treatments-xml/data/03/C2/9A/03C29A73FFC24105FF4BA76A0477FD6D.xml
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<document id="0E913F434C79AE038C9902BB12A6A934" ID-CLB-Dataset="7088" ID-DOI="10.11646/zootaxa.4963.3.11" ID-GBIF-Dataset="37bbbcdd-7455-4fd2-854c-49f010b3b27c" ID-ISSN="1175-5326" ID-Zenodo-Dep="4704278" ID-ZooBank="05C2E47F-426D-42EF-B07E-D586C63B6527" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618904250788" checkinUser="plazi" docAuthor="Liu, Hsing-Che &amp; Fikáček, Martin" docDate="2021" docId="03C29A73FFC24105FF4BA76A0477FD6D" docLanguage="en" docName="zootaxa.4963.3.11.pdf" docOrigin="Zootaxa 4963 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laccobius (Microlaccobius) hoi Liu &amp; Fikacek 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="594" masterDocId="FFFBE20BFFC74102FFDCA234052DFFA0" masterDocTitle="Review of the Taiwanese Laccobius Erichson, 1837 (Coleoptera: Hydrophilidae) with description of a new species" masterLastPageNumber="599" masterPageNumber="587" pageNumber="592" updateTime="1698937256683" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DB73AB1727138DADC282F1525F9671FC">Review of the Taiwanese Laccobius Erichson, 1837 (Coleoptera: Hydrophilidae) with description of a new species</mods:title>
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<mods:namePart id="B29EA232F414DADB21C7035D630772EF">Liu, Hsing-Che</mods:namePart>
<mods:affiliation id="1AF6D133F30EE765F14399124F13187F">Taiwanese Society of Insect Natural History, No. 8, Wufeng Rd., Hsinchu County 310, Taiwan.</mods:affiliation>
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<mods:namePart id="B8A17496D5B5BB54B69C1E3AFCFD0F02">Fikáček, Martin</mods:namePart>
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<mods:affiliation id="34A761EFC486A71B781ABD86EAB364A0">Department of Biological Sciences, National Sun Yat-sen University, No. 70 Lien-hai Rd., Kaohsiung City 80424, Taiwan. &amp; Department of Entomology, National Museum, Cirkusová 1740, CZ- 19100 Praha 9 - Horní Počernice, Czech Republic. &amp; Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 12844 Praha 2, Czech Republic. mfikacek @ gmail. com; https: // orcid. org / 0000 - 0002 - 2078 - 6798</mods:affiliation>
<mods:nameIdentifier id="C844968F18F4A5FD5784F3C519AA4C7B" type="email">mfikacek@gmail.com</mods:nameIdentifier>
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<heading id="D09C9C09FFC24107FF4BA76A06DFFAD7" bold="true" box="[151,1010,1374,1401]" fontSize="11" level="1" pageId="5" pageNumber="592" reason="1">
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<taxonomicName id="4C6B50E6FFC24107FF4BA76A07F8FAD9" ID-CoL="8Y53L" authority="Liu &amp; Fikacek" authorityName="Liu &amp; Fikacek" authorityYear="2021" box="[151,725,1374,1401]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="592" phylum="Arthropoda" rank="species" species="hoi" status="sp. nov." subGenus="Microlaccobius">
<emphasis id="B91FF777FFC24107FF4BA76A043EFAD9" bold="true" box="[151,275,1374,1401]" italics="true" pageId="5" pageNumber="592">Laccobius</emphasis>
(
<emphasis id="B91FF777FFC24107FEF8A76A04F2FAD9" bold="true" box="[292,479,1374,1401]" italics="true" pageId="5" pageNumber="592">Microlaccobius</emphasis>
)
<emphasis id="B91FF777FFC24107FE2DA76A0735FAD9" bold="true" box="[497,536,1374,1401]" italics="true" pageId="5" pageNumber="592">hoi</emphasis>
Liu &amp; Fikáček
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何氏flǿ牙ḃ
</emphasis>
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(
<figureCitation id="135037E0FFC24107FF43A7B70429FA3D" box="[159,260,1410,1437]" captionStart="FIGURES 15" captionStartId="2.[151,264,1806,1830]" captionTargetBox="[151,1435,181,1728]" captionTargetId="figure-24@2.[151,1435,181,1782]" captionTargetPageId="2" captionText="FIGURES 15. Laccobius (Microlaccobius) hoi sp. nov., male. 12: habitus of the holotype (1: dorsal, 2: ventral). 35: aedeagus (3: dorsal view, holotype, 4: ventral view, holotype, 5: lateral view, paratype). Photos 12 by Uitsiann Ong." figureDoi="http://doi.org/10.5281/zenodo.4704282" httpUri="https://zenodo.org/record/4704282/files/figure.png" pageId="5" pageNumber="592">Figs 15</figureCitation>
,
<figureCitation id="135037E0FFC24107FECCA7B70403FA3D" box="[272,302,1411,1437]" captionStart="FIGURES 1922" captionStartId="6.[151,264,984,1008]" captionTargetBox="[165,1406,237,940]" captionTargetId="figure-24@6.[151,1436,181,960]" captionTargetPageId="6" captionText="FIGURES 1922. Pronotum of Taiwanese Laccobius (Microlaccobius) species. 19: L. formosus Gentili, 1979. 20: L. hammondi Gentili, 1984. 21: L. hoi sp. nov. 22: L. roseiceps Régimbart, 1903." figureDoi="http://doi.org/10.5281/zenodo.4704295" httpUri="https://zenodo.org/record/4704295/files/figure.png" pageId="5" pageNumber="592">21</figureCitation>
,
<figureCitation id="135037E0FFC24107FEE6A7B6047BFA3D" box="[314,342,1410,1437]" captionStart="FIGURES 2326" captionStartId="8.[151,264,1431,1455]" captionTargetBox="[184,1404,189,1402]" captionTargetId="figure-24@8.[177,1411,181,1408]" captionTargetPageId="8" captionText="FIGURES 2326. Habitat of Taiwanese Laccobius species. 23: Nantou County, Puli: habitat of L. formosus. 24: Taichung City, Wufeng: habitat of L. hammondi and L. hoi sp. nov. 25: Pingtung County, Ligang: habitat of L. formosus, L. hoi sp. nov. and L. roseiceps. 26: New Taipei City, Pingxi: habitat of L. politus." figureDoi="http://doi.org/10.5281/zenodo.4704299" httpUri="https://zenodo.org/record/4704299/files/figure.png" pageId="5" pageNumber="592">25</figureCitation>
)
</paragraph>
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<typeStatus id="54D095C7FFC24107FF4BA7FF05FFFA45" box="[151,210,1483,1509]" pageId="5" pageNumber="592">Type</typeStatus>
locality.
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,
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,
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<emphasis id="B91FF777FFC24107FCB5A7FF06E6FA42" bold="true" box="[873,971,1483,1506]" pageId="5" pageNumber="592">[¶ffiė</emphasis>
],
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[
<emphasis id="B91FF777FFC24107FB73A7FF002EFA42" bold="true" box="[1199,1283,1483,1506]" pageId="5" pageNumber="592">二AEë</emphasis>
],
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a. s. l.
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</paragraph>
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<emphasis id="B91FF777FFC24107FF1BA4270459F98C" bold="true" box="[199,372,1554,1581]" pageId="5" pageNumber="592">Type material.</emphasis>
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<typeStatus id="54D095C7FFC24107FEA6A42704C6F98D" box="[378,491,1555,1581]" pageId="5" pageNumber="592" type="holotype">Holotype</typeStatus>
:
</emphasis>
male (
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):
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:
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:
</emphasis>
<collectingMunicipality id="6BB0B11FFFC24107FC7FA4270144F98D" box="[931,1129,1555,1581]" pageId="5" pageNumber="592">Ligang Township</collectingMunicipality>
,
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,
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,
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,
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<collectingDate id="EF91F44DFFC24107FEB4A40304CAF9F1" box="[360,487,1591,1617]" pageId="5" pageNumber="592" value="2020-04-26">26.IV.2020</collectingDate>
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.
<collectorName id="269E4EB3FFC24107FE2DA40307BAF9F1" box="[497,663,1591,1617]" pageId="5" pageNumber="592">Hsing-Che Liu</collectorName>
leg.
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<typeStatus id="54D095C7FFC24107FD11A4030664F9F1" box="[717,841,1591,1617]" pageId="5" pageNumber="592" type="paratype">Paratypes</typeStatus>
:
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:
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:
</emphasis>
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(
<collectionCode id="ED7AB3A0FFC24107FB71A40301D2F9F1" box="[1197,1279,1591,1617]" pageId="5" pageNumber="592">HCLC</collectionCode>
): same data as the holotype
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.
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<collectingRegion id="49AFE587FFC24107FEF3A46E0453F9D3" box="[303,382,1626,1652]" country="Taiwan" name="Chiayi" pageId="5" pageNumber="592">CHIAYI</collectingRegion>
:
</emphasis>
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,
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(
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):
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[
<emphasis id="B91FF777FFC24107FCA1A46F06FCF9D2" bold="true" box="[893,977,1627,1650]" pageId="5" pageNumber="592">k埔ė</emphasis>
],
<location id="8EB47DBEFFC24107FC3BA46F01A1F9D5" LSID="urn:lsid:plazi:treatment:03C29A73FFC24105FF4BA76A0477FD6D:8EB47DBEFFC24107FC3BA46F01A1F9D5" box="[999,1164,1627,1653]" country="Taiwan" municipality="Dapu Township" name="Zengwen Dam" pageId="5" pageNumber="592" stateProvince="Chiayi">Zengwen Dam</location>
[
<emphasis id="B91FF777FFC24107FB40A46F0021F9D2" bold="true" box="[1180,1292,1627,1650]" pageId="5" pageNumber="592">曾Ý水庫</emphasis>
],
<date id="FFD50DA5FFC24107FAFFA46F00B5F9D5" box="[1315,1432,1627,1653]" pageId="5" pageNumber="592" value="2020-05-17">
<collectingDate id="EF91F44DFFC24107FAFFA46F00B5F9D5" box="[1315,1432,1627,1653]" pageId="5" pageNumber="592" value="2020-05-17">17.V.2020</collectingDate>
</date>
.
<collectorName id="269E4EB3FFC24107FF4BA44B0428F939" box="[151,261,1663,1689]" pageId="5" pageNumber="592">Kunta Ho</collectorName>
leg.
</materialsCitation>
by light trap.
<materialsCitation id="3B032138FFC24107FE0BA44B0796F91D" ID-GBIF-Occurrence="3107229308" collectingDate="2020-06-24" collectionCode="HCLC" collectorName="Hsing-Che Liu &amp; Uitsiann Ong" country="Taiwan" location="Laonong River" municipality="Liouguei District" pageId="5" pageNumber="592" specimenCount="1" specimenCount-female="1" stateProvince="Kaohsiung" typeStatus="paratype">
<emphasis id="B91FF777FFC24107FE0BA44B074CF939" bold="true" box="[471,609,1663,1689]" pageId="5" pageNumber="592">
<collectingRegion id="49AFE587FFC24107FE0BA44B0775F937" box="[471,600,1663,1689]" country="Taiwan" name="Kaohsiung" pageId="5" pageNumber="592">KAOHSIUNG</collectingRegion>
:
</emphasis>
<specimenCount id="9D6DE0ECFFC24107FDB5A44B07E6F939" box="[617,715,1663,1689]" pageId="5" pageNumber="592" type="female">1 female</specimenCount>
(
<collectionCode id="ED7AB3A0FFC24107FD07A44B0600F939" box="[731,813,1663,1689]" pageId="5" pageNumber="592">HCLC</collectionCode>
):
<collectingMunicipality id="6BB0B11FFFC24107FCE3A44B012DF939" box="[831,1024,1663,1689]" pageId="5" pageNumber="592">Liouguei District</collectingMunicipality>
[
<emphasis id="B91FF777FFC24107FBCDA44B0148F936" bold="true" box="[1041,1125,1663,1686]" pageId="5" pageNumber="592">六AE區</emphasis>
],
<location id="8EB47DBEFFC24107FBA1A44B000AF939" LSID="urn:lsid:plazi:treatment:03C29A73FFC24105FF4BA76A0477FD6D:8EB47DBEFFC24107FBA1A44B000AF939" box="[1149,1319,1663,1689]" country="Taiwan" municipality="Liouguei District" name="Laonong River" pageId="5" pageNumber="592" stateProvince="Kaohsiung">Laonong River</location>
[
<emphasis id="B91FF777FFC24107FAE4A44B00A1F936" bold="true" box="[1336,1420,1663,1686]" pageId="5" pageNumber="592">Ž濃ë</emphasis>
],
<date id="FFD50DA5FFC24107FF4BA4970437F91D" box="[151,282,1699,1725]" pageId="5" pageNumber="592" value="2020-06-24">
<collectingDate id="EF91F44DFFC24107FF4BA4970437F91D" box="[151,282,1699,1725]" pageId="5" pageNumber="592" value="2020-06-24">24.VI.2020</collectingDate>
</date>
.
<collectorName id="269E4EB3FFC24107FEF9A49704E0F91D" box="[293,461,1699,1725]" pageId="5" pageNumber="592">Hsing-Che Liu</collectorName>
&amp;
<collectorName id="269E4EB3FFC24107FE2FA49707A4F91D" box="[499,649,1699,1725]" pageId="5" pageNumber="592">Uitsiann Ong</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B032138FFC24107FD1EA4970442F941" ID-GBIF-Occurrence="3107229312" collectingDate="2020-05-15" collectionCode="TARI" collectorName="Hsing-Che Liu" country="Taiwan" location="Wufeng District" pageId="5" pageNumber="592" specimenCount="3" stateProvince="Taichung" typeStatus="paratype">
<emphasis id="B91FF777FFC24107FD1EA4970617F91D" bold="true" box="[706,826,1699,1725]" pageId="5" pageNumber="592">
<collectingRegion id="49AFE587FFC24107FD1EA497061CF91B" box="[706,817,1699,1725]" country="Taiwan" name="Taichung" pageId="5" pageNumber="592">TAICHUNG</collectingRegion>
:
</emphasis>
<specimenCount id="9D6DE0ECFFC24107FC9EA49706BCF91D" box="[834,913,1699,1725]" pageId="5" pageNumber="592" type="generic">3 spec.</specimenCount>
(
<collectionCode id="ED7AB3A0FFC24107FC43A49706CFF91D" box="[927,994,1699,1725]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="5" pageNumber="592">TARI</collectionCode>
)
<location id="8EB47DBEFFC24107FC2DA497018BF91D" LSID="urn:lsid:plazi:treatment:03C29A73FFC24105FF4BA76A0477FD6D:8EB47DBEFFC24107FC2DA497018BF91D" box="[1009,1190,1699,1725]" country="Taiwan" name="Wufeng District" pageId="5" pageNumber="592" stateProvince="Taichung">Wufeng District</location>
[
<emphasis id="B91FF777FFC24107FB6BA4970026F91A" bold="true" box="[1207,1291,1699,1722]" pageId="5" pageNumber="592">Ȃ峰區</emphasis>
],
<date id="FFD50DA5FFC24107FAFFA49600B5F91D" box="[1315,1432,1698,1725]" pageId="5" pageNumber="592" value="2020-05-15">
<collectingDate id="EF91F44DFFC24107FAFFA49600B5F91D" box="[1315,1432,1698,1725]" pageId="5" pageNumber="592" value="2020-05-15">15.V.2020</collectingDate>
</date>
.
<collectorName id="269E4EB3FFC24107FF4BA4F30412F941" box="[151,319,1735,1761]" pageId="5" pageNumber="592">Hsing-Che Liu</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37178EEFFC24104FF1BA4DE0704F961" lastPageId="6" lastPageNumber="593" pageId="5" pageNumber="592" type="description">
<paragraph id="8BD42B65FFC24107FF1BA4DE046CF87D" blockId="5.[151,1437,1482,2013]" pageId="5" pageNumber="592">
<emphasis id="B91FF777FFC24107FF1BA4DE0734F8A5" bold="true" box="[199,537,1770,1797]" pageId="5" pageNumber="592">
Description.
<emphasis id="B91FF777FFC24107FE83A4DF0734F8A5" bold="true" box="[351,537,1770,1797]" italics="true" pageId="5" pageNumber="592">Form and Color</emphasis>
</emphasis>
(
<figureCitation id="135037E0FFC24107FDF4A4DF07BEF8A5" box="[552,659,1771,1797]" captionStart="FIGURES 15" captionStartId="2.[151,264,1806,1830]" captionTargetBox="[151,1435,181,1728]" captionTargetId="figure-24@2.[151,1435,181,1782]" captionTargetPageId="2" captionText="FIGURES 15. Laccobius (Microlaccobius) hoi sp. nov., male. 12: habitus of the holotype (1: dorsal, 2: ventral). 35: aedeagus (3: dorsal view, holotype, 4: ventral view, holotype, 5: lateral view, paratype). Photos 12 by Uitsiann Ong." figureDoi="http://doi.org/10.5281/zenodo.4704282" httpUri="https://zenodo.org/record/4704282/files/figure.png" pageId="5" pageNumber="592">Figs. 12</figureCitation>
). Body length
<quantity id="4C938680FFC24107FCEAA4DE0691F8A5" box="[822,956,1770,1797]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="5" pageNumber="592" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.52.7 mm</quantity>
(
<typeStatus id="54D095C7FFC24107FC16A4DF011FF8A5" box="[970,1074,1771,1797]" pageId="5" pageNumber="592" type="holotype">holotype</typeStatus>
:
<quantity id="4C938680FFC24107FBE7A4DF01B9F8A5" box="[1083,1172,1771,1797]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="592" unit="mm" value="2.6">2.6 mm</quantity>
), maximum body width
<quantity id="4C938680FFC24107FF4BA53B0432F889" box="[151,287,1807,1833]" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="5" pageNumber="592" unit="mm" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1.71.9 mm</quantity>
(
<typeStatus id="54D095C7FFC24107FEF1A53B04B8F889" box="[301,405,1807,1833]" pageId="5" pageNumber="592" type="holotype">holotype</typeStatus>
:
<quantity id="4C938680FFC24107FE7CA53B04D6F889" box="[416,507,1807,1833]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="592" unit="mm" value="1.9">1.9 mm</quantity>
). Body oval, moderately convex. Maxillary palpus light brownish yellow, blackish at the very apex of the last palpomere. Antennae yellowish to brown, antennal club slightly darker. Head black, with distinct preocular spots. Labrum and mentum black. Pronotum pale, with irregularly shaped central dark spot ca. as wide as three-fifths of pronotum width (
<figureCitation id="135037E0FFC24107FD8BA54F0787F835" box="[599,682,1915,1941]" captionStart="FIGURES 1922" captionStartId="6.[151,264,984,1008]" captionTargetBox="[165,1406,237,940]" captionTargetId="figure-24@6.[151,1436,181,960]" captionTargetPageId="6" captionText="FIGURES 1922. Pronotum of Taiwanese Laccobius (Microlaccobius) species. 19: L. formosus Gentili, 1979. 20: L. hammondi Gentili, 1984. 21: L. hoi sp. nov. 22: L. roseiceps Régimbart, 1903." figureDoi="http://doi.org/10.5281/zenodo.4704295" httpUri="https://zenodo.org/record/4704295/files/figure.png" pageId="5" pageNumber="592">Fig. 21</figureCitation>
). Elytra pale, with variable pattern of slightly irregular longitudinal series consisting of black spots. Whole scutellar shield black. Ventral surface black with yellowish pubescence. Leg pale yellowish.
</paragraph>
<caption id="DF147BEDFFC14104FF4BA1EC0613FBB5" ID-DOI="http://doi.org/10.5281/zenodo.4704295" ID-Zenodo-Dep="4704295" httpUri="https://zenodo.org/record/4704295/files/figure.png" pageId="6" pageNumber="593" startId="6.[151,264,984,1008]" targetBox="[165,1406,237,940]" targetPageId="6">
<paragraph id="8BD42B65FFC14104FF4BA1EC0613FBB5" blockId="6.[151,1436,984,1045]" pageId="6" pageNumber="593">
<emphasis id="B91FF777FFC14104FF4BA1EC047EFC51" bold="true" box="[151,339,984,1009]" pageId="6" pageNumber="593">FIGURES 1922.</emphasis>
Pronotum of Taiwanese
<taxonomicName id="4C6B50E6FFC14104FD97A1ED064BFC51" authority="" box="[587,870,985,1009]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="593" phylum="Arthropoda" rank="subGenus" subGenus="Microlaccobius">
<emphasis id="B91FF777FFC14104FD97A1ED079EFC51" box="[587,691,985,1009]" italics="true" pageId="6" pageNumber="593">Laccobius</emphasis>
(
<emphasis id="B91FF777FFC14104FD1CA1ED0673FC51" box="[704,862,985,1009]" italics="true" pageId="6" pageNumber="593">Microlaccobius</emphasis>
)
</taxonomicName>
species. 19:
<taxonomicName id="4C6B50E6FFC14104FC3AA1ED01C7FC51" authority="Gentili, 1979" authorityName="Gentili" authorityYear="1979" box="[998,1258,985,1009]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="593" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B91FF777FFC14104FC3AA1ED0170FC51" box="[998,1117,985,1009]" italics="true" pageId="6" pageNumber="593">L. formosus</emphasis>
<bibRefCitation id="EFFA5694FFC14104FBBEA1ED01C7FC51" author="Gentili, E." box="[1122,1258,985,1009]" pageId="6" pageNumber="593" pagination="27 - 50" refId="ref7501" refString="Gentili, E. (1979) I Laccobius della regione orientale (Coleoptera, Hydrophilidae). Annuario Osservatorio di Fisica Terrestre e Museo Antonio Stoppani del Seminario Arcivescoville di Milano (New Series), 1, 27 - 50." type="journal article" year="1979">Gentili, 1979</bibRefCitation>
</taxonomicName>
. 20:
<taxonomicName id="4C6B50E6FFC14104FAC4A1ED040DFBB5" authority="Gentili, 1984" authorityName="Gentili" authorityYear="1984" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="593" phylum="Arthropoda" rank="species" species="hammondi">
<emphasis id="B91FF777FFC14104FAC4A1ED00B1FC51" box="[1304,1436,985,1009]" italics="true" pageId="6" pageNumber="593">L. hammondi</emphasis>
Gentili, 1984
</taxonomicName>
. 21:
<taxonomicName id="4C6B50E6FFC14104FE8CA1C904A6FBB5" authority="Liu &amp; Fikáček, 2021" authorityName="Liu &amp; Fikáček" authorityYear="2021" box="[336,395,1021,1045]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="593" phylum="Arthropoda" rank="species" species="hoi" status="sp. nov.">
<emphasis id="B91FF777FFC14104FE8CA1C904A6FBB5" box="[336,395,1021,1045]" italics="true" pageId="6" pageNumber="593">L. hoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C4A0CFFC14104FE4EA1C904CCFBB4" box="[402,481,1021,1044]" pageId="6" pageNumber="593" rank="species">sp. nov.</taxonomicNameLabel>
22:
<taxonomicName id="4C6B50E6FFC14104FDD1A1C90617FBB5" authority="Regimbart, 1903" authorityName="Regimbart" authorityYear="1903" box="[525,826,1021,1045]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="593" phylum="Arthropoda" rank="species" species="roseiceps">
<emphasis id="B91FF777FFC14104FDD1A1C907ABFBB4" box="[525,646,1021,1044]" italics="true" pageId="6" pageNumber="593">L. roseiceps</emphasis>
Régimbart, 1903
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BD42B65FFC14104FF1BA676073FFAB1" blockId="6.[151,1437,1090,2017]" pageId="6" pageNumber="593">
<emphasis id="B91FF777FFC14104FF1BA6760421FBFD" bold="true" box="[199,268,1090,1117]" italics="true" pageId="6" pageNumber="593">Head.</emphasis>
Surface densely punctate, sagittal suture indistinct. Eyes moderately large, somewhat protruding, separated by ca. 2.0× the width of one eye. Labrum coarsely punctate, anterior margin not emarginate. Mentum glabrous, ca. 1.21.3× as wide as long. Antennae with 8 antennomeres, longer than maxillary palpus, intermediate segments (antennomeres III and IV) very short; each club antennomere of different shape, closely segmented, with long or short pubescence; antennomere VI largest, antennomere VII semicircular in shape, antennomere VIII longer than wide, tapering towards apex.
</paragraph>
<paragraph id="8BD42B65FFC14104FF1BA72E0443FA01" blockId="6.[151,1437,1090,2017]" pageId="6" pageNumber="593">
<emphasis id="B91FF777FFC14104FF1BA72E040EFA95" bold="true" box="[199,291,1306,1333]" italics="true" pageId="6" pageNumber="593">Thorax.</emphasis>
Pronotum ca. 2.6× wider than long; surface rough, loosely punctate. Surface of mesoventrite densely pubescent. Metanepisterna ca. 5.3× as long as wide, parallel-sided; metaventrite densely pubescent except along midline, slightly raised. Scutellar shield in form of equilateral triangle. Elytral surface with slightly irregular rows of serial punctures.
</paragraph>
<paragraph id="8BD42B65FFC14104FF1BA79F061EF9AD" blockId="6.[151,1437,1090,2017]" pageId="6" pageNumber="593">
<emphasis id="B91FF777FFC14104FF1BA79F0428FA65" bold="true" box="[199,261,1451,1477]" italics="true" pageId="6" pageNumber="593">Legs.</emphasis>
Profemora densely pubescent basally; protibiae with longest stiff setae situated in apical third. Mesofemora almost bare, with sparse punctation; mesotibiae with longitudinal rows of stiff setae. Metafemora smooth, wider than mesofemora; metatibiae slender, slightly bent inwards.
</paragraph>
<paragraph id="8BD42B65FFC14104FF1BA4220134F991" blockId="6.[151,1437,1090,2017]" box="[199,1049,1558,1585]" pageId="6" pageNumber="593">
<emphasis id="B91FF777FFC14104FF1BA4220411F991" bold="true" box="[199,316,1558,1585]" italics="true" pageId="6" pageNumber="593">Abdomen.</emphasis>
Ventrites IV sparsely pubescent; Ventrite VI densely pubescent.
</paragraph>
<paragraph id="8BD42B65FFC14104FF1BA40E0704F961" blockId="6.[151,1437,1090,2017]" pageId="6" pageNumber="593">
<emphasis id="B91FF777FFC14104FF1BA40E0443F9F5" bold="true" box="[199,366,1594,1621]" italics="true" pageId="6" pageNumber="593">Male genitalia</emphasis>
(
<figureCitation id="135037E0FFC14104FEA2A40F04C6F9F6" box="[382,491,1595,1622]" captionStart="FIGURES 15" captionStartId="2.[151,264,1806,1830]" captionTargetBox="[151,1435,181,1728]" captionTargetId="figure-24@2.[151,1435,181,1782]" captionTargetPageId="2" captionText="FIGURES 15. Laccobius (Microlaccobius) hoi sp. nov., male. 12: habitus of the holotype (1: dorsal, 2: ventral). 35: aedeagus (3: dorsal view, holotype, 4: ventral view, holotype, 5: lateral view, paratype). Photos 12 by Uitsiann Ong." figureDoi="http://doi.org/10.5281/zenodo.4704282" httpUri="https://zenodo.org/record/4704282/files/figure.png" pageId="6" pageNumber="593">Figs. 35</figureCitation>
). Aedeagus
<quantity id="4C938680FFC14104FDAFA40F07E4F9F5" box="[627,713,1595,1621]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="593" unit="mm" value="0.6">0.6 mm</quantity>
long, phallobase slightly narrowed and truncate basally. Median lobe very broad, slightly shorter than parameres, narrowing and membranous at apex, surface with lateral deep rugae. Parameres flattened laterally, very narrow and slightly bent inwards in dorsal and ventral views, strongly narrowing and acute in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C37178EEFFC14104FF1BA4FF06C3F839" pageId="6" pageNumber="593" type="diagnosis">
<paragraph id="8BD42B65FFC14104FF1BA4FF06C3F839" blockId="6.[151,1437,1090,2017]" pageId="6" pageNumber="593">
<emphasis id="B91FF777FFC14104FF1BA4FF04E1F945" bold="true" box="[199,460,1739,1765]" pageId="6" pageNumber="593">Differential diagnosis.</emphasis>
This species is very similar
<taxonomicName id="4C6B50E6FFC14104FCDFA4FF01AAF946" authority="Gentili, 2005" authorityName="Gentili" authorityYear="2005" box="[771,1159,1739,1766]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="593" phylum="Arthropoda" rank="species" species="philipinus">
<emphasis id="B91FF777FFC14104FCDFA4FF06C1F945" box="[771,1004,1739,1765]" italics="true" pageId="6" pageNumber="593">Laccobius philipinus</emphasis>
<bibRefCitation id="EFFA5694FFC14104FC2EA4FF01AAF946" author="Gentili, E." box="[1010,1159,1739,1766]" pageId="6" pageNumber="593" pagination="317 - 370" refId="ref7731" refString="Gentili, E. (2005) The genus Laccobius Erichson, 1837 in the Australian region (Coleoptera, Hydrophilidae). In: Daccordi, M. &amp; Giachino, P. M. (Eds.), Results of the zoological missions to Australia of the Regional Museum of Natural Sciences of Turin, Italy. Volume II. Monografie del Museo Regionale di Scienze Naturali Torino, 42, 317 - 370." type="journal article" year="2005">Gentili, 2005</bibRefCitation>
</taxonomicName>
, but can be distinguished from it by the pronotal dark spot almost as wide as the head (narrower than the head in
<taxonomicName id="4C6B50E6FFC14104FB85A4C401C8F8A9" authorityName="Gentili" authorityYear="2005" box="[1113,1253,1775,1801]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="593" phylum="Arthropoda" rank="species" species="philipinus">
<emphasis id="B91FF777FFC14104FB85A4C401C8F8A9" box="[1113,1253,1775,1801]" italics="true" pageId="6" pageNumber="593">L. philipinus</emphasis>
</taxonomicName>
;
<figureCitation id="135037E0FFC14104FB2FA4DB0018F8AA" box="[1267,1333,1775,1802]" captionStart="FIGURES 69" captionStartId="3.[151,264,1305,1329]" captionTargetBox="[192,1387,511,1264]" captionTargetId="figure-131@3.[177,1411,489,1281]" captionTargetPageId="3" captionText="FIGURES 69. Laccobius (Microlaccobius) philipinus Gentili, 2005. 6: habitus of the holotype, dorsal view. 79: aedeagus of the holotype (7: dorsal view, 8: lateral view, 9: ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4704287" httpUri="https://zenodo.org/record/4704287/files/figure.png" pageId="6" pageNumber="593">Fig. 6</figureCitation>
), median lobe very wide and almost of the same width throughout except at apex (median lobe moderately wide, indistinctly constricted subapically in
<taxonomicName id="4C6B50E6FFC14104FE65A50C076BF8F1" authorityName="Gentili" authorityYear="2005" box="[441,582,1847,1873]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="593" phylum="Arthropoda" rank="species" species="philipinus">
<emphasis id="B91FF777FFC14104FE65A50C076BF8F1" box="[441,582,1847,1873]" italics="true" pageId="6" pageNumber="593">L. philipinus</emphasis>
</taxonomicName>
), parameres very narrow, bent inwards, rounded at apex in dorsal and ventral views (parameres moderately wide, straight and pointed at apex in dorsal and ventral views in
<taxonomicName id="4C6B50E6FFC14104FB12A5680070F8D5" authorityName="Gentili" authorityYear="2005" box="[1230,1373,1883,1909]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="593" phylum="Arthropoda" rank="species" species="philipinus">
<emphasis id="B91FF777FFC14104FB12A5680070F8D5" box="[1230,1373,1883,1909]" italics="true" pageId="6" pageNumber="593">L. philipinus</emphasis>
</taxonomicName>
). For comparison with the aedeagus of the
<typeStatus id="54D095C7FFC14104FDE4A54B07B7F839" box="[568,666,1919,1945]" pageId="6" pageNumber="593" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C6B50E6FFC14104FD63A5B40660F839" authorityName="Gentili" authorityYear="2005" box="[703,845,1919,1945]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="593" phylum="Arthropoda" rank="species" species="philipinus">
<emphasis id="B91FF777FFC14104FD63A5B40660F839" box="[703,845,1919,1945]" italics="true" pageId="6" pageNumber="593">L. philipinus</emphasis>
</taxonomicName>
, see
<figureCitation id="135037E0FFC14104FC5AA54B06C7F839" box="[902,1002,1919,1945]" captionStart="FIGURES 69" captionStartId="3.[151,264,1305,1329]" captionTargetBox="[192,1387,511,1264]" captionTargetId="figure-131@3.[177,1411,489,1281]" captionTargetPageId="3" captionText="FIGURES 69. Laccobius (Microlaccobius) philipinus Gentili, 2005. 6: habitus of the holotype, dorsal view. 79: aedeagus of the holotype (7: dorsal view, 8: lateral view, 9: ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4704287" httpUri="https://zenodo.org/record/4704287/files/figure.png" pageId="6" pageNumber="593">Figs 79</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37178EEFFC14104FF1BA59705DCF841" pageId="6" pageNumber="593" type="etymology">
<paragraph id="8BD42B65FFC14104FF1BA59705DCF841" blockId="6.[151,1437,1090,2017]" pageId="6" pageNumber="593">
<emphasis id="B91FF777FFC14104FF1BA5970465F81D" bold="true" box="[199,328,1955,1981]" pageId="6" pageNumber="593">Etymology</emphasis>
. The species is named after Mr. Kunta Ho who first discovered this species in
<collectingRegion id="49AFE587FFC14104FB00A59700B5F81D" box="[1244,1432,1955,1981]" country="Taiwan" name="Pingtung" pageId="6" pageNumber="593">Pingtung county</collectingRegion>
,
<collectingCountry id="F37C6BF5FFC14104FF4BA5F305C0F841" box="[151,237,1991,2017]" name="Taiwan" pageId="6" pageNumber="593">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C37178EEFFC04105FF1BA2A201A0FEC5" pageId="7" pageNumber="594" type="distribution">
<paragraph id="8BD42B65FFC04105FF1BA2A205DAFF75" blockId="7.[151,1437,150,717]" pageId="7" pageNumber="594">
<emphasis id="B91FF777FFC04105FF1BA2A2044CFF10" bold="true" box="[199,353,150,176]" pageId="7" pageNumber="594">Distribution.</emphasis>
This species is known from central and southern
<collectingCountry id="F37C6BF5FFC04105FCA2A2A306FDFF11" box="[894,976,151,177]" name="Taiwan" pageId="7" pageNumber="594">Taiwan</collectingCountry>
(
<collectingRegion id="49AFE587FFC04105FC01A2A30101FF11" box="[989,1068,151,177]" country="Taiwan" name="Chiayi" pageId="7" pageNumber="594">Chiayi</collectingRegion>
,
<collectingRegion id="49AFE587FFC04105FBEAA2A3019FFF11" box="[1078,1202,151,177]" country="Taiwan" name="Kaohsiung" pageId="7" pageNumber="594">Kaohsiung</collectingRegion>
,
<collectingRegion id="49AFE587FFC04105FB60A2A30009FF11" box="[1212,1316,151,177]" country="Taiwan" name="Pingtung" pageId="7" pageNumber="594">Pingtung</collectingRegion>
,
<collectingRegion id="49AFE587FFC04105FAF1A2A300B4FF11" box="[1325,1433,151,177]" country="Taiwan" name="Taichung" pageId="7" pageNumber="594">Taichung</collectingRegion>
;
<figureCitation id="135037E0FFC04105FF4BA28F05C6FF75" box="[151,235,187,213]" captionStart="FIGURES 27" captionStartId="10.[151,264,1944,1968]" captionTargetBox="[151,1431,309,1911]" captionTargetId="figure-86@10.[151,1435,309,1921]" captionTargetPageId="10" captionText="FIGURES 27. Map of Taiwan showing the distribution of the Laccobius species." figureDoi="http://doi.org/10.5281/zenodo.4704305" httpUri="https://zenodo.org/record/4704305/files/figure.png" pageId="7" pageNumber="594">Fig. 27</figureCitation>
).
</paragraph>
<paragraph id="8BD42B65FFC04105FF1BA2EA01A0FEC5" blockId="7.[151,1437,150,717]" pageId="7" pageNumber="594">
<emphasis id="B91FF777FFC04105FF1BA2EA0466FF58" bold="true" box="[199,331,222,248]" pageId="7" pageNumber="594">Bionomics.</emphasis>
Specimens from
<collectingRegion id="49AFE587FFC04105FDCFA2EB0755FF59" box="[531,632,223,249]" country="Taiwan" name="Pingtung" pageId="7" pageNumber="594">Pingtung</collectingRegion>
and
<collectingRegion id="49AFE587FFC04105FD6DA2EB0607FF59" box="[689,810,223,249]" country="Taiwan" name="Kaohsiung" pageId="7" pageNumber="594">Kaohsiung</collectingRegion>
were collected from a lowland stream and co-occurred syntopically with
<taxonomicName id="4C6B50E6FFC04105FEBEA33704CAFEBD" authorityName="Gentili" authorityYear="1979" box="[354,487,259,285]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B91FF777FFC04105FEBEA33704CAFEBD" box="[354,487,259,285]" italics="true" pageId="7" pageNumber="594">L. formosus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B50E6FFC04105FDC3A337078BFEBC" authorityName="Regimbart" authorityYear="1903" box="[543,678,259,284]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="roseiceps">
<emphasis id="B91FF777FFC04105FDC3A337078BFEBC" box="[543,678,259,284]" italics="true" pageId="7" pageNumber="594">L. roseiceps</emphasis>
</taxonomicName>
(
<figureCitation id="135037E0FFC04105FD6AA3370626FEBD" box="[694,779,258,285]" captionStart="FIGURES 2326" captionStartId="8.[151,264,1431,1455]" captionTargetBox="[184,1404,189,1402]" captionTargetId="figure-24@8.[177,1411,181,1408]" captionTargetPageId="8" captionText="FIGURES 2326. Habitat of Taiwanese Laccobius species. 23: Nantou County, Puli: habitat of L. formosus. 24: Taichung City, Wufeng: habitat of L. hammondi and L. hoi sp. nov. 25: Pingtung County, Ligang: habitat of L. formosus, L. hoi sp. nov. and L. roseiceps. 26: New Taipei City, Pingxi: habitat of L. politus." figureDoi="http://doi.org/10.5281/zenodo.4704299" httpUri="https://zenodo.org/record/4704299/files/figure.png" pageId="7" pageNumber="594">Fig. 25</figureCitation>
). Specimens from
<collectingRegion id="49AFE587FFC04105FC03A3370107FEBD" box="[991,1066,259,285]" country="Taiwan" name="Chiayi" pageId="7" pageNumber="594">Chiayi</collectingRegion>
were collected in a light trap set near a water reservoir. Specimens from
<collectingRegion id="49AFE587FFC04105FD8AA3130793FEE1" box="[598,702,295,321]" country="Taiwan" name="Taichung" pageId="7" pageNumber="594">Taichung</collectingRegion>
were collected from a small stream and co-occurred syntopically with
<taxonomicName id="4C6B50E6FFC04105FF0CA37F0449FEC5" authorityName="Gentili" authorityYear="1984" box="[208,356,331,357]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="hammondi">
<emphasis id="B91FF777FFC04105FF0CA37F0449FEC5" box="[208,356,331,357]" italics="true" pageId="7" pageNumber="594">L. hammondi</emphasis>
</taxonomicName>
(
<figureCitation id="135037E0FFC04105FEAFA37F04EBFEC5" box="[371,454,331,357]" captionStart="FIGURES 2326" captionStartId="8.[151,264,1431,1455]" captionTargetBox="[184,1404,189,1402]" captionTargetId="figure-24@8.[177,1411,181,1408]" captionTargetPageId="8" captionText="FIGURES 2326. Habitat of Taiwanese Laccobius species. 23: Nantou County, Puli: habitat of L. formosus. 24: Taichung City, Wufeng: habitat of L. hammondi and L. hoi sp. nov. 25: Pingtung County, Ligang: habitat of L. formosus, L. hoi sp. nov. and L. roseiceps. 26: New Taipei City, Pingxi: habitat of L. politus." figureDoi="http://doi.org/10.5281/zenodo.4704299" httpUri="https://zenodo.org/record/4704299/files/figure.png" pageId="7" pageNumber="594">Fig. 24</figureCitation>
), but
<taxonomicName id="4C6B50E6FFC04105FDDAA37F076DFEC5" authority="Liu &amp; Fikáček, 2021" authorityName="Liu &amp; Fikáček" authorityYear="2021" box="[518,576,331,357]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="hoi" status="sp. nov.">
<emphasis id="B91FF777FFC04105FDDAA37F076DFEC5" box="[518,576,331,357]" italics="true" pageId="7" pageNumber="594">L hoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C4A0CFFC04105FD9AA37F07B0FEC5" box="[582,669,331,357]" pageId="7" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
was much less abundant than
<emphasis id="B91FF777FFC04105FC2DA37F01A0FEC5" box="[1009,1165,331,357]" italics="true" pageId="7" pageNumber="594">
<taxonomicName id="4C6B50E6FFC04105FC2DA37F01A4FEC5" authorityName="Gentili" authorityYear="1984" box="[1009,1161,331,357]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="hammondi">L. hammondi</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37178EEFFC04105FF1BA35A0477FD6D" pageId="7" pageNumber="594" type="discussion">
<paragraph id="8BD42B65FFC04105FF1BA35A0477FD6D" blockId="7.[151,1437,150,717]" pageId="7" pageNumber="594">
<emphasis id="B91FF777FFC04105FF1BA35A0462FE28" bold="true" box="[199,335,366,392]" pageId="7" pageNumber="594">Comments.</emphasis>
<bibRefCitation id="EFFA5694FFC04105FE85A35B04D1FE29" author="Gentili, E." box="[345,508,366,393]" pageId="7" pageNumber="594" pagination="317 - 370" refId="ref7731" refString="Gentili, E. (2005) The genus Laccobius Erichson, 1837 in the Australian region (Coleoptera, Hydrophilidae). In: Daccordi, M. &amp; Giachino, P. M. (Eds.), Results of the zoological missions to Australia of the Regional Museum of Natural Sciences of Turin, Italy. Volume II. Monografie del Museo Regionale di Scienze Naturali Torino, 42, 317 - 370." type="journal article" year="2005">Gentili (2005)</bibRefCitation>
described
<taxonomicName id="4C6B50E6FFC04105FDA0A35B0620FE29" authorityName="Gentili" authorityYear="2005" box="[636,781,367,393]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="philipinus">
<emphasis id="B91FF777FFC04105FDA0A35B0620FE29" box="[636,781,367,393]" italics="true" pageId="7" pageNumber="594">L. philipinus</emphasis>
</taxonomicName>
from the
<collectingCountry id="F37C6BF5FFC04105FC58A35B0128FE29" box="[900,1029,367,393]" name="Philippines" pageId="7" pageNumber="594">Philippines</collectingCountry>
, but included a single female from
<collectingCountry id="F37C6BF5FFC04105FF4BA3A705C4FE0D" box="[151,233,403,429]" name="Taiwan" pageId="7" pageNumber="594">Taiwan</collectingCountry>
in the type series. The Taiwanese specimen was collected in Liukui near
<collectingRegion id="49AFE587FFC04105FBC7A3A701B5FE0D" box="[1051,1176,403,429]" country="Taiwan" name="Kaohsiung" pageId="7" pageNumber="594">Kaohsiung</collectingRegion>
. The first author of this study (HCL) visited the area in 2020 and collected the
<taxonomicName id="4C6B50E6FFC04105FCCDA383069EFE71" box="[785,947,439,465]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="594" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
from streams and wet rocks in Liukui and surrounding areas in the
<collectingRegion id="49AFE587FFC04105FE7BA3EF070DFE55" box="[423,544,475,501]" country="Taiwan" name="Kaohsiung" pageId="7" pageNumber="594">Kaohsiung</collectingRegion>
county. Three species of
<taxonomicName id="4C6B50E6FFC04105FCE4A3EF0686FE55" authorityName="Erichson" authorityYear="1837" box="[824,939,475,501]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="594" phylum="Arthropoda" rank="genus">
<emphasis id="B91FF777FFC04105FCE4A3EF0686FE55" box="[824,939,475,501]" italics="true" pageId="7" pageNumber="594">Laccobius</emphasis>
</taxonomicName>
were found:
<taxonomicName id="4C6B50E6FFC04105FBE0A3EF01EDFE55" authorityName="Gentili" authorityYear="1979" box="[1084,1216,475,501]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B91FF777FFC04105FBE0A3EF01EDFE55" box="[1084,1216,475,501]" italics="true" pageId="7" pageNumber="594">L. formosus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6B50E6FFC04105FB10A3EF007FFE54" authorityName="Regimbart" authorityYear="1903" box="[1228,1362,475,500]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="roseiceps">
<emphasis id="B91FF777FFC04105FB10A3EF007FFE54" box="[1228,1362,475,500]" italics="true" pageId="7" pageNumber="594">L. roseiceps</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B50E6FFC04105FA5AA3EF0596FDB9" authority="Liu &amp; Fikáček, 2021" authorityName="Liu &amp; Fikáček" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="hoi" status="sp. nov.">
<emphasis id="B91FF777FFC04105FA5AA3EF0596FDB9" italics="true" pageId="7" pageNumber="594">L. hoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C4A0CFFC04105FF1EA3CB0436FDB9" box="[194,283,511,537]" pageId="7" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
, but no specimens of
<taxonomicName id="4C6B50E6FFC04105FDD2A3CB07B1FDB9" authorityName="Gentili" authorityYear="2005" box="[526,668,511,537]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="philipinus">
<emphasis id="B91FF777FFC04105FDD2A3CB07B1FDB9" box="[526,668,511,537]" italics="true" pageId="7" pageNumber="594">L. philipinus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C6B50E6FFC04105FD76A3CB06B9FDB9" authorityName="Gentili" authorityYear="2005" box="[682,916,511,537]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="philipinus">
<emphasis id="B91FF777FFC04105FD76A3CB06B9FDB9" box="[682,916,511,537]" italics="true" pageId="7" pageNumber="594">Laccobius philipinus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B50E6FFC04105FC17A3CB0121FDB9" authority="Liu &amp; Fikáček, 2021" authorityName="Liu &amp; Fikáček" authorityYear="2021" box="[971,1036,511,537]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="hoi" status="sp. nov.">
<emphasis id="B91FF777FFC04105FC17A3CB0121FDB9" box="[971,1036,511,537]" italics="true" pageId="7" pageNumber="594">L. hoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C4A0CFFC04105FBCFA3CB0147FDB9" box="[1043,1130,511,537]" pageId="7" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
are very similar externally, despite their male genitalia which differ obviously between the species (see differential diagnosis). The
<typeStatus id="54D095C7FFC04105FAE0A01700B1FD9D" box="[1340,1436,547,573]" pageId="7" pageNumber="594" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C6B50E6FFC04105FF6BA0730412FDC1" authorityName="Gentili" authorityYear="2005" box="[183,319,583,609]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="philipinus">
<emphasis id="B91FF777FFC04105FF6BA0730412FDC1" box="[183,319,583,609]" italics="true" pageId="7" pageNumber="594">L philipinus</emphasis>
</taxonomicName>
from
<collectingCountry id="F37C6BF5FFC04105FE5BA07304F4FDC1" box="[391,473,583,609]" name="Taiwan" pageId="7" pageNumber="594">Taiwan</collectingCountry>
designated by
<bibRefCitation id="EFFA5694FFC04105FD54A0730604FDC1" author="Gentili, E." box="[648,809,582,609]" pageId="7" pageNumber="594" pagination="317 - 370" refId="ref7731" refString="Gentili, E. (2005) The genus Laccobius Erichson, 1837 in the Australian region (Coleoptera, Hydrophilidae). In: Daccordi, M. &amp; Giachino, P. M. (Eds.), Results of the zoological missions to Australia of the Regional Museum of Natural Sciences of Turin, Italy. Volume II. Monografie del Museo Regionale di Scienze Naturali Torino, 42, 317 - 370." type="journal article" year="2005">Gentili (2005)</bibRefCitation>
is deposited in the collection of Elio Gentili in Museo Civico di Storia Naturale, Verona,
<collectingCountry id="F37C6BF5FFC04105FDFCA05F0779FD25" box="[544,596,619,645]" name="Italy" pageId="7" pageNumber="594">Italy</collectingCountry>
, and was not examined by us. It seems to correspond to
<taxonomicName id="4C6B50E6FFC04105FB0BA05F0034FD25" authority="Liu &amp; Fikáček, 2021" authorityName="Liu &amp; Fikáček" authorityYear="2021" box="[1239,1305,619,645]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="hoi" status="sp. nov.">
<emphasis id="B91FF777FFC04105FB0BA05F0034FD25" box="[1239,1305,619,645]" italics="true" pageId="7" pageNumber="594">L. hoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C4A0CFFC04105FAFDA05F0054FD25" box="[1313,1401,619,645]" pageId="7" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
by the size of the pronotal dark spot (E. Gentili, pers. comm.
<date id="FFD50DA5FFC04105FCFFA0BB06BBFD09" box="[803,918,655,681]" pageId="7" pageNumber="594" value="2020-07">July 2020</date>
). Therefore, we remove
<taxonomicName id="4C6B50E6FFC04105FB74A0BB0018FD09" authorityName="Gentili" authorityYear="2005" box="[1192,1333,655,681]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="594" phylum="Arthropoda" rank="species" species="philipinus">
<emphasis id="B91FF777FFC04105FB74A0BB0018FD09" box="[1192,1333,655,681]" italics="true" pageId="7" pageNumber="594">L. philipinus</emphasis>
</taxonomicName>
from the Taiwanese fauna.
</paragraph>
</subSubSection>
</treatment>
</document>