383 lines
42 KiB
XML
383 lines
42 KiB
XML
<document id="57B4D6A5C36FEFCC6142F61D05FC0EEA" ID="10.11646/zootaxa.4175.5.9" ID-DOI="10.11646/zootaxa.4175.5.9" ID-GBIF-Dataset="f697088f-e0b4-4152-a4ed-49748867fca1" ID-ISSN="1175-5326" ID-Zenodo-Dep="161496" ID-ZooBank="8A1428DB-F358-4D7B-92AB-52BCE65CA172" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1476774862764" checkinUser="plazi" docAuthor="Rossini, M., Vaz-De-Mello, F. Z. & Zunino, M." docDate="2016" docId="03C287F9E56C010225D5D839FCD0C350" docLanguage="en" docName="zootaxa.4175.5.9.pdf" docOrigin="Zootaxa 4175 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus striatulus subsp. striatulus (Palisot de Beauvois, 1809) (Howden & Cartwright 1963" docType="treatment" docVersion="8" lastPageNumber="499" masterDocId="FFFBFF81E56801072542DC48FFB1C67D" masterDocTitle="More transcontinental mislabeling in the genus Onthophagus: d’Orbigny’s “ African ” Group 27 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="500" masterPageNumber="494" pageNumber="498" updateTime="1698390084022" updateUser="plazi">
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<mods:titleInfo id="6EEDCDF805B333CDD85EE0C5B2A4B5B0">
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<mods:title id="FB908768EBFD8E25067385BD17B268C7">More transcontinental mislabeling in the genus Onthophagus: d’Orbigny’s “ African ” Group 27 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="DEB166F55A050EF43BC5893DA63FC554">Rossini, M.</mods:namePart>
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<mods:namePart id="AA5FDE4B5CB16B31C09BC2190A74A8C5">Vaz-De-Mello, F. Z.</mods:namePart>
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<mods:namePart id="EA5EFBC3CA18ACB6E6BD72F1C975F6EA">Zunino, M.</mods:namePart>
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<mods:title id="0787D5FA5C1B7BA2EB53FEBF5804064E">Zootaxa</mods:title>
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<mods:part id="400418E9E337763AB94A9FD03C82CA32">
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<mods:date id="69B2567F719CFF1784B8CEDBEB011CF5">2016</mods:date>
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<mods:number id="037E9605F3B640DD74472430086F28D3">4175</mods:number>
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<treatment id="03C287F9E56C010225D5D839FCD0C350" ID-DOI="http://doi.org/10.5281/zenodo.6087431" ID-GBIF-Taxon="124996649" ID-Zenodo-Dep="6087431" LSID="urn:lsid:plazi:treatment:03C287F9E56C010225D5D839FCD0C350" httpUri="http://treatment.plazi.org/id/03C287F9E56C010225D5D839FCD0C350" lastPageId="5" lastPageNumber="499" pageId="4" pageNumber="498">
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<subSubSection id="C3716564E56C010325D5D839FA2DC2F6" box="[151,1436,1137,1163]" pageId="4" pageNumber="498" type="nomenclature">
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<paragraph id="8BD436EFE56C010325D5D839FA2DC2F6" blockId="4.[151,1436,1137,2032]" box="[151,1436,1137,1163]" pageId="4" pageNumber="498">
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<heading id="D09C8183E56C010325D5D839FA2DC2F6" bold="true" box="[151,1436,1137,1163]" fontSize="11" level="1" pageId="4" pageNumber="498" reason="1">
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<emphasis id="B91FEAFDE56C010325D5D839FA2DC2F6" bold="true" box="[151,1436,1137,1163]" pageId="4" pageNumber="498">
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<taxonomicName id="4C6B4D6CE56C010325D5D839FAB2C2F6" ID-CoL="7ZCHG" authority="(Palisot de Beauvois, 1809) (Howden & Cartwright, 1963)" authorityName="(Palisot de Beauvois, 1809) (Howden & Cartwright" authorityYear="1963" box="[151,1283,1137,1163]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="subSpecies" species="striatulus" subSpecies="striatulus">
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<emphasis id="B91FEAFDE56C010325D5D839FD9FC2F6" bold="true" box="[151,558,1137,1163]" italics="true" pageId="4" pageNumber="498">Onthophagus striatulus striatulus</emphasis>
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(Palisot de Beauvois, 1809) (
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<bibRefCitation id="EFFA4B1EE56C010326D3D839FB4BC2F6" author="Howden" box="[913,1274,1137,1163]" pageId="4" pageNumber="498" refString="Howden, H. F. & Cartwright, O. L. (1963) Scarab beetles of the genus Onthophagus Latreille north of Mexico (Coleoptera: Scarabaeidae). Proceedings of the United States National Museum, 114 (Number 3467), 1 - 135. http: // dx. doi. org / 10.5479 / si. 00963801.114 - 3467.1" type="journal article" year="1963">Howden & Cartwright, 1963</bibRefCitation>
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)
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</taxonomicName>
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||
(
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<figureCitation id="13502A6AE56C01032050D839FAE9C2F6" box="[1298,1368,1137,1163]" captionStart="FIGURE 1" captionStartId="3.[151,244,1951,1973]" captionTargetBox="[166,1412,226,1912]" captionTargetId="figure@3.[150,1436,194,1928]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus and labels of the primary types. A, F: O. bidentatus and O. bicornis. B, G: O. striatulus. C, H: O. semichalcites. D, I: O. gibber. E, L: O. gibberosus. Scale bar: 1 mm." httpUri="https://zenodo.org/record/272490/files/figure.png" pageId="4" pageNumber="498">Fig. 1</figureCitation>
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B, G)
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3716564E56C010325D5D8DCFCC6C2B0" pageId="4" pageNumber="498" type="reference_group">
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<paragraph id="8BD436EFE56C010325D5D8DCFCF5C2D6" blockId="4.[151,1436,1137,2032]" box="[151,836,1172,1195]" pageId="4" pageNumber="498">
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<treatmentCitationGroup id="AB7B11C1E56C010325D5D8DCFCF5C2D6" box="[151,836,1172,1195]" pageId="4" pageNumber="498">
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<taxonomicName id="4C6B4D6CE56C010325D5D8DCFDEBC2D6" ID-CoL="7XN83" authority="d'Orbigny 1905: 493" authorityName="d'Orbigny" authorityPageNumber="493" authorityYear="1905" box="[151,602,1172,1195]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="species" species="gibber">
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<emphasis id="B91FEAFDE56C010325D5D8DCFEC3C2D6" box="[151,370,1172,1195]" italics="true" pageId="4" pageNumber="498">Onthophagus gibber</emphasis>
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<treatmentCitation id="0ACA10FEE56C0103243BD8DCFDEBC2D6" author="d'Orbigny" box="[377,602,1172,1195]" page="493" pageId="4" pageNumber="498" year="1905">
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<bibRefCitation id="EFFA4B1EE56C0103243BD8DCFD99C2D6" author="d'Orbigny" box="[377,552,1172,1195]" pageId="4" pageNumber="498" refString="d'Orbigny, H. (1905) Onthophagides provenant du voyage de M. Alluaud dans l'Afrique orientale et descriptions d'autres especes africains nouvelles. Annales de la Societe entomologique de France, 74 (3 - 4), 381 - 536." type="journal article" year="1905">d’Orbigny 1905</bibRefCitation>
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: 493
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</treatmentCitation>
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</taxonomicName>
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<taxonomicNameLabel id="A22C5786E56C01032723D8DCFD75C2D6" box="[609,708,1172,1195]" pageId="4" pageNumber="498">
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<emphasis id="B91FEAFDE56C01032723D8DCFD75C2D6" bold="true" box="[609,708,1172,1195]" pageId="4" pageNumber="498">syn. nov.</emphasis>
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</taxonomicNameLabel>
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(
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<figureCitation id="13502A6AE56C01032790D8DCFCBFC2D6" box="[722,782,1172,1195]" captionStart="FIGURE 1" captionStartId="3.[151,244,1951,1973]" captionTargetBox="[166,1412,226,1912]" captionTargetId="figure@3.[150,1436,194,1928]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus and labels of the primary types. A, F: O. bidentatus and O. bicornis. B, G: O. striatulus. C, H: O. semichalcites. D, I: O. gibber. E, L: O. gibberosus. Scale bar: 1 mm." httpUri="https://zenodo.org/record/272490/files/figure.png" pageId="4" pageNumber="498">Fig. 1</figureCitation>
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D, I)
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BD436EFE56C010325D5D8FEFCC6C2B0" blockId="4.[151,1436,1137,2032]" box="[151,887,1206,1229]" pageId="4" pageNumber="498">
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<treatmentCitationGroup id="AB7B11C1E56C010325D5D8FEFCC6C2B0" box="[151,887,1206,1229]" pageId="4" pageNumber="498">
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<taxonomicName id="4C6B4D6CE56C010325D5D8FEFD38C2B0" ID-CoL="7XN85" authority="d'Orbigny 1905: 492" authorityName="d'Orbigny" authorityPageNumber="492" authorityYear="1905" box="[151,649,1206,1229]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="species" species="gibberosus">
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<emphasis id="B91FEAFDE56C010325D5D8FEFE10C2B0" box="[151,417,1206,1229]" italics="true" pageId="4" pageNumber="498">Onthophagus gibberosus</emphasis>
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<treatmentCitation id="0ACA10FEE56C010324EAD8FEFD38C2B0" author="d'Orbigny" box="[424,649,1206,1229]" page="492" pageId="4" pageNumber="498" year="1905">
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<bibRefCitation id="EFFA4B1EE56C010324EAD8FEFDE7C2B0" author="d'Orbigny" box="[424,598,1206,1229]" pageId="4" pageNumber="498" refString="d'Orbigny, H. (1905) Onthophagides provenant du voyage de M. Alluaud dans l'Afrique orientale et descriptions d'autres especes africains nouvelles. Annales de la Societe entomologique de France, 74 (3 - 4), 381 - 536." type="journal article" year="1905">d’Orbigny 1905</bibRefCitation>
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: 492
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</treatmentCitation>
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</taxonomicName>
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<taxonomicNameLabel id="A22C5786E56C010327CDD8FEFD43C2B0" box="[655,754,1206,1229]" pageId="4" pageNumber="498">
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<emphasis id="B91FEAFDE56C010327CDD8FEFD43C2B0" bold="true" box="[655,754,1206,1229]" pageId="4" pageNumber="498">syn. nov.</emphasis>
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</taxonomicNameLabel>
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(
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<figureCitation id="13502A6AE56C01032643D8FEFC8DC2B0" box="[769,828,1206,1229]" captionStart="FIGURE 1" captionStartId="3.[151,244,1951,1973]" captionTargetBox="[166,1412,226,1912]" captionTargetId="figure@3.[150,1436,194,1928]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus and labels of the primary types. A, F: O. bidentatus and O. bicornis. B, G: O. striatulus. C, H: O. semichalcites. D, I: O. gibber. E, L: O. gibberosus. Scale bar: 1 mm." httpUri="https://zenodo.org/record/272490/files/figure.png" pageId="4" pageNumber="498">Fig. 1</figureCitation>
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E, L)
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3716564E56C010325D5D891FE64C28D" box="[151,469,1241,1264]" pageId="4" pageNumber="498" type="materials_examined">
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<paragraph id="8BD436EFE56C010325D5D891FE64C28D" blockId="4.[151,1436,1137,2032]" box="[151,469,1241,1264]" pageId="4" pageNumber="498">
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<materialsCitation id="3B033CB2E56C010325D5D891FE64C28D" ID-GBIF-Occurrence="1323867455" box="[151,469,1241,1264]" location="South Carolina" pageId="4" pageNumber="498" specimenCount="1" stateProvince="South Carolina" typeStatus="holotype">
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<emphasis id="B91FEAFDE56C010325D5D891FE97C28D" bold="true" box="[151,294,1241,1264]" pageId="4" pageNumber="498">
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<typeStatus id="54D0884DE56C010325D5D891FF7EC28D" box="[151,207,1241,1264]" pageId="4" pageNumber="498">Type</typeStatus>
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locality
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</emphasis>
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.
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<collectingRegion id="49AFF80DE56C01032476D891FE64C28D" box="[308,469,1241,1264]" country="United States of America" name="South Carolina" pageId="4" pageNumber="498">
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<collectingRegion id="49AFF80DE56C01032476D891FEC3C28D" box="[308,370,1241,1264]" country="Tanzania" name="Kusini Unguja" pageId="4" pageNumber="498">South</collectingRegion>
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Carolina
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</collectingRegion>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3716564E56C01032585D8B4FC0AC36E" box="[199,955,1276,1299]" pageId="4" pageNumber="498" type="distribution">
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<paragraph id="8BD436EFE56C01032585D8B4FC0AC36E" blockId="4.[151,1436,1137,2032]" box="[199,955,1276,1299]" pageId="4" pageNumber="498">
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<emphasis id="B91FEAFDE56C01032585D8B4FEE3C36E" bold="true" box="[199,338,1276,1299]" pageId="4" pageNumber="498">Distribution</emphasis>
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. Central and eastern
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<collectingCountry id="F37C767FE56C01032776D8B4FDD8C36E" box="[564,617,1276,1299]" name="United States of America" pageId="4" pageNumber="498">USA</collectingCountry>
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||
(
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<bibRefCitation id="EFFA4B1EE56C01032735D8B4FC1EC36E" author="Howden" box="[631,943,1276,1299]" pageId="4" pageNumber="498" refString="Howden, H. F. & Cartwright, O. L. (1963) Scarab beetles of the genus Onthophagus Latreille north of Mexico (Coleoptera: Scarabaeidae). Proceedings of the United States National Museum, 114 (Number 3467), 1 - 135. http: // dx. doi. org / 10.5479 / si. 00963801.114 - 3467.1" type="journal article" year="1963">Howden & Cartwright, 1963</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3716564E56C01022585D955FEE9C27B" lastPageId="5" lastPageNumber="499" pageId="4" pageNumber="498" type="discussion">
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<paragraph id="8BD436EFE56C01032585D955FA96C3A6" blockId="4.[151,1436,1137,2032]" pageId="4" pageNumber="498">
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<emphasis id="B91FEAFDE56C01032585D955FE84C349" bold="true" box="[199,309,1309,1332]" pageId="4" pageNumber="498">Remarks.</emphasis>
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Palisot de Beauvois (hereafter referred to as PB) was a French botanist and considered, together with F. V. Melsheimer, one of the first entomologists who collected and described insects in North America, especially in Hispaniola and
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<collectingCountry id="F37C767FE56C0103247FD928FEC7C30A" box="[317,374,1376,1399]" name="United States of America" pageId="4" pageNumber="498">USA</collectingCountry>
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. PB arrived in
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<collectingCountry id="F37C767FE56C01032753D928FDF7C30A" box="[529,582,1376,1399]" name="Haiti" pageId="4" pageNumber="498">Haiti</collectingCountry>
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in
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<date id="FFD5102FE56C01032725D928FD51C30A" box="[615,736,1376,1399]" pageId="4" pageNumber="498" value="1791" valueMax="1792">1791–1792</date>
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on board a slave ship coming from the Gulf of
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<collectingCountry id="F37C767FE56C01032191D928FA92C30A" box="[1235,1315,1376,1399]" name="Guinea-Bissau" pageId="4" pageNumber="498">Guinea</collectingCountry>
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. Indeed, he was hired as the main naturalist in the French expedition led by Captain Landolphe and which aimed to establish a new trading settlement in Oware, a territory between the current
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<collectingCountry id="F37C767FE56C01032659D9EAFCEAC3C4" box="[795,859,1442,1465]" name="Benin" pageId="4" pageNumber="498">Benin</collectingCountry>
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and
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<collectingCountry id="F37C767FE56C010326CDD9EAFC52C3C4" box="[911,995,1442,1465]" name="Nigeria" pageId="4" pageNumber="498">Nigeria</collectingCountry>
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. In 1791, the French colony was invaded by the British army and the spread of an epidemic of yellow fever compelled a debilitated PB to leave Africa.
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</paragraph>
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<paragraph id="8BD436EFE56C01032585D9ADFE36C0DE" blockId="4.[151,1436,1137,2032]" pageId="4" pageNumber="498">
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PB spent the next 12 years in North America, collecting in
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<collectingCountry id="F37C767FE56C0103260CD9ADFC37C381" box="[846,902,1509,1532]" name="Haiti" pageId="4" pageNumber="498">Haiti</collectingCountry>
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, St. Domingue and
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<collectingCountry id="F37C767FE56C0103211CD9ADFB27C381" box="[1118,1174,1509,1532]" name="United States of America" pageId="4" pageNumber="498">USA</collectingCountry>
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, and obtaining financial support by the French Minister Pierre Adet. He became a member of the American Philosophical Society of Philadelphia, which became an important means to communicate his own natural history observations. PB returned finally to
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<collectingCountry id="F37C767FE56C010325BDDA01FEF9C01D" box="[255,328,1609,1632]" name="France" pageId="4" pageNumber="498">France</collectingCountry>
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at the beginning of 19th century, when the revolutionary government decided to restore his citizenship, but on the way back his last collections, made during
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<date id="FFD5102FE56C0103279DDA22FCE8C0FC" box="[735,857,1642,1665]" pageId="4" pageNumber="498" value="1793" valueMax="1798">1793–1798</date>
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, were destroyed in a shipwreck off the coast of Nova Scotia (
|
||
<bibRefCitation id="EFFA4B1EE56C010325A5DAC4FECAC0DE" author="Merrill" box="[231,379,1676,1699]" pageId="4" pageNumber="498" refString="Merrill, E. D. (1936) Palisot de Beauvois as an overlooked American botanist. Proceeding of the American Philosophical Society, 76 (6), 899 - 920." type="journal article" year="1936">Merrill, 1936</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD436EFE56C01022585DAE5FE44C749" blockId="4.[151,1436,1137,2032]" lastBlockId="5.[151,1436,152,1325]" lastPageId="5" lastPageNumber="499" pageId="4" pageNumber="498">
|
||
Natural history specimens collected by PB are now very rare in public collections, as most of them have been lost over the years: “
|
||
<emphasis id="B91FEAFDE56C01032407DA86FF64C155" italics="true" pageId="4" pageNumber="498">
|
||
in his adventurous years that he spent away from
|
||
<collectingCountry id="F37C767FE56C01032617DA86FC13C098" box="[853,930,1742,1765]" name="France" pageId="4" pageNumber="498">France</collectingCountry>
|
||
he lost, through no fault of his own, three great collections of natural history material that he had laboriously assembled in Africa, in
|
||
<collectingCountry id="F37C767FE56C01032104DAB8FBCCC17A" box="[1094,1149,1776,1799]" name="Haiti" pageId="4" pageNumber="498">Haiti</collectingCountry>
|
||
and in the eastern
|
||
<collectingCountry id="F37C767FE56C01032011DAB8FF64C155" name="United States of America" pageId="4" pageNumber="498">United States</collectingCountry>
|
||
</emphasis>
|
||
” (
|
||
<bibRefCitation id="EFFA4B1EE56C010325B3DB59FE0FC155" author="Merrill" box="[241,446,1809,1832]" pageId="4" pageNumber="498" refString="Merrill, E. D. (1936) Palisot de Beauvois as an overlooked American botanist. Proceeding of the American Philosophical Society, 76 (6), 899 - 920." type="journal article" year="1936">Merrill, 1936: 899</bibRefCitation>
|
||
).
|
||
<collectingRegion id="49AFF80DE56C01032490DB59FE59C155" box="[466,488,1809,1832]" country="Nigeria" name="Rivers" pageId="4" pageNumber="498">A</collectingRegion>
|
||
. Chevrolat acquired part of the PB’s collection and received a second lot of specimens from
|
||
<collectingRegion id="49AFF80DE56C01032593DB7AFF56C134" box="[209,231,1842,1865]" country="Nigeria" name="Rivers" pageId="4" pageNumber="498">A</collectingRegion>
|
||
. Serville (Chevrolat, 1852). Importantly, among these specimens, it is very likely that Chevrolat obtained also the
|
||
<typeStatus id="54D0884DE56C010325D5DB1CFF74C116" box="[151,197,1876,1899]" pageId="4" pageNumber="498">type</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6B4D6CE56C010325ADDB1CFEC8C116" box="[239,377,1876,1899]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="species" species="striatulus">
|
||
<emphasis id="B91FEAFDE56C010325ADDB1CFEC8C116" box="[239,377,1876,1899]" italics="true" pageId="4" pageNumber="498">O. striatulus</emphasis>
|
||
</taxonomicName>
|
||
(see pp. 630 and 642). According to Horn
|
||
<emphasis id="B91FEAFDE56C01032621DB1CFC27C116" box="[867,918,1876,1899]" italics="true" pageId="4" pageNumber="498">et al</emphasis>
|
||
. (1990) part of the PB’s specimens have been incorporated in the Neervoort Van de Poll collection (via
|
||
<collectingRegion id="49AFF80DE56C01032656DB3DFC9BC1F1" box="[788,810,1909,1932]" country="Nigeria" name="Rivers" pageId="4" pageNumber="498">A</collectingRegion>
|
||
. Chevrolat), but it is also important to point out that the greater part of the dung beetles formerly included in the Chevrolat collection is now deposited at the Museum für Tierkunde of Dresden (Felsche collection). Again,
|
||
<bibRefCitation id="EFFA4B1EE56C01032780DBF0FB2CC1B2" author="Bousquet" box="[706,1181,1976,1999]" pageId="4" pageNumber="498" refString="Bousquet, Y. (2016) Litteratura Coleopterologica (1758 - 1900): a guide to selected books related to the taxonomy of Coleoptera with publication dates and notes. Zookeys, 583, 1 - 776. http: // dx. doi. org / 10.3897 / zookeys. 583.7084" type="journal article" year="2016">Bousquet (2016, and herein cited literature)</bibRefCitation>
|
||
states that some of the PB’s specimens might have been incorporated in the Dejean’s collection, who either bought or received this material from Palisot de Beauvois himself (see p. iii of the “Avertissement” of the Dejean’s Catalogue first edition, 1821). Over the years, Dejean’s collection was divided in lots and acquired by several Institutes and private collectors, and the MNHN of Paris currently holds the bulk of this natural history collection. Part of it was bought by M. Spinola and today housed at the Museo Regionale di Scienze Naturali of Turin. But no potential
|
||
<typeStatus id="54D0884DE56D01022694DCB4FBB5C76E" box="[982,1028,252,275]" pageId="5" pageNumber="499">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="4C6B4D6CE56D010221DCDCB4FA92C76E" box="[1182,1315,252,275]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="species" species="striatulus">
|
||
<emphasis id="B91FEAFDE56D010221DCDCB4FA92C76E" box="[1182,1315,252,275]" italics="true" pageId="5" pageNumber="499">O. striatulus</emphasis>
|
||
</taxonomicName>
|
||
seem to be deposited here (
|
||
<bibRefCitation id="EFFA4B1EE56D0102247DDD55FE58C749" author="Giachino" box="[319,489,285,308]" pageId="5" pageNumber="499" refString="Giachino, P. M. (1982) Collezione coleotterologica di Massimiliano Spinola. Cataloghi del Museo Regionale di Scienze Naturali di Torino, 3, 1 - 616." type="journal article" year="1982">Giachino, 1982</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD436EFE56D01022585DD76FCA0C467" blockId="5.[151,1436,152,1325]" pageId="5" pageNumber="499">
|
||
As already pointed out by
|
||
<bibRefCitation id="EFFA4B1EE56D010224A3DD76FC90C728" author="Howden" box="[481,801,318,341]" pageId="5" pageNumber="499" refString="Howden, H. F. & Cartwright, O. L. (1963) Scarab beetles of the genus Onthophagus Latreille north of Mexico (Coleoptera: Scarabaeidae). Proceedings of the United States National Museum, 114 (Number 3467), 1 - 135. http: // dx. doi. org / 10.5479 / si. 00963801.114 - 3467.1" type="journal article" year="1963">Howden & Cartwright (1963)</bibRefCitation>
|
||
, the colour pattern of
|
||
<taxonomicName id="4C6B4D6CE56D0102214EDD76FB20C728" box="[1036,1169,318,341]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="species" species="striatulus">
|
||
<emphasis id="B91FEAFDE56D0102214EDD76FB20C728" box="[1036,1169,318,341]" italics="true" pageId="5" pageNumber="499">O. striatulus</emphasis>
|
||
</taxonomicName>
|
||
is highly variable within its area of distribution, with populations having body single-coloured (dark green to reddish brown), while others have elytra bicolored (with distinct paler spots at the base and apex). Nevertheless, these variations were considered to occur within the same species (
|
||
<emphasis id="B91FEAFDE56D010224E5DDE9FDE4C7C5" box="[423,597,417,440]" italics="true" pageId="5" pageNumber="499">
|
||
i.e.
|
||
<taxonomicName id="4C6B4D6CE56D0102248DDDE9FDE4C7C5" box="[463,597,417,440]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="species" species="striatulus">O. striatulus</taxonomicName>
|
||
</emphasis>
|
||
). In the same work, according to morphological and biogeographic evidence,
|
||
<taxonomicName id="4C6B4D6CE56D010225D5DD89FE67C7A5" authority="Blatchley, 1928" authorityName="Blatchley" authorityYear="1928" box="[151,470,449,472]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="species" species="floridanus">
|
||
<emphasis id="B91FEAFDE56D010225D5DD89FE97C7A5" box="[151,294,449,472]" italics="true" pageId="5" pageNumber="499">O. floridanus</emphasis>
|
||
Blatchley, 1928
|
||
</taxonomicName>
|
||
is downgraded to subspecific rank under
|
||
<taxonomicName id="4C6B4D6CE56D010226D9DD89FB91C7A5" box="[923,1056,449,472]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="species" species="striatulus">
|
||
<emphasis id="B91FEAFDE56D010226D9DD89FB91C7A5" box="[923,1056,449,472]" italics="true" pageId="5" pageNumber="499">O. striatulus</emphasis>
|
||
</taxonomicName>
|
||
. Indeed, although morphologically very similar to
|
||
<taxonomicName id="4C6B4D6CE56D0102247BDDAAFE0FC784" box="[313,446,482,505]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="species" species="striatulus">
|
||
<emphasis id="B91FEAFDE56D0102247BDDAAFE0FC784" box="[313,446,482,505]" italics="true" pageId="5" pageNumber="499">O. striatulus</emphasis>
|
||
</taxonomicName>
|
||
, specimens collected in
|
||
<collectingRegion id="49AFF80DE56D01022786DDAAFCA5C784" box="[708,788,482,505]" country="United States of America" name="Florida" pageId="5" pageNumber="499">Florida</collectingRegion>
|
||
, Atlantic coast of
|
||
<collectingRegion id="49AFF80DE56D01022694DDAAFB9AC784" box="[982,1067,482,505]" country="United States of America" name="Georgia" pageId="5" pageNumber="499">
|
||
<collectingCountry id="F37C767FE56D01022694DDAAFB9AC784" box="[982,1067,482,505]" name="Georgia" pageId="5" pageNumber="499">Georgia</collectingCountry>
|
||
</collectingRegion>
|
||
and southern
|
||
<collectingRegion id="49AFF80DE56D01022180DDAAFAD2C784" box="[1218,1379,482,505]" country="United States of America" name="South Carolina" pageId="5" pageNumber="499">
|
||
<collectingRegion id="49AFF80DE56D01022180DDAAFAB1C784" box="[1218,1280,482,505]" country="Tanzania" name="Kusini Unguja" pageId="5" pageNumber="499">South</collectingRegion>
|
||
Carolina
|
||
</collectingRegion>
|
||
have body completely black and elytral sculpture not alutaceous.
|
||
</paragraph>
|
||
<paragraph id="8BD436EFE56D01022585DE6CFE92C401" blockId="5.[151,1436,152,1325]" pageId="5" pageNumber="499">
|
||
We carefully examined the external morphology and male genital organs of
|
||
<taxonomicName id="4C6B4D6CE56D01022178DE6CFB77C446" box="[1082,1222,548,571]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="species" species="striatulus">
|
||
<emphasis id="B91FEAFDE56D01022178DE6CFB77C446" box="[1082,1222,548,571]" italics="true" pageId="5" pageNumber="499">O. striatulus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B4D6CE56D01022045DE6CFA2AC446" box="[1287,1435,548,571]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="species" species="floridanus">
|
||
<emphasis id="B91FEAFDE56D01022045DE6CFA2AC446" box="[1287,1435,548,571]" italics="true" pageId="5" pageNumber="499">O. floridanus</emphasis>
|
||
</taxonomicName>
|
||
(including the primary
|
||
<typeStatus id="54D0884DE56D010224E4DE0CFE65C426" box="[422,468,580,603]" pageId="5" pageNumber="499">type</typeStatus>
|
||
of the latter) and we decided to maintain the Howden & Cartwright’s taxonomic arrangement.
|
||
</paragraph>
|
||
<paragraph id="8BD436EFE56D01022585DECEFCC9C4A2" blockId="5.[151,1436,152,1325]" pageId="5" pageNumber="499">
|
||
Over the last two years, we carried out intensive researches and visits to all the European museums mentioned above where it was likely to locate Palisot de Beauvois’ specimens, but none were found. This leads us to believe the
|
||
<typeStatus id="54D0884DE56D01022004DEEFFACFC4C3" box="[1350,1406,679,702]" pageId="5" pageNumber="499">types</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6B4D6CE56D010225D5DE80FEADC4A2" box="[151,284,712,735]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="species" species="striatulus">
|
||
<emphasis id="B91FEAFDE56D010225D5DE80FEADC4A2" box="[151,284,712,735]" italics="true" pageId="5" pageNumber="499">O. striatulus</emphasis>
|
||
</taxonomicName>
|
||
may have been lost with many other of PB’s specimens.
|
||
</paragraph>
|
||
<paragraph id="8BD436EFE56D01022585DEA1FE07C5B9" blockId="5.[151,1436,152,1325]" pageId="5" pageNumber="499">
|
||
Therefore, because of the need to define objectively a nominal taxon for
|
||
<taxonomicName id="4C6B4D6CE56D01022155DEA1FEF9C55D" authority="Palisot de Beauvois, 1809" authorityName="Palisot de Beauvois" authorityYear="1809" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="species" species="striatulus">
|
||
<emphasis id="B91FEAFDE56D01022155DEA1FAA9C57D" box="[1047,1304,745,768]" italics="true" pageId="5" pageNumber="499">Onthophagus striatulus</emphasis>
|
||
(Palisot de Beauvois, 1809)
|
||
</taxonomicName>
|
||
and preserve the stability of its zoological nomenclature, we designated a male from
|
||
<collectingRegion id="49AFF80DE56D010221BADF41FA2AC55D" box="[1272,1435,777,800]" country="United States of America" name="South Carolina" pageId="5" pageNumber="499">
|
||
<collectingRegion id="49AFF80DE56D010221BADF41FA87C55D" box="[1272,1334,777,800]" country="Tanzania" name="Kusini Unguja" pageId="5" pageNumber="499">South</collectingRegion>
|
||
Carolina
|
||
</collectingRegion>
|
||
(label details below) as a
|
||
<typeStatus id="54D0884DE56D0102248DDF62FD92C53C" box="[463,547,810,833]" pageId="5" pageNumber="499" type="neotype">neotype</typeStatus>
|
||
(
|
||
<bibRefCitation id="EFFA4B1EE56D01022778DF62FC83C53C" author="ICZN" box="[570,818,810,833]" pageId="5" pageNumber="499" refString="ICZN (International Commission on Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN, 1999: Art. 75</bibRefCitation>
|
||
). This
|
||
<typeStatus id="54D0884DE56D010226C9DF62FC6EC53C" box="[907,991,810,833]" pageId="5" pageNumber="499" type="neotype">neotype</typeStatus>
|
||
provides characters to distinguish the nominotypical subspecies from
|
||
<emphasis id="B91FEAFDE56D010224B7DF03FD4DC51F" box="[501,764,843,866]" italics="true" pageId="5" pageNumber="499">
|
||
<taxonomicName id="4C6B4D6CE56D010224B7DF03FD49C51F" box="[501,760,843,866]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="subSpecies" species="striatulus" subSpecies="floridanus">O. striatulus floridanus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
such as elytra with paler spots at the base and apex, elytral surface opaque and weakly shining, elytral striae shallow, interstriae flat (
|
||
<figureCitation id="13502A6AE56D010226ADDF24FB82C5FE" box="[1007,1075,876,899]" captionStart="FIGURE 1" captionStartId="3.[151,244,1951,1973]" captionTargetBox="[166,1412,226,1912]" captionTargetId="figure@3.[150,1436,194,1928]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus and labels of the primary types. A, F: O. bidentatus and O. bicornis. B, G: O. striatulus. C, H: O. semichalcites. D, I: O. gibber. E, L: O. gibberosus. Scale bar: 1 mm." httpUri="https://zenodo.org/record/272490/files/figure.png" pageId="5" pageNumber="499">Fig. 1</figureCitation>
|
||
B). (
|
||
<bibRefCitation id="EFFA4B1EE56D01022129DF24FA21C5FE" author="ICZN" box="[1131,1424,876,899]" pageId="5" pageNumber="499" refString="ICZN (International Commission on Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN, 1999: Art. 75.3.2;</bibRefCitation>
|
||
). Furthermore, the
|
||
<typeStatus id="54D0884DE56D01022412DFC4FE15C5DE" box="[336,420,908,931]" pageId="5" pageNumber="499" type="neotype">neotype</typeStatus>
|
||
eventually defines a
|
||
<typeStatus id="54D0884DE56D010227C7DFC4FD02C5DE" box="[645,691,908,931]" pageId="5" pageNumber="499">type</typeStatus>
|
||
locality for
|
||
<taxonomicName id="4C6B4D6CE56D01022677DFC4FB97C5DE" box="[821,1062,908,931]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="subSpecies" species="striatulus" subSpecies="striatulus">
|
||
<emphasis id="B91FEAFDE56D01022677DFC4FB97C5DE" box="[821,1062,908,931]" italics="true" pageId="5" pageNumber="499">O. striatulus striatulus</emphasis>
|
||
</taxonomicName>
|
||
(Art. 75.3.6), that is “Dovehaven”,
|
||
<quantity id="4C939B0AE56D010225D5DFE5FF76C5B9" box="[151,199,941,964]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="5" pageNumber="499" unit="mi" value="7.0">7 mi</quantity>
|
||
north-east of Pickens.
|
||
</paragraph>
|
||
<paragraph id="8BD436EFE56D01022585DF86FEE9C27B" blockId="5.[151,1436,152,1325]" pageId="5" pageNumber="499">
|
||
<materialsCitation id="3B033CB2E56D01022585DF86FEE9C27B" ID-GBIF-Occurrence="1323867456" collectionCode="MNHN" location="Arthropods" pageId="5" pageNumber="499" specimenCount="1" stateProvince="Florida" typeStatus="neotype">
|
||
The
|
||
<typeStatus id="54D0884DE56D010225BADF86FEFDC598" box="[248,332,974,997]" pageId="5" pageNumber="499" type="neotype">neotype</typeStatus>
|
||
has been selected from the
|
||
<collectingRegion id="49AFF80DE56D0102273ADF86FD74C598" box="[632,709,974,997]" country="United States of America" name="Florida" pageId="5" pageNumber="499">Florida</collectingRegion>
|
||
State Collection of
|
||
<location id="8EB46034E56D010226DEDF86FBA7C598" LSID="urn:lsid:plazi:treatment:03C287F9E56C010225D5D839FCD0C350:8EB46034E56D010226DEDF86FBA7C598" box="[924,1046,974,997]" name="Arthropods" pageId="5" pageNumber="499" stateProvince="Florida">Arthropods</location>
|
||
and is now deposited in the
|
||
<collectionCode id="ED7AAE2AE56D01022008DF86FA2AC598" box="[1354,1435,974,997]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="499">MNHN</collectionCode>
|
||
of Paris (
|
||
<figureCitation id="13502A6AE56D010225BADFA7FE85C27B" box="[248,308,1007,1030]" captionStart="FIGURE 1" captionStartId="3.[151,244,1951,1973]" captionTargetBox="[166,1412,226,1912]" captionTargetId="figure@3.[150,1436,194,1928]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus and labels of the primary types. A, F: O. bidentatus and O. bicornis. B, G: O. striatulus. C, H: O. semichalcites. D, I: O. gibber. E, L: O. gibberosus. Scale bar: 1 mm." httpUri="https://zenodo.org/record/272490/files/figure.png" pageId="5" pageNumber="499">Fig. 1</figureCitation>
|
||
B).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3716564E56D01022585D858FB1BC2B6" pageId="5" pageNumber="499" type="materials_examined">
|
||
<paragraph id="8BD436EFE56D01022585D858FC95C215" blockId="5.[151,1436,152,1325]" pageId="5" pageNumber="499">
|
||
<emphasis id="B91FEAFDE56D01022585D858FE46C25A" bold="true" box="[199,503,1040,1063]" pageId="5" pageNumber="499">
|
||
<typeStatus id="54D0884DE56D01022585D858FF4EC25A" box="[199,255,1040,1063]" pageId="5" pageNumber="499">Type</typeStatus>
|
||
specimens examined.
|
||
</emphasis>
|
||
Of
|
||
<taxonomicName id="4C6B4D6CE56D01022766D858FD1BC25A" box="[548,682,1040,1063]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="species" species="striatulus">
|
||
<emphasis id="B91FEAFDE56D01022766D858FD1BC25A" box="[548,682,1040,1063]" italics="true" pageId="5" pageNumber="499">O. striatulus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<typeStatus id="54D0884DE56D010227F8D858FCA6C25A" box="[698,791,1040,1063]" pageId="5" pageNumber="499" type="neotype">
|
||
<emphasis id="B91FEAFDE56D010227F8D858FCA6C25A" bold="true" box="[698,791,1040,1063]" pageId="5" pageNumber="499">Neotype</emphasis>
|
||
</typeStatus>
|
||
(
|
||
<specimenCount id="9D6DFD66E56D0102266AD858FC8CC25A" box="[808,829,1040,1063]" pageId="5" pageNumber="499" type="male">♂</specimenCount>
|
||
<collectionCode id="ED7AAE2AE56D01022607D858FC2DC25A" box="[837,924,1040,1063]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="499">MNHN</collectionCode>
|
||
): 1: “Dovehaven”,
|
||
<quantity id="4C939B0AE56D01022129D858FB2CC25A" box="[1131,1181,1040,1063]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="5" pageNumber="499" unit="mi" value="7.0">7 mi</quantity>
|
||
N.E. of Pickens, S. C.,
|
||
<date id="FFD5102FE56D010225D5D879FEB2C235" box="[151,259,1073,1096]" pageId="5" pageNumber="499" value="1983-09-27">9.27.1983</date>
|
||
, H. L. Dozier (printed and handwritten on white label). 2:
|
||
<typeStatus id="54D0884DE56D010226DBD879FBA4C235" box="[921,1045,1073,1096]" pageId="5" pageNumber="499" type="neotype">NEOTYPE</typeStatus>
|
||
,
|
||
<taxonomicName id="4C6B4D6CE56D01022161D879FEF7C215" authority="Palisot de Beauvois, 1809" authorityName="Palisot de Beauvois" authorityYear="1809" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="species" species="striatulus">Onthophagus striatulus (Palisot de Beauvois, 1809)</taxonomicName>
|
||
des.
|
||
<bibRefCitation id="EFFA4B1EE56D0102243FD819FD85C215" author="Rossini" box="[381,564,1105,1128]" pageId="5" pageNumber="499" refString="Rossini, M. & Vaz-de-Mello F. Z. (2016) On the identity of Onthophagus columbianus Boucomont, 1932 (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa, 4061 (3), 296 - 300. http: // dx. doi. org / 10.11646 / zootaxa. 4061.3.10" type="journal article" year="2016">M. Rossini, 2016</bibRefCitation>
|
||
(printed on red label).
|
||
</paragraph>
|
||
<paragraph id="8BD436EFE56D01022585D83AFB1BC2B6" blockId="5.[151,1436,152,1325]" pageId="5" pageNumber="499">
|
||
Of
|
||
<taxonomicName id="4C6B4D6CE56D010225A8D83AFEFEC2F4" box="[234,335,1138,1161]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="species" species="gibber">
|
||
<emphasis id="B91FEAFDE56D010225A8D83AFEFEC2F4" box="[234,335,1138,1161]" italics="true" pageId="5" pageNumber="499">O. gibber</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<typeStatus id="54D0884DE56D0102241CD83AFE77C2F4" box="[350,454,1138,1161]" pageId="5" pageNumber="499" type="holotype">
|
||
<emphasis id="B91FEAFDE56D0102241CD83AFE77C2F4" bold="true" box="[350,454,1138,1161]" pageId="5" pageNumber="499">Holotype</emphasis>
|
||
</typeStatus>
|
||
(
|
||
<specimenCount id="9D6DFD66E56D01022497D83AFE58C2F4" box="[469,489,1138,1161]" pageId="5" pageNumber="499" type="male">♂</specimenCount>
|
||
<collectionCode id="ED7AAE2AE56D010224B2D83AFD81C2F4" box="[496,560,1138,1161]" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="5" pageNumber="499">MTD</collectionCode>
|
||
): 1:
|
||
<taxonomicName id="4C6B4D6CE56D0102271FD83AFD10C2F4" box="[605,673,1138,1161]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="species" species="gibber" status="sp. nov.">gibber</taxonomicName>
|
||
<taxonomicNameLabel id="A22C5786E56D010227EAD83AFD50C2F4" box="[680,737,1138,1161]" pageId="5" pageNumber="499" rank="species">n. sp.</taxonomicNameLabel>
|
||
d’Orb. (handwritten in italic on cream label). 2: Coll. C. Felsche, Kauf 20, 1918 (printed on blue label). 3: Grahamstown (printed on blue label). 4: Staatl. Museum für Tierkunde, Dresden (printed on white label). 5:
|
||
<taxonomicName id="4C6B4D6CE56D010224F8D8FCFDAEC2B6" box="[442,543,1204,1227]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="species" species="gibber">O. gibber</taxonomicName>
|
||
(handwritten in italic blue on white label with black border).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3716564E56D01022585D89DFCD0C350" pageId="5" pageNumber="499" type="description">
|
||
<paragraph id="8BD436EFE56D01022585D89DFCD0C350" blockId="5.[151,1436,152,1325]" pageId="5" pageNumber="499">
|
||
Of
|
||
<taxonomicName id="4C6B4D6CE56D010225ADD89DFE3BC291" box="[239,394,1237,1260]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="species" species="gibberosus">
|
||
<emphasis id="B91FEAFDE56D010225ADD89DFE3BC291" box="[239,394,1237,1260]" italics="true" pageId="5" pageNumber="499">O. gibberosus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<typeStatus id="54D0884DE56D010224DCD89DFDB7C291" box="[414,518,1237,1260]" pageId="5" pageNumber="499" type="holotype">
|
||
<emphasis id="B91FEAFDE56D010224DCD89DFDB7C291" bold="true" box="[414,518,1237,1260]" pageId="5" pageNumber="499">Holotype</emphasis>
|
||
</typeStatus>
|
||
(
|
||
<specimenCount id="9D6DFD66E56D01022759D89DFD9EC291" box="[539,559,1237,1260]" pageId="5" pageNumber="499" type="male">♂</specimenCount>
|
||
MTD): 1:
|
||
<collectingCountry id="F37C767FE56D010227F7D89DFCB3C291" box="[693,770,1237,1260]" name="Liberia" pageId="5" pageNumber="499">Liberia</collectingCountry>
|
||
(handwritten on cream label). 2:
|
||
<taxonomicName id="4C6B4D6CE56D010221C8D89DFB4CC291" box="[1162,1277,1237,1260]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="499" phylum="Arthropoda" rank="species" species="gibberosus" status="sp. nov.">gibberosus</taxonomicName>
|
||
<taxonomicNameLabel id="A22C5786E56D01022048D89DFAF8C291" box="[1290,1353,1237,1260]" pageId="5" pageNumber="499" rank="species">n. sp.</taxonomicNameLabel>
|
||
d’Orb. (handwritten on cream label). 3: Coll. C. Felsche, Kauf 20, 1918 (printed on blue label). 4:
|
||
<typeStatus id="54D0884DE56D01022128D8BDFB1AC371" box="[1130,1195,1269,1292]" pageId="5" pageNumber="499">Typus</typeStatus>
|
||
(printed on red label). 5: Staatl. Museum für Tierkunde, Dresden (printed on white label).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |