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<document id="BB980E87B8D18C12020273C3BA7D2AE4" ID-CLB-Dataset="182444" ID-DOI="10.11646/zootaxa.5202.1.1" ID-GBIF-Dataset="45ba25c1-47a9-4001-93d9-294815bf37e7" ID-ISSN="1175-5326" ID-Zenodo-Dep="7285600" ID-ZooBank="BF4831D2-D98B-4265-9138-03DB8607B826" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1667554680289" checkinUser="plazi" docAuthor="Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta &amp; Riva, Ignacio De La" docDate="2022" docId="03C287E8FFBCFF8DFF4BFDBFFABCE687" docLanguage="en" docName="zootaxa.5202.1.1.pdf" docOrigin="Zootaxa 5202 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5202.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ancylodactylus koehleri" docType="treatment" docVersion="9" lastPageNumber="31" masterDocId="FFFBFF90FFA2FF93FFDCFF9DFFB5E232" masterDocTitle="Synopsis of the terrestrial Reptiles of Equatorial Guinea" masterLastPageNumber="197" masterPageNumber="1" pageNumber="31" updateTime="1699409742602" updateUser="ExternalLinkService">
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<mods:title id="E58DC0C4FC48A69A0DD10781ABD00BB7">Synopsis of the terrestrial Reptiles of Equatorial Guinea</mods:title>
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<treatment id="03C287E8FFBCFF8DFF4BFDBFFABCE687" ID-DOI="http://doi.org/10.5281/zenodo.7293391" ID-GBIF-Taxon="204305625" ID-Zenodo-Dep="7293391" LSID="urn:lsid:plazi:treatment:03C287E8FFBCFF8DFF4BFDBFFABCE687" httpUri="http://treatment.plazi.org/id/03C287E8FFBCFF8DFF4BFDBFFABCE687" lastPageNumber="31" pageId="30" pageNumber="31">
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<paragraph id="8BD436FEFFBCFF8DFF4BFDBFFD33E00E" blockId="30.[151,646,546,609]" box="[151,646,546,573]" pageId="30" pageNumber="31">
<heading id="D09C8192FFBCFF8DFF4BFDBFFD33E00E" bold="true" box="[151,646,546,573]" fontSize="11" level="2" pageId="30" pageNumber="31" reason="4">
<taxonomicName id="4C6B4D7DFFBCFF8DFF4BFDBFFD33E00E" authority="(Mertens, 1937)" baseAuthorityName="Mertens" baseAuthorityYear="1937" box="[151,646,546,573]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="koehleri">
<emphasis id="B91FEAECFFBCFF8DFF4BFDBFFD33E00E" bold="true" box="[151,646,546,573]" pageId="30" pageNumber="31">
<emphasis id="B91FEAECFFBCFF8DFF4BFDBFFE0FE00F" bold="true" box="[151,442,546,573]" italics="true" pageId="30" pageNumber="31">Ancylodactylus koehleri</emphasis>
(
<bibRefCitation id="EFFA4B0FFFBCFF8DFE15FDBFFDCBE00E" author="Mertens, R." box="[457,638,546,572]" pageId="30" pageNumber="31" pagination="381 - 384" refId="ref84630" refString="Mertens, R. (1937) Eine neue, tiergeographisch bemerkenswerte Eidechse aus Kamerun. Senckenbergiana, 19, 381 - 384." type="journal article" year="1937">Mertens, 1937</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3716575FFBCFF8DFF4BFDDAFEBDE053" box="[151,264,583,609]" pageId="30" pageNumber="31" type="description">
<paragraph id="8BD436FEFFBCFF8DFF4BFDDAFEBDE053" blockId="30.[151,646,546,609]" box="[151,264,583,609]" pageId="30" pageNumber="31">
<figureCitation id="13502A7BFFBCFF8DFF4BFDDAFEBDE053" box="[151,264,583,609]" captionStart="FIGURE 7" captionStartId="33.[151,250,1918,1942]" captionTargetBox="[153,1434,191,1894]" captionTargetId="figure-18@33.[153,1434,181,1894]" captionTargetPageId="33" captionText="FIGURE 7. From left to right: dorsal, lateral, and ventral views of the head of: Agama lebretoni (AB), male (A: MNCN 4725) and female (B: MNCN 4726); and dorsal and ventral views of Ancylodactylus koehleri (C) (R-0014927). Photos: AS-V (AB) and Cornell University Museum of Vertebrates (C)." figureDoi="http://doi.org/10.5281/zenodo.7285629" httpUri="https://zenodo.org/record/7285629/files/figure.png" pageId="30" pageNumber="31">Figure 7C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFBCFF8DFF4BFD11FC8DE0D3" pageId="30" pageNumber="31" type="reference_group">
<paragraph id="8BD436FEFFBCFF8DFF4BFD11FC88E091" blockId="30.[151,829,651,737]" box="[151,829,651,676]" pageId="30" pageNumber="31">
<taxonomicName id="4C6B4D7DFFBCFF8DFF4BFD11FE45E091" authority="Mertens, 1937" authorityName="Mertens" authorityYear="1937" box="[151,496,651,675]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="kohleri">
<emphasis id="B91FEAECFFBCFF8DFF4BFD11FEE0E091" box="[151,341,651,675]" italics="true" pageId="30" pageNumber="31">Cnemaspis köhleri</emphasis>
<bibRefCitation id="EFFA4B0FFFBCFF8DFE87FD11FE45E091" author="Mertens, R." box="[347,496,651,675]" pageId="30" pageNumber="31" pagination="381 - 384" refId="ref84630" refString="Mertens, R. (1937) Eine neue, tiergeographisch bemerkenswerte Eidechse aus Kamerun. Senckenbergiana, 19, 381 - 384." type="journal article" year="1937">Mertens, 1937</bibRefCitation>
</taxonomicName>
.
<typeStatus id="54D0885CFFBCFF8DFE25FD11FD9EE096" box="[505,555,652,676]" pageId="30" pageNumber="31">Type</typeStatus>
locality:
<location id="8EB46025FFBCFF8DFD57FD16FD77E091" LSID="urn:lsid:plazi:treatment:03C287E8FFBCFF8DFF4BFDBFFABCE687:8EB46025FFBCFF8DFD57FD16FD77E091" box="[651,706,651,675]" country="Cameroon" name="Buea" pageId="30" pageNumber="31">Buéa</location>
,
<collectingCountry id="F37C766EFFBCFF8DFD10FD16FC8CE091" box="[716,825,651,675]" name="Cameroon" pageId="30" pageNumber="31">Cameroun</collectingCountry>
.
</paragraph>
<paragraph id="8BD436FEFFBCFF8DFF4BFD37FC9AE0F0" blockId="30.[151,829,651,737]" box="[151,815,682,706]" pageId="30" pageNumber="31">
<treatmentCitationGroup id="AB7B11D0FFBCFF8DFF4BFD37FC9AE0F0" box="[151,815,682,706]" pageId="30" pageNumber="31">
<taxonomicName id="4C6B4D7DFFBCFF8DFF4BFD37FDE2E0F0" authority="Mertens, 1937" authorityName="Mertens" authorityYear="1937" box="[151,599,682,706]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="subSpecies" species="africanus" subSpecies="kohleri">
<emphasis id="B91FEAECFFBCFF8DFF4BFD37FE0EE0F0" box="[151,443,682,706]" italics="true" pageId="30" pageNumber="31">Cnemaspis africanus köhleri</emphasis>
<bibRefCitation id="EFFA4B0FFFBCFF8DFE1DFD37FDE2E0F0" author="Mertens, R." box="[449,599,682,706]" pageId="30" pageNumber="31" pagination="381 - 384" refId="ref84630" refString="Mertens, R. (1937) Eine neue, tiergeographisch bemerkenswerte Eidechse aus Kamerun. Senckenbergiana, 19, 381 - 384." type="journal article" year="1937">Mertens, 1937</bibRefCitation>
</taxonomicName>
:
<treatmentCitation id="0ACA10EFFFBCFF8DFDBDFD37FC9EE0F0" author="Loveridge, A." box="[609,811,682,706]" page="90" pageId="30" pageNumber="31" year="1947">
<bibRefCitation id="EFFA4B0FFFBCFF8DFDBDFD37FC9EE0F0" author="Loveridge, A." box="[609,811,682,706]" pageId="30" pageNumber="31" pagination="1 - 469" refId="ref83041" refString="Loveridge, A. (1947) Revision of the African lizards of the family Gekkonidae. Bulletin of the Museum of Comparative Zoology, 98, 1 - 469." type="journal article" year="1947">Loveridge 1947: 90</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD436FEFFBCFF8DFF4BFD55FC8DE0D3" blockId="30.[151,829,651,737]" box="[151,824,712,737]" pageId="30" pageNumber="31">
<taxonomicName id="4C6B4D7DFFBCFF8DFF4BFD55FD8CE0D3" authority="(Mertens, 1937)" baseAuthorityName="Mertens" baseAuthorityYear="1937" box="[151,569,712,737]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="koehleri">
<emphasis id="B91FEAECFFBCFF8DFF4BFD55FE3EE0D2" box="[151,395,712,736]" italics="true" pageId="30" pageNumber="31">Ancylodactylus koehleri</emphasis>
(
<bibRefCitation id="EFFA4B0FFFBCFF8DFE45FD54FD84E0D3" author="Mertens, R." box="[409,561,713,737]" pageId="30" pageNumber="31" pagination="381 - 384" refId="ref84630" refString="Mertens, R. (1937) Eine neue, tiergeographisch bemerkenswerte Eidechse aus Kamerun. Senckenbergiana, 19, 381 - 384." type="journal article" year="1937">Mertens, 1937</bibRefCitation>
)
</taxonomicName>
:
<bibRefCitation id="EFFA4B0FFFBCFF8DFD9EFD55FC81E0D3" author="Malonza, P. K. &amp; Bauer, A. M." box="[578,820,712,737]" pageId="30" pageNumber="31" pagination="101 - 139" refId="ref83696" refString="Malonza, P. K. &amp; Bauer, A. M. (2022) Resurrection of the African gecko genus Ancylodactylus Muller, 1907 (Squamata: Gekkonidae) and description of six new species from Kenya. Zootaxa, 5141 (2), 101 - 139. https: // doi. org / 10.11646 / zootaxa. 5141.2.1" type="journal article" year="2022">Malonza &amp; Bauer 2022</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFBCFF8DFF4BFC93FB98E143" pageId="30" pageNumber="31" type="distribution">
<paragraph id="8BD436FEFFBCFF8DFF4BFC93FB98E143" blockId="30.[151,1437,782,1205]" pageId="30" pageNumber="31">
<emphasis id="B91FEAECFFBCFF8DFF4BFC93FE84E11A" bold="true" box="[151,305,782,808]" pageId="30" pageNumber="31">Distribution.</emphasis>
This species occurs in
<collectingCountry id="F37C766EFFBCFF8DFDE6FC92FD1AE11B" box="[570,687,783,809]" name="Cameroon" pageId="30" pageNumber="31">Cameroon</collectingCountry>
and
<collectingCountry id="F37C766EFFBCFF8DFD3BFC92FC0DE11B" box="[743,952,783,809]" name="Equatorial Guinea" pageId="30" pageNumber="31">Equatorial Guinea</collectingCountry>
. In
<collectingCountry id="F37C766EFFBCFF8DFC3FFC92FB05E11B" box="[995,1200,783,809]" name="Equatorial Guinea" pageId="30" pageNumber="31">Equatorial Guinea</collectingCountry>
it has been recorded from
<collectingCountry id="F37C766EFFBCFF8DFF04FCAEFEFDE17F" box="[216,328,819,845]" name="Equatorial Guinea" pageId="30" pageNumber="31">Río Muni</collectingCountry>
at Benito River based on a single female specimen held at the British Museum (
<bibRefCitation id="EFFA4B0FFFBCFF8DFB21FCAEFEC5E143" author="Thys van den Audenaerde, D. F. E." pageId="30" pageNumber="31" pagination="163 - 177" refId="ref90661" refString="Thys van den Audenaerde, D. F. E. (1967) Les Gekkonidae de l'Afrique centrale. Revue de Zoologie et de Botanique Africaines, 76 (1 - 2), 163 - 177." type="journal article" year="1967">Thys van den Audenaerde (1967)</bibRefCitation>
[cited as
<taxonomicName id="4C6B4D7DFFBCFF8DFE00FCCAFCA7E143" authority="(Werner, 1896)" baseAuthorityName="Werner" baseAuthorityYear="1896" box="[476,786,855,881]" class="Reptilia" family="Trionychidae" genus="Fordia" kingdom="Animalia" order="Testudines" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="africana">
<emphasis id="B91FEAECFFBCFF8DFE00FCCAFDE9E143" box="[476,604,855,881]" italics="true" pageId="30" pageNumber="31">C. africana</emphasis>
(Werner, 1896)
</taxonomicName>
];
<bibRefCitation id="EFFA4B0FFFBCFF8DFCF8FCCAFC12E143" author="Perret, J. - L." box="[804,935,855,881]" pageId="30" pageNumber="31" pagination="457 - 505" refId="ref86076" refString="Perret, J. - L. (1986) Revision des especes africaines du genre Cnemaspis Strauch, sous-genre Ancylodactylus Muller (Lacertilia, Gekkonidae), avec la description de quatre especes nouvelles. Revue Suisse de Zoologie, 93 (2), 457 - 505. https: // doi. org / 10.5962 / bhl. part. 79705" type="journal article" year="1986">Perret 1986</bibRefCitation>
) (
<figureCitation id="13502A7BFFBCFF8DFC62FCCAFB95E143" box="[958,1056,855,881]" captionStart="MAPS 6" captionStartId="32.[151,223,1943,1967]" captionTargetBox="[203,1383,191,1906]" captionTargetId="figure-19@32.[161,1427,181,1919]" captionTargetPageId="32" captionText="MAPS 6 AC. Distribution maps for Equatorial Guinean records of (A) Ancylodactylus koehleri; (B) Hemidactylus angulatus (question mark represents a dubious record; see species account); (C) Hemidactylus aporus." figureDoi="http://doi.org/10.5281/zenodo.7304641" httpUri="https://zenodo.org/record/7304641/files/figure.png" pageId="30" pageNumber="31">Map 6A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFBCFF8DFF1BFCE7FABCE687" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8BD436FEFFBCFF8DFF1BFCE7FE1FE6A3" blockId="30.[151,1437,782,1205]" pageId="30" pageNumber="31">
<emphasis id="B91FEAECFFBCFF8DFF1BFCE7FEFAE1A6" bold="true" box="[199,335,890,916]" pageId="30" pageNumber="31">Comments.</emphasis>
The genus
<taxonomicName id="4C6B4D7DFFBCFF8DFE0FFCE6FDCBE1A7" authorityName="Muller" authorityYear="1907" box="[467,638,891,917]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="genus">
<emphasis id="B91FEAECFFBCFF8DFE0FFCE6FDCBE1A7" box="[467,638,891,917]" italics="true" pageId="30" pageNumber="31">Ancylodactylus</emphasis>
</taxonomicName>
has been recently proposed to include all the African species formerly treated within
<taxonomicName id="4C6B4D7DFFBCFF8DFEE9FC02FD60E18B" authority="(Malonda &amp; Bauer, 2022)" baseAuthorityName="Malonda &amp; Bauer" baseAuthorityYear="2022" box="[309,725,927,953]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="genus">
<emphasis id="B91FEAECFFBCFF8DFEE9FC02FE05E18B" box="[309,432,927,953]" italics="true" pageId="30" pageNumber="31">Cnemaspis</emphasis>
(Malonda &amp; Bauer, 2022)
</taxonomicName>
.
<bibRefCitation id="EFFA4B0FFFBCFF8DFD01FC02FBFFE18B" author="Thys van den Audenaerde, D. F. E." box="[733,1098,927,953]" pageId="30" pageNumber="31" pagination="163 - 177" refId="ref90661" refString="Thys van den Audenaerde, D. F. E. (1967) Les Gekkonidae de l'Afrique centrale. Revue de Zoologie et de Botanique Africaines, 76 (1 - 2), 163 - 177." type="journal article" year="1967">Thys van den Audenaerde (1967)</bibRefCitation>
recorded the specimen housed in the British Museum as
<taxonomicName id="4C6B4D7DFFBCFF8DFE67FC5EFD8EE1EF" baseAuthorityName="Werner" baseAuthorityYear="1896" box="[443,571,963,989]" class="Reptilia" family="Trionychidae" genus="Fordia" kingdom="Animalia" order="Testudines" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="africana">
<emphasis id="B91FEAECFFBCFF8DFE67FC5EFD8EE1EF" box="[443,571,963,989]" italics="true" pageId="30" pageNumber="31">C. africana</emphasis>
</taxonomicName>
, noticing that it was distinct from
<taxonomicName id="4C6B4D7DFFBCFF8DFC1CFC5EFB89E1EF" baseAuthorityName="Mertens" baseAuthorityYear="1937" box="[960,1084,963,989]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="koehleri">
<emphasis id="B91FEAECFFBCFF8DFC1CFC5EFB89E1EF" box="[960,1084,963,989]" italics="true" pageId="30" pageNumber="31">A. koehleri</emphasis>
</taxonomicName>
. However, after the revision of the genus
<taxonomicName id="4C6B4D7DFFBCFF8DFED5FC7AFE31E633" baseAuthorityName="Malonda &amp; Bauer" baseAuthorityYear="2022" box="[265,388,999,1025]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="genus">
<emphasis id="B91FEAECFFBCFF8DFED5FC7AFE31E633" box="[265,388,999,1025]" italics="true" pageId="30" pageNumber="31">Cnemaspis</emphasis>
</taxonomicName>
of Africa by
<bibRefCitation id="EFFA4B0FFFBCFF8DFDC5FC7AFD1AE633" author="Perret, J. - L." box="[537,687,999,1025]" pageId="30" pageNumber="31" pagination="457 - 505" refId="ref86076" refString="Perret, J. - L. (1986) Revision des especes africaines du genre Cnemaspis Strauch, sous-genre Ancylodactylus Muller (Lacertilia, Gekkonidae), avec la description de quatre especes nouvelles. Revue Suisse de Zoologie, 93 (2), 457 - 505. https: // doi. org / 10.5962 / bhl. part. 79705" type="journal article" year="1986">Perret (1986)</bibRefCitation>
, this specimen was treated as
<taxonomicName id="4C6B4D7DFFBCFF8DFC23FC7AFBCEE633" baseAuthorityName="Mertens" baseAuthorityYear="1937" box="[1023,1147,999,1025]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="koehleri">
<emphasis id="B91FEAECFFBCFF8DFC23FC7AFBCEE633" box="[1023,1147,999,1025]" italics="true" pageId="30" pageNumber="31">C. koehleri</emphasis>
</taxonomicName>
. A single specimen of the genus
<taxonomicName id="4C6B4D7DFFBCFF8DFF03FB96FE3FE617" authorityName="Muller" authorityYear="1907" box="[223,394,1035,1061]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="genus">
<emphasis id="B91FEAECFFBCFF8DFF03FB96FE3FE617" box="[223,394,1035,1061]" italics="true" pageId="30" pageNumber="31">Ancylodactylus</emphasis>
</taxonomicName>
was collected on Bioko at Caldera de Luba, Hormigas Camp, by Rayna Bell in 2013 and held at the Herpetological Collection of the Cornell University Museum of Vertebrates (catalog number: R-0014927). Based on the morphological traits and the phenotype exhibited (
<figureCitation id="13502A7BFFBCFF8DFCBEFBCEFC62E65F" box="[866,983,1107,1133]" captionStart="FIGURE 7" captionStartId="33.[151,250,1918,1942]" captionTargetBox="[153,1434,191,1894]" captionTargetId="figure-18@33.[153,1434,181,1894]" captionTargetPageId="33" captionText="FIGURE 7. From left to right: dorsal, lateral, and ventral views of the head of: Agama lebretoni (AB), male (A: MNCN 4725) and female (B: MNCN 4726); and dorsal and ventral views of Ancylodactylus koehleri (C) (R-0014927). Photos: AS-V (AB) and Cornell University Museum of Vertebrates (C)." figureDoi="http://doi.org/10.5281/zenodo.7285629" httpUri="https://zenodo.org/record/7285629/files/figure.png" pageId="30" pageNumber="31">Figure 7C</figureCitation>
), we identify it as
<taxonomicName id="4C6B4D7DFFBCFF8DFB79FBCEFA95E65F" baseAuthorityName="Mertens" baseAuthorityYear="1937" box="[1189,1312,1107,1133]" class="Reptilia" family="Gekkonidae" genus="Ancylodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="koehleri">
<emphasis id="B91FEAECFFBCFF8DFB79FBCEFA95E65F" box="[1189,1312,1107,1133]" italics="true" pageId="30" pageNumber="31">A. koehleri</emphasis>
</taxonomicName>
, extending its distribution to Bioko.
</paragraph>
<paragraph id="8BD436FEFFBCFF8DFF1BFB07FABCE687" blockId="30.[151,1437,782,1205]" box="[199,1289,1178,1205]" pageId="30" pageNumber="31">
<emphasis id="B91FEAECFFBCFF8DFF1BFB07FE73E686" bold="true" box="[199,454,1178,1204]" pageId="30" pageNumber="31">Specimens examined.</emphasis>
No specimens of this taxon were found by us in the examined collections.
</paragraph>
</subSubSection>
</treatment>
</document>