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<mods:title id="0F8170D4715CC3D22D5AF34CCEDC3A43">Cryptic diversity in the Azorean beetle genus Tarphius Erichson, 1845 (Coleoptera: Zopheridae): An integrative taxonomic approach with description of four new species</mods:title>
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<treatment id="03C287C3C52FDB07FF4CFF7DDFC40AFE" ID-DOI="http://doi.org/10.5281/zenodo.5697942" ID-GBIF-Taxon="127709852" ID-Zenodo-Dep="5697942" LSID="urn:lsid:plazi:treatment:03C287C3C52FDB07FF4CFF7DDFC40AFE" httpUri="http://treatment.plazi.org/id/03C287C3C52FDB07FF4CFF7DDFC40AFE" lastPageId="20" lastPageNumber="421" pageId="17" pageNumber="418">
<subSubSection id="C371655EC52FDB02FF4CFF7DDBC80FED" pageId="17" pageNumber="418" type="nomenclature">
<paragraph id="8BD436D5C52FDB02FF4CFF7DD9C30F8C" blockId="17.[151,755,152,211]" box="[151,755,152,178]" pageId="17" pageNumber="418">
<heading id="D09C81B9C52FDB02FF4CFF7DD9C30F8C" bold="true" box="[151,755,152,178]" fontSize="11" level="1" pageId="17" pageNumber="418" reason="1">
<emphasis id="B91FEAC7C52FDB02FF4CFF7DD9C30F8C" bold="true" box="[151,755,152,178]" pageId="17" pageNumber="418">
<taxonomicName id="4C6B4D56C52FDB02FF4CFF7DD9670F8C" authority="Borges &amp; Serrano" authorityName="Borges &amp; Serrano" box="[151,599,152,178]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="furtadoi" status="sp. nov.">
<emphasis id="B91FEAC7C52FDB02FF4CFF7DDA5B0F8C" bold="true" box="[151,363,152,178]" italics="true" pageId="17" pageNumber="418">Tarphius furtadoi</emphasis>
Borges &amp; Serrano
</taxonomicName>
,
<taxonomicNameLabel id="A22C57BCC52FDB02FDB8FF7DD9C30F8C" box="[611,755,152,178]" pageId="17" pageNumber="418" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="8BD436D5C52FDB02FF4CFF5FDBC80FED" blockId="17.[151,755,152,211]" box="[151,248,186,211]" pageId="17" pageNumber="418">
(
<figureCitation id="13502A50C52FDB02FF44FF5FDBDF0FED" box="[159,239,186,211]" captionStart="FIGURE 11" captionStartId="18.[151,250,1887,1909]" captionTargetBox="[217,1340,290,1756]" captionTargetId="figure@18.[151,1436,193,1865]" captionTargetPageId="18" captionText="FIGURE 11. Habitus of Tarphius furtadoi sp. nov.. Scale 0.5 mm. (Photo: Erno-Endre Gergely)." httpUri="https://zenodo.org/record/322241/files/figure.png" pageId="17" pageNumber="418">Fig. 11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C371655EC52FDB02FF4CFEE7D9CC0D75" pageId="17" pageNumber="418" type="reference_group">
<paragraph id="8BD436D5C52FDB02FF4CFEE7D9E00E26" blockId="17.[151,992,258,587]" box="[151,720,258,280]" pageId="17" pageNumber="418">
<taxonomicName id="4C6B4D56C52FDB02FF4CFEE7D9100E26" authority="in Gillerfors" authorityName="in Gillerfors" authorityYear="1986" box="[151,544,258,280]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="tornvalli">
<emphasis id="B91FEAC7C52FDB02FF4CFEE7DA7D0E26" box="[151,333,258,280]" italics="true" pageId="17" pageNumber="418">Tarphius tornvalli</emphasis>
in
<bibRefCitation id="EFFA4B24C52FDB02FEB5FEE7D9100E26" author="Gillerfors" box="[366,544,258,280]" pageId="17" pageNumber="418" refString="Gillerfors, G. (1986 b) Contribution to the coleopterous fauna of the Azores. Boletim do Museu Municipal do Funchal, 38, 16 - 27." type="journal article" year="1986" yearSuffix="b">Gillerfors (1986b</bibRefCitation>
</taxonomicName>
, p. 21) (Pico Isl.)
</paragraph>
<paragraph id="8BD436D5C52FDB02FF4CFEC4D9840E09" blockId="17.[151,992,258,587]" box="[151,692,289,311]" pageId="17" pageNumber="418">
<taxonomicName id="4C6B4D56C52FDB02FF4CFEC4D97A0E09" authority="in Borges" authorityName="in Borges" authorityYear="1990" box="[151,586,289,311]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="tornvalli">
<emphasis id="B91FEAC7C52FDB02FF4CFEC4DA7D0E09" box="[151,333,289,311]" italics="true" pageId="17" pageNumber="418">Tarphius tornvalli</emphasis>
in
<bibRefCitation id="EFFA4B24C52FDB02FEB5FEC7D97A0E09" author="Borges" box="[366,586,289,311]" pageId="17" pageNumber="418" refString="Borges, P. A. V. (1990) A checklist of Coleoptera from the Azores with some systematic and biogeographic comments. Boletim do Museu Municipal do Funchal, 42, 87 - 136." type="journal article" year="1990" yearSuffix="p">Borges (1990, p. 112)</bibRefCitation>
</taxonomicName>
(Pico Isl.)
</paragraph>
<paragraph id="8BD436D5C52FDB02FF4CFEA5D9AC0E68" blockId="17.[151,992,258,587]" box="[151,668,320,342]" pageId="17" pageNumber="418">
<taxonomicName id="4C6B4D56C52FDB02FF4CFEA5D9020E68" authority="in Borges" authorityName="in Borges" authorityYear="1991" box="[151,562,320,342]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="tornvalli">
<emphasis id="B91FEAC7C52FDB02FF4CFEA5DA7D0E68" box="[151,333,320,342]" italics="true" pageId="17" pageNumber="418">Tarphius tornvalli</emphasis>
in
<bibRefCitation id="EFFA4B24C52FDB02FEB5FEA4D9020E68" author="Borges" box="[366,562,320,342]" pageId="17" pageNumber="418" refString="Borges, P. A. V. (1991) Two new species of Tarphius Erichson, 1848 (Coleoptera, Colydiidae) from the Azores. Bocagiana, 143, 1 - 11." type="journal article" year="1991" yearSuffix="p">Borges (1991, p. 2)</bibRefCitation>
</taxonomicName>
(Pico Isl.)
</paragraph>
<paragraph id="8BD436D5C52FDB02FF4CFEBBD82D0ED1" blockId="17.[151,992,258,587]" pageId="17" pageNumber="418">
<taxonomicName id="4C6B4D56C52FDB02FF4CFEBBDA7D0E4A" box="[151,333,350,372]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="tornvalli">
<emphasis id="B91FEAC7C52FDB02FF4CFEBBDA7D0E4A" box="[151,333,350,372]" italics="true" pageId="17" pageNumber="418">Tarphius tornvalli</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B4D56C52FDB02FEA5FEBBD8140E4A" authority="in Borges et al." authorityName="in Borges et al." authorityYear="2005" box="[382,804,350,373]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="azoricus">
<emphasis id="B91FEAC7C52FDB02FEA5FEBBD9040E4B" box="[382,564,350,373]" italics="true" pageId="17" pageNumber="418">Tarphius azoricus</emphasis>
in
<bibRefCitation id="EFFA4B24C52FDB02FD8FFEBAD8140E4A" author="Borges" box="[596,804,350,373]" pageId="17" pageNumber="418" refString="Borges, P. A. V., Vieira, V., Dinis, F., Jarroca, S., Aguiar, C., Amaral, J., Aarvik, L., Ashmole, P., Ashmole, M., Amorim, I. R., Andre, G., Argente, M. C., Arraiol, A., Cabrera, A., Diaz, S., Enghoff, H., Gaspar, C., Mendonca, E. P., Gisbert, H. M., Goncalves, P., Lopes, D. H., Melo, C., Mota, J. A., Oliveira, O., Oromi, P., Pereira, F., Pombo, D. T., Quartau, J. A., Ribeiro, S. P., Rodrigues, A. C., Santos, A. M. C., Serrano, A. R. M., Simoes, A. M., Soares, A. O., Sousa, A. B., Vieira, L., Vitorino, A. &amp; Wunderlich, J. (2005 b) List of arthropods (Arthropoda) In: Borges, P. A. V., Cunha, R., Gabriel, R., Martins, A. M. F., Silva, L. &amp; Vieira, V. (Eds.), A list of the terrestrial fauna (Mollusca and Arthropoda) and flora (Bryophyta, Pteridophyta and Spermatophyta) from the Azores. Direccao Regional de Ambiente and Universidade dos Acores, Horta, Angra do Heroismo and Ponta Delgada, pp. 163 - 221." type="book chapter" year="2005" yearSuffix="b">
Borges
<emphasis id="B91FEAC7C52FDB02FD79FE85D9E50E4A" box="[674,725,350,373]" italics="true" pageId="17" pageNumber="418">et al.</emphasis>
(2005b
</bibRefCitation>
</taxonomicName>
, p. 207) (Pico Isl.)
<taxonomicName id="4C6B4D56C52FDB02FF4CFE98DA7D0EAD" box="[151,333,381,403]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="tornvalli">
<emphasis id="B91FEAC7C52FDB02FF4CFE98DA7D0EAD" box="[151,333,381,403]" italics="true" pageId="17" pageNumber="418">Tarphius tornvalli</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B4D56C52FDB02FEA5FE98D8230EAD" authority="in Oromi et al." authorityName="in Oromi et al." authorityYear="2010" box="[382,787,381,403]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="azoricus">
<emphasis id="B91FEAC7C52FDB02FEA5FE98D9040EAD" box="[382,564,381,403]" italics="true" pageId="17" pageNumber="418">Tarphius azoricus</emphasis>
in
<bibRefCitation id="EFFA4B24C52FDB02FD8FFE98D8230EAD" author="Oromi" box="[596,787,381,403]" pageId="17" pageNumber="418" refString="Oromi, P., Serrano, A. R. M. &amp; Borges, P. A. V. (2010) Coleoptera (Coordination) In: Borges, P. A. V., Costa, A., Cunha, R., Gabriel, R., Goncalves, V., Martins, A. F., Melo, I., Parente, M., Raposeiro, P., Rodrigues, P., Santos, R. S., Silva, L., Vieira, P. &amp; Vieira, V. (Eds.), A list of the terrestrial and marine biota from the Azores. Principia, Cascais, pp. 222 - 232. https: // doi. org / 10.1093 / molbev / msn 083" type="book chapter" year="2010">
Oromí
<emphasis id="B91FEAC7C52FDB02FD47FE9BD9F90EAD" box="[668,713,381,403]" italics="true" pageId="17" pageNumber="418">et al</emphasis>
. (2010
</bibRefCitation>
</taxonomicName>
, p. 232) (Pico Isl.)
<taxonomicName id="4C6B4D56C52FDB02FF4CFE79DA7D0E8C" box="[151,333,412,434]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="tornvalli">
<emphasis id="B91FEAC7C52FDB02FF4CFE79DA7D0E8C" box="[151,333,412,434]" italics="true" pageId="17" pageNumber="418">Tarphius tornvalli</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B4D56C52FDB02FEA5FE79D9D40E8C" authority="in Amorim et al." authorityName="in Amorim et al." box="[382,740,412,434]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="azoricus">
<emphasis id="B91FEAC7C52FDB02FEA5FE79D9040E8C" box="[382,564,412,434]" italics="true" pageId="17" pageNumber="418">Tarphius azoricus</emphasis>
in Amorim
<emphasis id="B91FEAC7C52FDB02FD6BFE78D9D40E8C" box="[688,740,412,434]" italics="true" pageId="17" pageNumber="418">et al.</emphasis>
</taxonomicName>
(2012,
<figureCitation id="13502A50C52FDB02FCEAFE79D85C0E8C" box="[817,876,412,434]" captionStart="FIGURE 2" captionStartId="5.[151,250,1679,1701]" captionTargetBox="[163,1436,229,1622]" captionTargetId="figure@5.[151,1436,217,1629]" captionTargetPageId="5" captionText="FIGURE 2. Bayesian inference tree for Tarphius beetles of the Azores based on 915 bp of nuclear DNA (Elongation Factor 1 alpha). Clade credibility values are shown for nodes with a posteror probability ± 0.95." httpUri="https://zenodo.org/record/322233/files/figure.png" pageId="17" pageNumber="418">Fig. 2</figureCitation>
) (Pico Isl.)
<taxonomicName id="4C6B4D56C52FDB02FF4CFE5FD90D0EEE" authority="in Borges et al." authorityName="in Borges et al." authorityYear="2005" box="[151,573,442,464]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="azoricus">
<emphasis id="B91FEAC7C52FDB02FF4CFE5FDA7C0EEE" box="[151,332,442,464]" italics="true" pageId="17" pageNumber="418">Tarphius azoricus</emphasis>
in
<bibRefCitation id="EFFA4B24C52FDB02FEB7FE5ED90D0EEE" author="Borges" box="[364,573,442,464]" pageId="17" pageNumber="418" refString="Borges, P. A. V., Vieira, V., Dinis, F., Jarroca, S., Aguiar, C., Amaral, J., Aarvik, L., Ashmole, P., Ashmole, M., Amorim, I. R., Andre, G., Argente, M. C., Arraiol, A., Cabrera, A., Diaz, S., Enghoff, H., Gaspar, C., Mendonca, E. P., Gisbert, H. M., Goncalves, P., Lopes, D. H., Melo, C., Mota, J. A., Oliveira, O., Oromi, P., Pereira, F., Pombo, D. T., Quartau, J. A., Ribeiro, S. P., Rodrigues, A. C., Santos, A. M. C., Serrano, A. R. M., Simoes, A. M., Soares, A. O., Sousa, A. B., Vieira, L., Vitorino, A. &amp; Wunderlich, J. (2005 b) List of arthropods (Arthropoda) In: Borges, P. A. V., Cunha, R., Gabriel, R., Martins, A. M. F., Silva, L. &amp; Vieira, V. (Eds.), A list of the terrestrial fauna (Mollusca and Arthropoda) and flora (Bryophyta, Pteridophyta and Spermatophyta) from the Azores. Direccao Regional de Ambiente and Universidade dos Acores, Horta, Angra do Heroismo and Ponta Delgada, pp. 163 - 221." type="book chapter" year="2005" yearSuffix="b">
Borges
<emphasis id="B91FEAC7C52FDB02FE61FE5EDADD0EEE" box="[442,493,442,464]" italics="true" pageId="17" pageNumber="418">et al.</emphasis>
(2005b
</bibRefCitation>
</taxonomicName>
, p. 207) (São Jorge Isl.)
<taxonomicName id="4C6B4D56C52FDB02FF4CFE3CD91B0ED1" authority="in Oromi et al." authorityName="in Oromi et al." authorityYear="2010" box="[151,555,473,495]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="azoricus">
<emphasis id="B91FEAC7C52FDB02FF4CFE3CDA7C0ED1" box="[151,332,473,495]" italics="true" pageId="17" pageNumber="418">Tarphius azoricus</emphasis>
in
<bibRefCitation id="EFFA4B24C52FDB02FEB7FE3CD91B0ED1" author="Oromi" box="[364,555,473,495]" pageId="17" pageNumber="418" refString="Oromi, P., Serrano, A. R. M. &amp; Borges, P. A. V. (2010) Coleoptera (Coordination) In: Borges, P. A. V., Costa, A., Cunha, R., Gabriel, R., Goncalves, V., Martins, A. F., Melo, I., Parente, M., Raposeiro, P., Rodrigues, P., Santos, R. S., Silva, L., Vieira, P. &amp; Vieira, V. (Eds.), A list of the terrestrial and marine biota from the Azores. Principia, Cascais, pp. 222 - 232. https: // doi. org / 10.1093 / molbev / msn 083" type="book chapter" year="2010">
Oromí
<emphasis id="B91FEAC7C52FDB02FE6EFE3FDAD10ED1" box="[437,481,473,495]" italics="true" pageId="17" pageNumber="418">et al</emphasis>
. (2010
</bibRefCitation>
</taxonomicName>
, p. 232) (São Jorge Isl.)
</paragraph>
<paragraph id="8BD436D5C52FDB02FF4CFE1DD9DD0D12" blockId="17.[151,992,258,587]" pageId="17" pageNumber="418">
<taxonomicName id="4C6B4D56C52FDB02FF4CFE1DDACC0D30" authority="in Amorim et al." authorityName="in Amorim et al." box="[151,508,504,526]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="azoricus">
<emphasis id="B91FEAC7C52FDB02FF4CFE1DDA7C0D30" box="[151,332,504,526]" italics="true" pageId="17" pageNumber="418">Tarphius azoricus</emphasis>
in Amorim
<emphasis id="B91FEAC7C52FDB02FE13FE1CDACC0D30" box="[456,508,504,526]" italics="true" pageId="17" pageNumber="418">et al.</emphasis>
</taxonomicName>
(2012,
<figureCitation id="13502A50C52FDB02FD92FE1DD9B40D30" box="[585,644,504,526]" captionStart="FIGURE 2" captionStartId="5.[151,250,1679,1701]" captionTargetBox="[163,1436,229,1622]" captionTargetId="figure@5.[151,1436,217,1629]" captionTargetPageId="5" captionText="FIGURE 2. Bayesian inference tree for Tarphius beetles of the Azores based on 915 bp of nuclear DNA (Elongation Factor 1 alpha). Clade credibility values are shown for nodes with a posteror probability ± 0.95." httpUri="https://zenodo.org/record/322233/files/figure.png" pageId="17" pageNumber="418">Fig. 2</figureCitation>
) (São Jorge Isl.)
<taxonomicName id="4C6B4D56C52FDB02FF4CFDF3D91B0D12" authority="in Oromi et al." authorityName="in Oromi et al." authorityYear="2010" box="[151,555,534,557]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="azoricus">
<emphasis id="B91FEAC7C52FDB02FF4CFDF3DA7C0D13" box="[151,332,534,557]" italics="true" pageId="17" pageNumber="418">Tarphius azoricus</emphasis>
in
<bibRefCitation id="EFFA4B24C52FDB02FEB7FDF3D91B0D12" author="Oromi" box="[364,555,534,557]" pageId="17" pageNumber="418" refString="Oromi, P., Serrano, A. R. M. &amp; Borges, P. A. V. (2010) Coleoptera (Coordination) In: Borges, P. A. V., Costa, A., Cunha, R., Gabriel, R., Goncalves, V., Martins, A. F., Melo, I., Parente, M., Raposeiro, P., Rodrigues, P., Santos, R. S., Silva, L., Vieira, P. &amp; Vieira, V. (Eds.), A list of the terrestrial and marine biota from the Azores. Principia, Cascais, pp. 222 - 232. https: // doi. org / 10.1093 / molbev / msn 083" type="book chapter" year="2010">
Oromí
<emphasis id="B91FEAC7C52FDB02FE6EFDFDDAD10D12" box="[437,481,534,557]" italics="true" pageId="17" pageNumber="418">et al</emphasis>
. (2010
</bibRefCitation>
</taxonomicName>
, p. 232) (Faial Isl.)
</paragraph>
<paragraph id="8BD436D5C52FDB02FF4CFDD0D9CC0D75" blockId="17.[151,992,258,587]" box="[151,764,565,587]" pageId="17" pageNumber="418">
<taxonomicName id="4C6B4D56C52FDB02FF4CFDD0DACC0D75" authority="in Amorim et al." authorityName="in Amorim et al." box="[151,508,565,587]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="azoricus">
<emphasis id="B91FEAC7C52FDB02FF4CFDD0DA7C0D75" box="[151,332,565,587]" italics="true" pageId="17" pageNumber="418">Tarphius azoricus</emphasis>
in Amorim
<emphasis id="B91FEAC7C52FDB02FE13FDD3DACC0D75" box="[456,508,565,587]" italics="true" pageId="17" pageNumber="418">et al.</emphasis>
</taxonomicName>
(2012,
<figureCitation id="13502A50C52FDB02FD92FDD0D9B40D75" box="[585,644,565,587]" captionStart="FIGURE 2" captionStartId="5.[151,250,1679,1701]" captionTargetBox="[163,1436,229,1622]" captionTargetId="figure@5.[151,1436,217,1629]" captionTargetPageId="5" captionText="FIGURE 2. Bayesian inference tree for Tarphius beetles of the Azores based on 915 bp of nuclear DNA (Elongation Factor 1 alpha). Clade credibility values are shown for nodes with a posteror probability ± 0.95." httpUri="https://zenodo.org/record/322233/files/figure.png" pageId="17" pageNumber="418">Fig. 2</figureCitation>
) (Faial Isl.)
</paragraph>
</subSubSection>
<subSubSection id="C371655EC52FDB02FF4CFD97DF4F0BD1" pageId="17" pageNumber="418" type="materials_examined">
<paragraph id="8BD436D5C52FDB02FF4CFD97D82E0DB5" blockId="17.[151,1436,626,1983]" box="[151,798,626,651]" pageId="17" pageNumber="418">
<materialsCitation id="3B033C88C52FDB02FF4CFD97D82E0DB5" ID-GBIF-Occurrence="1437079342" box="[151,798,626,651]" country="Iceland" location="Azores" pageId="17" pageNumber="418" specimenCount="1" stateProvince="Azores" typeStatus="holotype">
<emphasis id="B91FEAC7C52FDB02FF4CFD97DA050DB5" bold="true" box="[151,309,626,651]" pageId="17" pageNumber="418">
<typeStatus id="54D08877C52FDB02FF4CFD97DBE20DB5" box="[151,210,626,651]" pageId="17" pageNumber="418">Type</typeStatus>
locality.
</emphasis>
The
<collectingRegion id="49AFF837C52FDB02FEB5FD96DAF00DB5" box="[366,448,627,651]" country="Portugal" name="Acores" pageId="17" pageNumber="418">Azores</collectingRegion>
, Pico
<collectingCountry id="F37C7645C52FDB02FDDEFD97D97C0DB5" box="[517,588,626,651]" name="Iceland" pageId="17" pageNumber="418">Island</collectingCountry>
, Lagoa do Caiado.
</materialsCitation>
</paragraph>
<paragraph id="8BD436D5C52FDB02FF1CFD73DF150DED" blockId="17.[151,1436,626,1983]" pageId="17" pageNumber="418">
<materialsCitation id="3B033C88C52FDB02FF1CFD73DF150DED" ID-GBIF-Occurrence="1437079355" collectingDate="1990-03-04" collectingDateMax="1990-03-19" collectingDateMin="1990-03-04" collectionCode="DTPC" collectorName="Paulo A. V." location="Pico" municipality="Lagoa do Caiado" pageId="17" pageNumber="418" specimenCount="2" stateProvince="Acores" typeStatus="holotype">
<emphasis id="B91FEAC7C52FDB02FF1CFD73DA480D91" bold="true" box="[199,376,662,687]" pageId="17" pageNumber="418">
<typeStatus id="54D08877C52FDB02FF1CFD73DA320D91" box="[199,258,662,687]" pageId="17" pageNumber="418">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54D08877C52FDB02FE59FD72D9290D91" box="[386,537,663,687]" pageId="17" pageNumber="418" type="holotype">HOLOTYPE</typeStatus>
, male, deposited at
<collectionCode id="ED7AAE10C52FDB02FCD9FD72D87D0D91" box="[770,845,663,687]" pageId="17" pageNumber="418">DTPC</collectionCode>
, labeled:
<collectingMunicipality id="6BB0ACAFC52FDB02FC65FD72DFB60D91" box="[958,1158,662,687]" pageId="17" pageNumber="418">Lagoa do Caiado</collectingMunicipality>
,
<location id="8EB4600EC52FDB02FB4EFD73DFFA0D91" LSID="urn:lsid:plazi:treatment:03C287C3C52FDB07FF4CFF7DDFC40AFE:8EB4600EC52FDB02FB4EFD73DFFA0D91" box="[1173,1226,662,687]" municipality="Lagoa do Caiado" name="Pico" pageId="17" pageNumber="418" stateProvince="Acores">Pico</location>
,
<collectingRegion id="49AFF837C52FDB02FB03FD72DE1A0D91" box="[1240,1322,663,687]" country="Portugal" name="Acores" pageId="17" pageNumber="418">Açores</collectingRegion>
,
<date id="FFD51015C52FDB02FAE2FD72DBFF0DED" pageId="17" pageNumber="418" value="1990-03-04" valueMax="1990-03-19" valueMin="1990-03-04">
<collectingDate id="EF91E9FDC52FDB02FAE2FD72DBFF0DED" pageId="17" pageNumber="418" value="1990-03-04" valueMax="1990-03-19" valueMin="1990-03-04">419/03/ 1990</collectingDate>
</date>
(UTM 26S 390266, 4257172);
<collectorName id="269E5303C52FDB02FDE8FD5FD9840DEC" box="[563,692,698,723]" pageId="17" pageNumber="418">Paulo A. V.</collectorName>
Borges &amp; Fernando Pereira leg..
</materialsCitation>
</paragraph>
<paragraph id="8BD436D5C52FDB02FF1CFD3ADF4F0BD1" blockId="17.[151,1436,626,1983]" pageId="17" pageNumber="418">
<typeStatus id="54D08877C52FDB02FF1CFD3ADA550DC9" box="[199,357,735,759]" pageId="17" pageNumber="418" type="paratype">
<collectionCode id="ED7AAE10C52FDB02FF1CFD3ADA550DC9" box="[199,357,735,759]" pageId="17" pageNumber="418">PARATYPES</collectionCode>
</typeStatus>
,
<collectionCode id="ED7AAE10C52FDB02FEADFD3ADA860DC9" box="[374,438,735,759]" pageId="17" pageNumber="418">PICO</collectionCode>
<collectingCountry id="F37C7645C52FDB02FE1FFD3AD91F0DC9" box="[452,559,735,759]" name="Iceland" pageId="17" pageNumber="418">
<collectionCode id="ED7AAE10C52FDB02FE1FFD3AD91F0DC9" box="[452,559,735,759]" pageId="17" pageNumber="418">ISLAND</collectionCode>
</collectingCountry>
: 280 specimens—Chão Verde Superior (T01) (Natural Forest Reserve of Mistério da Prainha),
<date id="FFD51015C52FDB02FE45FCE6DACA0C25" box="[414,506,771,795]" pageId="17" pageNumber="418" value="1999-09">IX.1999</date>
(4 exx),
<date id="FFD51015C52FDB02FDA8FCE6D9E90C25" box="[627,729,771,795]" pageId="17" pageNumber="418" value="2010-07">VII.2010</date>
(1 ex.) (UTM 26S 388838, 4259774); Lagoa do Caiado - Euphorbias (T02) (Natural Forest Reserve of Lagoa do Caiado),
<date id="FFD51015C52FDB02FCADFCC2D8E20C01" box="[886,978,807,831]" pageId="17" pageNumber="418" value="1999-09">IX.1999</date>
(36 exx) (UTM 26S 390266, 4257172); Chão Verde Inferior (T03) (Natural Forest Reserve of Mistério da Prainha)
<date id="FFD51015C52FDB02FC2AFCAEDF7D0C5D" box="[1009,1101,843,867]" pageId="17" pageNumber="418" value="1999-09">IX.1999</date>
(39 exx) (UTM 26S 388946, 4260693); Picos do Caveiro (T08) (Natural Forest Reserve of Caveiro),
<date id="FFD51015C52FDB02FC1BFC8ADF2C0CB9" box="[960,1052,879,903]" pageId="17" pageNumber="418" value="1999-09">IX.1999</date>
(6 exx),
<date id="FFD51015C52FDB02FB59FC8ADFD90CB9" box="[1154,1257,879,903]" pageId="17" pageNumber="418" value="2010-07">VII.2010</date>
(23 exx) (UTM 26S 395274, 4255409); Caveiro Base (T09) (Natural Forest Reserve of Caveiro),
<date id="FFD51015C52FDB02FBEDFC76DFA20C95" box="[1078,1170,915,939]" pageId="17" pageNumber="418" value="1999-09">IX.1999</date>
(19 exx),
<date id="FFD51015C52FDB02FAD2FC76DE5E0C95" box="[1289,1390,915,939]" pageId="17" pageNumber="418" value="2010-07">VII.2010</date>
(23 exx) (UTM 26S 394341, 4255080); Chão Verde Inferior Cima (T14) (Natural Forest Reserve of Mistério da Prainha),
<date id="FFD51015C52FDB02FED3FC3EDA5F0CCD" box="[264,367,987,1011]" pageId="17" pageNumber="418" value="2000-07">VII.2000</date>
(8 exx) (UTM 26S 389041, 4260364); Chão Verde Superior Perpendicular (T15) (Natural Forest Reserve of Mistério da Prainha),
<date id="FFD51015C52FDB02FD83FC1AD98D0B29" box="[600,701,1023,1047]" pageId="17" pageNumber="418" value="2000-07">VII.2000</date>
(2 exx) (UTM 26S 389035, 4259508); Lagoa do Caiado—Frente a Lagoa (T16) (Natural Forest Reserve of Lagoa do Caiado),
<date id="FFD51015C52FDB02FC86FBC6D8FC0B05" box="[861,972,1059,1083]" pageId="17" pageNumber="418" value="2000-08">VIII.2000</date>
(21 exx) (UTM 26S 390537, 4257057); Lagoa do Caiado—Frente a Euphorbias (T17) (Natural Forest Reserve of Lagoa do Caiado),
<date id="FFD51015C52FDB02FB1FFBA2DE020B61" box="[1220,1330,1095,1119]" pageId="17" pageNumber="418" value="2000-08">VIII.2000</date>
(28 exx) (UTM 26S 390661, 4257058); Caveiro, Prado (T20) (Natural Forest Reserve of Caveiro),
<date id="FFD51015C52FDB02FB48FB8EDE320BBD" box="[1171,1282,1131,1155]" pageId="17" pageNumber="418" value="2000-08">VIII.2000</date>
(1 ex) (UTM 26S 394533, 4255179); Caveiro, Morro (T21) (Natural Forest Reserve of Caveiro),
<date id="FFD51015C52FDB02FB9DFB6ADF840B99" box="[1094,1204,1167,1191]" pageId="17" pageNumber="418" value="2000-08">VIII.2000</date>
(47 exx) (UTM 26S 394055, 4255376); Caveiro, Meio da Reserva (T22) (Natural Forest Reserve of Caveiro),
<date id="FFD51015C52FDB02FB65FB56DE1D0BF5" box="[1214,1325,1203,1227]" pageId="17" pageNumber="418" value="2000-08">VIII.2000</date>
(22 exx) (UTM 26S 396643, 4255074) (collected with pitfall traps; Paulo A. V. Borges
<emphasis id="B91FEAC7C52FDB02FBDCFB32DF0F0BD1" box="[1031,1087,1238,1263]" italics="true" pageId="17" pageNumber="418">et al.</emphasis>
leg.).
</paragraph>
</subSubSection>
<subSubSection id="C371655EC52FDB02FF1CFB1ED9B509A1" pageId="17" pageNumber="418" type="description">
<paragraph id="8BD436D5C52FDB02FF1CFB1EDFC90A9D" blockId="17.[151,1436,626,1983]" pageId="17" pageNumber="418">
<materialsCitation id="3B033C88C52FDB02FF1CFB1EDAA20A09" ID-GBIF-Occurrence="1437079367" collectingDate="2009-09" collectionCode="FAIAL" collectorName="Fernando Pereira" country="Iceland" location="Trinta" pageId="17" pageNumber="418" specimenCount="1">
<collectionCode id="ED7AAE10C52FDB02FF1CFB1EDA270A2D" box="[199,279,1275,1299]" pageId="17" pageNumber="418">FAIAL</collectionCode>
<collectingCountry id="F37C7645C52FDB02FEFDFB1EDAA10A2D" box="[294,401,1275,1299]" name="Iceland" pageId="17" pageNumber="418">ISLAND</collectingCountry>
: 142 specimens—Cabeço dos
<location id="8EB4600EC52FDB02FCDEFB1FD87B0A2D" LSID="urn:lsid:plazi:treatment:03C287C3C52FDB07FF4CFF7DDFC40AFE:8EB4600EC52FDB02FCDEFB1FD87B0A2D" box="[773,843,1274,1299]" country="Iceland" name="Trinta" pageId="17" pageNumber="418">Trinta</location>
,
<date id="FFD51015C52FDB02FC85FB1ED88A0A2D" box="[862,954,1275,1299]" pageId="17" pageNumber="418" value="2009-09">
<collectingDate id="EF91E9FDC52FDB02FC85FB1ED88A0A2D" box="[862,954,1275,1299]" pageId="17" pageNumber="418" value="2009-09">IX.2009</collectingDate>
</date>
(3 exx) (UTM 26S 348327, 4271670),
<collectorName id="269E5303C52FDB02FF4CFAFBDA6A0A09" box="[151,346,1310,1335]" pageId="17" pageNumber="418">Fernando Pereira</collectorName>
, leg.
</materialsCitation>
;
<materialsCitation id="3B033C88C52FDB02FE45FAFBDFC90A9D" ID-GBIF-Occurrence="1437079379" collectingDate="2010-09" collectorName="Paulo A. V." country="Iceland" county="Natural Forest" location="Cabeco do Fogo" municipality="Natural Forest" pageId="17" pageNumber="418" specimenCount="1">
<location id="8EB4600EC52FDB02FE45FAFBD9670A09" LSID="urn:lsid:plazi:treatment:03C287C3C52FDB07FF4CFF7DDFC40AFE:8EB4600EC52FDB02FE45FAFBD9670A09" box="[414,599,1310,1335]" country="Iceland" county="Natural Forest" municipality="Natural Forest" name="Cabeco do Fogo" pageId="17" pageNumber="418">Cabeço do Fogo</location>
(T03) (
<location id="8EB4600EC52FDB02FD77FAFBD8800A08" LSID="urn:lsid:plazi:treatment:03C287C3C52FDB07FF4CFF7DDFC40AFE:8EB4600EC52FDB02FD77FAFBD8800A08" box="[684,944,1310,1335]" country="Iceland" county="Natural Forest" municipality="Natural Forest" name="Natural Forest Reserve" pageId="17" pageNumber="418">
<location id="8EB4600EC52FDB02FD77FAFBD87F0A08" LSID="urn:lsid:plazi:treatment:03C287C3C52FDB07FF4CFF7DDFC40AFE:8EB4600EC52FDB02FD77FAFBD87F0A08" box="[684,847,1310,1335]" country="Iceland" county="Natural Forest" municipality="Natural Forest" name="Natural Forest" pageId="17" pageNumber="418">Natural Forest</location>
Reserve
</location>
of
<location id="8EB4600EC52FDB02FC0CFAFBDFA40A09" LSID="urn:lsid:plazi:treatment:03C287C3C52FDB07FF4CFF7DDFC40AFE:8EB4600EC52FDB02FC0CFAFBDFA40A09" box="[983,1172,1310,1335]" country="Iceland" county="Natural Forest" municipality="Natural Forest" name="Cabeco do Fogo" pageId="17" pageNumber="418">Cabeço do Fogo</location>
),
<date id="FFD51015C52FDB02FB7CFAFADE340A09" box="[1191,1284,1311,1335]" pageId="17" pageNumber="418" value="2010-09">
<collectingDate id="EF91E9FDC52FDB02FB7CFAFADE340A09" box="[1191,1284,1311,1335]" pageId="17" pageNumber="418" value="2010-09">IX.2010</collectingDate>
</date>
(1 ex) (UTM 26S 346239, 4272261); Caldeira do Faial (T10) (
<collectingMunicipality id="6BB0ACAFC52FDB02FD12FAA7D85D0A64" box="[713,877,1346,1371]" pageId="17" pageNumber="418">Natural Forest</collectingMunicipality>
Reserve of Caldeira do Faial),
<date id="FFD51015C52FDB02FB0CFAA6DE040A65" box="[1239,1332,1347,1371]" pageId="17" pageNumber="418" value="2010-09">
<collectingDate id="EF91E9FDC52FDB02FB0CFAA6DE040A65" box="[1239,1332,1347,1371]" pageId="17" pageNumber="418" value="2010-09">IX.2010</collectingDate>
</date>
(10 exx) (UTM 26S 351110, 4271917); Caldeira do Faial (TB26) (
<collectingCounty id="62B54E59C52FDB02FCF6FA83D8E20A40" box="[813,978,1382,1407]" pageId="17" pageNumber="418">Natural Forest</collectingCounty>
Reserve of Caldeira do Faial),
<date id="FFD51015C52FDB02FA9BFA82DEAC0A41" box="[1344,1436,1383,1407]" pageId="17" pageNumber="418" value="2010-09">
<collectingDate id="EF91E9FDC52FDB02FA9BFA82DEAC0A41" box="[1344,1436,1383,1407]" pageId="17" pageNumber="418" value="2010-09">IX.2010</collectingDate>
</date>
(128 exx) (UTM 26S 350857, 4272272), (
<collectingMethod id="522A4EC2C52FDB02FDB5FA6FD8A80A9C" box="[622,920,1418,1443]" pageId="17" pageNumber="418">collected with pitfall traps</collectingMethod>
;
<collectorName id="269E5303C52FDB02FC7FFA6FDF150A9C" box="[932,1061,1418,1443]" pageId="17" pageNumber="418">Paulo A. V.</collectorName>
Borges
<emphasis id="B91FEAC7C52FDB02FB5AFA69DF890A9D" box="[1153,1209,1418,1443]" italics="true" pageId="17" pageNumber="418">et al.</emphasis>
leg.).
</materialsCitation>
</paragraph>
<paragraph id="8BD436D5C52FDB02FF1CFA4BDA960945" blockId="17.[151,1436,626,1983]" pageId="17" pageNumber="418">
<materialsCitation id="3B033C88C52FDB02FF1CFA4BD8A30AD5" ID-GBIF-Occurrence="1437079389" collectingDate="2003-08" collectionCode="SAO, JORGE" collectorName="Isabel R. &amp; A. V. Borges" country="Iceland" location="Seca" pageId="17" pageNumber="418" specimenCount="1">
<collectionCode id="ED7AAE10C52FDB02FF1CFA4BDBCF0AF9" box="[199,255,1454,1479]" country="Switzerland" httpUri="http://grbio.org/cool/2pta-s8y1" name="Sammlung Oberli" pageId="17" pageNumber="418">SÃO</collectionCode>
<collectionCode id="ED7AAE10C52FDB02FED7FA4ADA530AF9" box="[268,355,1455,1479]" pageId="17" pageNumber="418">JORGE</collectionCode>
<collectingCountry id="F37C7645C52FDB02FEABFA4ADAEB0AF9" box="[368,475,1455,1479]" name="Iceland" pageId="17" pageNumber="418">ISLAND</collectingCountry>
: 52 specimens—Ribeira
<location id="8EB4600EC52FDB02FCDEFA4AD80D0AF9" LSID="urn:lsid:plazi:treatment:03C287C3C52FDB07FF4CFF7DDFC40AFE:8EB4600EC52FDB02FCDEFA4AD80D0AF9" box="[773,829,1455,1479]" country="Iceland" name="Seca" pageId="17" pageNumber="418">Seca</location>
,
<date id="FFD51015C52FDB02FC95FA4AD88D0AF9" box="[846,957,1455,1479]" pageId="17" pageNumber="418" value="2003-08">
<collectingDate id="EF91E9FDC52FDB02FC95FA4AD88D0AF9" box="[846,957,1455,1479]" pageId="17" pageNumber="418" value="2003-08">VIII.2003</collectingDate>
</date>
(32 exx) (UTM 26S 415675, 4273358) (collected by hand;
<collectorName id="269E5303C52FDB02FEA6FA37DAD30AD4" box="[381,483,1490,1515]" pageId="17" pageNumber="418">Isabel R.</collectorName>
Amorim &amp; Paulo
<collectorName id="269E5303C52FDB02FD64FA36D8640AD5" box="[703,852,1491,1515]" pageId="17" pageNumber="418">A. V. Borges</collectorName>
leg.)
</materialsCitation>
;
<materialsCitation id="3B033C88C52FDB02FC79FA37DA960945" ID-GBIF-Occurrence="1437079405" collectingDate="2004-08" collectingDateMax="2010-09" collectingDateMin="2004-08" collectorName="Paulo A. V." country="Iceland" county="Natural Forest" location="Pico Pinheiro Inferior" municipality="Natural Forest" pageId="17" pageNumber="418" specimenCount="1">
<location id="8EB4600EC52FDB02FC79FA37DFAB0AD5" LSID="urn:lsid:plazi:treatment:03C287C3C52FDB07FF4CFF7DDFC40AFE:8EB4600EC52FDB02FC79FA37DFAB0AD5" box="[930,1179,1490,1515]" country="Iceland" county="Natural Forest" municipality="Natural Forest" name="Pico Pinheiro Inferior" pageId="17" pageNumber="418">Pico Pinheiro Inferior</location>
(T13) (
<location id="8EB4600EC52FDB02FB2DFA37DBC10930" LSID="urn:lsid:plazi:treatment:03C287C3C52FDB07FF4CFF7DDFC40AFE:8EB4600EC52FDB02FB2DFA37DBC10930" country="Iceland" county="Natural Forest" municipality="Natural Forest" name="Natural Forest Reserve" pageId="17" pageNumber="418">
<location id="8EB4600EC52FDB02FB2DFA37DEAC0AD4" LSID="urn:lsid:plazi:treatment:03C287C3C52FDB07FF4CFF7DDFC40AFE:8EB4600EC52FDB02FB2DFA37DEAC0AD4" box="[1270,1436,1490,1515]" country="Iceland" county="Natural Forest" municipality="Natural Forest" name="Natural Forest" pageId="17" pageNumber="418">Natural Forest</location>
Reserve
</location>
of
<location id="8EB4600EC52FDB02FEC4FA13DAF00931" LSID="urn:lsid:plazi:treatment:03C287C3C52FDB07FF4CFF7DDFC40AFE:8EB4600EC52FDB02FEC4FA13DAF00931" box="[287,448,1526,1551]" country="Iceland" county="Natural Forest" municipality="Natural Forest" name="Pico Pinheiro" pageId="17" pageNumber="418">Pico Pinheiro</location>
),
<date id="FFD51015C52FDB02FE0CFA12D9760931" box="[471,582,1527,1551]" pageId="17" pageNumber="418" value="2004-08">
<collectingDate id="EF91E9FDC52FDB02FE0CFA12D9760931" box="[471,582,1527,1551]" pageId="17" pageNumber="418" value="2004-08">VIII.2004</collectingDate>
</date>
(14 exx) (UTM 26S 409256, 4278677); Topo
<taxonomicName id="4C6B4D56C52FDB02FBA8FA13DFD80931" box="[1139,1256,1526,1551]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="17" pageNumber="418" phylum="Tracheophyta" rank="genus">Euphorbia</taxonomicName>
(T16) (
<collectingMunicipality id="6BB0ACAFC52FDB02FA9DFA13DBED090C" pageId="17" pageNumber="418">Natural Forest</collectingMunicipality>
Reserve of Topo),
<date id="FFD51015C52FDB02FE1EF9FED904090D" box="[453,564,1563,1587]" pageId="17" pageNumber="418" value="2004-08">
<collectingDate id="EF91E9FDC52FDB02FE1EF9FED904090D" box="[453,564,1563,1587]" pageId="17" pageNumber="418" value="2004-08">VIII.2004</collectingDate>
</date>
(1 ex) (UTM 26S 422871, 4271075); Pico Pinheiro Erical (T09) (
<collectingCounty id="62B54E59C52FDB02FA9DF9FFDBED0968" pageId="17" pageNumber="418">Natural Forest</collectingCounty>
Reserve of Pico Pinheiro),
<date id="FFD51015C52FDB02FDCAF9DAD95E0969" box="[529,622,1599,1623]" pageId="17" pageNumber="418" value="2010-09">
<collectingDate id="EF91E9FDC52FDB02FDCAF9DAD95E0969" box="[529,622,1599,1623]" pageId="17" pageNumber="418" value="2010-09">IX.2010</collectingDate>
</date>
(5 exx) (UTM 26S 408602, 4277888) (
<collectingMethod id="522A4EC2C52FDB02FBFCF9DBDE610968" box="[1063,1361,1598,1623]" pageId="17" pageNumber="418">collected with pitfall traps</collectingMethod>
;
<collectorName id="269E5303C52FDB02FA87F9DBDBE10944" pageId="17" pageNumber="418">Paulo A. V.</collectorName>
Borges
<emphasis id="B91FEAC7C52FDB02FEF6F981DA560945" box="[301,358,1634,1659]" italics="true" pageId="17" pageNumber="418">et al.</emphasis>
leg.).
</materialsCitation>
</paragraph>
<paragraph id="8BD436D5C52FDB02FF1CF963D9B509A1" blockId="17.[151,1436,626,1983]" box="[199,645,1670,1695]" pageId="17" pageNumber="418">Deposited at FCULC, SNM and DTPC.</paragraph>
</subSubSection>
<subSubSection id="C371655EC52FDB02FF1CF94FD8F709D9" pageId="17" pageNumber="418" type="etymology">
<paragraph id="8BD436D5C52FDB02FF1CF94FD8F709D9" blockId="17.[151,1436,626,1983]" pageId="17" pageNumber="418">
<emphasis id="B91FEAC7C52FDB02FF1CF94FDA7E09FD" bold="true" box="[199,334,1706,1731]" pageId="17" pageNumber="418">Etymology.</emphasis>
The species is named after the Azorean naturalist Francisco Arruda Furtado (
<date id="FFD51015C52FDB02FB70F94FDE1B09FD" box="[1195,1323,1706,1731]" pageId="17" pageNumber="418" value="1854" valueMax="1887">18541887</date>
) who was one of the first Portuguese followers of the Darwin's theory of evolution.
</paragraph>
</subSubSection>
<subSubSection id="C371655EC52FDB07FF1CF917DAA70E23" lastPageId="20" lastPageNumber="421" pageId="17" pageNumber="418" type="diagnosis">
<paragraph id="8BD436D5C52FDB02FF1CF917D9F30881" blockId="17.[151,1436,626,1983]" pageId="17" pageNumber="418">
<emphasis id="B91FEAC7C52FDB02FF1CF917DA800835" bold="true" box="[199,432,1778,1803]" pageId="17" pageNumber="418">
Diagnosis (
<figureCitation id="13502A50C52FDB02FE97F917DA930835" box="[332,419,1778,1803]" captionStart="FIGURE 11" captionStartId="18.[151,250,1887,1909]" captionTargetBox="[217,1340,290,1756]" captionTargetId="figure@18.[151,1436,193,1865]" captionTargetPageId="18" captionText="FIGURE 11. Habitus of Tarphius furtadoi sp. nov.. Scale 0.5 mm. (Photo: Erno-Endre Gergely)." httpUri="https://zenodo.org/record/322241/files/figure.png" pageId="17" pageNumber="418">Fig. 11</figureCitation>
).
</emphasis>
Small to medium species (
<quantity id="4C939B30C52FDB02FD34F917D8AA0834" box="[751,922,1778,1803]" metricMagnitude="-3" metricUnit="m" metricValue="3.42" metricValueMax="4.24" metricValueMin="2.6" pageId="17" pageNumber="418" unit="mm" value="3.42" valueMax="4.24" valueMin="2.6">2.604.24 mm</quantity>
; mean: 3.23±
<quantity id="4C939B30C52FDB02FBE3F916DF930835" box="[1080,1187,1779,1803]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="17" pageNumber="418" unit="mm" value="0.41">0.41 mm</quantity>
); body dorsal surface uniform reddish to reddish-brown, antennae, mouth-parts and legs yellowish-brown; body arched (convex); pronotum small and narrower than elytra, transverse, On average 1.45 times as broad as long (1.29x to 1.62x), disc with dense, large rounded granules, each one with a yellowish long, thin, erect rigid long and sub-obtuse seta, with lateral sides slightly rounded and maximum width at middle (
<figureCitation id="13502A50C52FDB02FCBBF867D89708A5" box="[864,935,1922,1947]" captionStart="FIGURE 4" captionStartId="8.[151,250,1787,1809]" captionTargetBox="[151,1436,916,1766]" captionTargetId="figure@8.[151,1436,916,1766]" captionTargetPageId="8" captionText="FIGURE 4. Details of setae in elytra for the species in complex “ tornvalli ”: T. tornvalli (restricted to São Miguel) (A); Tarphius relictus sp. nov. (restricted to Terceira) (B); and Tarphius furtadoi sp. nov. (restricted to São Jorge, Pico and Faial) with a specimen of São Jorge (C) and Faial (D). Scale 0.05 mm (Photos: Erno-Endre Gergely)." httpUri="https://zenodo.org/record/322235/files/figure.png" pageId="17" pageNumber="418">Fig. 4</figureCitation>
C). Differs from the most related species in some morphometrics (see Appendix S5), namely:
</paragraph>
<caption id="DF14665DC52CDB01FF4CF8BADF5F084B" box="[151,1135,1887,1909]" httpUri="https://zenodo.org/record/322241/files/figure.png" pageId="18" pageNumber="419" targetBox="[217,1340,290,1756]" targetPageId="18">
<paragraph id="8BD436D5C52CDB01FF4CF8BADF5F084B" blockId="18.[151,1135,1887,1909]" box="[151,1135,1887,1909]" pageId="18" pageNumber="419">
<emphasis id="B91FEAC7C52CDB01FF4CF8BADA2E084A" bold="true" box="[151,286,1887,1909]" pageId="18" pageNumber="419">FIGURE 11.</emphasis>
Habitus of
<taxonomicName id="4C6B4D56C52CDB01FE4FF8BAD975084B" box="[404,581,1887,1909]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="furtadoi" status="sp. nov.">
<emphasis id="B91FEAC7C52CDB01FE4FF8BAD975084B" box="[404,581,1887,1909]" italics="true" pageId="18" pageNumber="419">Tarphius furtadoi</emphasis>
</taxonomicName>
<emphasis id="B91FEAC7C52CDB01FD97F8BAD9AC084A" bold="true" box="[588,668,1887,1908]" pageId="18" pageNumber="419">
<taxonomicNameLabel id="A22C57BCC52CDB01FD97F8BAD9AC084A" box="[588,668,1887,1908]" pageId="18" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. Scale 0.5 mm. (Photo: Erno-Endre Gergely).
</paragraph>
</caption>
<caption id="DF14665DC52DDB00FF4CF854D95208DB" httpUri="https://zenodo.org/record/322242/files/figure.png" pageId="19" pageNumber="420" targetBox="[170,1406,193,1922]" targetPageId="19">
<paragraph id="8BD436D5C52DDB00FF4CF854D95208DB" blockId="19.[151,1435,1969,2021]" pageId="19" pageNumber="420">
<emphasis id="B91FEAC7C52DDB00FF4CF854DA2F08F8" bold="true" box="[151,287,1969,1991]" pageId="19" pageNumber="420">FIGURE 12.</emphasis>
Detail on the number of seta on the external row of the lateral margin of pronotum in
<taxonomicName id="4C6B4D56C52DDB00FB57F854DE0E08F9" box="[1164,1342,1969,1991]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="420" phylum="Arthropoda" rank="species" species="furtadoi" status="sp. nov.">
<emphasis id="B91FEAC7C52DDB00FB57F854DE0E08F9" box="[1164,1342,1969,1991]" italics="true" pageId="19" pageNumber="420">Tarphius furtadoi</emphasis>
</taxonomicName>
<emphasis id="B91FEAC7C52DDB00FA9FF854DEA508F8" bold="true" box="[1348,1429,1969,1990]" pageId="19" pageNumber="420">
<taxonomicNameLabel id="A22C57BCC52DDB00FA9FF854DEA508F8" box="[1348,1429,1969,1990]" pageId="19" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. Scale 0.25 mm (Photo: Erno-Endre Gergely).
</paragraph>
</caption>
<paragraph id="8BD436D5C52ADB07FF4CFF7DD96F0FEB" blockId="20.[151,1436,151,285]" pageId="20" pageNumber="421">
- differs from
<taxonomicName id="4C6B4D56C52ADB07FE82FF7CDAE70F8E" box="[345,471,151,176]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="tornvalli">
<emphasis id="B91FEAC7C52ADB07FE82FF7CDAE70F8E" box="[345,471,151,176]" italics="true" pageId="20" pageNumber="421">T. tornvalli</emphasis>
</taxonomicName>
in having smaller W elytra/pronotum and L elytra/pronotum ratios, and larger W/L elytra and W/L pronotum ratios; and
</paragraph>
<paragraph id="8BD436D5C52ADB07FF4CFF05DAA70E23" blockId="20.[151,1436,151,285]" pageId="20" pageNumber="421">
- differs from
<taxonomicName id="4C6B4D56C52ADB07FE88FF04DAF10FC6" box="[339,449,223,248]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="relictus" status="sp. nov.">
<emphasis id="B91FEAC7C52ADB07FE88FF04DAF10FC6" box="[339,449,223,248]" italics="true" pageId="20" pageNumber="421">T. relictus</emphasis>
</taxonomicName>
<emphasis id="B91FEAC7C52ADB07FE1CFF3AD9100FC6" bold="true" box="[455,544,223,248]" pageId="20" pageNumber="421">
<taxonomicNameLabel id="A22C57BCC52ADB07FE1CFF3AD9100FC6" box="[455,544,223,248]" pageId="20" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by its larger size and in having smaller L elytra/ Lpronotum, W/L elytra and W/ L pronotum ratios.
</paragraph>
</subSubSection>
<subSubSection id="C371655EC52ADB07FF1CFEA9DFC40AFE" pageId="20" pageNumber="421" type="description">
<paragraph id="8BD436D5C52ADB07FF1CFEA9D8130E93" blockId="20.[151,1437,332,1472]" pageId="20" pageNumber="421">
<emphasis id="B91FEAC7C52ADB07FF1CFEA9DA690E5B" bold="true" box="[199,345,332,357]" pageId="20" pageNumber="421">Description.</emphasis>
Body length of
<typeStatus id="54D08877C52ADB07FDF7FEA9D9A20E5B" box="[556,658,332,357]" pageId="20" pageNumber="421" type="holotype">holotype</typeStatus>
:
<quantity id="4C939B30C52ADB07FD7EFEA9D8230E5A" box="[677,787,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="3.18" pageId="20" pageNumber="421" unit="mm" value="3.18">3.18 mm</quantity>
; small to medium size (length
<quantity id="4C939B30C52ADB07FB4CFEA9DE750E5A" box="[1175,1349,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="3.42" metricValueMax="4.24" metricValueMin="2.6" pageId="20" pageNumber="421" unit="mm" value="3.42" valueMax="4.24" valueMin="2.6">2.604.24 mm</quantity>
; mean: 3.23±
<quantity id="4C939B30C52ADB07FF0CFE95DA0F0EB6" box="[215,319,368,392]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="20" pageNumber="421" unit="mm" value="0.41">0.41 mm</quantity>
); width
<quantity id="4C939B30C52ADB07FE40FE95DACF0EB6" box="[411,511,368,392]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="20" pageNumber="421" unit="mm" value="1.42">1.42 mm</quantity>
<quantity id="4C939B30C52ADB07FDD6FE95D9410EB9" box="[525,625,368,392]" metricMagnitude="-3" metricUnit="m" metricValue="2.17" pageId="20" pageNumber="421" unit="mm" value="2.17">2.17 mm</quantity>
(mean: 1.71±
<quantity id="4C939B30C52ADB07FCD1FE95D8450EB6" box="[778,885,368,392]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="421" unit="mm" value="0.2">0.20 mm</quantity>
), body convex and subquadrate-ovate, reddish or reddish-brown, setose, with rigid setae and large granules.
</paragraph>
<paragraph id="8BD436D5C52ADB07FF1CFE52DFD50DBB" blockId="20.[151,1437,332,1472]" pageId="20" pageNumber="421">
<emphasis id="B91FEAC7C52ADB07FF1CFE52DA370EEE" bold="true" box="[199,263,439,464]" pageId="20" pageNumber="421">Head</emphasis>
: Genae parallel; anterior clypeal margin straight; vertices protrude under the anterior margin of pronotum; surface brown or reddish brown, lighter in clypeus, with dense medium granules, except in clypeus which is more or less smooth, each one with one semi-erect fine hair-like seta slightly directed anteriorly; eyes clearly protruding, glabrous; antennae with segment II smaller than III, segment III 1.3x longer than IV, IV and V equal length, segments VIIIX as long as wide, compact club 2-segmented (segments X and XI); mouth-parts with mandibles brown, the other pieces lighter showing the general morphological pattern of the genus.
</paragraph>
<paragraph id="8BD436D5C52ADB07FF1CFD6ADFD60CD3" blockId="20.[151,1437,332,1472]" pageId="20" pageNumber="421">
<emphasis id="B91FEAC7C52ADB07FF1CFD6ADA700D96" bold="true" box="[199,320,655,680]" pageId="20" pageNumber="421">Pronotum</emphasis>
: Transverse, on average 1.45 times as broad as long, but quite variable (1.29x to 1.62x); surface dark brown, lateral parts reddish, widened in the middle; in general, shape of pronotum as rounded as
<taxonomicName id="4C6B4D56C52ADB07FAC4FD50DEAB0DF3" box="[1311,1435,692,717]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="tornvalli">
<emphasis id="B91FEAC7C52ADB07FAC4FD50DEAB0DF3" box="[1311,1435,692,717]" italics="true" pageId="20" pageNumber="421">T. tornvalli</emphasis>
</taxonomicName>
(cf.
<figureCitation id="13502A50C52ADB07FF12FD32DA230DD1" box="[201,275,727,752]" captionStart="FIGURE 7" captionStartId="11.[151,250,2013,2035]" captionText="FIGURE 7. Habitus of Tarphius tornvalli Gillerfors. Scale 0.5 mm. (Photo: Erno-Endre Gergely)." pageId="20" pageNumber="421">Figs 7</figureCitation>
and
<figureCitation id="13502A50C52ADB07FE88FD3DDA5E0DCE" box="[339,366,728,752]" captionStart="FIGURE 11" captionStartId="18.[151,250,1887,1909]" captionTargetBox="[217,1340,290,1756]" captionTargetId="figure@18.[151,1436,193,1865]" captionTargetPageId="18" captionText="FIGURE 11. Habitus of Tarphius furtadoi sp. nov.. Scale 0.5 mm. (Photo: Erno-Endre Gergely)." httpUri="https://zenodo.org/record/322241/files/figure.png" pageId="20" pageNumber="421">11</figureCitation>
) anterior margin deeply bisinuate, anterior angles strongly protruding and acuminate; hind margin largely bisinuate, in middle broadly produced rearwards; lateral margins arcuate with two irregular rows of long, fine and acuminate setae (2025 each one) (
<figureCitation id="13502A50C52ADB07FD0FFCFAD8190C06" box="[724,809,799,824]" captionStart="FIGURE 12" captionStartId="19.[151,250,1969,1991]" captionTargetBox="[170,1406,193,1922]" captionTargetId="figure@19.[170,1406,193,1922]" captionTargetPageId="19" captionText="FIGURE 12. Detail on the number of seta on the external row of the lateral margin of pronotum in Tarphius furtadoi sp. nov.. Scale 0.25 mm (Photo: Erno-Endre Gergely)." httpUri="https://zenodo.org/record/322242/files/figure.png" pageId="20" pageNumber="421">Fig. 12</figureCitation>
), slightly or without sinuation just before the posterior angles which are prominent; disc with a clear median longitudinal sulcus, half latero-posterior region with a slight nodule; a distinct transverse channel just before the extreme base between the two lateral notches; lateral surface variable in terms of flatness; dorsal upper surface with dense distinct large rounded granules, gradually diminishing in diameter to the sides, each one with one rigid erect sub-obtuse yellowish or golden seta (
<figureCitation id="13502A50C52ADB07FB70FC4ADFCF0CF6" box="[1195,1279,943,968]" captionStart="FIGURE 12" captionStartId="19.[151,250,1969,1991]" captionTargetBox="[170,1406,193,1922]" captionTargetId="figure@19.[170,1406,193,1922]" captionTargetPageId="19" captionText="FIGURE 12. Detail on the number of seta on the external row of the lateral margin of pronotum in Tarphius furtadoi sp. nov.. Scale 0.25 mm (Photo: Erno-Endre Gergely)." httpUri="https://zenodo.org/record/322242/files/figure.png" pageId="20" pageNumber="421">Fig. 12</figureCitation>
); ratios width pronotum/width elytra 0.91±0.03 and length pronotum/length elytra 0.58±0.05 (see Appendix S6).
</paragraph>
<paragraph id="8BD436D5C52ADB07FF1CFC12DF940BFA" blockId="20.[151,1437,332,1472]" pageId="20" pageNumber="421">
<emphasis id="B91FEAC7C52ADB07FF1CFC12DA2C0B2E" bold="true" box="[199,284,1015,1040]" pageId="20" pageNumber="421">Elytra:</emphasis>
On average 0.92 times as broad as long, quite variable (0.84x to 1.09x); in general 1.10 times (mean) broader than pronotum, but less variable than in
<taxonomicName id="4C6B4D56C52ADB07FD6DFBF8D8030B0B" box="[694,819,1052,1077]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="tornvalli">
<emphasis id="B91FEAC7C52ADB07FD6DFBF8D8030B0B" box="[694,819,1052,1077]" italics="true" pageId="20" pageNumber="421">T. tornvalli</emphasis>
</taxonomicName>
(1.03x to 1.18x); disc reddish-brown, reddish laterally, convex, rugosely seriate-punctate, and wrinkled transversally on dorsal surface which is covered with dense granules each one with an erect rigid long and sub-obtuse yellowish or golden seta, with lateral sides slightly rounded and maximum width at middle (
<figureCitation id="13502A50C52ADB07FDA6FB62D9FF0B9E" box="[637,719,1159,1184]" captionStart="FIGURE 4" captionStartId="8.[151,250,1787,1809]" captionTargetBox="[151,1436,916,1766]" captionTargetId="figure@8.[151,1436,916,1766]" captionTargetPageId="8" captionText="FIGURE 4. Details of setae in elytra for the species in complex “ tornvalli ”: T. tornvalli (restricted to São Miguel) (A); Tarphius relictus sp. nov. (restricted to Terceira) (B); and Tarphius furtadoi sp. nov. (restricted to São Jorge, Pico and Faial) with a specimen of São Jorge (C) and Faial (D). Scale 0.05 mm (Photos: Erno-Endre Gergely)." httpUri="https://zenodo.org/record/322235/files/figure.png" pageId="20" pageNumber="421">Figs. 4</figureCitation>
C and D); nodules or gibbosities very distinct with a pattern formula 2, 3, 2, 1, covered with setae more aggregate which confers a pom-pom appearance.
</paragraph>
<paragraph id="8BD436D5C52ADB07FF1CFB2AD9300A32" blockId="20.[151,1437,332,1472]" pageId="20" pageNumber="421">
<emphasis id="B91FEAC7C52ADB07FF1CFB2ADA6E0BD6" bold="true" box="[199,350,1231,1256]" pageId="20" pageNumber="421">Ventral side</emphasis>
. Prosternum as granulose as mesosternum and metasternum; grooves on anterior half of prosternal sides slightly present.
</paragraph>
<paragraph id="8BD436D5C52ADB07FF1CFAF2D89A0A6A" blockId="20.[151,1437,332,1472]" pageId="20" pageNumber="421">
<emphasis id="B91FEAC7C52ADB07FF1CFAF2DBCF0A0E" bold="true" box="[199,255,1303,1328]" pageId="20" pageNumber="421">Legs</emphasis>
. Tibia normal not expanded apically; tarsi 4-segmented, simple, first three tarsomeres of all legs and in both sexes covered ventrally with several very long hyaline fine setae.
</paragraph>
<paragraph id="8BD436D5C52ADB07FF1CFABAD8A80A46" blockId="20.[151,1437,332,1472]" box="[199,920,1375,1400]" pageId="20" pageNumber="421">
<emphasis id="B91FEAC7C52ADB07FF1CFABADA0A0A46" bold="true" box="[199,314,1375,1400]" pageId="20" pageNumber="421">Aedeagus</emphasis>
: Similar to
<taxonomicName id="4C6B4D56C52ADB07FE64FA84D9090A46" box="[447,569,1375,1400]" class="Insecta" family="Zopheridae" genus="Tarphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="tornvalli">
<emphasis id="B91FEAC7C52ADB07FE64FA84D9090A46" box="[447,569,1375,1400]" italics="true" pageId="20" pageNumber="421">T. tornvalli</emphasis>
</taxonomicName>
(see
<figureCitation id="13502A50C52ADB07FDADFABAD9870A46" box="[630,695,1375,1400]" captionStart="FIGURE 6" captionStartId="10.[151,250,1914,1936]" captionTargetBox="[175,1418,661,1880]" captionTargetId="figure@10.[174,1418,661,1880]" captionTargetPageId="10" captionText="FIGURE 6. Details on the number of seta on the external row of the lateral margin of pronotum in the complex “ azoricuswollastoni-depressus ”: T. azoricus (restricted to São Miguel and Flores) (A); Tarphius wollastoni (restricted to Faial) (B); Tarphius gabrielae sp. nov. (restricted to Pico) (C); Tarphius floresensis sp. nov. (restricted to Flores) (D); and T. depressus (restricted to S. Maria) (E). Scales 0.25 mm (Photos: Erno-Endre Gergely)." httpUri="https://zenodo.org/record/322237/files/figure.png" pageId="20" pageNumber="421">
Fig.
<quantity id="4C939B30C52ADB07FD72FABAD9870A46" box="[681,695,1375,1400]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="20" pageNumber="421" unit="in" value="6.0">6</quantity>
</figureCitation>
in
<bibRefCitation id="EFFA4B24C52ADB07FD00FABAD8BB0A46" author="Gillerfors" box="[731,907,1375,1400]" pageId="20" pageNumber="421" refString="Gillerfors, G. (1985) Two new species of the genus Tarphius Erichson from the Azores and redescription of Tarphius wollastoni Crotch (Coleoptera, Colydiidae) Bocagiana, 85, 1 - 7." type="journal article" year="1985">Gillerfors 1985</bibRefCitation>
).
</paragraph>
<paragraph id="8BD436D5C52ADB07FF1CFA61DFC40AFE" blockId="20.[151,1437,332,1472]" pageId="20" pageNumber="421">
<emphasis id="B91FEAC7C52ADB07FF1CFA61DA7A0AA3" bold="true" box="[199,330,1412,1437]" pageId="20" pageNumber="421">Bionomics.</emphasis>
Most specimens were collected in pitfall traps (
<bibRefCitation id="EFFA4B24C52ADB07FCADFA61DF6C0AA3" author="Borges" box="[886,1116,1412,1437]" pageId="20" pageNumber="421" refString="Borges, P. A. V., Aguiar, C., Amaral, J., Amorim, I. R., Andre, G., Arraiol, A., Baz, A., Dinis, F., Enghoff, H., Gaspar, C., Ilharco, F., Mahnert, V., Melo, C., Pereira, F., Quartau, J. A., Ribeiro, S., Ribes, J., Serrano, A. R. M., Sousa, A. B., Strassen, R. Z., Vieira, L., Vieira, V., Vitorino, A. &amp; Wunderlich, J. (2005 a) Ranking protected areas in the Azores using standardized sampling of soil epigean arthropods. Biodiversity and Conservation, 14, 2029 - 2060. http: // dx. doi. org / 10.1007 / s 10531 - 004 - 4283 - y" type="journal article" year="2005" yearSuffix="a">
Borges
<emphasis id="B91FEAC7C52ADB07FC14FA60DF330AA3" box="[975,1027,1412,1437]" italics="true" pageId="20" pageNumber="421">et al</emphasis>
. 2005a
</bibRefCitation>
), but also under the bark of endemic trees and under the bark of dead trunks of exotic trees (
<emphasis id="B91FEAC7C52ADB07FCBCFA4DDF8D0A81" box="[871,1213,1448,1472]" italics="true" pageId="20" pageNumber="421">
<taxonomicName id="4C6B4D56C52ADB07FCBCFA4DDF510AFE" box="[871,1121,1448,1472]" class="Insecta" family="Noctuidae" genus="Cryptomeria" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="japonica">Cryptomeria japonica</taxonomicName>
;
<taxonomicName id="4C6B4D56C52ADB07FBB4FA4DDF8D0A81" box="[1135,1213,1448,1471]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="20" pageNumber="421" phylum="Tracheophyta" rank="genus">Acacia</taxonomicName>
</emphasis>
sp.).
</paragraph>
</subSubSection>
</treatment>
</document>