180 lines
19 KiB
XML
180 lines
19 KiB
XML
<document id="6DE3518D0148CCA097801081921E347F" ID-CLB-Dataset="21156" ID-DOI="10.11646/zootaxa.4661.2.3" ID-GBIF-Dataset="6c12aca1-7350-4265-9ded-6d415152f181" ID-ISSN="1175-5326" ID-Zenodo-Dep="3998528" ID-ZooBank="95722A8A-E02C-47E5-AA9B-777B6CA226E6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1598306200512" checkinUser="felipe" docAuthor="Fukuda, Yuki, Ogawa, Ryo & Hori, Michio" docDate="2019" docId="1D1C8783DF698E156986311C8B1CF823" docLanguage="en" docName="zootaxa.4661.2.3.pdf.imf" docOrigin="Zootaxa 4661 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sophiodela okinawana Fukuda & Ogawa & Hori 2019, comb. nov." docType="treatment" docVersion="6" lastPageNumber="284" masterDocId="E125FFFBDF648E18691134438958FFF1" masterDocTitle="The reclassification of Sophiodela and other tiger beetles (Coleoptera, Cicindelidae) based on the structure of the everted internal sac of the male genitalia" masterLastPageNumber="308" masterPageNumber="271" pageNumber="284" updateTime="1698845354460" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="E2642AD42BD74B5A3EE784E6793F9A03" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4E77BC8F5412106114608B0F0E9950A5">
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<mods:title id="BF9138633B50ABBF991319843E162413">The reclassification of Sophiodela and other tiger beetles (Coleoptera, Cicindelidae) based on the structure of the everted internal sac of the male genitalia</mods:title>
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<mods:name id="679EC400568B4BB55646A5CEF561D631" type="personal">
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<mods:role id="829595AA206E120A580EE9A74FEB50E0">
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<mods:roleTerm id="543FC4C4841A2E69713BD38CA2A70A48">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="751A399A57D94FCDA62B4000761BFCEB">Fukuda, Yuki</mods:namePart>
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<mods:affiliation id="BCB28DEEE7FC6F68676EA334DAD6C8DA">Kurayoshi, 111 - 8, Kamogawa-town, Tottori, 682 - 0923 Japan E-mail: yuki 03 go 3 @ gmail. com</mods:affiliation>
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<mods:nameIdentifier id="D6F49E0A838E2F7EB85E5CE2497C0FCF" type="email">yuki03go3@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="D05DFB4CD1C581658F1E2C8B35FCFBF9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B5D87C026EF325CCAA1ECDBE73548E86">Ogawa, Ryo</mods:namePart>
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<mods:affiliation id="83DD277C67C8B6C8C4693B8E14520095">Matsuyama, 6 - 5 - 8, Hakusuidai, Ehime, 791 - 0113 Japan E-mail: ailmasbl 0854 @ yahoo. co. jp</mods:affiliation>
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</mods:name>
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<mods:name id="A4FD451A20FB180C9B495E1C01446198" type="personal">
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<mods:roleTerm id="3B59A612BC09F70D919C35F24A9F1543">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="614341095D2F0381740096A0699EC19D">Hori, Michio</mods:namePart>
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<mods:affiliation id="103A3F623AA280A2CD45ABB6E5DD3CEB">Wakayama, 6 - 1 - 2 Takajyomachi, Wakayama, 640 - 8135 Japan E-mail: hori @ terra. zool. kyoto-u. ac. jp</mods:affiliation>
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<mods:typeOfResource id="3F77CA24FC703CBAD4E66E0EF6C6B3E7">text</mods:typeOfResource>
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<mods:title id="7B6F280D08E74443BB45F7D2196AEA57">Zootaxa</mods:title>
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<mods:part id="9806BFC7CEE2E9F5F0FCAA2EC984EA1B">
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<mods:date id="751F91B2F99238B962A52DF284FEA747">2019</mods:date>
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<mods:number id="C1C045ECFD1FC7D0E9CA5D9BF20FABF1">2019-08-28</mods:number>
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<mods:detail id="9781D7B08AFF31615AE934ED899F05F1" type="volume">
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<mods:number id="0080F9847C07D1BCA0A8263E52995E52">4661</mods:number>
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<mods:detail id="FB1159A3F81AFD7E712628826EB949E1" type="issue">
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<mods:number id="C6B96FA67B2C596A6129A3A8DB42B289">2</mods:number>
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<mods:start id="D74AEEF56EE13D86C92CA82904AA61D6">271</mods:start>
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<mods:classification id="0E965F3D33D884C64080059ABBC92520">journal article</mods:classification>
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<mods:identifier id="983D80D44AD19681AFD46647AE6ADC03" type="DOI">10.11646/zootaxa.4661.2.3</mods:identifier>
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<treatment id="1D1C8783DF698E156986311C8B1CF823" ID-DOI="http://doi.org/10.5281/zenodo.4337419" ID-GBIF-Taxon="167293615" ID-Zenodo-Dep="4337419" LSID="urn:lsid:plazi:treatment:1D1C8783DF698E156986311C8B1CF823" httpUri="http://treatment.plazi.org/id/1D1C8783DF698E156986311C8B1CF823" lastPageNumber="284" pageId="13" pageNumber="284">
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<subSubSection id="DDAF651EDF698E156986311C8BA5FA88" box="[151,765,1375,1401]" pageId="13" pageNumber="284" type="nomenclature">
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<paragraph id="950A3695DF698E156986311C8BA5FA88" blockId="13.[151,765,1375,1437]" box="[151,765,1375,1401]" pageId="13" pageNumber="284">
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<heading id="CE4281F9DF698E156986311C8BA5FA88" box="[151,765,1375,1401]" level="2" pageId="13" pageNumber="284">
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<taxonomicName id="52B54D16DF698E156986311C8B31FA88" ID-CoL="8M8PF" authority="(Nakane, 1957)" authorityName="Fukuda & Ogawa & Hori" authorityYear="2019" baseAuthorityName="Nakane" baseAuthorityYear="1957" box="[151,617,1375,1401]" class="Insecta" family="Carabidae" genus="Sophiodela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="284" phylum="Arthropoda" rank="species" species="okinawana" status="comb. nov.">
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<emphasis id="A7C1EA87DF698E156986311C88FCFA88" box="[151,420,1375,1401]" italics="true" pageId="13" pageNumber="284">Sophiodela okinawana</emphasis>
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<emphasis id="A7C1EA87DF698E1568BA311C8B31FA88" bold="true" box="[427,617,1375,1401]" pageId="13" pageNumber="284">
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(
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<bibRefCitation id="F1244B64DF698E1568A2311C8B38FA88" author="Nakane, T." box="[435,608,1375,1401]" pageId="13" pageNumber="284" pagination="41 - 43" refId="ref19657" refString="Nakane, T. (1957) New or little-known Coleoptera from Japan and its adjacent regions, XIV. The Scientific Report of the Saikyo University, 2 (4), 41 - 43." type="journal article" year="1957">Nakane, 1957</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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<emphasis id="A7C1EA87DF698E156B78311C8BA5FA88" bold="true" box="[617,765,1375,1401]" pageId="13" pageNumber="284">
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,
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<taxonomicNameLabel id="BCF257FCDF698E156B67311C8BA5FA88" box="[630,765,1375,1401]" pageId="13" pageNumber="284" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="DDAF651EDF698E15698631C08866FA6C" box="[151,318,1411,1437]" pageId="13" pageNumber="284" type="description">
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<paragraph id="950A3695DF698E15698631C08866FA6C" blockId="13.[151,765,1375,1437]" box="[151,318,1411,1437]" pageId="13" pageNumber="284">
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(
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<figureCitation id="0D8E2A10DF698E15698E31C089A2FA6C" box="[159,250,1411,1437]" captionStart="FIGURE 7" captionStartId="12.[151,250,1675,1700]" captionTargetBox="[153,1433,194,1642]" captionTargetId="figure@12.[151,1435,181,1650]" captionTargetPageId="12" captionText="FIGURE 7. Habitus of Sophiodela and Calochroa. A, Sophiodela chinensis chinensis (DeGeer) stat. nov.; B, Sophiodela japonica (Thunberg) stat. nov.; C, Sophiodela okinawana (Nakane) stat. nov.; D, Sophiodela ferriei (Fleutiaux) stat. nov.; E, Sophiodela chinensis flammifera (Horn) stat. nov.; F, Calochroa octonotata (Wiedemann); G, Calochroa carissima (Fleutiaux)." figureDoi="http://doi.org/10.5281/zenodo.3998542" httpUri="https://zenodo.org/record/3998542/files/figure.png" pageId="13" pageNumber="284">Figs 7C</figureCitation>
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,
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<figureCitation id="0D8E2A10DF698E15681731C0886EFA6C" box="[262,310,1411,1437]" captionStart="FIGURE 12" captionStartId="27.[151,250,1896,1921]" captionTargetBox="[151,1435,181,1870]" captionTargetId="figure@27.[151,1435,181,1870]" captionTargetPageId="27" captionText="FIGURE 12. Everted internal sacs of Sophiodela. A, Sophiodela ferriei (Fleutiaux) stat. nov.; B, S. japonica (Thunberg) stat. nov.; C, S. okinawana (Nakane) stat. nov.; D, S. chinensis chinensis (DeGeer) stat. nov.; E, S. chinensis flammifera (Horn) stat. nov. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3998552" httpUri="https://zenodo.org/record/3998552/files/figure.png" pageId="13" pageNumber="284">12C</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="DDAF651EDF698E15698631848844F99B" pageId="13" pageNumber="284" type="reference_group">
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<paragraph id="950A3695DF698E15698631848A4FFA11" blockId="13.[151,791,1479,1534]" box="[151,791,1479,1504]" pageId="13" pageNumber="284">
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<taxonomicName id="52B54D16DF698E15698631848A4CFA11" ID-CoL="8M8PF" authority="Nakane, 1957: 41" authorityName="Nakane" authorityPageNumber="41" authorityYear="1957" box="[151,788,1479,1504]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="284" phylum="Arthropoda" rank="subSpecies" species="chinensis" subGenus="Sophiodela" subSpecies="okinawana">
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<emphasis id="A7C1EA87DF698E156986318489A2FA2E" box="[151,250,1479,1503]" italics="true" pageId="13" pageNumber="284">Cicindela</emphasis>
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(
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<emphasis id="A7C1EA87DF698E15681931848822FA2E" box="[264,378,1479,1503]" italics="true" pageId="13" pageNumber="284">Sophiodela</emphasis>
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)
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<emphasis id="A7C1EA87DF698E15689831848B05FA2E" box="[393,605,1479,1503]" italics="true" pageId="13" pageNumber="284">chinensis okinawana</emphasis>
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<bibRefCitation id="F1244B64DF698E156B7231848A4CFA11" author="Nakane, T." box="[611,788,1479,1504]" pageId="13" pageNumber="284" pagination="41 - 43" refId="ref19657" refString="Nakane, T. (1957) New or little-known Coleoptera from Japan and its adjacent regions, XIV. The Scientific Report of the Saikyo University, 2 (4), 41 - 43." type="journal article" year="1957">Nakane, 1957:41</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="950A3695DF698E15698631A58BA0FA0F" blockId="13.[151,791,1479,1534]" box="[151,760,1509,1534]" pageId="13" pageNumber="284">
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<taxonomicName id="52B54D16DF698E15698631A58BACFA0F" ID-CoL="8M8PF" authority=": Fukuda et al. 2015: 273" authorityName="Fukuda" authorityPageNumber="273" authorityYear="2015" box="[151,756,1509,1534]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="284" phylum="Arthropoda" rank="species" species="okinawana" subGenus="Sophiodela">
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<emphasis id="A7C1EA87DF698E15698631A589A2FA0F" box="[151,250,1510,1534]" italics="true" pageId="13" pageNumber="284">Cicindela</emphasis>
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(
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<emphasis id="A7C1EA87DF698E15681931A58822FA0F" box="[264,378,1510,1534]" italics="true" pageId="13" pageNumber="284">Sophiodela</emphasis>
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)
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<emphasis id="A7C1EA87DF698E15689831A588A0FA0F" box="[393,504,1510,1534]" italics="true" pageId="13" pageNumber="284">okinawana</emphasis>
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:
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<bibRefCitation id="F1244B64DF698E156B1731A68BACFA0F" author="Fukuda, Y. & Yamasako, J. & Ogawa, R. & Sakai, M." box="[518,756,1509,1534]" pageId="13" pageNumber="284" pagination="269 - 280" refId="ref18294" refString="Fukuda, Y., Yamasako, J., Ogawa, R. & Sakai, M. (2015) Review of the Japanese species of the genus Cicindela (Sophiodela) (Coleoptera, Carabidae, Cicindelinae) based on the characters of internal sac. Elytra, New Series, 5 (2), 269 - 280." type="journal article" year="2015">
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Fukuda
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<emphasis id="A7C1EA87DF698E156B4931A48BDDFA0F" box="[600,645,1510,1534]" italics="true" pageId="13" pageNumber="284">et al</emphasis>
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. 2015:273
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</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="950A3695DF698E156986326F8844F99B" blockId="13.[151,1437,1580,2003]" pageId="13" pageNumber="284">
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<emphasis id="A7C1EA87DF698E156986326F88C0F9B6" bold="true" box="[151,408,1580,1607]" pageId="13" pageNumber="284">Specimens examined.</emphasis>
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(N = 1)
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<materialsCitation id="25DD3CC8DF698E1568EE326E8844F99B" ID-GBIF-Occurrence="2848419320" collectingDate="2015-03-17" collectorName="Y. Fukuda" country="Japan" location="Yonakunigami-village" municipality="Yona-Field" pageId="13" pageNumber="284" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa">
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<emphasis id="A7C1EA87DF698E1568EE326E8B02F9B6" bold="true" box="[511,602,1581,1607]" pageId="13" pageNumber="284">
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[
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<collectingCountry id="EDA27605DF698E156B17326E8B0CF9B6" box="[518,596,1581,1607]" name="Japan" pageId="13" pageNumber="284">Japan</collectingCountry>
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]
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</emphasis>
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<specimenCount id="83B3FD1CDF698E156B72326E8BDEF9B6" box="[611,646,1581,1607]" pageId="13" pageNumber="284" type="male">1♂</specimenCount>
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(
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<collectionCode id="F3A4AE50DF698E156B86326E8B93F9B6" box="[663,715,1581,1607]" pageId="13" pageNumber="284">YFJ</collectionCode>
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),
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<collectingMunicipality id="756EACEFDF698E156BCF326F8A06F9B7" box="[734,862,1580,1606]" pageId="13" pageNumber="284">Yona-Field</collectingMunicipality>
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,
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<location id="906A604EDF698E156A7B326F8D3EF9B7" LSID="urn:lsid:plazi:treatment:1D1C8783DF698E156986311C8B1CF823:906A604EDF698E156A7B326F8D3EF9B7" box="[874,1126,1580,1606]" country="Japan" municipality="Yona-Field" name="Yonakunigami-village" pageId="13" pageNumber="284" stateProvince="Okinawa">Yonakunigami-village</location>
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,
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<collectingRegion id="5771F877DF698E156D63326F8D82F9B7" box="[1138,1242,1580,1606]" country="Japan" name="Okinawa" pageId="13" pageNumber="284">Okinawa</collectingRegion>
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,
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<date id="E10B1055DF698E156DF6326E8C21F9B6" box="[1255,1401,1580,1607]" pageId="13" pageNumber="284" value="2015-03-17">
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<collectingDate id="F14FE9BDDF698E156DF6326E8C21F9B6" box="[1255,1401,1580,1607]" pageId="13" pageNumber="284" value="2015-03-17">17. III. 2015</collectingDate>
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</date>
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,
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<collectorName id="38405343DF698E156C94326E89B3F99B" pageId="13" pageNumber="284">Y. Fukuda</collectorName>
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leg.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="DDAF651EDF698E1569D632368B1CF823" pageId="13" pageNumber="284" type="description">
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<paragraph id="950A3695DF698E1569D632368A59F943" blockId="13.[151,1437,1580,2003]" pageId="13" pageNumber="284">
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<emphasis id="A7C1EA87DF698E1569D63236880BF97F" box="[199,339,1653,1678]" italics="true" pageId="13" pageNumber="284">Description.</emphasis>
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Internal sac everted dorsally in lateral view. DA largest in the bladders: DLL the second largest: VA as large as B: A exist: VLR, BLR and VB not exist.
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</paragraph>
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<paragraph id="950A3695DF698E1569D632FE8AE2F8B2" blockId="13.[151,1437,1580,2003]" pageId="13" pageNumber="284">VA short and roundly or triangular everted apically. DA everted basally. DA everted basally. B oval swollen right laterally, with well sclerotized spines, which are forming an oblong band. DLL swollen dorsally, covered numerously with well sclerotized spines, which are a forming band. Flagellum well short, curved and formed S shape. Medial tooth short, curved right ventro-laterally, tapered gently at apex.</paragraph>
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<paragraph id="950A3695DF698E1569D6330F8B1CF823" blockId="13.[151,1437,1580,2003]" pageId="13" pageNumber="284">
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<emphasis id="A7C1EA87DF698E1569D6330F8871F897" box="[199,297,1868,1894]" italics="true" pageId="13" pageNumber="284">Remarks</emphasis>
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. This species was treated as
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<taxonomicName id="52B54D16DF698E156B60330F8AA2F897" box="[625,1018,1868,1895]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="284" phylum="Arthropoda" rank="species" species="okinawana" subGenus="Sophiodela">
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<emphasis id="A7C1EA87DF698E156B60330F8B86F897" box="[625,734,1868,1894]" italics="true" pageId="13" pageNumber="284">Cicindela</emphasis>
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(
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<emphasis id="A7C1EA87DF698E156BE1330F8A36F897" box="[752,878,1868,1894]" italics="true" pageId="13" pageNumber="284">Sophiodela</emphasis>
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)
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<emphasis id="A7C1EA87DF698E156A6E330F8AA2F897" box="[895,1018,1868,1894]" italics="true" pageId="13" pageNumber="284">okinawana</emphasis>
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</taxonomicName>
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by
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<bibRefCitation id="F1244B64DF698E156D37330F8C48F896" author="Fukuda, Y. & Yamasako, J. & Ogawa, R. & Sakai, M." box="[1062,1296,1868,1895]" pageId="13" pageNumber="284" pagination="269 - 280" refId="ref18294" refString="Fukuda, Y., Yamasako, J., Ogawa, R. & Sakai, M. (2015) Review of the Japanese species of the genus Cicindela (Sophiodela) (Coleoptera, Carabidae, Cicindelinae) based on the characters of internal sac. Elytra, New Series, 5 (2), 269 - 280." type="journal article" year="2015">
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Fukuda
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<emphasis id="A7C1EA87DF698E156D93330E8DE3F897" box="[1154,1211,1868,1894]" italics="true" pageId="13" pageNumber="284">et al.</emphasis>
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(2015)
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</bibRefCitation>
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, but the formal taxonomic treatment had not been accepted. As mentioned earlier, this species is clearly distinguished from the congener species of
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<taxonomicName id="52B54D16DF698E1568B733D78B7CF85F" authorityName="Fukuda & Ogawa & Hori" authorityYear="2019" box="[422,548,1940,1966]" class="Insecta" family="Carabidae" genus="Sophiodela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="284" phylum="Arthropoda" rank="genus">
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<emphasis id="A7C1EA87DF698E1568B733D78B7CF85F" box="[422,548,1940,1966]" italics="true" pageId="13" pageNumber="284">Sophiodela</emphasis>
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</taxonomicName>
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by the structures of the internal sac. Therefore, we reassigned
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<taxonomicName id="52B54D16DF698E156DF933D68C27F85F" box="[1256,1407,1940,1967]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="284" phylum="Arthropoda" rank="species" species="okinawana">
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<emphasis id="A7C1EA87DF698E156DF933D68C27F85F" box="[1256,1407,1940,1967]" italics="true" pageId="13" pageNumber="284">S. okinawana</emphasis>
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</taxonomicName>
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in
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<taxonomicName id="52B54D16DF698E15698633FB884DF823" authorityName="Fukuda & Ogawa & Hori" authorityYear="2019" box="[151,277,1976,2002]" class="Insecta" family="Carabidae" genus="Sophiodela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="284" phylum="Arthropoda" rank="genus">
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<emphasis id="A7C1EA87DF698E15698633FB884DF823" box="[151,277,1976,2002]" italics="true" pageId="13" pageNumber="284">Sophiodela</emphasis>
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</taxonomicName>
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as an independent species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |