505 lines
56 KiB
XML
505 lines
56 KiB
XML
<document id="C375478C2D11C780E69FDED41F7EA7C2" ID="10.11646/zootaxa.4175.5.9" ID-DOI="10.11646/zootaxa.4175.5.9" ID-GBIF-Dataset="f697088f-e0b4-4152-a4ed-49748867fca1" ID-ISSN="1175-5326" ID-Zenodo-Dep="161496" ID-ZooBank="8A1428DB-F358-4D7B-92AB-52BCE65CA172" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1476774862764" checkinUser="plazi" docAuthor="Rossini, M., Vaz-De-Mello, F. Z. & Zunino, M." docDate="2016" docId="03C287F9E56A010325D5DDBAFB65C57A" docLanguage="en" docName="zootaxa.4175.5.9.pdf" docOrigin="Zootaxa 4175 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus bidentatus Drapiez 1819" docType="treatment" docVersion="8" lastPageNumber="498" masterDocId="FFFBFF81E56801072542DC48FFB1C67D" masterDocTitle="More transcontinental mislabeling in the genus Onthophagus: d’Orbigny’s “ African ” Group 27 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="500" masterPageNumber="494" pageNumber="496" updateTime="1698390084022" updateUser="plazi">
|
||
<mods:mods id="CCC9AE73A817E22E59FED3CA5B011D1C" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="6E9EEB368A6F5A0D78CD06F4809188B4">
|
||
<mods:title id="56E20600F809A3831D945574FF528C9A">More transcontinental mislabeling in the genus Onthophagus: d’Orbigny’s “ African ” Group 27 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="088EB70630EB17DDE108B4F8F0F3884C" type="personal">
|
||
<mods:role id="F94FCB1E0FF4D1558F04D60B5DA25B09">
|
||
<mods:roleTerm id="DDA4292EE3FCF057608B62BF6F7D1AB7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="04ED156BE25A1D4439E113A27B2344C8">Rossini, M.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="9C0A3B8689228039352745700E5978B5" type="personal">
|
||
<mods:role id="0AA83D55069B569CB2F5F1F7A8EC5DF5">
|
||
<mods:roleTerm id="6457FF9622BEC46903F4EF175F2FF19F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="13FBB2C536C87D967F711ED768CB19FE">Vaz-De-Mello, F. Z.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="7D3FA235762269ED844ECC0189CA08D2" type="personal">
|
||
<mods:role id="4D1027DF2B30744872F9C6B656FBA71A">
|
||
<mods:roleTerm id="975ABF9B91D34C082825F4E2922CFF79">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="AD7118B4A89AFE2EE181F29A3DD65EAF">Zunino, M.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="CDFD3103A8FFF179BEA80EB89321ABBE">text</mods:typeOfResource>
|
||
<mods:relatedItem id="6C45F90BD631FF6041FC6841B87F4B91" type="host">
|
||
<mods:titleInfo id="D229BCC9F92919E14E3B3176F233BE62">
|
||
<mods:title id="4D32911E302F2C651EE2258C09770726">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="EC8B5214AF02AC47A2CA0E75D229D7B4">
|
||
<mods:date id="12BF5265AC339BAE7187D0330A2C6E79">2016</mods:date>
|
||
<mods:detail id="C392B8FBFF8497CA4D6CCFA448E40CEE" type="volume">
|
||
<mods:number id="A35AAD5EE44CE9B5C54CB2B85879F020">4175</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="129DE036EA30776D125568821DC4BEF7" type="issue">
|
||
<mods:number id="ADEBB1625DB8E0BEBF991F9F271F7418">5</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="BF31314D598C0A0A022651D3156FE388" unit="page">
|
||
<mods:start id="DAF87F74405ED53EE9BB53E956271E73">494</mods:start>
|
||
<mods:end id="F951945096D3CF6EBB88E924101921B3">500</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="D765591913EF27E4D98231DAB84A6DFC">journal article</mods:classification>
|
||
<mods:identifier id="53E30C0E45D35B939224ED0A7F3962D4" type="DOI">10.11646/zootaxa.4175.5.9</mods:identifier>
|
||
<mods:identifier id="C5C4543757CA9E4815C86CF9AA3677AF" type="GBIF-Dataset">f697088f-e0b4-4152-a4ed-49748867fca1</mods:identifier>
|
||
<mods:identifier id="022813D65E0830FF97E2CF2CF7B7B8DD" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="E335FCF10A86833C53BB005860DD401F" type="Zenodo-Dep">161496</mods:identifier>
|
||
<mods:identifier id="A74927B4D5E2334BC58BD509E589BB10" type="ZooBank">8A1428DB-F358-4D7B-92AB-52BCE65CA172</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03C287F9E56A010325D5DDBAFB65C57A" ID-DOI="http://doi.org/10.5281/zenodo.6087427" ID-GBIF-Taxon="124996651" ID-Zenodo-Dep="6087427" LSID="urn:lsid:plazi:treatment:03C287F9E56A010325D5DDBAFB65C57A" httpUri="http://treatment.plazi.org/id/03C287F9E56A010325D5DDBAFB65C57A" lastPageId="4" lastPageNumber="498" pageId="2" pageNumber="496">
|
||
<subSubSection id="C3716564E56A010525D5DDBAFCF6C471" box="[151,839,498,524]" pageId="2" pageNumber="496" type="nomenclature">
|
||
<paragraph id="8BD436EFE56A010525D5DDBAFCF6C471" blockId="2.[151,1436,498,2020]" box="[151,839,498,524]" pageId="2" pageNumber="496">
|
||
<heading id="D09C8183E56A010525D5DDBAFCF6C471" bold="true" box="[151,839,498,524]" fontSize="11" level="1" pageId="2" pageNumber="496" reason="1">
|
||
<treatmentCitationGroup id="AB7B11C1E56A010525D5DDBAFCF6C471" box="[151,839,498,524]" pageId="2" pageNumber="496">
|
||
<emphasis id="B91FEAFDE56A010525D5DDBAFCF6C471" bold="true" box="[151,839,498,524]" pageId="2" pageNumber="496">
|
||
<taxonomicName id="4C6B4D6CE56A010525D5DDBAFD00C471" ID-CoL="7XMGC" authority="Drapiez, 1819: 134" authorityName="Drapiez" authorityPageNumber="134" authorityYear="1819" box="[151,689,498,524]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
<emphasis id="B91FEAFDE56A010525D5DDBAFE71C471" bold="true" box="[151,448,498,524]" italics="true" pageId="2" pageNumber="496">Onthophagus bidentatus</emphasis>
|
||
<treatmentCitation id="0ACA10FEE56A01052485DDBAFD00C471" author="Drapiez" box="[455,689,498,524]" page="134" pageId="2" pageNumber="496" year="1819">
|
||
<bibRefCitation id="EFFA4B1EE56A01052485DDBAFDC9C471" author="Drapiez" box="[455,632,498,524]" pageId="2" pageNumber="496" refString="Drapiez, M. (1819) Descriptions de quelques insectes nouveaux. Annales Generales des Sciences Physiques, 1 (45 - 55), 130 - 138." type="journal article" year="1819">Drapiez, 1819</bibRefCitation>
|
||
: 134
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13502A6AE56A01052782DDBAFCB7C471" box="[704,774,498,524]" captionStart="FIGURE 1" captionStartId="3.[151,244,1951,1973]" captionTargetBox="[166,1412,226,1912]" captionTargetId="figure@3.[150,1436,194,1928]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus and labels of the primary types. A, F: O. bidentatus and O. bicornis. B, G: O. striatulus. C, H: O. semichalcites. D, I: O. gibber. E, L: O. gibberosus. Scale bar: 1 mm." httpUri="https://zenodo.org/record/272490/files/figure.png" pageId="2" pageNumber="496">Fig. 1</figureCitation>
|
||
A, F)
|
||
</emphasis>
|
||
</treatmentCitationGroup>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3716564E56A010525D5DE5DFCA2C432" pageId="2" pageNumber="496" type="reference_group">
|
||
<paragraph id="8BD436EFE56A010525D5DE5DFC15C451" blockId="2.[151,1436,498,2020]" box="[151,932,533,556]" pageId="2" pageNumber="496">
|
||
<treatmentCitationGroup id="AB7B11C1E56A010525D5DE5DFC15C451" box="[151,932,533,556]" pageId="2" pageNumber="496">
|
||
<taxonomicName id="4C6B4D6CE56A010525D5DE5DFDD1C451" ID-CoL="7XMSY" authority="Castelnau, 1840: 87" authorityName="Castelnau" authorityPageNumber="87" authorityYear="1840" box="[151,608,533,556]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bicornis">
|
||
<emphasis id="B91FEAFDE56A010525D5DE5DFE35C451" box="[151,388,533,556]" italics="true" pageId="2" pageNumber="496">Onthophagus bicornis</emphasis>
|
||
<treatmentCitation id="0ACA10FEE56A010524C9DE5DFDD1C451" author="Castelnau" box="[395,608,533,556]" page="87" pageId="2" pageNumber="496" year="1840">
|
||
<bibRefCitation id="EFFA4B1EE56A010524C9DE5DFD8AC451" author="Castelnau" box="[395,571,533,556]" pageId="2" pageNumber="496" refString="Castelnau, F. L. de (1840) Histoire naturelle des Animaux articules, annelides, crustaces, arachnides, myriapodes et insectes. Histoire naturelle des Insectes Coleopteres. Tome deuxieme. P. Dumenil, Paris, 563 pp., 38 pls." type="book" year="1840">Castelnau, 1840</bibRefCitation>
|
||
: 87
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B91FEAFDE56A0105272DDE5DFD26C451" box="[623,663,533,556]" italics="true" pageId="2" pageNumber="496">fide</emphasis>
|
||
<treatmentCitation id="0ACA10FEE56A010527DCDE5DFC2DC451" author="Blackwelder" box="[670,924,533,556]" page="211" pageId="2" pageNumber="496" year="1944">
|
||
<bibRefCitation id="EFFA4B1EE56A010527DCDE5DFCDBC451" author="Blackwelder" box="[670,874,533,556]" pageId="2" pageNumber="496" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. Bulletin of the United States National Museum, 185, 189 - 341." type="journal article" year="1944">Blackwelder, 1944</bibRefCitation>
|
||
: 211
|
||
</treatmentCitation>
|
||
)
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BD436EFE56A010525D5DE70FCA2C432" blockId="2.[151,1436,498,2020]" box="[151,787,568,591]" pageId="2" pageNumber="496">
|
||
<treatmentCitationGroup id="AB7B11C1E56A010525D5DE70FCA2C432" box="[151,787,568,591]" pageId="2" pageNumber="496">
|
||
<taxonomicName id="4C6B4D6CE56A010525D5DE70FD18C432" ID-CoL="7XNXG" authority="d'Orbigny, 1902: 149" authorityName="d'Orbigny" authorityPageNumber="149" authorityYear="1902" box="[151,681,568,591]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="semichalcites">
|
||
<emphasis id="B91FEAFDE56A010525D5DE70FE0CC432" box="[151,445,568,591]" italics="true" pageId="2" pageNumber="496">Onthophagus semichalcites</emphasis>
|
||
<treatmentCitation id="0ACA10FEE56A01052481DE70FD18C432" author="d'Orbigny" box="[451,681,568,591]" page="149" pageId="2" pageNumber="496" year="1902">
|
||
<bibRefCitation id="EFFA4B1EE56A01052481DE70FDC6C432" author="d'Orbigny" box="[451,631,568,591]" pageId="2" pageNumber="496" refString="d'Orbigny, H. (1902) Memoire sur les Onthophagides d'Afrique. Annales de la Societe Entomologique de France. Paris 71 (1 - 2), 1 - 324." type="journal article" year="1902">d’Orbigny, 1902</bibRefCitation>
|
||
: 149
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
<emphasis id="B91FEAFDE56A010527F2DE70FCA2C432" bold="true" box="[688,787,568,591]" pageId="2" pageNumber="496">
|
||
<taxonomicNameLabel id="A22C5786E56A010527F2DE70FCA2C432" box="[688,787,568,591]" pageId="2" pageNumber="496">syn. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3716564E56A010525D5DE12FB0EC20A" pageId="2" pageNumber="496" type="materials_examined">
|
||
<paragraph id="8BD436EFE56A010525D5DE12FDFCC40C" blockId="2.[151,1436,498,2020]" box="[151,589,602,625]" pageId="2" pageNumber="496">
|
||
<materialsCitation id="3B033CB2E56A010525D5DE12FDFCC40C" ID-GBIF-Occurrence="1323867453" box="[151,589,602,625]" country="Guyana" location="Guyana" pageId="2" pageNumber="496" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B91FEAFDE56A010525D5DE12FE99C40C" bold="true" box="[151,296,602,625]" pageId="2" pageNumber="496">
|
||
<typeStatus id="54D0884DE56A010525D5DE12FF7EC40C" box="[151,207,602,625]" pageId="2" pageNumber="496">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
: French
|
||
<collectingCountry id="F37C767FE56A010524C5DE12FE6FC40C" box="[391,478,602,625]" name="Guyana" pageId="2" pageNumber="496">Guyana</collectingCountry>
|
||
: Cayenne.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD436EFE56A01052585DE35FE01C4CA" blockId="2.[151,1436,498,2020]" pageId="2" pageNumber="496">
|
||
<emphasis id="B91FEAFDE56A01052585DE35FDE3C4E9" bold="true" box="[199,594,637,660]" pageId="2" pageNumber="496">Notes on ecology and distribution</emphasis>
|
||
. This species appears to be very common in open habitats, rarely found in transitional forested areas.
|
||
</paragraph>
|
||
<paragraph id="8BD436EFE56A01052585DE8AFB00C5FA" blockId="2.[151,1436,498,2020]" pageId="2" pageNumber="496">
|
||
<bibRefCitation id="EFFA4B1EE56A01052585DE8AFD93C4A4" author="Pulido-Herrera" box="[199,546,706,729]" pageId="2" pageNumber="496" refString="Pulido-Herrera, L. A. & Zunino, M. (2007) Catalogo preliminar de los Onthophagini de America (Coleoptera: Scarabaeinae). In: Zunino, M. & Melic, A. (Eds.), Escarabajos, diversidad y conservacion biologica. Ensayos enhomenaje a Gonzalo Halffter. m 3 m - Monografia Tercer Milenio. Vo l. 7. S. E. A., Zaragoza, pp. 93 - 129." type="journal article" year="2007">Pulido-Herrera & Zunino (2007)</bibRefCitation>
|
||
mentioned
|
||
<taxonomicName id="4C6B4D6CE56A010527E2DE8AFC9EC4A4" box="[672,815,706,729]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
<emphasis id="B91FEAFDE56A010527E2DE8AFC9EC4A4" box="[672,815,706,729]" italics="true" pageId="2" pageNumber="496">O. bidentatus</emphasis>
|
||
</taxonomicName>
|
||
of Guadalupe, Colombia, Venezuela, Guiana, Brazil, Peru and Argentina. In our opinion, the Guadalupe specimens mentioned in
|
||
<bibRefCitation id="EFFA4B1EE56A010526CCDEADFBD2C481" author="Blackwelder" box="[910,1123,741,764]" pageId="2" pageNumber="496" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. Bulletin of the United States National Museum, 185, 189 - 341." type="journal article" year="1944">Blackwelder (1944)</bibRefCitation>
|
||
may belong to
|
||
<taxonomicName id="4C6B4D6CE56A01052048DEADFE93C562" authority="Arrow, 1903" authorityName="Arrow" authorityYear="1903" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="antillarum">
|
||
<emphasis id="B91FEAFDE56A01052048DEADFA2DC481" box="[1290,1436,741,764]" italics="true" pageId="2" pageNumber="496">O. antillarum</emphasis>
|
||
Arrow, 1903
|
||
</taxonomicName>
|
||
, which is undoubtedly a close relative of
|
||
<emphasis id="B91FEAFDE56A010527ADDF40FC39C562" box="[751,904,776,799]" italics="true" pageId="2" pageNumber="496">
|
||
<taxonomicName id="4C6B4D6CE56A010527ADDF40FC35C562" box="[751,900,776,799]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bidentatus">O. bidentatus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Furthermore, as already questioned by
|
||
<bibRefCitation id="EFFA4B1EE56A01052079DF40FF51C53C" author="Martinez" pageId="2" pageNumber="496" refString="Martinez, A. (1959) Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales " Bernardino Rivadavia ", Ciencias Zoologicas, 5 (1), 1 - 126." type="journal article" year="1959">Martínez (1959)</bibRefCitation>
|
||
, the occurrence of
|
||
<taxonomicName id="4C6B4D6CE56A010524F2DF62FDF0C53C" box="[432,577,810,833]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
<emphasis id="B91FEAFDE56A010524F2DF62FDF0C53C" box="[432,577,810,833]" italics="true" pageId="2" pageNumber="496">O. bidentatus</emphasis>
|
||
</taxonomicName>
|
||
in Argentina (Jujuy) (
|
||
<bibRefCitation id="EFFA4B1EE56A01052671DF62FC0BC53C" author="Bruch" box="[819,954,810,833]" pageId="2" pageNumber="496" refString="Bruch, C. (1911) Catalogo sistematico de los Coleopteros de la Republica Argentina. Pars IV: Lucanidae, Scarabaeidae (Coprini- Cetonini), Passalidae. Revista del Museo de La Plata, 17, 181 - 225." type="journal article" year="1911">Bruch, 1911</bibRefCitation>
|
||
) is still very doubtful. According to Bruch’s catalogue, the specimen/s is/are deposited in his personal collection at the Museo Argentino de Ciencias Naturales “Bernardino Rivadavia” (marked with an asterisk), but to date we could not examine this material.
|
||
</paragraph>
|
||
<paragraph id="8BD436EFE56A01052585DFDAFC88C26C" blockId="2.[151,1436,498,2020]" pageId="2" pageNumber="496">
|
||
<bibRefCitation id="EFFA4B1EE56A01052585DFDAFE22C5D4" author="Boucomont" box="[199,403,914,937]" pageId="2" pageNumber="496" refString="Boucomont, A. (1932) Synopsis des Onthophagus d'Amerique du Sud (Col. Scarab). Annales de la Societe Entomologique de France, 101 (4), 293 - 332." type="journal article" year="1932">Boucomont (1932)</bibRefCitation>
|
||
mentioned
|
||
<taxonomicName id="4C6B4D6CE56A01052756DFDAFD14C5D4" box="[532,677,914,937]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
<emphasis id="B91FEAFDE56A01052756DFDAFD14C5D4" box="[532,677,914,937]" italics="true" pageId="2" pageNumber="496">O. bidentatus</emphasis>
|
||
</taxonomicName>
|
||
of Callanga and
|
||
<collectingRegion id="49AFF80DE56A01052620DFDAFC17C5D4" box="[866,934,914,937]" country="Peru" name="Cusco" pageId="2" pageNumber="496">Cuzco</collectingRegion>
|
||
(
|
||
<collectingCountry id="F37C767FE56A010526F5DFDAFC5CC5D4" box="[951,1005,914,937]" name="Peru" pageId="2" pageNumber="496">Peru</collectingCountry>
|
||
:
|
||
<collectingRegion id="49AFF80DE56A010526BADFDAFB8FC5D4" box="[1016,1086,914,937]" country="Peru" name="Cusco" pageId="2" pageNumber="496">Cuzco</collectingRegion>
|
||
) and the recent study of several natural history collections led us to confirm the occurrence of this species in Callanga (two males,
|
||
<collectionCode id="ED7AAE2AE56A010521AEDFFDFA8DC5B1" box="[1260,1340,949,972]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="496">NMPC</collectionCode>
|
||
), Machu Picchu (one male, MZc) and
|
||
<collectingRegion id="49AFF80DE56A0105248CDF90FDB4C592" box="[462,517,984,1007]" country="Peru" name="Puno" pageId="2" pageNumber="496">Puno</collectingRegion>
|
||
(one male, MZc). Although we could not directly prove the presence of
|
||
<taxonomicName id="4C6B4D6CE56A0105204EDF90FA2DC592" box="[1292,1436,984,1007]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
<emphasis id="B91FEAFDE56A0105204EDF90FA2DC592" box="[1292,1436,984,1007]" italics="true" pageId="2" pageNumber="496">O. bidentatus</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingRegion id="49AFF80DE56A010525F0DFB2FF48C26C" box="[178,249,1018,1041]" country="Peru" name="Cusco" pageId="2" pageNumber="496">Cuzco</collectingRegion>
|
||
, it seems quite reasonable to confirm this locality too.
|
||
</paragraph>
|
||
<paragraph id="8BD436EFE56A01052585D855FB0EC20A" blockId="2.[151,1436,498,2020]" pageId="2" pageNumber="496">
|
||
<materialsCitation id="3B033CB2E56A01052585D855FB2DC249" ID-GBIF-Occurrence="1323867449" box="[199,1180,1053,1076]" country="Panama" location="Veraguas" pageId="2" pageNumber="496" specimenCount="1" stateProvince="Veraguas">
|
||
New geographic records for
|
||
<taxonomicName id="4C6B4D6CE56A0105274ED855FD10C249" box="[524,673,1053,1076]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
<emphasis id="B91FEAFDE56A0105274ED855FD10C249" box="[524,673,1053,1076]" italics="true" pageId="2" pageNumber="496">O. bidentatus</emphasis>
|
||
</taxonomicName>
|
||
are
|
||
<collectingCountry id="F37C767FE56A0105279BD855FC9CC249" box="[729,813,1053,1076]" name="Panama" pageId="2" pageNumber="496">Panama</collectingCountry>
|
||
(
|
||
<collectingRegion id="49AFF80DE56A01052602D855FC17C249" box="[832,934,1053,1076]" country="Panama" name="Veraguas" pageId="2" pageNumber="496">Veraguas</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37C767FE56A010526F4D855FBBBC249" box="[950,1034,1053,1076]" name="Panama" pageId="2" pageNumber="496">Panama</collectingCountry>
|
||
and
|
||
<collectingRegion id="49AFF80DE56A0105210BD855FB27C249" box="[1097,1174,1053,1076]" country="Panama" name="Darien" pageId="2" pageNumber="496">Darién</collectingRegion>
|
||
)
|
||
</materialsCitation>
|
||
, Trinidad and Tobago,
|
||
<materialsCitation id="3B033CB2E56A010525D5D876FB0EC20A" ID-GBIF-Occurrence="1323867448" country="Suriname" location="Napo" pageId="2" pageNumber="496" specimenCount="1" stateProvince="Napo">
|
||
<collectingCountry id="F37C767FE56A010525D5D876FF4CC228" box="[151,253,1086,1109]" name="Suriname" pageId="2" pageNumber="496">Suriname</collectingCountry>
|
||
(Paramaribo) and
|
||
<collectingCountry id="F37C767FE56A01052480D876FDAAC228" box="[450,539,1086,1109]" name="Ecuador" pageId="2" pageNumber="496">Ecuador</collectingCountry>
|
||
(
|
||
<collectingRegion id="49AFF80DE56A0105276AD876FDD2C228" box="[552,611,1086,1109]" country="Ecuador" name="Napo" pageId="2" pageNumber="496">Napo</collectingRegion>
|
||
and
|
||
<collectingRegion id="49AFF80DE56A010527D4D876FCD4C228" box="[662,869,1086,1109]" country="Ecuador" name="Zamora-Chinchipe" pageId="2" pageNumber="496">Zamora-Chinchipe</collectingRegion>
|
||
). More details about the geographic distribution of
|
||
<taxonomicName id="4C6B4D6CE56A010520C0D876FEB7C20A" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
<emphasis id="B91FEAFDE56A010520C0D876FEB7C20A" italics="true" pageId="2" pageNumber="496">O. bidentatus</emphasis>
|
||
</taxonomicName>
|
||
will soon be published in a revision of the
|
||
<emphasis id="B91FEAFDE56A01052790D828FC99C20A" box="[722,808,1120,1143]" italics="true" pageId="2" pageNumber="496">hirculus</emphasis>
|
||
group of the American
|
||
<taxonomicName id="4C6B4D6CE56A0105216AD828FB09C20A" box="[1064,1208,1120,1143]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAFDE56A0105216AD828FB09C20A" box="[1064,1208,1120,1143]" italics="true" pageId="2" pageNumber="496">Onthophagus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3716564E56A01032585D8CAFBF0C728" lastPageId="4" lastPageNumber="498" pageId="2" pageNumber="496" type="discussion">
|
||
<paragraph id="8BD436EFE56A01052585D8CAFA9CC31F" blockId="2.[151,1436,498,2020]" pageId="2" pageNumber="496">
|
||
<emphasis id="B91FEAFDE56A01052585D8CAFE84C2E4" bold="true" box="[199,309,1154,1177]" pageId="2" pageNumber="496">Remarks.</emphasis>
|
||
<bibRefCitation id="EFFA4B1EE56A0105247FD8CAFE50C2E4" author="Drapiez" box="[317,481,1154,1177]" pageId="2" pageNumber="496" refString="Drapiez, M. (1819) Descriptions de quelques insectes nouveaux. Annales Generales des Sciences Physiques, 1 (45 - 55), 130 - 138." type="journal article" year="1819">Drapiez (1819)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4C6B4D6CE56A01052714D8CAFD56C2E4" box="[598,743,1154,1177]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
<emphasis id="B91FEAFDE56A01052714D8CAFD56C2E4" box="[598,743,1154,1177]" italics="true" pageId="2" pageNumber="496">O. bidentatus</emphasis>
|
||
</taxonomicName>
|
||
from two specimens collected in Cayenne, French
|
||
<collectingCountry id="F37C767FE56A01052056D8CAFADBC2E4" box="[1300,1386,1154,1177]" name="Guyana" pageId="2" pageNumber="496">Guyana</collectingCountry>
|
||
, and received by M. J. Baudet-Lafarge (hereafter referred to as BL), who used to exchange natural history specimens with entomologists and amateurs around the World. It is likely that BL received these specimens from Mr Banon, a pharmacist settled in French
|
||
<collectingCountry id="F37C767FE56A01052495D8AEFD9BC280" box="[471,554,1254,1277]" name="Guyana" pageId="2" pageNumber="496">Guyana</collectingCountry>
|
||
with whom he had correspondence (short essay from Baudet-Lafarge’s original manuscript of his collection of
|
||
<taxonomicName id="4C6B4D6CE56A010524BBD940FDC2C362" box="[505,627,1288,1311]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
|
||
). According to
|
||
<bibRefCitation id="EFFA4B1EE56A01052666D940FC7CC362" author="Drapiez" box="[804,973,1288,1311]" pageId="2" pageNumber="496" refString="Drapiez, M. (1819) Descriptions de quelques insectes nouveaux. Annales Generales des Sciences Physiques, 1 (45 - 55), 130 - 138." type="journal article" year="1819">Drapiez (1819)</bibRefCitation>
|
||
, BL named these specimens
|
||
<emphasis id="B91FEAFDE56A0105205AD940FF58C33C" italics="true" pageId="2" pageNumber="496">Onthophage bicorne</emphasis>
|
||
(then latinized into
|
||
<taxonomicName id="4C6B4D6CE56A01052482D962FD1FC33C" box="[448,686,1322,1345]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bicornis">
|
||
<emphasis id="B91FEAFDE56A01052482D962FD1FC33C" box="[448,686,1322,1345]" italics="true" pageId="2" pageNumber="496">Onthophagus bicornis</emphasis>
|
||
</taxonomicName>
|
||
), although the French author never published even a short description of this species name, unless it was included in his rare manuscript (Description des insectes de ma collection).
|
||
</paragraph>
|
||
<paragraph id="8BD436EFE56A01052585D925FD2DC051" blockId="2.[151,1436,498,2020]" pageId="2" pageNumber="496">
|
||
Since the absence of cephalic horns to justify the epithet proposed by BL, Drapiez decided to replace this name with
|
||
<emphasis id="B91FEAFDE56A010525D5D9C6FEC3C3D8" box="[151,370,1422,1445]" italics="true" pageId="2" pageNumber="496">Onthophage bidenté</emphasis>
|
||
(then latinized into
|
||
<taxonomicName id="4C6B4D6CE56A01052714D9C6FCEFC3D8" box="[598,862,1422,1445]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
<emphasis id="B91FEAFDE56A01052714D9C6FCEFC3D8" box="[598,862,1422,1445]" italics="true" pageId="2" pageNumber="496">Onthophagus bidentatus</emphasis>
|
||
</taxonomicName>
|
||
) because of the characters observed in the pronotal protuberance: “
|
||
<emphasis id="B91FEAFDE56A0105247FD9F8FAEAC3BA" box="[317,1371,1456,1479]" italics="true" pageId="2" pageNumber="496">… nous n’avons point observé à la tête, d’organes qui justifiassent pleinement le nom de bicorne</emphasis>
|
||
,
|
||
<emphasis id="B91FEAFDE56A01052028D9F8FB50C394" italics="true" pageId="2" pageNumber="496">nous nous sommes permis de lui substituer celui de bidenté, tire du caractère du corselet qui paraÎt constant;</emphasis>
|
||
” [… we have not observed organs that fully justified the name
|
||
<emphasis id="B91FEAFDE56A01052737D9BBFD7CC077" box="[629,717,1523,1546]" italics="true" pageId="2" pageNumber="496">bicorne,</emphasis>
|
||
so we permitted to replace it with
|
||
<emphasis id="B91FEAFDE56A0105217ED9BBFB3BC077" box="[1084,1162,1523,1546]" italics="true" pageId="2" pageNumber="496">bidenté</emphasis>
|
||
, as the pronotal character seems to be constant] (
|
||
<bibRefCitation id="EFFA4B1EE56A010524CBDA5DFD3FC051" author="Drapiez" box="[393,654,1557,1580]" pageId="2" pageNumber="496" refString="Drapiez, M. (1819) Descriptions de quelques insectes nouveaux. Annales Generales des Sciences Physiques, 1 (45 - 55), 130 - 138." type="journal article" year="1819">Drapiez, 1819: 134–135</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD436EFE56A01052585DA7EFE86C0EC" blockId="2.[151,1436,498,2020]" pageId="2" pageNumber="496">
|
||
The name
|
||
<taxonomicName id="4C6B4D6CE56A0105247DDA7EFD5CC030" authority="Beaudet-Lafarge" authorityName="Beaudet-Lafarge" box="[319,749,1590,1613]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bicornis">
|
||
<emphasis id="B91FEAFDE56A0105247DDA7EFD81C030" box="[319,560,1590,1613]" italics="true" pageId="2" pageNumber="496">Onthophagus bicornis</emphasis>
|
||
Beaudet-Lafarge
|
||
</taxonomicName>
|
||
(misspelling of the author name as mentioned in the original paper) is reported (with its
|
||
<taxonomicName id="4C6B4D6CE56A010524FCDA10FD7CC012" authority="Lacordaire" authorityName="Lacordaire" box="[446,717,1624,1647]" pageId="2" pageNumber="496" rank="variety" variety="violaceus">
|
||
var.
|
||
<emphasis id="B91FEAFDE56A010524AFDA10FDE0C012" box="[493,593,1624,1647]" italics="true" pageId="2" pageNumber="496">violaceus</emphasis>
|
||
Lacordaire
|
||
</taxonomicName>
|
||
and its synonyms
|
||
<emphasis id="B91FEAFDE56A010526DADA10FC4FC012" box="[920,1022,1624,1647]" italics="true" pageId="2" pageNumber="496">inauratus</emphasis>
|
||
Lacordaire and
|
||
<emphasis id="B91FEAFDE56A010521F2DA10FB44C012" box="[1200,1269,1624,1647]" italics="true" pageId="2" pageNumber="496">ovinus</emphasis>
|
||
Lacordaire) by
|
||
<bibRefCitation id="EFFA4B1EE56A010525D5DA32FE82C0EC" author="Dejean" box="[151,307,1658,1681]" pageId="2" pageNumber="496" refString="Dejean, P. F. M. A. (1836) Catalogue des Coleopteres de la collection de M. le Comte Dejean. III ed. Mequignon, Marvis, 443 pp." type="book" year="1836">Dejean (1836)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD436EFE56A01052585DAD3FB0BC127" blockId="2.[151,1436,498,2020]" pageId="2" pageNumber="496">
|
||
In 1840, Laporte de Castelnau included
|
||
<taxonomicName id="4C6B4D6CE56A01052735DAD3FB91C0CF" authority="Beaudet-Lafarge" authorityName="Beaudet-Lafarge" box="[631,1056,1691,1714]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bicornis">
|
||
<emphasis id="B91FEAFDE56A01052735DAD3FCD4C0CF" box="[631,869,1691,1714]" italics="true" pageId="2" pageNumber="496">Onthophagus bicornis</emphasis>
|
||
Beaudet-Lafarge
|
||
</taxonomicName>
|
||
(misspelling of the author name as mentioned in the original paper) in his “Histoire Naturelle des Insectes Coléoptères”, along with a short description and bibliographic reference that refer undoubtedly to Drapiez’s work (“
|
||
<emphasis id="B91FEAFDE56A01052626DA96FC70C088" box="[868,961,1758,1781]" italics="true" pageId="2" pageNumber="496">Ann. des</emphasis>
|
||
.
|
||
<emphasis id="B91FEAFDE56A0105268CDA96FA30C088" box="[974,1409,1758,1781]" italics="true" pageId="2" pageNumber="496">Sc. phys., t. 1, p. 134, n. 13, pl. 7, Fig. 5.</emphasis>
|
||
”). Nonetheless, it is not clear whether Castelnau examined the
|
||
<typeStatus id="54D0884DE56A01052669DB48FCE8C16A" box="[811,857,1792,1815]" pageId="2" pageNumber="496">type</typeStatus>
|
||
specimens of this species or just used the diagnostic characters provided by
|
||
<bibRefCitation id="EFFA4B1EE56A010524D6DB6AFD89C144" author="Drapiez" box="[404,568,1826,1849]" pageId="2" pageNumber="496" refString="Drapiez, M. (1819) Descriptions de quelques insectes nouveaux. Annales Generales des Sciences Physiques, 1 (45 - 55), 130 - 138." type="journal article" year="1819">Drapiez (1819)</bibRefCitation>
|
||
to describe
|
||
<taxonomicName id="4C6B4D6CE56A010527FFDB6AFC87C144" box="[701,822,1826,1849]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bicornis">
|
||
<emphasis id="B91FEAFDE56A010527FFDB6AFC87C144" box="[701,822,1826,1849]" italics="true" pageId="2" pageNumber="496">O. bicornis</emphasis>
|
||
</taxonomicName>
|
||
. Anyhow, as formerly proposed by
|
||
<bibRefCitation id="EFFA4B1EE56A010521FCDB6AFF6FC127" author="Boucomont" pageId="2" pageNumber="496" refString="Boucomont, A. & Gillet, J. J. E. (1927) Scarabaeidae: Coprinae II, Termitotroginae. In: S. Schenkling (Eds.) Coleopterorum Catalogus, W. Junk, 90, pp. 103 - 263." type="journal article" year="1927">Boucomont & Gillet (1927)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFFA4B1EE56A01052450DB0BFE59C127" author="Blackwelder" box="[274,488,1859,1882]" pageId="2" pageNumber="496" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. Bulletin of the United States National Museum, 185, 189 - 341." type="journal article" year="1944">Blackwelder (1944)</bibRefCitation>
|
||
, there is no doubt about the authorship of Castelnau on
|
||
<emphasis id="B91FEAFDE56A01052179DB0BFB0BC127" box="[1083,1210,1859,1882]" italics="true" pageId="2" pageNumber="496">
|
||
<taxonomicName id="4C6B4D6CE56A01052179DB0BFB07C127" box="[1083,1206,1859,1882]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bicornis">O. bicornis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BD436EFE56A01052585DB2DFC81C199" blockId="2.[151,1436,498,2020]" pageId="2" pageNumber="496">
|
||
According to this information, it is conceivable to believe that
|
||
<taxonomicName id="4C6B4D6CE56A010526CBDB2DFBADC101" box="[905,1052,1893,1916]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
<emphasis id="B91FEAFDE56A010526CBDB2DFBADC101" box="[905,1052,1893,1916]" italics="true" pageId="2" pageNumber="496">O. bidentatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B4D6CE56A0105211ADB2DFB62C101" box="[1112,1235,1893,1916]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bicornis">
|
||
<emphasis id="B91FEAFDE56A0105211ADB2DFB62C101" box="[1112,1235,1893,1916]" italics="true" pageId="2" pageNumber="496">O. bicornis</emphasis>
|
||
</taxonomicName>
|
||
refer to the same biological entity, as the respective descriptions are based on specimens belonging to the same type series. Thus,
|
||
<taxonomicName id="4C6B4D6CE56A010525D5DBE2FDE1C1BC" authority="Drapiez, 1819" authorityName="Drapiez" authorityYear="1819" box="[151,592,1962,1985]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
<emphasis id="B91FEAFDE56A010525D5DBE2FE15C1BC" box="[151,420,1962,1985]" italics="true" pageId="2" pageNumber="496">Onthophagus bidentatus</emphasis>
|
||
<bibRefCitation id="EFFA4B1EE56A010524F0DBE2FDE1C1BC" author="Drapiez" box="[434,592,1962,1985]" pageId="2" pageNumber="496" refString="Drapiez, M. (1819) Descriptions de quelques insectes nouveaux. Annales Generales des Sciences Physiques, 1 (45 - 55), 130 - 138." type="journal article" year="1819">Drapiez, 1819</bibRefCitation>
|
||
</taxonomicName>
|
||
possesses all the requirements to be accepted as valid name instead of
|
||
<taxonomicName id="4C6B4D6CE56A010525D5DB85FD86C199" authority="Castelnau, 1840" authorityName="Castelnau" authorityYear="1840" box="[151,567,1997,2020]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="496" phylum="Arthropoda" rank="species" species="bicornis">
|
||
<emphasis id="B91FEAFDE56A010525D5DB85FE35C199" box="[151,388,1997,2020]" italics="true" pageId="2" pageNumber="496">Onthophagus bicornis</emphasis>
|
||
<bibRefCitation id="EFFA4B1EE56A010524C9DB85FD86C199" author="Castelnau" box="[395,567,1997,2020]" pageId="2" pageNumber="496" refString="Castelnau, F. L. de (1840) Histoire naturelle des Animaux articules, annelides, crustaces, arachnides, myriapodes et insectes. Histoire naturelle des Insectes Coleopteres. Tome deuxieme. P. Dumenil, Paris, 563 pp., 38 pls." type="book" year="1840">Castelnau, 1840</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFFA4B1EE56A01052707DB85FC95C199" author="ICZN" box="[581,804,1997,2020]" pageId="2" pageNumber="496" refString="ICZN (International Commission on Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN, 1999: Art. 23</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF146667E56B010425D5DBD7FC9DC1AF" httpUri="https://zenodo.org/record/272490/files/figure.png" pageId="3" pageNumber="497" targetBox="[166,1412,226,1912]" targetPageId="3">
|
||
<paragraph id="8BD436EFE56B010425D5DBD7FC9DC1AF" blockId="3.[151,1436,1951,2003]" pageId="3" pageNumber="497">
|
||
<emphasis id="B91FEAFDE56B010425D5DBD7FEA1C1C8" bold="true" box="[151,272,1951,1973]" pageId="3" pageNumber="497">FIGURE 1.</emphasis>
|
||
Dorsal habitus and labels of the primary types.
|
||
<emphasis id="B91FEAFDE56B010427B9DBD7FC9BC1C8" bold="true" box="[763,810,1951,1973]" pageId="3" pageNumber="497">A, F</emphasis>
|
||
:
|
||
<taxonomicName id="4C6B4D6CE56B01042678DBE8FC0EC1C8" box="[826,959,1952,1973]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="497" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
<emphasis id="B91FEAFDE56B01042678DBE8FC0EC1C8" box="[826,959,1952,1973]" italics="true" pageId="3" pageNumber="497">O. bidentatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B91FEAFDE56B010426B7DBE8FBDAC1C8" box="[1013,1131,1952,1973]" italics="true" pageId="3" pageNumber="497">
|
||
<taxonomicName id="4C6B4D6CE56B010426B7DBE8FBD6C1C8" box="[1013,1127,1952,1973]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="497" phylum="Arthropoda" rank="species" species="bicornis">O. bicornis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B91FEAFDE56B01042136DBD7FB14C1C8" bold="true" box="[1140,1189,1951,1973]" pageId="3" pageNumber="497">B, G</emphasis>
|
||
:
|
||
<emphasis id="B91FEAFDE56B010421F4DBE8FA89C1C8" box="[1206,1336,1952,1973]" italics="true" pageId="3" pageNumber="497">
|
||
<taxonomicName id="4C6B4D6CE56B010421F4DBE8FA84C1C8" box="[1206,1333,1952,1973]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="497" phylum="Arthropoda" rank="species" species="striatulus">O. striatulus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B91FEAFDE56B01042003DBD7FAC4C1C8" bold="true" box="[1345,1397,1951,1973]" pageId="3" pageNumber="497">C, H</emphasis>
|
||
:
|
||
<emphasis id="B91FEAFDE56B010420C6DBE8FEAEC1AF" italics="true" pageId="3" pageNumber="497">
|
||
<taxonomicName id="4C6B4D6CE56B010420C6DBE8FEAAC1AF" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="497" phylum="Arthropoda" rank="species" species="semichalcites">O. semichalcites</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B91FEAFDE56B01042466DBF5FEFBC1AE" bold="true" box="[292,330,1981,2003]" pageId="3" pageNumber="497">D, I</emphasis>
|
||
:
|
||
<taxonomicName id="4C6B4D6CE56B01042415DBF5FE00C1AF" box="[343,433,1981,2002]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="497" phylum="Arthropoda" rank="species" species="gibber">
|
||
<emphasis id="B91FEAFDE56B01042415DBF5FE00C1AF" box="[343,433,1981,2002]" italics="true" pageId="3" pageNumber="497">O. gibber</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B91FEAFDE56B010424FCDBF5FE5BC1AE" bold="true" box="[446,490,1981,2003]" pageId="3" pageNumber="497">E, L</emphasis>
|
||
:
|
||
<taxonomicName id="4C6B4D6CE56B010424B4DBF5FDCAC1AF" box="[502,635,1981,2002]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="497" phylum="Arthropoda" rank="species" species="gibberosus">
|
||
<emphasis id="B91FEAFDE56B010424B4DBF5FDCAC1AF" box="[502,635,1981,2002]" italics="true" pageId="3" pageNumber="497">O. gibberosus</emphasis>
|
||
</taxonomicName>
|
||
. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD436EFE56C01032585DCD0FBF0C728" blockId="4.[151,1436,152,775]" pageId="4" pageNumber="498">
|
||
<materialsCitation id="3B033CB2E56C01032585DCD0FBF0C728" ID-GBIF-Occurrence="1323867452" collectionCode="MNHN, ICZN" collectorName="R. Oberthur & Art." county="During" location="It" municipality="Drapiez" pageId="4" pageNumber="498" specimenCount="2" stateProvince="A" typeStatus="lectotype">
|
||
<collectingCounty id="62B54E63E56C01032585DCD0FEA3C6D2" box="[199,274,152,175]" pageId="4" pageNumber="498">During</collectingCounty>
|
||
a recent visit to the
|
||
<collectionCode id="ED7AAE2AE56C0103274BDCD0FDEBC6D2" box="[521,602,152,175]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="498">MNHN</collectionCode>
|
||
we found a specimen whose labels and morphology matched exactly the diagnosis provided by
|
||
<bibRefCitation id="EFFA4B1EE56C010324C8DCF1FD9CC6AD" author="Drapiez" box="[394,557,185,208]" pageId="4" pageNumber="498" refString="Drapiez, M. (1819) Descriptions de quelques insectes nouveaux. Annales Generales des Sciences Physiques, 1 (45 - 55), 130 - 138." type="journal article" year="1819">
|
||
<collectingMunicipality id="6BB0AC95E56C010324C8DCF1FE6FC6AD" box="[394,478,185,208]" pageId="4" pageNumber="498">Drapiez</collectingMunicipality>
|
||
(1819)
|
||
</bibRefCitation>
|
||
(
|
||
<figureCitation id="13502A6AE56C01032779DCF1FDC6C6AD" box="[571,631,185,208]" captionStart="FIGURE 1" captionStartId="3.[151,244,1951,1973]" captionTargetBox="[166,1412,226,1912]" captionTargetId="figure@3.[150,1436,194,1928]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus and labels of the primary types. A, F: O. bidentatus and O. bicornis. B, G: O. striatulus. C, H: O. semichalcites. D, I: O. gibber. E, L: O. gibberosus. Scale bar: 1 mm." httpUri="https://zenodo.org/record/272490/files/figure.png" pageId="4" pageNumber="498">Fig. 1</figureCitation>
|
||
<collectingRegion id="49AFF80DE56C01032735DCF1FD3EC6AD" box="[631,655,185,208]" country="Nigeria" name="Rivers" pageId="4" pageNumber="498">A</collectingRegion>
|
||
, F).
|
||
<location id="8EB46034E56C01032783DCF1FD60C6AD" LSID="urn:lsid:plazi:treatment:03C287F9E56A010325D5DDBAFB65C57A:8EB46034E56C01032783DCF1FD60C6AD" box="[705,721,185,208]" county="During" municipality="Drapiez" name="It" pageId="4" pageNumber="498" stateProvince="A">It</location>
|
||
is very likely that this female is one of the two specimens donated by
|
||
<location id="8EB46034E56C01032583DC92FED9C68C" LSID="urn:lsid:plazi:treatment:03C287F9E56A010325D5DDBAFB65C57A:8EB46034E56C01032583DC92FED9C68C" box="[193,360,218,241]" county="During" municipality="Drapiez" name="Baudet-Lafarge" pageId="4" pageNumber="498" stateProvince="A">Baudet-Lafarge</location>
|
||
to the
|
||
<location id="8EB46034E56C0103248EDC92FDAEC68C" LSID="urn:lsid:plazi:treatment:03C287F9E56A010325D5DDBAFB65C57A:8EB46034E56C0103248EDC92FDAEC68C" box="[460,543,218,241]" county="During" municipality="Drapiez" name="Belgian" pageId="4" pageNumber="498" stateProvince="A">Belgian</location>
|
||
naturalist at the beginning of the 19th century.
|
||
<location id="8EB46034E56C0103212ADC92FB65C68C" LSID="urn:lsid:plazi:treatment:03C287F9E56A010325D5DDBAFB65C57A:8EB46034E56C0103212ADC92FB65C68C" box="[1128,1236,218,241]" county="During" municipality="Drapiez" name="Therefore" pageId="4" pageNumber="498" stateProvince="A">Therefore</location>
|
||
, considering that
|
||
<taxonomicName id="4C6B4D6CE56C010325D5DCB4FD8EC76E" authority="Drapiez, 1819" authorityName="Drapiez" authorityYear="1819" box="[151,575,252,275]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
<emphasis id="B91FEAFDE56C010325D5DCB4FE2FC76E" box="[151,414,252,275]" italics="true" pageId="4" pageNumber="498">Onthophagus bidentatus</emphasis>
|
||
<bibRefCitation id="EFFA4B1EE56C010324E4DCB4FD8EC76E" author="Drapiez" box="[422,575,252,275]" pageId="4" pageNumber="498" refString="Drapiez, M. (1819) Descriptions de quelques insectes nouveaux. Annales Generales des Sciences Physiques, 1 (45 - 55), 130 - 138." type="journal article" year="1819">Drapiez, 1819</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B4D6CE56C01032735DCB4FBADC76E" authority="Castelnau, 1840" authorityName="Castelnau" authorityYear="1840" box="[631,1052,252,275]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="species" species="bicornis">
|
||
<emphasis id="B91FEAFDE56C01032735DCB4FCD7C76E" box="[631,870,252,275]" italics="true" pageId="4" pageNumber="498">Onthophagus bicornis</emphasis>
|
||
<bibRefCitation id="EFFA4B1EE56C0103262DDCB4FBADC76E" author="Castelnau" box="[879,1052,252,275]" pageId="4" pageNumber="498" refString="Castelnau, F. L. de (1840) Histoire naturelle des Animaux articules, annelides, crustaces, arachnides, myriapodes et insectes. Histoire naturelle des Insectes Coleopteres. Tome deuxieme. P. Dumenil, Paris, 563 pp., 38 pls." type="book" year="1840">Castelnau, 1840</bibRefCitation>
|
||
</taxonomicName>
|
||
have been described on specimens belonging to the same
|
||
<typeStatus id="54D0884DE56C010324D0DD55FE71C749" box="[402,448,285,308]" pageId="4" pageNumber="498">type</typeStatus>
|
||
series, we decided to designate the
|
||
<typeStatus id="54D0884DE56C01032612DD55FC03C749" box="[848,946,285,308]" pageId="4" pageNumber="498" type="lectotype">lectotype</typeStatus>
|
||
of both species names on this single female specimen deposited in the
|
||
<collectorName id="269E5339E56C010324F3DD76FD80C728" box="[433,561,318,341]" pageId="4" pageNumber="498">R. Oberthür</collectorName>
|
||
collection (
|
||
<collectionCode id="ED7AAE2AE56C010327ECDD76FCB2C728" box="[686,771,318,341]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="498">MNHN</collectionCode>
|
||
) (
|
||
<bibRefCitation id="EFFA4B1EE56C0103265ADD76FB85C728" author="ICZN" box="[792,1076,318,341]" pageId="4" pageNumber="498" refString="ICZN (International Commission on Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">
|
||
<collectionCode id="ED7AAE2AE56C0103265ADD76FCEBC728" box="[792,858,318,341]" pageId="4" pageNumber="498">ICZN</collectionCode>
|
||
, 1999:
|
||
<collectorName id="269E5339E56C010326EBDD76FC65C728" box="[937,980,318,341]" pageId="4" pageNumber="498">Art.</collectorName>
|
||
72.6; 74
|
||
</bibRefCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3716564E56C01032585DD28FF5CC460" pageId="4" pageNumber="498" type="materials_examined">
|
||
<paragraph id="8BD436EFE56C01032585DD28FF5CC460" blockId="4.[151,1436,152,775]" pageId="4" pageNumber="498">
|
||
<emphasis id="B91FEAFDE56C01032585DD28FE44C70A" bold="true" box="[199,501,352,375]" pageId="4" pageNumber="498">
|
||
<typeStatus id="54D0884DE56C01032585DD28FF4EC70A" box="[199,255,352,375]" pageId="4" pageNumber="498">Type</typeStatus>
|
||
specimens examined.
|
||
</emphasis>
|
||
Of
|
||
<taxonomicName id="4C6B4D6CE56C0103275DDD28FD01C70A" box="[543,688,352,375]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
<emphasis id="B91FEAFDE56C0103275DDD28FD01C70A" box="[543,688,352,375]" italics="true" pageId="4" pageNumber="498">O. bidentatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B4D6CE56C010327A6DD28FCEDC70A" box="[740,860,352,375]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="species" species="bicornis">
|
||
<emphasis id="B91FEAFDE56C010327A6DD28FCEDC70A" box="[740,860,352,375]" italics="true" pageId="4" pageNumber="498">O. bicornis</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<typeStatus id="54D0884DE56C0103262EDD28FC6CC70A" box="[876,989,352,375]" pageId="4" pageNumber="498" type="lectotype">
|
||
<emphasis id="B91FEAFDE56C0103262EDD28FC6CC70A" bold="true" box="[876,989,352,375]" pageId="4" pageNumber="498">Lectotype</emphasis>
|
||
</typeStatus>
|
||
(
|
||
<specimenCount id="9D6DFD66E56C010326ACDD28FBB1C70A" box="[1006,1024,352,375]" pageId="4" pageNumber="498" type="female">♀</specimenCount>
|
||
<collectionCode id="ED7AAE2AE56C01032145DD28FBEFC70A" box="[1031,1118,352,375]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="498">MNHN</collectionCode>
|
||
): 1:
|
||
<collectionCode id="ED7AAE2AE56C010321CEDD28FAE4C70A" box="[1164,1365,352,375]" pageId="4" pageNumber="498">MUSÉUM PARIS</collectionCode>
|
||
, 1952, coll. R. OBERTHÜR (printed on white label with black border). 2:
|
||
<taxonomicName id="4C6B4D6CE56C010326C4DDC9FC51C7E5" box="[902,992,385,408]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="species" species="bicornis">Bicornis</taxonomicName>
|
||
Bd-Laf, Cayen.,
|
||
<typeStatus id="54D0884DE56C010321E0DDC9FB59C7E5" box="[1186,1256,385,408]" pageId="4" pageNumber="498">Typus</typeStatus>
|
||
! (handwritten in italic on cream label with black line at the bottom). 3: Cayenna (handwritten in italic on cream label). 4:
|
||
<typeStatus id="54D0884DE56C010321BEDDEAFA26C7C4" box="[1276,1431,418,441]" pageId="4" pageNumber="498" type="lectotype">LECTOTYPE</typeStatus>
|
||
, female symbol,
|
||
<taxonomicName id="4C6B4D6CE56C01032404DD8CFDC5C7A6" authority="Cast." authorityName="Cast." box="[326,628,452,475]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="species" species="bicornis">Onthophagus bicornis Cast.</taxonomicName>
|
||
, des.
|
||
<bibRefCitation id="EFFA4B1EE56C010327F6DD8CFC0CC7A6" author="Vaz-de-Mello" box="[692,957,452,475]" pageId="4" pageNumber="498" refString="Rossini, M., Vaz-de-Mello, F. Z. & Mann, D. J. (2014) Onthophagus cervicornis Kirby, 1825, new synonym under Onthophagus dama (Fabricius, 1798) (Coleoptera: Scarabaeidae: Scarabaeinae). ZooKeys, 419, 111 - 115. http: // dx. doi. org / 10.3897 / zookeys. 419.7849" type="journal article" year="2014">F.Z. Vaz-de-Mello, 2014</bibRefCitation>
|
||
(printed and handwritten, on red label with black border). 5:
|
||
<typeStatus id="54D0884DE56C0103241CDDADFE48C781" box="[350,505,485,508]" pageId="4" pageNumber="498" type="lectotype">LECTOTYPE</typeStatus>
|
||
,
|
||
<taxonomicName id="4C6B4D6CE56C0103274BDDADFB97C781" authority="Drap., M. Rossini 2016" authorityName="Drap., M. Rossini" authorityYear="2016" box="[521,1062,485,508]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="species" species="bidentatus">
|
||
Onthophagus bidentatus Drap.,
|
||
<bibRefCitation id="EFFA4B1EE56C01032629DDADFB97C781" author="Rossini" box="[875,1062,485,508]" pageId="4" pageNumber="498" refString="Rossini, M. & Vaz-de-Mello F. Z. (2016) On the identity of Onthophagus columbianus Boucomont, 1932 (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa, 4061 (3), 296 - 300. http: // dx. doi. org / 10.11646 / zootaxa. 4061.3.10" type="journal article" year="2016">M. Rossini 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
(printed on red label with black border).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3716564E56C01032585DE60FB65C57A" pageId="4" pageNumber="498" type="reference_group">
|
||
<paragraph id="8BD436EFE56C01032585DE60FB65C57A" blockId="4.[151,1436,152,775]" pageId="4" pageNumber="498">
|
||
<materialsCitation id="3B033CB2E56C01032585DE60FE7FC41D" ID-GBIF-Occurrence="1323867450" collectionCode="MNHN" country="Benin" location="Benin" pageId="4" pageNumber="498" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
|
||
Of
|
||
<taxonomicName id="4C6B4D6CE56C010325ADDE60FE14C442" ID-CoL="7XNXG" box="[239,421,552,575]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="species" species="semichalcites">
|
||
<emphasis id="B91FEAFDE56C010325ADDE60FE14C442" box="[239,421,552,575]" italics="true" pageId="4" pageNumber="498">O. semichalcites</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<typeStatus id="54D0884DE56C010324FBDE60FD98C442" box="[441,553,552,575]" pageId="4" pageNumber="498" type="lectotype">
|
||
<emphasis id="B91FEAFDE56C010324FBDE60FD98C442" bold="true" box="[441,553,552,575]" pageId="4" pageNumber="498">Lectotype</emphasis>
|
||
</typeStatus>
|
||
(
|
||
<specimenCount id="9D6DFD66E56C0103277CDE60FDE3C442" box="[574,594,552,575]" pageId="4" pageNumber="498" type="male">♂</specimenCount>
|
||
<collectionCode id="ED7AAE2AE56C0103271DDE60FD07C442" box="[607,694,552,575]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="498">MNHN</collectionCode>
|
||
): 1:
|
||
<collectingCountry id="F37C767FE56C010327ADDE60FC9EC442" box="[751,815,552,575]" name="Benin" pageId="4" pageNumber="498">Benin</collectingCountry>
|
||
(handwritten on cream label). 2:
|
||
<taxonomicName id="4C6B4D6CE56C010321F7DE60FA2DC442" ID-CoL="7XNXG" authority="d'Orb." authorityName="d'Orb." box="[1205,1436,552,575]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="495" phylum="Arthropoda" rank="species" species="semichalcites">Semichalcites d’Orb.</taxonomicName>
|
||
(handwritten on cream label)
|
||
</materialsCitation>
|
||
. TYPE (printed on red label). 3: MUSEUM PARIS, Coll. H. d’ORBIGNY 1915 (printed on green label). 4: Espèce Américaine, Y. CAMBEFORT DET. (printed and handwritten on white label with black border). 5: LECTOTYPE, male symbol,
|
||
<taxonomicName id="4C6B4D6CE56C01032744DEC4FC1CC4DE" ID-CoL="7XNXG" authority="d'Orbigny" authorityName="d'Orbigny" box="[518,941,652,675]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="species" species="semichalcites">Onthophagus semichalcites d’Orbigny</taxonomicName>
|
||
, des.
|
||
<bibRefCitation id="EFFA4B1EE56C010326B7DEC4FAB6C4DE" author="Vaz-de-Mello" box="[1013,1287,652,675]" pageId="4" pageNumber="498" refString="Rossini, M., Vaz-de-Mello, F. Z. & Mann, D. J. (2014) Onthophagus cervicornis Kirby, 1825, new synonym under Onthophagus dama (Fabricius, 1798) (Coleoptera: Scarabaeidae: Scarabaeinae). ZooKeys, 419, 111 - 115. http: // dx. doi. org / 10.3897 / zookeys. 419.7849" type="journal article" year="2014">F.Z. Vaz-de-Mello, 2014</bibRefCitation>
|
||
(printed and handwritten on red label).
|
||
<materialsCitation id="3B033CB2E56C010324FADEE5FAABC4B9" ID-GBIF-Occurrence="1323867451" box="[440,1306,685,708]" collectionCode="MNHN" country="Benin" location="Benin" pageId="4" pageNumber="498" specimenCount="6" specimenCount-female="5" specimenCount-male="1" typeStatus="paralectotype">
|
||
<typeStatus id="54D0884DE56C010324FADEE5FDEEC4B9" box="[440,607,685,708]" pageId="4" pageNumber="498" type="paralectotype">
|
||
<emphasis id="B91FEAFDE56C010324FADEE5FDEEC4B9" bold="true" box="[440,607,685,708]" pageId="4" pageNumber="498">Paralectotypes</emphasis>
|
||
</typeStatus>
|
||
(
|
||
<specimenCount id="9D6DFD66E56C01032731DEE5FD2CC4B9" box="[627,669,685,708]" pageId="4" pageNumber="498" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D6DFD66E56C010327E8DEE5FD59C4B9" box="[682,744,685,708]" pageId="4" pageNumber="498" type="female">5 ♀♀</specimenCount>
|
||
<collectionCode id="ED7AAE2AE56C010327B2DEE5FCF6C4B9" box="[752,839,685,708]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="498">MNHN</collectionCode>
|
||
): 1:
|
||
<collectingCountry id="F37C767FE56C0103263ADEE5FC09C4B9" box="[888,952,685,708]" name="Benin" pageId="4" pageNumber="498">Benin</collectingCountry>
|
||
(handwritten on cream label). 2
|
||
</materialsCitation>
|
||
: MUSEUM PARIS, Coll. H. d’ORBIGNY 1915 (printed on blue label). 3: PARALECTOTYPE,
|
||
<taxonomicName id="4C6B4D6CE56C0103212DDE86FEB6C57A" ID-CoL="7XNXG" authority="d'Orbigny" authorityName="d'Orbigny" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="498" phylum="Arthropoda" rank="species" species="semichalcites">Onthophagus semichalcites d’Orbigny</taxonomicName>
|
||
, des.
|
||
<bibRefCitation id="EFFA4B1EE56C01032400DEB8FDF9C57A" author="Vaz-de-Mello" box="[322,584,752,775]" pageId="4" pageNumber="498" refString="Rossini, M., Vaz-de-Mello, F. Z. & Mann, D. J. (2014) Onthophagus cervicornis Kirby, 1825, new synonym under Onthophagus dama (Fabricius, 1798) (Coleoptera: Scarabaeidae: Scarabaeinae). ZooKeys, 419, 111 - 115. http: // dx. doi. org / 10.3897 / zookeys. 419.7849" type="journal article" year="2014">F.Z. Vaz-de-Mello, 2014</bibRefCitation>
|
||
(printed and handwritten on yellow label with black border).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |