscernible. Phallus, in lateral view shorter than segment IX, irregular; in ventral view posterior half slightly wider than anterior half.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="additional information">
<paragraph pageId="0" pageNumber="49">Additional information.</paragraph>
<paragraph pageId="0" pageNumber="49">
This species was referred to as &quot;sp. 53&quot; in
<bibRefCitation DOI="https://doi.org/10.1111/j.1420-9101.2010.02072.x" author="Espeland, M" journalOrPublisher="Journal of Evolutionary Biology" pageId="0" pageNumber="49" pagination="2112 - 2122" refId="B4" refString="Espeland, M, Johanson, KA, 2010a. The diversity and radiation of the largest monophyletic animal group on New Caledonia (Trichoptera: Ecnomidae: Agmina). Journal of Evolutionary Biology 23: 2112 - 2122, DOI: https://doi.org/10.1111/j.1420-9101.2010.02072.x" title="The diversity and radiation of the largest monophyletic animal group on New Caledonia (Trichoptera: Ecnomidae: Agmina)." url="https://doi.org/10.1111/j.1420-9101.2010.02072.x" volume="23" year="2010 a">Espeland and Johanson (2010a)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>