235 lines
46 KiB
XML
235 lines
46 KiB
XML
<document id="107B2F6C0EBA12F596659744DF01F5D4" ID-DOI="10.1111/zoj.12114" ID-ISSN="0024-4082" ID-Zenodo-Dep="10273948" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630109995688" checkinUser="marcus" docAuthor="Fikáček, Martin, Prokin, Alexander, Yan, Evgeny, Yue, Yanli, Wang, Bo, Ren, Dong & Beattie, Robert" docDate="2014" docId="03C287ACFF88D67EFE9ABBDC40359B39" docLanguage="en" docName="zoj12114.pdf" docOrigin="Zoological Journal of the Linnean Society 170 (4)" docSource="http://doi.wiley.com/10.1111/zoj.12114" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Baissalarva Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie 2014, GEN. NOV." docType="treatment" docVersion="3" lastPageNumber="718" masterDocId="FFFBFFD4FF80D676FFD0B961454A9D19" masterDocTitle="Modern hydrophilid clades present and widespread in the Late Jurassic and Early Cretaceous (Coleoptera: Hydrophiloidea: Hydrophilidae)" masterLastPageNumber="734" masterPageNumber="710" pageNumber="718" updateTime="1701826641646" updateUser="ExternalLinkService">
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<mods:title id="F1F91D5AC852D0DC3F3E102D064B50A4">Modern hydrophilid clades present and widespread in the Late Jurassic and Early Cretaceous (Coleoptera: Hydrophiloidea: Hydrophilidae)</mods:title>
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<mods:namePart id="8BE11C1680EB9202721CCFD42B6650AF">Fikáček, Martin</mods:namePart>
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<mods:namePart id="8B0D46CD680908789BA906C501736CCA">Prokin, Alexander</mods:namePart>
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<mods:namePart id="749BFFB1C8BF01AF8366470527D8D112">Yan, Evgeny</mods:namePart>
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<mods:name id="FED9DEB2F561F9FAC5A60C0D85670021" type="personal">
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<mods:namePart id="9A8007B7E192524BD8C7A10C55B87368">Yue, Yanli</mods:namePart>
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<mods:name id="2220C1411E2F3DAAF4ABD9FF7EDE9C78" type="personal">
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<mods:roleTerm id="CF42C86233F626C643BFE8EBFF99780C">Author</mods:roleTerm>
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<mods:namePart id="7949B634ED11A84F6B137AFD199F9E39">Wang, Bo</mods:namePart>
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<mods:namePart id="1EE20572DB5FBA44FA468CE25A33A84C">Ren, Dong</mods:namePart>
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<mods:namePart id="1A53F4513CCD536820F681E017EE5986">Beattie, Robert</mods:namePart>
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<mods:title id="7483BCDADD68BCD4D733BE7BEFC67CB9">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="D224C29CEEE0F219AEAF315D2B6B46D7">
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<mods:date id="108EDB1CDC069C923944FA2A88D907D7">2014</mods:date>
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<mods:number id="31FD5FDC918A03A60E68CBC5DEE61E6F">2014-04-30</mods:number>
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<mods:number id="D716133E37055D39341112EFE2FFE627">170</mods:number>
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<mods:number id="DDAF0589B36F9C2FE640E7A1C75EF8AD">4</mods:number>
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<mods:start id="BD04F4D61C3D9A5A888A7DB3A5180491">710</mods:start>
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<mods:end id="25964FCFE3ED6C08305AB9CA70AD09DB">734</mods:end>
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<mods:url id="CEA030B363911EAE254F7684A90DE572">http://doi.wiley.com/10.1111/zoj.12114</mods:url>
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<mods:classification id="59B94932B05A676E188D85EAAA8994E4">journal article</mods:classification>
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<mods:identifier id="C7F84E3DE42F7B473A41D8907C4CFED8" type="DOI">10.1111/zoj.12114</mods:identifier>
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<mods:identifier id="4AF1D51F0F3DF58F4A002EF4E839641B" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="D37DFBEBECB0FCB776D2ED7471D6BF44" type="Zenodo-Dep">10273948</mods:identifier>
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<treatment id="03C287ACFF88D67EFE9ABBDC40359B39" LSID="urn:lsid:plazi:treatment:03C287ACFF88D67EFE9ABBDC40359B39" httpUri="http://treatment.plazi.org/id/03C287ACFF88D67EFE9ABBDC40359B39" lastPageNumber="718" pageId="8" pageNumber="718">
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<subSubSection id="C3716531FF88D67EFE9ABBDC472F9FC8" box="[330,613,700,722]" pageId="8" pageNumber="718" type="nomenclature">
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<paragraph id="8BD436BAFF88D67EFE9ABBDC472F9FC8" blockId="8.[330,613,700,722]" box="[330,613,700,722]" pageId="8" pageNumber="718">
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<heading id="D09C81D6FF88D67EFE9ABBDC472F9FC8" box="[330,613,700,722]" centered="true" fontSize="9" level="2" pageId="8" pageNumber="718" reason="4">
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<taxonomicName id="4C6B4D39FF88D67EFE9ABBDC44AF9FC8" ID-CoL="8XJVS" authority="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie, 2014" authorityName="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie" authorityYear="2014" box="[330,485,701,722]" class="Insecta" family="Hydrophilidae" genus="Baissalarva" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="genus" status="gen. nov.">BAISSALARVA</taxonomicName>
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<emphasis id="B91FEAA8FF88D67EFE3FBBA1472F9FC8" bold="true" box="[495,613,700,721]" pageId="8" pageNumber="718">
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<taxonomicNameLabel id="A22C57D3FF88D67EFE3FBBA1472F9FC8" box="[495,613,700,721]" pageId="8" pageNumber="718" rank="genus">GEN. NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3716531FF88D67EFF74BB8540359B39" pageId="8" pageNumber="718" type="materials_examined">
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<paragraph id="8BD436BAFF88D67EFF74BB8544C59E0E" blockId="8.[164,779,739,791]" pageId="8" pageNumber="718">
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<emphasis id="B91FEAA8FF88D67EFF74BB85473A9FE1" box="[164,624,739,761]" italics="true" pageId="8" pageNumber="718">
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<typeStatus id="54D08818FF88D67EFF74BB8545929FE0" box="[164,216,740,761]" pageId="8" pageNumber="718">Type</typeStatus>
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species:
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<taxonomicName id="4C6B4D39FF88D67EFE99BB82473A9FE1" authority="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie, 2014" authorityName="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie" authorityYear="2014" box="[329,624,739,760]" class="Insecta" family="Hydrophilidae" genus="Baissalarva" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="species" species="hydrobioides" status="sp. nov.">Baissalarva hydrobioides</taxonomicName>
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</emphasis>
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<taxonomicNameLabel id="A22C57D3FF88D67EFDAFBB8247949FE1" box="[639,734,739,760]" pageId="8" pageNumber="718" rank="species">sp. nov.</taxonomicNameLabel>
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, by present designation.
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</paragraph>
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<paragraph id="8BD436BAFF88D67EFF74BA5A474C9E49" blockId="8.[164,518,826,848]" box="[164,518,826,848]" pageId="8" pageNumber="718">
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<emphasis id="B91FEAA8FF88D67EFF74BA5A44679E49" box="[164,301,827,848]" italics="true" pageId="8" pageNumber="718">Time range:</emphasis>
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Early Cretaceous.
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</paragraph>
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<paragraph id="8BD436BAFF88D67EFF74BA15449199C0" blockId="8.[164,779,883,1242]" pageId="8" pageNumber="718">
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<emphasis id="B91FEAA8FF88D67EFF74BA1544519E90" box="[164,283,884,905]" italics="true" pageId="8" pageNumber="718">Diagnosis:</emphasis>
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Labrum fused with clypeus; labroclypeus with five large teeth with slightly more isolated left lateral-most one (
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<figureCitation id="13502A3FFF88D67EFEA7BAD0449B9EDF" box="[375,465,945,967]" captionStart="Figure 3" captionStartId="9.[144,223,1669,1688]" captionTargetBox="[259,1302,206,1598]" captionTargetId="figure-0@9.[258,1308,195,1640]" captionTargetPageId="9" captionText="Figure 3. Early Cretaceous representatives of the Hydrobiusini. Cretoxenus australis gen. nov., sp. nov., Koonwarra, Australia, NMVP 103312, piece and counterpiece (A, B). Baissalarva hydrobioides gen. nov., sp. nov. from Baissa, Russia: paratype PIN 3063/6977 (C); holotype PIN 3063/6975: piece, detail of the head (D), counterpiece, detail of the head (E), whole specimen, counterpiece (F). Abbreviations: absc, abdominal dorsal sclerite; absc4, absc7, abdominal dorsal sclerite of segment 4 or 7; aes3, metanepisternum; apls, anapleural suture; cersc, cervical sclerite; dpl8, dorsal plate on abdominal segment 8; epl, epistomal lobe; md, mandible; men, mentum; mssc, mesoscutum; mstr1, mesotarsomere 1; msv, mesoventrite; mxp, maxillary palpus; mtsc, metascutum; ns, nasale; occf, occipital foramen; pros, prosternum; sbm, submentum; scsh, scutellar shield; sstr, sutural stria; ttr, tracheal trunk." figureDoi="http://doi.org/10.5281/zenodo.10273956" httpUri="https://zenodo.org/record/10273956/files/figure.png" pageId="8" pageNumber="718">Figs 3E</figureCitation>
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,
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<figureCitation id="13502A3FFF88D67EFE32BAD0474E9EDE" box="[482,516,945,967]" captionStart="Figure 6" captionStartId="14.[164,243,1707,1726]" captionTargetBox="[246,1359,194,1677]" captionTargetId="figure-0@14.[246,1360,194,1677]" captionTargetPageId="14" captionText="Figure 6. Fossil and recent representatives of the Hydrobiusini and Hydrophilini. Baissalarva hydrobioides sp. nov.: holotype PIN 3063/6975, anterior part of the piece (A), detail of the head of the piece (B), detail of the head of the counterpiece (C); paratype PIN 3063/6977 (D). Larva of extant Limnoxenus niger (Hydrobiusini): whole larva in dorsal view (E), detail of head (G). Larva of extant Sternolophus rufipes (Hydrophilini): whole larva in dorsal view (F), detail of head (H). Holotype of Cretoxenus australis sp. nov. NMVP 103312, piece and counterpiece (I, J). Extant hydrobiusine Limnoxenus zealandicus in ventral view, whole beetle (K) and detail of meso- and metaventrite (L). Abbreviations: md, mandible; mtvpr, metaventral process." figureDoi="http://doi.org/10.5281/zenodo.10273966" httpUri="https://zenodo.org/record/10273966/files/figure.png" pageId="8" pageNumber="718">6B</figureCitation>
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); epistomal lobes low, symmetrical (
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<figureCitation id="13502A3FFF88D67EFE96BAAE44F69EFC" box="[326,444,975,997]" captionStart="Figure 3" captionStartId="9.[144,223,1669,1688]" captionTargetBox="[259,1302,206,1598]" captionTargetId="figure-0@9.[258,1308,195,1640]" captionTargetPageId="9" captionText="Figure 3. Early Cretaceous representatives of the Hydrobiusini. Cretoxenus australis gen. nov., sp. nov., Koonwarra, Australia, NMVP 103312, piece and counterpiece (A, B). Baissalarva hydrobioides gen. nov., sp. nov. from Baissa, Russia: paratype PIN 3063/6977 (C); holotype PIN 3063/6975: piece, detail of the head (D), counterpiece, detail of the head (E), whole specimen, counterpiece (F). Abbreviations: absc, abdominal dorsal sclerite; absc4, absc7, abdominal dorsal sclerite of segment 4 or 7; aes3, metanepisternum; apls, anapleural suture; cersc, cervical sclerite; dpl8, dorsal plate on abdominal segment 8; epl, epistomal lobe; md, mandible; men, mentum; mssc, mesoscutum; mstr1, mesotarsomere 1; msv, mesoventrite; mxp, maxillary palpus; mtsc, metascutum; ns, nasale; occf, occipital foramen; pros, prosternum; sbm, submentum; scsh, scutellar shield; sstr, sutural stria; ttr, tracheal trunk." figureDoi="http://doi.org/10.5281/zenodo.10273956" httpUri="https://zenodo.org/record/10273956/files/figure.png" pageId="8" pageNumber="718">Fig. 3D–E</figureCitation>
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); occipital foramen situated dorsally (
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<figureCitation id="13502A3FFF88D67EFEF6BA8F44CA991A" box="[294,384,1006,1028]" captionStart="Figure 3" captionStartId="9.[144,223,1669,1688]" captionTargetBox="[259,1302,206,1598]" captionTargetId="figure-0@9.[258,1308,195,1640]" captionTargetPageId="9" captionText="Figure 3. Early Cretaceous representatives of the Hydrobiusini. Cretoxenus australis gen. nov., sp. nov., Koonwarra, Australia, NMVP 103312, piece and counterpiece (A, B). Baissalarva hydrobioides gen. nov., sp. nov. from Baissa, Russia: paratype PIN 3063/6977 (C); holotype PIN 3063/6975: piece, detail of the head (D), counterpiece, detail of the head (E), whole specimen, counterpiece (F). Abbreviations: absc, abdominal dorsal sclerite; absc4, absc7, abdominal dorsal sclerite of segment 4 or 7; aes3, metanepisternum; apls, anapleural suture; cersc, cervical sclerite; dpl8, dorsal plate on abdominal segment 8; epl, epistomal lobe; md, mandible; men, mentum; mssc, mesoscutum; mstr1, mesotarsomere 1; msv, mesoventrite; mxp, maxillary palpus; mtsc, metascutum; ns, nasale; occf, occipital foramen; pros, prosternum; sbm, submentum; scsh, scutellar shield; sstr, sutural stria; ttr, tracheal trunk." figureDoi="http://doi.org/10.5281/zenodo.10273956" httpUri="https://zenodo.org/record/10273956/files/figure.png" pageId="8" pageNumber="718">Figs 3E</figureCitation>
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,
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<figureCitation id="13502A3FFF88D67EFE74BA8F448C991D" box="[420,454,1006,1028]" captionStart="Figure 6" captionStartId="14.[164,243,1707,1726]" captionTargetBox="[246,1359,194,1677]" captionTargetId="figure-0@14.[246,1360,194,1677]" captionTargetPageId="14" captionText="Figure 6. Fossil and recent representatives of the Hydrobiusini and Hydrophilini. Baissalarva hydrobioides sp. nov.: holotype PIN 3063/6975, anterior part of the piece (A), detail of the head of the piece (B), detail of the head of the counterpiece (C); paratype PIN 3063/6977 (D). Larva of extant Limnoxenus niger (Hydrobiusini): whole larva in dorsal view (E), detail of head (G). Larva of extant Sternolophus rufipes (Hydrophilini): whole larva in dorsal view (F), detail of head (H). Holotype of Cretoxenus australis sp. nov. NMVP 103312, piece and counterpiece (I, J). Extant hydrobiusine Limnoxenus zealandicus in ventral view, whole beetle (K) and detail of meso- and metaventrite (L). Abbreviations: md, mandible; mtvpr, metaventral process." figureDoi="http://doi.org/10.5281/zenodo.10273966" httpUri="https://zenodo.org/record/10273966/files/figure.png" pageId="8" pageNumber="718">6A</figureCitation>
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); mandible with three retinacular teeth (
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<figureCitation id="13502A3FFF88D67EFEA8BD6D4499993B" box="[376,467,1036,1058]" captionStart="Figure 3" captionStartId="9.[144,223,1669,1688]" captionTargetBox="[259,1302,206,1598]" captionTargetId="figure-0@9.[258,1308,195,1640]" captionTargetPageId="9" captionText="Figure 3. Early Cretaceous representatives of the Hydrobiusini. Cretoxenus australis gen. nov., sp. nov., Koonwarra, Australia, NMVP 103312, piece and counterpiece (A, B). Baissalarva hydrobioides gen. nov., sp. nov. from Baissa, Russia: paratype PIN 3063/6977 (C); holotype PIN 3063/6975: piece, detail of the head (D), counterpiece, detail of the head (E), whole specimen, counterpiece (F). Abbreviations: absc, abdominal dorsal sclerite; absc4, absc7, abdominal dorsal sclerite of segment 4 or 7; aes3, metanepisternum; apls, anapleural suture; cersc, cervical sclerite; dpl8, dorsal plate on abdominal segment 8; epl, epistomal lobe; md, mandible; men, mentum; mssc, mesoscutum; mstr1, mesotarsomere 1; msv, mesoventrite; mxp, maxillary palpus; mtsc, metascutum; ns, nasale; occf, occipital foramen; pros, prosternum; sbm, submentum; scsh, scutellar shield; sstr, sutural stria; ttr, tracheal trunk." figureDoi="http://doi.org/10.5281/zenodo.10273956" httpUri="https://zenodo.org/record/10273956/files/figure.png" pageId="8" pageNumber="718">Figs 3D</figureCitation>
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,
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<figureCitation id="13502A3FFF88D67EFE30BD6D4748993B" box="[480,514,1036,1058]" captionStart="Figure 6" captionStartId="14.[164,243,1707,1726]" captionTargetBox="[246,1359,194,1677]" captionTargetId="figure-0@14.[246,1360,194,1677]" captionTargetPageId="14" captionText="Figure 6. Fossil and recent representatives of the Hydrobiusini and Hydrophilini. Baissalarva hydrobioides sp. nov.: holotype PIN 3063/6975, anterior part of the piece (A), detail of the head of the piece (B), detail of the head of the counterpiece (C); paratype PIN 3063/6977 (D). Larva of extant Limnoxenus niger (Hydrobiusini): whole larva in dorsal view (E), detail of head (G). Larva of extant Sternolophus rufipes (Hydrophilini): whole larva in dorsal view (F), detail of head (H). Holotype of Cretoxenus australis sp. nov. NMVP 103312, piece and counterpiece (I, J). Extant hydrobiusine Limnoxenus zealandicus in ventral view, whole beetle (K) and detail of meso- and metaventrite (L). Abbreviations: md, mandible; mtvpr, metaventral process." figureDoi="http://doi.org/10.5281/zenodo.10273966" httpUri="https://zenodo.org/record/10273966/files/figure.png" pageId="8" pageNumber="718">6C</figureCitation>
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); prothorax completely sclerotized dorsally (
|
||
<figureCitation id="13502A3FFF88D67EFE4CBD4A47529958" box="[412,536,1067,1089]" captionStart="Figure 3" captionStartId="9.[144,223,1669,1688]" captionTargetBox="[259,1302,206,1598]" captionTargetId="figure-0@9.[258,1308,195,1640]" captionTargetPageId="9" captionText="Figure 3. Early Cretaceous representatives of the Hydrobiusini. Cretoxenus australis gen. nov., sp. nov., Koonwarra, Australia, NMVP 103312, piece and counterpiece (A, B). Baissalarva hydrobioides gen. nov., sp. nov. from Baissa, Russia: paratype PIN 3063/6977 (C); holotype PIN 3063/6975: piece, detail of the head (D), counterpiece, detail of the head (E), whole specimen, counterpiece (F). Abbreviations: absc, abdominal dorsal sclerite; absc4, absc7, abdominal dorsal sclerite of segment 4 or 7; aes3, metanepisternum; apls, anapleural suture; cersc, cervical sclerite; dpl8, dorsal plate on abdominal segment 8; epl, epistomal lobe; md, mandible; men, mentum; mssc, mesoscutum; mstr1, mesotarsomere 1; msv, mesoventrite; mxp, maxillary palpus; mtsc, metascutum; ns, nasale; occf, occipital foramen; pros, prosternum; sbm, submentum; scsh, scutellar shield; sstr, sutural stria; ttr, tracheal trunk." figureDoi="http://doi.org/10.5281/zenodo.10273956" httpUri="https://zenodo.org/record/10273956/files/figure.png" pageId="8" pageNumber="718">Figs 3C, F</figureCitation>
|
||
,
|
||
<figureCitation id="13502A3FFF88D67EFDFBBD4A473F9958" box="[555,629,1067,1089]" captionStart="Figure 6" captionStartId="14.[164,243,1707,1726]" captionTargetBox="[246,1359,194,1677]" captionTargetId="figure-0@14.[246,1360,194,1677]" captionTargetPageId="14" captionText="Figure 6. Fossil and recent representatives of the Hydrobiusini and Hydrophilini. Baissalarva hydrobioides sp. nov.: holotype PIN 3063/6975, anterior part of the piece (A), detail of the head of the piece (B), detail of the head of the counterpiece (C); paratype PIN 3063/6977 (D). Larva of extant Limnoxenus niger (Hydrobiusini): whole larva in dorsal view (E), detail of head (G). Larva of extant Sternolophus rufipes (Hydrophilini): whole larva in dorsal view (F), detail of head (H). Holotype of Cretoxenus australis sp. nov. NMVP 103312, piece and counterpiece (I, J). Extant hydrobiusine Limnoxenus zealandicus in ventral view, whole beetle (K) and detail of meso- and metaventrite (L). Abbreviations: md, mandible; mtvpr, metaventral process." figureDoi="http://doi.org/10.5281/zenodo.10273966" httpUri="https://zenodo.org/record/10273966/files/figure.png" pageId="8" pageNumber="718">6A, D</figureCitation>
|
||
); meso- and metathorax with large transverse sclerites (
|
||
<figureCitation id="13502A3FFF88D67EFD7CBD2B45FE9967" captionStart="Figure 3" captionStartId="9.[144,223,1669,1688]" captionTargetBox="[259,1302,206,1598]" captionTargetId="figure-0@9.[258,1308,195,1640]" captionTargetPageId="9" captionText="Figure 3. Early Cretaceous representatives of the Hydrobiusini. Cretoxenus australis gen. nov., sp. nov., Koonwarra, Australia, NMVP 103312, piece and counterpiece (A, B). Baissalarva hydrobioides gen. nov., sp. nov. from Baissa, Russia: paratype PIN 3063/6977 (C); holotype PIN 3063/6975: piece, detail of the head (D), counterpiece, detail of the head (E), whole specimen, counterpiece (F). Abbreviations: absc, abdominal dorsal sclerite; absc4, absc7, abdominal dorsal sclerite of segment 4 or 7; aes3, metanepisternum; apls, anapleural suture; cersc, cervical sclerite; dpl8, dorsal plate on abdominal segment 8; epl, epistomal lobe; md, mandible; men, mentum; mssc, mesoscutum; mstr1, mesotarsomere 1; msv, mesoventrite; mxp, maxillary palpus; mtsc, metascutum; ns, nasale; occf, occipital foramen; pros, prosternum; sbm, submentum; scsh, scutellar shield; sstr, sutural stria; ttr, tracheal trunk." figureDoi="http://doi.org/10.5281/zenodo.10273956" httpUri="https://zenodo.org/record/10273956/files/figure.png" pageId="8" pageNumber="718">Figs 3C, F</figureCitation>
|
||
,
|
||
<figureCitation id="13502A3FFF88D67EFF12BD0944439967" box="[194,265,1128,1150]" captionStart="Figure 6" captionStartId="14.[164,243,1707,1726]" captionTargetBox="[246,1359,194,1677]" captionTargetId="figure-0@14.[246,1360,194,1677]" captionTargetPageId="14" captionText="Figure 6. Fossil and recent representatives of the Hydrobiusini and Hydrophilini. Baissalarva hydrobioides sp. nov.: holotype PIN 3063/6975, anterior part of the piece (A), detail of the head of the piece (B), detail of the head of the counterpiece (C); paratype PIN 3063/6977 (D). Larva of extant Limnoxenus niger (Hydrobiusini): whole larva in dorsal view (E), detail of head (G). Larva of extant Sternolophus rufipes (Hydrophilini): whole larva in dorsal view (F), detail of head (H). Holotype of Cretoxenus australis sp. nov. NMVP 103312, piece and counterpiece (I, J). Extant hydrobiusine Limnoxenus zealandicus in ventral view, whole beetle (K) and detail of meso- and metaventrite (L). Abbreviations: md, mandible; mtvpr, metaventral process." figureDoi="http://doi.org/10.5281/zenodo.10273966" httpUri="https://zenodo.org/record/10273966/files/figure.png" pageId="8" pageNumber="718">6A, D</figureCitation>
|
||
); abdominal segments largely membranous, with small dorsal circular sclerites only (
|
||
<figureCitation id="13502A3FFF88D67EFD5DBDE6464C9985" box="[653,774,1159,1181]" captionStart="Figure 3" captionStartId="9.[144,223,1669,1688]" captionTargetBox="[259,1302,206,1598]" captionTargetId="figure-0@9.[258,1308,195,1640]" captionTargetPageId="9" captionText="Figure 3. Early Cretaceous representatives of the Hydrobiusini. Cretoxenus australis gen. nov., sp. nov., Koonwarra, Australia, NMVP 103312, piece and counterpiece (A, B). Baissalarva hydrobioides gen. nov., sp. nov. from Baissa, Russia: paratype PIN 3063/6977 (C); holotype PIN 3063/6975: piece, detail of the head (D), counterpiece, detail of the head (E), whole specimen, counterpiece (F). Abbreviations: absc, abdominal dorsal sclerite; absc4, absc7, abdominal dorsal sclerite of segment 4 or 7; aes3, metanepisternum; apls, anapleural suture; cersc, cervical sclerite; dpl8, dorsal plate on abdominal segment 8; epl, epistomal lobe; md, mandible; men, mentum; mssc, mesoscutum; mstr1, mesotarsomere 1; msv, mesoventrite; mxp, maxillary palpus; mtsc, metascutum; ns, nasale; occf, occipital foramen; pros, prosternum; sbm, submentum; scsh, scutellar shield; sstr, sutural stria; ttr, tracheal trunk." figureDoi="http://doi.org/10.5281/zenodo.10273956" httpUri="https://zenodo.org/record/10273956/files/figure.png" pageId="8" pageNumber="718">Figs 3C, F</figureCitation>
|
||
,
|
||
<figureCitation id="13502A3FFF88D67EFF74BDC445A099A2" box="[164,234,1189,1211]" captionStart="Figure 6" captionStartId="14.[164,243,1707,1726]" captionTargetBox="[246,1359,194,1677]" captionTargetId="figure-0@14.[246,1360,194,1677]" captionTargetPageId="14" captionText="Figure 6. Fossil and recent representatives of the Hydrobiusini and Hydrophilini. Baissalarva hydrobioides sp. nov.: holotype PIN 3063/6975, anterior part of the piece (A), detail of the head of the piece (B), detail of the head of the counterpiece (C); paratype PIN 3063/6977 (D). Larva of extant Limnoxenus niger (Hydrobiusini): whole larva in dorsal view (E), detail of head (G). Larva of extant Sternolophus rufipes (Hydrophilini): whole larva in dorsal view (F), detail of head (H). Holotype of Cretoxenus australis sp. nov. NMVP 103312, piece and counterpiece (I, J). Extant hydrobiusine Limnoxenus zealandicus in ventral view, whole beetle (K) and detail of meso- and metaventrite (L). Abbreviations: md, mandible; mtvpr, metaventral process." figureDoi="http://doi.org/10.5281/zenodo.10273966" httpUri="https://zenodo.org/record/10273966/files/figure.png" pageId="8" pageNumber="718">6A, D</figureCitation>
|
||
); tracheal system with a pair of large tracheal trunks (
|
||
<figureCitation id="13502A3FFF88D67EFED2BDA5443099C0" box="[258,378,1220,1242]" captionStart="Figure 3" captionStartId="9.[144,223,1669,1688]" captionTargetBox="[259,1302,206,1598]" captionTargetId="figure-0@9.[258,1308,195,1640]" captionTargetPageId="9" captionText="Figure 3. Early Cretaceous representatives of the Hydrobiusini. Cretoxenus australis gen. nov., sp. nov., Koonwarra, Australia, NMVP 103312, piece and counterpiece (A, B). Baissalarva hydrobioides gen. nov., sp. nov. from Baissa, Russia: paratype PIN 3063/6977 (C); holotype PIN 3063/6975: piece, detail of the head (D), counterpiece, detail of the head (E), whole specimen, counterpiece (F). Abbreviations: absc, abdominal dorsal sclerite; absc4, absc7, abdominal dorsal sclerite of segment 4 or 7; aes3, metanepisternum; apls, anapleural suture; cersc, cervical sclerite; dpl8, dorsal plate on abdominal segment 8; epl, epistomal lobe; md, mandible; men, mentum; mssc, mesoscutum; mstr1, mesotarsomere 1; msv, mesoventrite; mxp, maxillary palpus; mtsc, metascutum; ns, nasale; occf, occipital foramen; pros, prosternum; sbm, submentum; scsh, scutellar shield; sstr, sutural stria; ttr, tracheal trunk." figureDoi="http://doi.org/10.5281/zenodo.10273956" httpUri="https://zenodo.org/record/10273956/files/figure.png" pageId="8" pageNumber="718">Figs 3C, F</figureCitation>
|
||
,
|
||
<figureCitation id="13502A3FFF88D67EFE58BDA5448599C0" box="[392,463,1220,1242]" captionStart="Figure 6" captionStartId="14.[164,243,1707,1726]" captionTargetBox="[246,1359,194,1677]" captionTargetId="figure-0@14.[246,1360,194,1677]" captionTargetPageId="14" captionText="Figure 6. Fossil and recent representatives of the Hydrobiusini and Hydrophilini. Baissalarva hydrobioides sp. nov.: holotype PIN 3063/6975, anterior part of the piece (A), detail of the head of the piece (B), detail of the head of the counterpiece (C); paratype PIN 3063/6977 (D). Larva of extant Limnoxenus niger (Hydrobiusini): whole larva in dorsal view (E), detail of head (G). Larva of extant Sternolophus rufipes (Hydrophilini): whole larva in dorsal view (F), detail of head (H). Holotype of Cretoxenus australis sp. nov. NMVP 103312, piece and counterpiece (I, J). Extant hydrobiusine Limnoxenus zealandicus in ventral view, whole beetle (K) and detail of meso- and metaventrite (L). Abbreviations: md, mandible; mtvpr, metaventral process." figureDoi="http://doi.org/10.5281/zenodo.10273966" httpUri="https://zenodo.org/record/10273966/files/figure.png" pageId="8" pageNumber="718">6A, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD436BAFF88D67EFF74BD9C47779876" blockId="8.[164,779,1277,1391]" pageId="8" pageNumber="718">
|
||
<emphasis id="B91FEAA8FF88D67EFF74BD9C4468980B" box="[164,290,1277,1298]" italics="true" pageId="8" pageNumber="718">Etymology:</emphasis>
|
||
The generic name consists of the name of the
|
||
<typeStatus id="54D08818FF88D67EFF0ABC7D44419828" box="[218,267,1308,1329]" pageId="8" pageNumber="718">type</typeStatus>
|
||
locality in which the genus was found (Baissa) and the ending
|
||
<emphasis id="B91FEAA8FF88D67EFE11BC5B474F9856" box="[449,517,1338,1359]" italics="true" pageId="8" pageNumber="718">-larva</emphasis>
|
||
indicating that it represents the larval stage. Feminine.
|
||
</paragraph>
|
||
<paragraph id="8BD436BAFF88D67EFF74BCF344B19A0E" blockId="8.[164,779,1426,1815]" pageId="8" pageNumber="718">
|
||
<emphasis id="B91FEAA8FF88D67EFF74BCF344CF98BE" box="[164,389,1426,1447]" italics="true" pageId="8" pageNumber="718">Family assignment:</emphasis>
|
||
The combination of dorsally situated occipital foramen of the head (present due to the hyperprognathous head allowing it to hold the prey above the water surface when it is processed preorally) and the anterior margin of the head bearing series of teeth on labroclypeus reliably assigns the fossils to the family
|
||
<taxonomicName id="4C6B4D39FF88D67EFDF9BF28479A9B46" box="[553,720,1609,1631]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
|
||
. The family assignment is also supported by the general habitus, which closely corresponds to those of larvae of extant
|
||
<taxonomicName id="4C6B4D39FF88D67EFEF1BFC4448F9BA2" box="[289,453,1701,1723]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
|
||
(see, e.g.
|
||
<figureCitation id="13502A3FFF88D67EFD9DBFC4478E9BA2" box="[589,708,1701,1723]" captionStart="Figure 6" captionStartId="14.[164,243,1707,1726]" captionTargetBox="[246,1359,194,1677]" captionTargetId="figure-0@14.[246,1360,194,1677]" captionTargetPageId="14" captionText="Figure 6. Fossil and recent representatives of the Hydrobiusini and Hydrophilini. Baissalarva hydrobioides sp. nov.: holotype PIN 3063/6975, anterior part of the piece (A), detail of the head of the piece (B), detail of the head of the counterpiece (C); paratype PIN 3063/6977 (D). Larva of extant Limnoxenus niger (Hydrobiusini): whole larva in dorsal view (E), detail of head (G). Larva of extant Sternolophus rufipes (Hydrophilini): whole larva in dorsal view (F), detail of head (H). Holotype of Cretoxenus australis sp. nov. NMVP 103312, piece and counterpiece (I, J). Extant hydrobiusine Limnoxenus zealandicus in ventral view, whole beetle (K) and detail of meso- and metaventrite (L). Abbreviations: md, mandible; mtvpr, metaventral process." figureDoi="http://doi.org/10.5281/zenodo.10273966" httpUri="https://zenodo.org/record/10273966/files/figure.png" pageId="8" pageNumber="718">Fig. 6E–H</figureCitation>
|
||
), and by the presence of massive tracheal trunks in the abdomen (an adaptation for breathing atmospheric oxygen in submerged larvae).
|
||
</paragraph>
|
||
<paragraph id="8BD436BAFF88D67EFF74BE5B40EB9F17" blockId="8.[164,779,1850,1903]" lastBlockId="8.[826,1442,198,526]" pageId="8" pageNumber="718">
|
||
<emphasis id="B91FEAA8FF88D67EFF74BE5B47689A56" box="[164,546,1850,1871]" italics="true" pageId="8" pageNumber="718">Results of phylogenetic analysis:</emphasis>
|
||
The unconstrained analysis resulted in seven most parsimonious trees 207 steps in length. Their strict consensus tree (
|
||
<figureCitation id="13502A3FFF88D67EFC92B98546DF9DE3" box="[834,917,228,250]" captionStart="Figure 4" captionStartId="10.[164,243,1310,1329]" captionTargetBox="[164,1444,194,1279]" captionTargetId="figure-205@10.[164,1444,194,1280]" captionTargetPageId="10" captionText="Figure 4. Results of the phylogenetic analyses of the position of Baissalarva hydrobioides. Unconstrained analysis, strict consensus of seven most-parsimonious trees of 207 steps (A); single most parsimonious tree of length 240 steps resulting from the constrained analysis (B)." figureDoi="http://doi.org/10.5281/zenodo.10273960" httpUri="https://zenodo.org/record/10273960/files/figure.png" pageId="8" pageNumber="718">Fig. 4A</figureCitation>
|
||
) is largely unresolved, but
|
||
<taxonomicName id="4C6B4D39FF88D67EFB1BB98540199DE0" authorityName="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie" authorityYear="2014" box="[1227,1363,228,249]" class="Insecta" family="Hydrophilidae" genus="Baissalarva" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA8FF88D67EFB1BB98540199DE0" box="[1227,1363,228,249]" italics="true" pageId="8" pageNumber="718">Baissalarva</emphasis>
|
||
</taxonomicName>
|
||
is still recognized to form a group with all included genera of the
|
||
<taxonomicName id="4C6B4D39FF88D67EFC4AB84041659C2E" baseAuthorityName="Mulsant" baseAuthorityYear="1844" box="[922,1071,289,311]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="tribe" tribe="Hydrobiusini">Hydrobiusini</taxonomicName>
|
||
and the hydrophiline genus
|
||
<taxonomicName id="4C6B4D39FF88D67EFCEAB821469A9C4C" authorityName="Solier" authorityYear="1834" box="[826,976,320,341]" class="Insecta" family="Hydrophilidae" genus="Sternolophus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA8FF88D67EFCEAB821469A9C4C" box="[826,976,320,341]" italics="true" pageId="8" pageNumber="718">Sternolophus</emphasis>
|
||
</taxonomicName>
|
||
based on the nasale with five teeth and the left-most one distinctly separated from the other. In the constrained analysis using the molecules-based backbone tree, a single most parsimonious tree of length 240 steps was found (
|
||
<figureCitation id="13502A3FFF88D67EFB5FB8DB41AF9CC8" box="[1167,1253,442,465]" captionStart="Figure 4" captionStartId="10.[164,243,1310,1329]" captionTargetBox="[164,1444,194,1279]" captionTargetId="figure-205@10.[164,1444,194,1280]" captionTargetPageId="10" captionText="Figure 4. Results of the phylogenetic analyses of the position of Baissalarva hydrobioides. Unconstrained analysis, strict consensus of seven most-parsimonious trees of 207 steps (A); single most parsimonious tree of length 240 steps resulting from the constrained analysis (B)." figureDoi="http://doi.org/10.5281/zenodo.10273960" httpUri="https://zenodo.org/record/10273960/files/figure.png" pageId="8" pageNumber="718">Fig. 4B</figureCitation>
|
||
).
|
||
<taxonomicName id="4C6B4D39FF88D67EFB2BB8DB40C99CD6" authorityName="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie" authorityYear="2014" box="[1275,1411,442,463]" class="Insecta" family="Hydrophilidae" genus="Baissalarva" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA8FF88D67EFB2BB8DB40C99CD6" box="[1275,1411,442,463]" italics="true" pageId="8" pageNumber="718">Baissalarva</emphasis>
|
||
</taxonomicName>
|
||
is placed as a sister taxon to the extant representatives of the
|
||
<taxonomicName id="4C6B4D39FF88D67EFC52B899415D9F17" baseAuthorityName="Mulsant" baseAuthorityYear="1844" box="[898,1047,504,526]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="tribe" tribe="Hydrobiusini">Hydrobiusini</taxonomicName>
|
||
based on characters of the nasale.
|
||
</paragraph>
|
||
<paragraph id="8BD436BAFF88D67EFCEABB5940359B39" blockId="8.[826,1441,568,1569]" pageId="8" pageNumber="718">
|
||
<emphasis id="B91FEAA8FF88D67EFCEABB5941EC9F57" box="[826,1190,568,590]" italics="true" pageId="8" pageNumber="718">Comparison with other genera:</emphasis>
|
||
The presence of five teeth of labroclypeus with the left one slightly more separated from the others is very characteristic for the modern larvae of the tribe
|
||
<taxonomicName id="4C6B4D39FF88D67EFB06BBF540219FB3" baseAuthorityName="Mulsant" baseAuthorityYear="1844" box="[1238,1387,660,682]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="tribe" tribe="Hydrobiusini">Hydrobiusini</taxonomicName>
|
||
and the hydrophiline genus
|
||
<taxonomicName id="4C6B4D39FF88D67EFBB9BBD340EB9FDE" authority="Solier, 1834" authorityName="Solier" authorityYear="1834" box="[1129,1441,690,712]" class="Insecta" family="Hydrophilidae" genus="Sternolophus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA8FF88D67EFBB9BBD341B59FDE" box="[1129,1279,690,711]" italics="true" pageId="8" pageNumber="718">Sternolophus</emphasis>
|
||
<bibRefCitation id="EFFA4B4BFF88D67EFAC1BBD340EB9FDE" author="Solier AJJ" box="[1297,1441,690,712]" pageId="8" pageNumber="718" pagination="299 - 318" refId="ref14938" refString="Solier AJJ. 1834. Observations sur la tribu des Hydrophiliens, et principalement sur le genre Hydrophilus de Fabricius. Annales de la Societe Entomologique de France 3: 299 - 318." type="journal article" year="1834">Solier, 1834</bibRefCitation>
|
||
</taxonomicName>
|
||
(Archangelsky, 1997, 2004;
|
||
<bibRefCitation id="EFFA4B4BFF88D67EFB59BBB046359E1C" author="Minoshima Y & Hayashi M" pageId="8" pageNumber="718" pagination="1 - 118" refId="ref14257" refString="Minoshima Y, Hayashi M. 2011 a. Larval morphology of the Japanese species of the tribes Acidocerini, Hydrobiusini and Hydrophilini (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 51 (Supplementum): 1 - 118." type="journal article" year="2011">Minoshima & Hayashi, 2011a</bibRefCitation>
|
||
, b, 2012b). The form of the nasale (i.e. characters 5 and 6) is also responsible for grouping
|
||
<taxonomicName id="4C6B4D39FF88D67EFCEABA4C46889E5B" authorityName="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie" authorityYear="2014" box="[826,962,813,834]" class="Insecta" family="Hydrophilidae" genus="Baissalarva" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA8FF88D67EFCEABA4C46889E5B" box="[826,962,813,834]" italics="true" pageId="8" pageNumber="718">Baissalarva</emphasis>
|
||
</taxonomicName>
|
||
with these taxa in both phylogenetic analyses performed. Comparison of
|
||
<taxonomicName id="4C6B4D39FF88D67EFB0CBA2A402E9E79" authorityName="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie" authorityYear="2014" box="[1244,1380,843,864]" class="Insecta" family="Hydrophilidae" genus="Baissalarva" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA8FF88D67EFB0CBA2A402E9E79" box="[1244,1380,843,864]" italics="true" pageId="8" pageNumber="718">Baissalarva</emphasis>
|
||
</taxonomicName>
|
||
with modern representatives of the
|
||
<taxonomicName id="4C6B4D39FF88D67EFB70BA0B407F9E99" baseAuthorityName="Mulsant" baseAuthorityYear="1844" box="[1184,1333,874,896]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="tribe" tribe="Hydrobiusini">Hydrobiusini</taxonomicName>
|
||
(
|
||
<figureCitation id="13502A3FFF88D67EFA97BA0B46049E86" captionStart="Figure 6" captionStartId="14.[164,243,1707,1726]" captionTargetBox="[246,1359,194,1677]" captionTargetId="figure-0@14.[246,1360,194,1677]" captionTargetPageId="14" captionText="Figure 6. Fossil and recent representatives of the Hydrobiusini and Hydrophilini. Baissalarva hydrobioides sp. nov.: holotype PIN 3063/6975, anterior part of the piece (A), detail of the head of the piece (B), detail of the head of the counterpiece (C); paratype PIN 3063/6977 (D). Larva of extant Limnoxenus niger (Hydrobiusini): whole larva in dorsal view (E), detail of head (G). Larva of extant Sternolophus rufipes (Hydrophilini): whole larva in dorsal view (F), detail of head (H). Holotype of Cretoxenus australis sp. nov. NMVP 103312, piece and counterpiece (I, J). Extant hydrobiusine Limnoxenus zealandicus in ventral view, whole beetle (K) and detail of meso- and metaventrite (L). Abbreviations: md, mandible; mtvpr, metaventral process." figureDoi="http://doi.org/10.5281/zenodo.10273966" httpUri="https://zenodo.org/record/10273966/files/figure.png" pageId="8" pageNumber="718">Fig. 6E, G</figureCitation>
|
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) and
|
||
<taxonomicName id="4C6B4D39FF88D67EFC5FBAE9416F9E84" authorityName="Solier" authorityYear="1834" box="[911,1061,904,925]" class="Insecta" family="Hydrophilidae" genus="Sternolophus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA8FF88D67EFC5FBAE9416F9E84" box="[911,1061,904,925]" italics="true" pageId="8" pageNumber="718">Sternolophus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13502A3FFF88D67EFBE5BAE941ED9E87" box="[1077,1191,904,926]" captionStart="Figure 6" captionStartId="14.[164,243,1707,1726]" captionTargetBox="[246,1359,194,1677]" captionTargetId="figure-0@14.[246,1360,194,1677]" captionTargetPageId="14" captionText="Figure 6. Fossil and recent representatives of the Hydrobiusini and Hydrophilini. Baissalarva hydrobioides sp. nov.: holotype PIN 3063/6975, anterior part of the piece (A), detail of the head of the piece (B), detail of the head of the counterpiece (C); paratype PIN 3063/6977 (D). Larva of extant Limnoxenus niger (Hydrobiusini): whole larva in dorsal view (E), detail of head (G). Larva of extant Sternolophus rufipes (Hydrophilini): whole larva in dorsal view (F), detail of head (H). Holotype of Cretoxenus australis sp. nov. NMVP 103312, piece and counterpiece (I, J). Extant hydrobiusine Limnoxenus zealandicus in ventral view, whole beetle (K) and detail of meso- and metaventrite (L). Abbreviations: md, mandible; mtvpr, metaventral process." figureDoi="http://doi.org/10.5281/zenodo.10273966" httpUri="https://zenodo.org/record/10273966/files/figure.png" pageId="8" pageNumber="718">Fig. 6F, H</figureCitation>
|
||
) shows that the fossil resembles the hydrobiusine larvae (mandibles wide with wider retinacular teeth, head widely quadratic, anterior margin of the occipital foramen arcuate, cervical sclerites wide, abdominal segments 1–8 with two pairs of isolated sclerites). At the same time, it clearly differs from the larva of
|
||
<taxonomicName id="4C6B4D39FF88D67EFB66BD214006994C" authorityName="Solier" authorityYear="1834" box="[1206,1356,1088,1109]" class="Insecta" family="Hydrophilidae" genus="Sternolophus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA8FF88D67EFB66BD214006994C" box="[1206,1356,1088,1109]" italics="true" pageId="8" pageNumber="718">Sternolophus</emphasis>
|
||
</taxonomicName>
|
||
, which has long slender mandibles and retinacular teeth, elongate head, triangular anterior margin of the occipital foramen, very narrow cervical sclerites, and mesonotum with both pairs of tergites fused). All these characters are shared by all other genera of the
|
||
<taxonomicName id="4C6B4D39FF88D67EFCEABD9646869814" authorityName="Latreille" authorityYear="1802" box="[826,972,1271,1293]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="tribe" tribe="Hydrophilini">Hydrophilini</taxonomicName>
|
||
except
|
||
<taxonomicName id="4C6B4D39FF88D67EFB93BD9640D79814" authority="Geoffroy, 1762" authorityName="Geoffroy" authorityYear="1762" box="[1091,1437,1271,1293]" class="Liliopsida" family="Restionaceae" genus="Hydrophilus" kingdom="Plantae" order="Poales" pageId="8" pageNumber="718" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B91FEAA8FF88D67EFB93BD9641989815" box="[1091,1234,1271,1292]" italics="true" pageId="8" pageNumber="718">Hydrophilus</emphasis>
|
||
<bibRefCitation id="EFFA4B4BFF88D67EFB39BD9640D79814" author="Geoffroy EL" box="[1257,1437,1271,1293]" pageId="8" pageNumber="718" refId="ref13670" refString="Geoffroy EL. 1762. Histoire abregee des Insectes: qui se trouvent aux environs de Paris: dans laquelle ces animaux sont ranges suivant un ordre methodique. Tome premier. Paris: Durand." type="book" year="1762">Geoffroy, 1762</bibRefCitation>
|
||
</taxonomicName>
|
||
. Comparison of the fossils with modern larvae hence corresponds well with the results of our constrained analysis, and we consider
|
||
<taxonomicName id="4C6B4D39FF88D67EFBA9BC32404B9871" authorityName="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie" authorityYear="2014" box="[1145,1281,1363,1384]" class="Insecta" family="Hydrophilidae" genus="Baissalarva" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA8FF88D67EFBA9BC32404B9871" box="[1145,1281,1363,1384]" italics="true" pageId="8" pageNumber="718">Baissalarva</emphasis>
|
||
</taxonomicName>
|
||
as belonging to the tribe
|
||
<taxonomicName id="4C6B4D39FF88D67EFC01BC1341209891" baseAuthorityName="Mulsant" baseAuthorityYear="1844" box="[977,1130,1394,1416]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="tribe" tribe="Hydrobiusini">Hydrobiusini</taxonomicName>
|
||
. Knowledge of the extant larvae of the
|
||
<taxonomicName id="4C6B4D39FF88D67EFC08BCF1412798BF" baseAuthorityName="Mulsant" baseAuthorityYear="1844" box="[984,1133,1424,1446]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="tribe" tribe="Hydrobiusini">Hydrobiusini</taxonomicName>
|
||
is still rather limited and the fossils lack the generic diagnostic characters (chaetotaxy, detailed morphology of mouthparts and antenna). For this reason we prefer to assign the fossil to a new fossil genus,
|
||
<taxonomicName id="4C6B4D39FF88D67EFB57BF6A40459B39" authority="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie, 2014" authorityName="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie" authorityYear="2014" box="[1159,1295,1547,1568]" class="Insecta" family="Hydrophilidae" genus="Baissalarva" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="718" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA8FF88D67EFB57BF6A40459B39" box="[1159,1295,1547,1568]" italics="true" pageId="8" pageNumber="718">Baissalarva</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22C57D3FF88D67EFAC9BF6A40359B39" box="[1305,1407,1547,1568]" pageId="8" pageNumber="718" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |