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<document id="16FD2251265F7D65816B91059FBFF185" ID-DOI="10.21248/contrib.entomol.57.2.367-396" ID-ISSN="0005-805X" ID-Zenodo-Dep="5364164" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630242723631" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2007" docId="03C28789FFFCFF8D9544FAEE544B18F1" docLanguage="en" docName="ContribEntomol.2007.57.2.367-396.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 57 (2)" docSource="https://www.contributions-to-entomology.org/article/view/1710" docStyle="DocumentStyle:DA3631FDA31ED51D1CF99ECA6F54F23F.3:ContribEntomol.2005-2012.journal_article" docStyleId="DA3631FDA31ED51D1CF99ECA6F54F23F" docStyleName="ContribEntomol.2005-2012.journal_article" docStyleVersion="3" docTitle="Pronomaea khnzoriani SEMENOV" docType="treatment" docVersion="3" lastPageNumber="390" masterDocId="FFFBFFF1FFEAFF9A944DFFD3565D1C3A" masterDocTitle="A revision of the species of Pronomaea E of the Western Palaearctic region, including Middle Asia (Coleoptera: Staphylinidae: Aleocharinae: Pronomaeini)" masterLastPageNumber="396" masterPageNumber="367" pageNumber="389" updateTime="1699196480509" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="FF6584B1557656BA6AE2C3B4EF81EC8A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="077F3A0FE90348719230B6896CB2E515">A revision of the species of Pronomaea E of the Western Palaearctic region, including Middle Asia (Coleoptera: Staphylinidae: Aleocharinae: Pronomaeini)</mods:title>
</mods:titleInfo>
<mods:name id="1C1B98445C26E3AE50BAB73B58133CE8" type="personal">
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<mods:namePart id="FEE29606041DEE984BD878492FC8F816">Assing, Volker</mods:namePart>
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<mods:typeOfResource id="C5E726332AF761381252DDE40DDFC6EF">text</mods:typeOfResource>
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<mods:title id="AA7BD85BD1E155B3F0248A0F33C09A6E">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="B9C106DDF1DEDDB2FCC435ED271A0D30">
<mods:date id="8525743998DFF4DBCAC126061171CA01">2007</mods:date>
<mods:detail id="FDD92AFA5B2B7D55303B6EDC4F294CE1" type="pubDate">
<mods:number id="3D5ECE3691859A69E5F4AB825E6C27DA">2007-12-21</mods:number>
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<mods:number id="622F15F6578935494A91ACFE907EE75A">57</mods:number>
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<mods:url id="A3949DC92C7DD37E33E4EE01377CB2E0">https://www.contributions-to-entomology.org/article/view/1710</mods:url>
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<mods:classification id="F250E0861AC3E94C89D8B9B41A3D12F5">journal article</mods:classification>
<mods:identifier id="2746CD52D64EA9E175DC97C13DF91B2E" type="DOI">10.21248/contrib.entomol.57.2.367-396</mods:identifier>
<mods:identifier id="7AB0DFFAFBD69BE00CDB2476787C983F" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="2066FFE85507AE3D1425F82D2AAA80D3" type="Zenodo-Dep">5364164</mods:identifier>
</mods:mods>
<treatment id="03C28789FFFCFF8D9544FAEE544B18F1" LSID="urn:lsid:plazi:treatment:03C28789FFFCFF8D9544FAEE544B18F1" httpUri="http://treatment.plazi.org/id/03C28789FFFCFF8D9544FAEE544B18F1" lastPageId="23" lastPageNumber="390" pageId="22" pageNumber="389">
<subSubSection id="C3716514FFFCFF8C9544FAEE55FD196F" box="[265,928,1340,1365]" pageId="22" pageNumber="389" type="nomenclature">
<paragraph id="8BD4369FFFFCFF8C9544FAEE55FD196F" blockId="22.[265,928,1340,1365]" box="[265,928,1340,1365]" pageId="22" pageNumber="389">
<emphasis id="B91FEA8DFFFCFF8C9544FAEE549F1968" bold="true" box="[265,706,1341,1362]" pageId="22" pageNumber="389">
<taxonomicName id="4C6B4D1CFFFCFF8C9544FAEE54201975" ID-CoL="789NX" authority="SEMENOV" authorityName="SEMENOV" box="[265,637,1341,1359]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="389" phylum="Arthropoda" rank="species" species="khnzoriani">
<emphasis id="B91FEA8DFFFCFF8C9544FAEE54531975" bold="true" box="[265,526,1341,1359]" italics="true" pageId="22" pageNumber="389">Pronomaea khnzoriani</emphasis>
SEMENOV
</taxonomicName>
,, 2003
</emphasis>
(
<figureCitation id="13502A1AFFFCFF8C969FFAEF551A196E" box="[722,839,1340,1364]" captionStart="Figs 44-51" captionStartId="21.[112,149,1338,1360]" captionTargetBox="[116,1075,184,1323]" captionTargetId="figure-76@21.[111,1092,174,1336]" captionTargetPageId="21" captionText="Figs 44-51: Pronomaea anatolica FAGEL (44-47), P. khnzoriani SEMENOV (48-50), and P. flavirostris SEMENOV (51): median lobe of aedeagus in lateral view (44-46, 48-51) of holotype (44) and of males from Muğla (45-46), Iran (48-49), Amasya (50), and Tajikistan (51); spermatheca (47). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364182" httpUri="https://zenodo.org/record/5364182/files/figure.png" pageId="22" pageNumber="389">Figs 48-50</figureCitation>
,
<figureCitation id="13502A1AFFFCFF8C971EFAEE55C4196F" box="[851,921,1341,1365]" captionStart="Map 4" captionStartId="22.[195,239,844,865]" captionTargetBox="[195,1178,174,836]" captionTargetId="figure-325@22.[195,1178,174,836]" captionTargetPageId="22" captionText="Map 4: Distributions of Pronomaea anatolica FAGEL (filled circles) and P. khnzoriani SEMENOV (open circles), based on examined and literature records." figureDoi="http://doi.org/10.5281/zenodo.5364184" httpUri="https://zenodo.org/record/5364184/files/figure.png" pageId="22" pageNumber="389">Map 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3716514FFFCFF8C94F0FAB854051947" box="[189,600,1387,1405]" pageId="22" pageNumber="389" type="reference_group">
<paragraph id="8BD4369FFFFCFF8C94F0FAB854051947" blockId="22.[189,600,1387,1405]" box="[189,600,1387,1405]" pageId="22" pageNumber="389">
<treatmentCitationGroup id="AB7B11B1FFFCFF8C94F0FAB854051947" box="[189,600,1387,1405]" pageId="22" pageNumber="389">
<treatmentCitation id="0ACA108EFFFCFF8C94F0FAB854081947" author="SEMENOV, V. B." box="[189,597,1387,1405]" page="201" pageId="22" pageNumber="389" year="2003">
<taxonomicName id="4C6B4D1CFFFCFF8C94F0FAB854081947" ID-CoL="789NX" authority="SEMENOV, 2003: 201" authorityName="SEMENOV" authorityPageNumber="201" authorityYear="2003" box="[189,597,1387,1405]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="389" phylum="Arthropoda" rank="species" species="khnzoriani">
<emphasis id="B91FEA8DFFFCFF8C94F0FAB857D51941" box="[189,392,1387,1403]" italics="true" pageId="22" pageNumber="389">Pronomaea khnzoriani</emphasis>
<bibRefCitation id="EFFA4B6EFFFCFF8C95C2FAB854081947" author="SEMENOV, V. B." box="[399,597,1387,1405]" pageId="22" pageNumber="389" pagination="199 - 202" refId="ref17954" refString="SEMENOV, V. B. 2003: Die kaukasischen und mittelasiatischen Arten der Gattung Pronomaea ERICHSON, 1837 (Coleoptera: Staphylinidae: Aleocharinae). - Russian Entomological Journal 12: 199 - 202." type="journal article" year="2003">SEMENOV, 2003: 201</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3716514FFFCFF8C94F3FA4455A81A79" pageId="22" pageNumber="389" type="discussion">
<paragraph id="8BD4369FFFFCFF8C94F3FA4457721993" blockId="22.[190,303,1431,1449]" box="[190,303,1431,1449]" pageId="22" pageNumber="389">
<emphasis id="B91FEA8DFFFCFF8C94F3FA4457721993" bold="true" box="[190,303,1431,1449]" pageId="22" pageNumber="389">Comment:</emphasis>
</paragraph>
<paragraph id="8BD4369FFFFCFF8C94F0FA6F551A1A2B" blockId="22.[189,1179,1468,1554]" pageId="22" pageNumber="389">
<materialsCitation id="3B033CC2FFFCFF8C94F0FA6F551E1A2B" country="Armenia" location="The" pageId="22" pageNumber="389" specimenCount="1">
The original description of this recently described species is based on
<specimenCount id="9D6DFD16FFFCFF8C97DAFA6F55AD19EF" box="[919,1008,1468,1493]" pageId="22" pageNumber="389" type="generic" typeStatus="type">five type</specimenCount>
specimens from
<collectingCountry id="F37C760FFFFCFF8C94F0FA08574419C9" box="[189,281,1499,1523]" name="Armenia" pageId="22" pageNumber="389">Armenia</collectingCountry>
.
<location id="8EB46044FFFCFF8C956CFA08571619C9" LSID="urn:lsid:plazi:treatment:03C28789FFFCFF8D9544FAEE544B18F1:8EB46044FFFCFF8C956CFA08571619C9" box="[289,331,1499,1523]" country="Armenia" name="The" pageId="22" pageNumber="389">The</location>
<typeStatus id="54D0883DFFFCFF8C951FFA0F572319CE" box="[338,382,1500,1524]" pageId="22" pageNumber="389">type</typeStatus>
material was not studied, but the detailed figures of the aedeagus in
<bibRefCitation id="EFFA4B6EFFFCFF8C9077FA0F575E1A28" author="SEMENOV, V. B." pageId="22" pageNumber="389" pagination="199 - 202" refId="ref17954" refString="SEMENOV, V. B. 2003: Die kaukasischen und mittelasiatischen Arten der Gattung Pronomaea ERICHSON, 1837 (Coleoptera: Staphylinidae: Aleocharinae). - Russian Entomological Journal 12: 199 - 202." type="journal article" year="2003">SEMENOV (2003)</bibRefCitation>
leave no doubt that the present interpretation is correct
</materialsCitation>
.
</paragraph>
<paragraph id="8BD4369FFFFCFF8C94F0F9FD55A81A79" blockId="22.[189,1180,1581,1661]" box="[189,1013,1581,1603]" pageId="22" pageNumber="389">
<emphasis id="B91FEA8DFFFCFF8C94F0F9FD57811A04" bold="true" box="[189,476,1582,1598]" pageId="22" pageNumber="389">Additional material examined:</emphasis>
<materialsCitation id="3B033CC2FFFCFF8C95AEF9FE55AC1A79" box="[483,1009,1581,1603]" collectionCode="NHMW" collectorName="Korb" country="Turkey" location="Amasya" pageId="22" pageNumber="389" specimenCount="2" stateProvince="Amasya">
<emphasis id="B91FEA8DFFFCFF8C95AEF9FE54771A78" bold="true" box="[483,554,1581,1602]" pageId="22" pageNumber="389">
<collectingCountry id="F37C760FFFFCFF8C95AEF9FE547B1A78" box="[483,550,1581,1602]" name="Turkey" pageId="22" pageNumber="389">Turkey</collectingCountry>
:
</emphasis>
<specimenCount id="9D6DFD16FFFCFF8C967CF9FE54351A79" box="[561,616,1581,1603]" pageId="22" pageNumber="389" type="generic">2 exs.</specimenCount>
,
<collectingRegion id="49AFF87DFFFCFF8C963CF9FE54E61A78" box="[625,699,1581,1602]" country="Turkey" name="Amasya" pageId="22" pageNumber="389">Amasya</collectingRegion>
, 1888, leg.
<collectorName id="269E5349FFFCFF8C976AF9FE550B1A79" box="[807,854,1581,1603]" pageId="22" pageNumber="389">Korb</collectorName>
(
<collectionCode id="ED7AAE5AFFFCFF8C972EF9FE55EA1A78" box="[867,951,1581,1602]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="389" type="Museum">NHMW</collectionCode>
, cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3716514FFFCFF8D94F3F99956931DD9" lastPageId="23" lastPageNumber="390" pageId="22" pageNumber="389" type="materials_examined">
<paragraph id="8BD4369FFFFCFF8C94F3F99952C11A46" blockId="22.[189,1180,1581,1661]" pageId="22" pageNumber="389">
<materialsCitation id="3B033CC2FFFCFF8C94F3F99955011A5A" box="[190,860,1610,1632]" collectingDate="1989-06-26" collectorName="Putz" country="Georgia" location="Uplichziche" municipality="Gori" pageId="22" pageNumber="389" specimenCount="1">
<emphasis id="B91FEA8DFFFCFF8C94F3F99957501A5A" bold="true" box="[190,269,1610,1632]" pageId="22" pageNumber="389">
<collectingCountry id="F37C760FFFFCFF8C94F3F99957541A5A" box="[190,265,1610,1632]" name="Georgia" pageId="22" pageNumber="389">Georgia</collectingCountry>
:
</emphasis>
<specimenCount id="9D6DFD16FFFCFF8C9555F999571A1A5A" box="[280,327,1610,1632]" pageId="22" pageNumber="389" type="generic">1 ex.</specimenCount>
,
<collectingMunicipality id="6BB0ACE5FFFCFF8C951EF99957221A5A" box="[339,383,1610,1632]" pageId="22" pageNumber="389">Gori</collectingMunicipality>
env.,
<location id="8EB46044FFFCFF8C95F0F99954731A5A" LSID="urn:lsid:plazi:treatment:03C28789FFFCFF8D9544FAEE544B18F1:8EB46044FFFCFF8C95F0F99954731A5A" box="[445,558,1610,1632]" country="Georgia" municipality="Gori" name="Uplichziche" pageId="22" pageNumber="389">Uplichziche</location>
,
<date id="FFD5105FFFFCFF8C9677F99954F41A5A" box="[570,681,1610,1632]" pageId="22" pageNumber="389" value="1989-06-26">
<collectingDate id="EF91E9B7FFFCFF8C9677F99954F41A5A" box="[570,681,1610,1632]" pageId="22" pageNumber="389" value="1989-06-26">26.VI.1989</collectingDate>
</date>
, leg.
<collectorName id="269E5349FFFCFF8C96ADF99855541A5A" box="[736,777,1611,1632]" pageId="22" pageNumber="389">Pütz</collectorName>
(cWun)
</materialsCitation>
;
<materialsCitation id="3B033CC2FFFCFF8C9724F99955891A47" collectingDate="1985-07-19" collectionCode="DEI" collectorName="Wrase" country="Georgia" latitude="41.733334" location="Tbilisskoye" longLatPrecision="1154" longitude="44.833332" pageId="22" pageNumber="389" specimenCount="10" stateProvince="Tbilisi">
<specimenCount id="9D6DFD16FFFCFF8C9724F99955F21A5A" box="[873,943,1610,1632]" pageId="22" pageNumber="389" type="generic">10 exs.</specimenCount>
,
<collectingRegion id="49AFF87DFFFCFF8C97F7F99955AB1A5A" box="[954,1014,1610,1632]" country="Georgia" name="Tbilisi" pageId="22" pageNumber="389">Tbilisi</collectingRegion>
env.,
<location id="8EB46044FFFCFF8C907EF99952C61A5A" LSID="urn:lsid:plazi:treatment:03C28789FFFCFF8D9544FAEE544B18F1:8EB46044FFFCFF8C907EF99952C61A5A" box="[1075,1179,1610,1632]" country="Georgia" latitude="41.733334" longLatPrecision="1154" longitude="44.833332" name="Tbilisskoye" pageId="22" pageNumber="389" stateProvince="Tbilisi">Tbilisskoye</location>
ozero [&quot; Tbiliskoje osero &quot;;
<geoCoordinate id="EE5F5058FFFCFF8C95FAF9B454551A40" box="[439,520,1639,1658]" degrees="41" direction="north" minutes="44" orientation="latitude" pageId="22" pageNumber="389" precision="925" value="41.733334">41°44 'N</geoCoordinate>
,
<geoCoordinate id="EE5F5058FFFCFF8C9658F9B4543E1A47" box="[533,611,1639,1661]" degrees="44" direction="east" minutes="50" orientation="longitude" pageId="22" pageNumber="389" precision="925" value="44.833332">44°50 'E</geoCoordinate>
],
<date id="FFD5105FFFFCFF8C963BF9B454B11A47" box="[630,748,1639,1661]" pageId="22" pageNumber="389" value="1985-07-19">
<collectingDate id="EF91E9B7FFFCFF8C963BF9B454B11A47" box="[630,748,1639,1661]" pageId="22" pageNumber="389" value="1985-07-19">19.VII.1985</collectingDate>
</date>
, leg.
<collectorName id="269E5349FFFCFF8C976CF9B455041A46" box="[801,857,1639,1660]" pageId="22" pageNumber="389">Wrase</collectorName>
(
<collectionCode id="ED7AAE5AFFFCFF8C9724F9B455C81A46" box="[873,917,1639,1660]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="22" pageNumber="389" type="Other type of research institution/biorepository">DEI</collectionCode>
, cAss)
</materialsCitation>
;
<specimenCount id="9D6DFD16FFFCFF8C97ADF9B4524A1A46" box="[992,1047,1639,1660]" pageId="22" pageNumber="389" type="generic">3 exs.</specimenCount>
, W-Caucasus,
</paragraph>
<paragraph id="8BD4369FFFFDFF8D9421FF7C55841CFF" blockId="23.[106,1097,175,483]" box="[108,985,175,197]" pageId="23" pageNumber="390">
&quot;
<materialsCitation id="3B033CC2FFFDFF8D9439FF7C55881CFF" box="[116,981,175,197]" collectionCode="NHMW" collectorName="Leder" pageId="23" pageNumber="390" specimenCount="1">
Landschaft Letschgum &quot;, leg.
<collectorName id="269E5349FFFDFF8D95CCFF7C57E81CFF" box="[385,437,175,197]" pageId="23" pageNumber="390">Leder</collectorName>
(
<collectionCode id="ED7AAE5AFFFDFF8D958FFF7C544B1CFE" box="[450,534,175,196]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="390" type="Museum">NHMW</collectionCode>
, cAss);
<specimenCount id="9D6DFD16FFFDFF8D9616FF7C54D71CFE" box="[603,650,175,196]" pageId="23" pageNumber="390" type="generic">1 ex.</specimenCount>
, &quot; Swanetien &quot;, leg.
<collectorName id="269E5349FFFDFF8D977BFF7C55371CFF" box="[822,874,175,197]" pageId="23" pageNumber="390">Leder</collectorName>
(
<collectionCode id="ED7AAE5AFFFDFF8D9735FF7C55931CFF" box="[888,974,175,197]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="390" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD4369FFFFDFF8D9426FF1F54A51CD8" blockId="23.[106,1097,175,483]" box="[107,760,204,226]" pageId="23" pageNumber="390">
<emphasis id="B91FEA8DFFFDFF8D9426FF1F569A1CDB" bold="true" box="[107,199,204,225]" pageId="23" pageNumber="390">Dagestan:</emphasis>
<materialsCitation id="3B033CC2FFFDFF8D9482FF1F54A81CD8" box="[207,757,204,226]" collectionCode="NHMW" collectorName="Leder &amp; Reitter" pageId="23" pageNumber="390" specimenCount="1">
<specimenCount id="9D6DFD16FFFDFF8D9482FF1F56A01CDB" box="[207,253,204,225]" pageId="23" pageNumber="390" type="generic">1 ex.</specimenCount>
, locality not specified, leg.
<collectorName id="269E5349FFFDFF8D95BAFF1F54761CD8" box="[503,555,204,226]" pageId="23" pageNumber="390">Leder</collectorName>
&amp;
<collectorName id="269E5349FFFDFF8D9606FF1F54D71CD8" box="[587,650,204,226]" pageId="23" pageNumber="390">Reitter</collectorName>
(
<collectionCode id="ED7AAE5AFFFDFF8D96DAFF1F54B01CD8" box="[663,749,204,226]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="390" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD4369FFFFDFF8D9427FF3A57F01D0D" blockId="23.[106,1097,175,483]" pageId="23" pageNumber="390">
<materialsCitation id="3B033CC2FFFDFF8D9427FF3A56C11D26" collectingDate="1975-09-18" collectorName="Hieke" country="Turkmenistan" location="Kopet Dag" municipality="Firyuza" pageId="23" pageNumber="390" specimenCount="3">
<emphasis id="B91FEA8DFFFDFF8D9427FF3A56AE1CC3" bold="true" box="[106,243,233,249]" pageId="23" pageNumber="390">
<collectingCountry id="F37C760FFFFDFF8D9427FF3A56B21CC3" box="[106,239,233,249]" name="Turkmenistan" pageId="23" pageNumber="390">Turkmenistan</collectingCountry>
:
</emphasis>
<specimenCount id="9D6DFD16FFFDFF8D94B0FF3A57691CC4" box="[253,308,233,254]" pageId="23" pageNumber="390" type="generic">3 exs.</specimenCount>
,
<location id="8EB46044FFFDFF8D9572FF3A57F51CC4" LSID="urn:lsid:plazi:treatment:03C28789FFFCFF8D9544FAEE544B18F1:8EB46044FFFDFF8D9572FF3A57F51CC4" box="[319,424,233,254]" country="Turkmenistan" municipality="Firyuza" name="Kopet Dag" pageId="23" pageNumber="390">Kopet Dag</location>
, stream
<locationDeviation id="9A662FBDFFFDFF8D95B7FF3B54FD1CC4" box="[506,672,232,254]" location="Tschuli" pageId="23" pageNumber="390">
<quantity id="4C939B7AFFFDFF8D95B7FF3B54711CC4" box="[506,556,232,254]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="23" pageNumber="390" unit="km" value="6.0">6 km</quantity>
NE Tschuli
</locationDeviation>
near
<collectingMunicipality id="6BB0ACE5FFFDFF8D9695FF3B55421CC4" box="[728,799,232,254]" pageId="23" pageNumber="390">Firyuza</collectingMunicipality>
,
<date id="FFD5105FFFFDFF8D9766FF3B55C51CC4" box="[811,920,232,254]" pageId="23" pageNumber="390" value="1975-09-18">
<collectingDate id="EF91E9B7FFFDFF8D9766FF3B55C51CC4" box="[811,920,232,254]" pageId="23" pageNumber="390" value="1975-09-18">18.IX.1975</collectingDate>
</date>
, leg.
<collectorName id="269E5349FFFDFF8D9780FF3B52581CC4" box="[973,1029,232,254]" pageId="23" pageNumber="390">Hieke</collectorName>
(cVog, cAss)
</materialsCitation>
;
<materialsCitation id="3B033CC2FFFDFF8D94E5FED6525E1D20" box="[168,1027,261,283]" collectionCode="NHMW" collectorName="W Ashkhabad &amp; Sulyukly" country="Turkmenistan" latitude="38.033333" longLatPrecision="1178" longitude="57.416668" pageId="23" pageNumber="390" specimenCount="4">
<specimenCount id="9D6DFD16FFFDFF8D94E5FED656821D21" box="[168,223,261,283]" pageId="23" pageNumber="390" type="generic">4 exs.</specimenCount>
,
<collectorName id="269E5349FFFDFF8D94A4FED5572F1D21" box="[233,370,261,283]" pageId="23" pageNumber="390">W Ashkhabad</collectorName>
,
<collectorName id="269E5349FFFDFF8D9530FED657901D21" box="[381,461,261,283]" pageId="23" pageNumber="390">Sulyukly</collectorName>
[&quot; Transcaspia, Neu-Saratow &quot;,
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,
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] (
<collectionCode id="ED7AAE5AFFFDFF8D97E9FED555A61D20" box="[932,1019,262,282]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="390" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CC2FFFDFF8D9043FED557F41D0D" collectionCode="NHMW" collectorName="Lotfabad" country="Turkmenistan" pageId="23" pageNumber="390" specimenCount="2">
<specimenCount id="9D6DFD16FFFDFF8D9043FED552181D20" box="[1038,1093,262,282]" pageId="23" pageNumber="390" type="generic">2 exs.</specimenCount>
,
<collectorName id="269E5349FFFDFF8D9426FEF156E01D02" box="[107,189,290,312]" pageId="23" pageNumber="390">Loţfābād</collectorName>
[&quot; Ljutfabad &quot;] (
<collectionCode id="ED7AAE5AFFFDFF8D9501FEF057FF1D0D" box="[332,418,291,311]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="390" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD4369FFFFDFF8D9426FEEC56931DD9" blockId="23.[106,1097,175,483]" pageId="23" pageNumber="390">
<materialsCitation id="3B033CC2FFFDFF8D9426FEEC55EA1D6F" box="[107,951,319,341]" collectingDate="1973-07-22" collectionCode="MHNG" collectorName="Senglet" country="Iran" latitude="36.566666" location="Sari" longLatPrecision="1186" longitude="53.15" pageId="23" pageNumber="390" specimenCount="2" stateProvince="Mazandaran">
<emphasis id="B91FEA8DFFFDFF8D9426FEEC56CB1D75" bold="true" box="[107,150,319,335]" pageId="23" pageNumber="390">
<collectingCountry id="F37C760FFFFDFF8D9426FEEC56CF1D75" box="[107,146,319,335]" name="Iran" pageId="23" pageNumber="390">Iran</collectingCountry>
:
</emphasis>
<specimenCount id="9D6DFD16FFFDFF8D94EFFEEC56841D68" box="[162,217,319,338]" pageId="23" pageNumber="390" type="generic">2 exs.</specimenCount>
,
<collectingRegion id="49AFF87DFFFDFF8D94ABFEEC57061D68" box="[230,347,319,338]" country="Iran" name="Mazandaran" pageId="23" pageNumber="390">Mazanderan</collectingRegion>
,
<location id="8EB46044FFFDFF8D9527FEEC57CD1D68" LSID="urn:lsid:plazi:treatment:03C28789FFFCFF8D9544FAEE544B18F1:8EB46044FFFDFF8D9527FEEC57CD1D68" box="[362,400,319,338]" country="Iran" latitude="36.566666" longLatPrecision="1186" longitude="53.15" name="Sari" pageId="23" pageNumber="390" stateProvince="Mazandaran">Sari</location>
,
<geoCoordinate id="EE5F5058FFFDFF8D95D3FEEC57B21D68" box="[414,495,319,338]" degrees="36" direction="north" minutes="34" orientation="latitude" pageId="23" pageNumber="390" precision="925" value="36.566666">36°34 'N</geoCoordinate>
,
<geoCoordinate id="EE5F5058FFFDFF8D95B3FEEC54161D6E" box="[510,587,319,340]" degrees="53" direction="east" minutes="09" orientation="longitude" pageId="23" pageNumber="390" precision="925" value="53.15">53°09 'E</geoCoordinate>
,
<date id="FFD5105FFFFDFF8D9614FEEC548D1D6E" box="[601,720,319,340]" pageId="23" pageNumber="390" value="1973-07-22">
<collectingDate id="EF91E9B7FFFDFF8D9614FEEC548D1D6E" box="[601,720,319,340]" pageId="23" pageNumber="390" value="1973-07-22">22.VII.1973</collectingDate>
</date>
, leg.
<collectorName id="269E5349FFFDFF8D9747FEEC55101D6F" box="[778,845,319,341]" pageId="23" pageNumber="390">Senglet</collectorName>
(
<collectionCode id="ED7AAE5AFFFDFF8D9713FEEC55ED1D6F" box="[862,944,319,341]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="23" pageNumber="390" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CC2FFFDFF8D9788FEEC55641D48" collectingDate="1974-08-08" collectionCode="MHNG" collectorName="Senglet" country="Iran" latitude="36.466667" location="Tarazan-Lowshan" longLatPrecision="1187" longitude="49.516666" pageId="23" pageNumber="390" specimenCount="1" stateProvince="Tehran">
<specimenCount id="9D6DFD16FFFDFF8D9788FEEC55A91D6E" box="[965,1012,319,340]" pageId="23" pageNumber="390" type="generic">1 ex.</specimenCount>
,
<collectingRegion id="49AFF87DFFFDFF8D904CFEEC52181D6F" box="[1025,1093,319,341]" country="Iran" name="Tehran" pageId="23" pageNumber="390">Tehran</collectingRegion>
,
<location id="8EB46044FFFDFF8D9427FE8F57521D55" LSID="urn:lsid:plazi:treatment:03C28789FFFCFF8D9544FAEE544B18F1:8EB46044FFFDFF8D9427FE8F57521D55" box="[106,271,348,367]" country="Iran" latitude="36.466667" longLatPrecision="1187" longitude="49.516666" name="Tarazan-Lowshan" pageId="23" pageNumber="390" stateProvince="Tehran">Tarazan-Lowshan</location>
,
<geoCoordinate id="EE5F5058FFFDFF8D9553FE8F57321D55" box="[286,367,348,367]" degrees="36" direction="north" minutes="28" orientation="latitude" pageId="23" pageNumber="390" precision="925" value="36.466667">36°28 'N</geoCoordinate>
,
<geoCoordinate id="EE5F5058FFFDFF8D95CDFE8F57901D4B" box="[384,461,348,369]" degrees="49" direction="east" minutes="31" orientation="longitude" pageId="23" pageNumber="390" precision="925" value="49.516666">49°31 'E</geoCoordinate>
,
<date id="FFD5105FFFFDFF8D9591FE8F54121D4B" box="[476,591,348,369]" pageId="23" pageNumber="390" value="1974-08-08">
<collectingDate id="EF91E9B7FFFDFF8D9591FE8F54121D4B" box="[476,591,348,369]" pageId="23" pageNumber="390" value="1974-08-08">8.VIII.1974</collectingDate>
</date>
, leg.
<collectorName id="269E5349FFFDFF8D96C6FE8F54931D48" box="[651,718,348,370]" pageId="23" pageNumber="390">Senglet</collectorName>
(
<collectionCode id="ED7AAE5AFFFDFF8D96ACFE8F556E1D48" box="[737,819,348,370]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="23" pageNumber="390" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CC2FFFDFF8D9704FE8F54381DB5" collectingDate="1974-07-06" collectionCode="MHNG" collectorName="Senglet" country="Iran" latitude="36.316666" location="Agha Baba" longLatPrecision="1188" longitude="49.816666" pageId="23" pageNumber="390" specimenCount="1" stateProvince="Tehran">
<specimenCount id="9D6DFD16FFFDFF8D9704FE8F55251D4B" box="[841,888,348,369]" pageId="23" pageNumber="390" type="generic">1 ex.</specimenCount>
,
<collectingRegion id="49AFF87DFFFDFF8D97CBFE8F55971D48" box="[902,970,348,370]" country="Iran" name="Tehran" pageId="23" pageNumber="390">Tehran</collectingRegion>
,
<location id="8EB46044FFFDFF8D9794FE8F52191D48" LSID="urn:lsid:plazi:treatment:03C28789FFFCFF8D9544FAEE544B18F1:8EB46044FFFDFF8D9794FE8F52191D48" box="[985,1092,348,370]" country="Iran" latitude="36.316666" longLatPrecision="1188" longitude="49.816666" name="Agha Baba" pageId="23" pageNumber="390" stateProvince="Tehran">Agha Baba</location>
,
<geoCoordinate id="EE5F5058FFFDFF8D9426FEAA56E61DB6" box="[107,187,377,396]" degrees="36" direction="north" minutes="19" orientation="latitude" pageId="23" pageNumber="390" precision="925" value="36.316666">36°19 'N</geoCoordinate>
,
<geoCoordinate id="EE5F5058FFFDFF8D948BFEAA57491DB4" box="[198,276,377,398]" degrees="49" direction="east" minutes="49" orientation="longitude" pageId="23" pageNumber="390" precision="925" value="49.816666">49°49 'E</geoCoordinate>
,
<date id="FFD5105FFFFDFF8D9553FEAB57D41DB4" box="[286,393,376,398]" pageId="23" pageNumber="390" value="1974-07-06">
<collectingDate id="EF91E9B7FFFDFF8D9553FEAB57D41DB4" box="[286,393,376,398]" pageId="23" pageNumber="390" value="1974-07-06">6.VII.1974</collectingDate>
</date>
, leg.
<collectorName id="269E5349FFFDFF8D95F6FEAB57A31DB4" box="[443,510,376,398]" pageId="23" pageNumber="390">Senglet</collectorName>
(
<collectionCode id="ED7AAE5AFFFDFF8D9641FEAA54031DB5" box="[524,606,377,399]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="23" pageNumber="390" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CC2FFFDFF8D9622FEAA57F71D91" collectingDate="1975-06-22" collectionCode="MHNG" collectorName="Senglet" country="Iran" latitude="35.466667" location="Sanandajd" longLatPrecision="1193" longitude="47.016666" pageId="23" pageNumber="390" specimenCount="1" stateProvince="Kordestan">
<specimenCount id="9D6DFD16FFFDFF8D9622FEAA54C31DB4" box="[623,670,377,398]" pageId="23" pageNumber="390" type="generic">1 ex.</specimenCount>
,
<collectingRegion id="49AFF87DFFFDFF8D96E5FEAB55541DB4" box="[680,777,376,398]" country="Iran" name="Kordestan" pageId="23" pageNumber="390">Kordestan</collectingRegion>
, N
<location id="8EB46044FFFDFF8D9760FEAB55CD1DB4" LSID="urn:lsid:plazi:treatment:03C28789FFFCFF8D9544FAEE544B18F1:8EB46044FFFDFF8D9760FEAB55CD1DB4" box="[813,912,376,398]" country="Iran" latitude="35.466667" longLatPrecision="1193" longitude="47.016666" name="Sanandajd" pageId="23" pageNumber="390" stateProvince="Kordestan">Sanandajd</location>
,
<geoCoordinate id="EE5F5058FFFDFF8D97D6FEAB55B11DB6" box="[923,1004,376,398]" degrees="35" direction="north" minutes="28" orientation="latitude" pageId="23" pageNumber="390" precision="925" value="35.466667">35°28 'N</geoCoordinate>
,
<geoCoordinate id="EE5F5058FFFDFF8D97BAFEAA52181DB6" box="[1015,1093,377,396]" degrees="47" direction="east" minutes="01" orientation="longitude" pageId="23" pageNumber="390" precision="925" value="47.016666">47°01 'E</geoCoordinate>
,
<date id="FFD5105FFFFDFF8D9427FE4656851D91" box="[106,216,405,427]" pageId="23" pageNumber="390" value="1975-06-22">
<collectingDate id="EF91E9B7FFFDFF8D9427FE4656851D91" box="[106,216,405,427]" pageId="23" pageNumber="390" value="1975-06-22">22.VI.1975</collectingDate>
</date>
, leg.
<collectorName id="269E5349FFFDFF8D954EFE46571B1D91" box="[259,326,405,427]" pageId="23" pageNumber="390">Senglet</collectorName>
(
<collectionCode id="ED7AAE5AFFFDFF8D951CFE4657FE1D91" box="[337,419,405,427]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="23" pageNumber="390" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CC2FFFDFF8D95FCFE4657661DF2" collectingDate="1975-07-29" collectionCode="MHNG" collectorName="Senglet" country="Iran" elevation="1200" latitude="37.2" location="Esfayaren" longLatPrecision="1183" longitude="57.45" pageId="23" pageNumber="390" specimenCount="1" stateProvince="Khorasan-e Shomali">
<specimenCount id="9D6DFD16FFFDFF8D95FCFE4657BD1D91" box="[433,480,405,427]" pageId="23" pageNumber="390" type="generic">1 ex.</specimenCount>
,
<collectingRegion id="49AFF87DFFFDFF8D95ABFE46541C1D91" box="[486,577,405,427]" country="Iran" name="Khorasan-e Shomali" pageId="23" pageNumber="390">Khorasan</collectingRegion>
, NE
<location id="8EB46044FFFDFF8D9621FE46549A1D91" LSID="urn:lsid:plazi:treatment:03C28789FFFCFF8D9544FAEE544B18F1:8EB46044FFFDFF8D9621FE46549A1D91" box="[620,711,405,427]" country="Iran" latitude="37.2" longLatPrecision="1183" longitude="57.45" name="Esfayaren" pageId="23" pageNumber="390" stateProvince="Khorasan-e Shomali">Esfayaren</location>
,
<geoCoordinate id="EE5F5058FFFDFF8D9683FE4655421D93" box="[718,799,405,426]" degrees="37" direction="north" minutes="12" orientation="latitude" pageId="23" pageNumber="390" precision="925" value="37.2">37°12 'N</geoCoordinate>
,
<geoCoordinate id="EE5F5058FFFDFF8D976AFE4555291D93" box="[807,884,406,425]" degrees="57" direction="east" minutes="27" orientation="longitude" pageId="23" pageNumber="390" precision="925" value="57.45">57°27 'E</geoCoordinate>
,
<quantity id="4C939B7AFFFDFF8D9736FE45559B1D93" box="[891,966,406,425]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="23" pageNumber="390" unit="m" value="1200.0">
<elevation id="0046D1ACFFFDFF8D9736FE45559B1D93" box="[891,966,406,425]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="23" pageNumber="390" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<date id="FFD5105FFFFDFF8D9783FE4552191D93" box="[974,1092,406,425]" pageId="23" pageNumber="390" value="1975-07-29">
<collectingDate id="EF91E9B7FFFDFF8D9783FE4552191D93" box="[974,1092,406,425]" pageId="23" pageNumber="390" value="1975-07-29">29.VII.1975</collectingDate>
</date>
, leg.
<collectorName id="269E5349FFFDFF8D94DCFE6156891DF2" box="[145,212,434,456]" pageId="23" pageNumber="390">Senglet</collectorName>
(
<collectionCode id="ED7AAE5AFFFDFF8D94AFFE6157691DF2" box="[226,308,434,456]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="23" pageNumber="390" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CC2FFFDFF8D950BFE6156971DD9" collectingDate="1973-07-04" collectionCode="MHNG" collectorName="Senglet" country="Iran" latitude="37.516666" location="Bandar Pahlevi" longLatPrecision="1181" longitude="49.316666" municipality="Galugah" pageId="23" pageNumber="390" specimenCount="1" stateProvince="Gilan">
<specimenCount id="9D6DFD16FFFDFF8D950BFE6157291DF2" box="[326,372,434,456]" pageId="23" pageNumber="390" type="generic">1 ex.</specimenCount>
,
<collectingRegion id="49AFF87DFFFDFF8D9532FE6157E81DF2" box="[383,437,434,456]" country="Iran" name="Gilan" pageId="23" pageNumber="390">Gilan</collectingRegion>
,
<collectingMunicipality id="6BB0ACE5FFFDFF8D958DFE61544C1DF2" box="[448,529,434,456]" pageId="23" pageNumber="390">Galugah</collectingMunicipality>
,
<location id="8EB46044FFFDFF8D9656FE6154F71DF2" LSID="urn:lsid:plazi:treatment:03C28789FFFCFF8D9544FAEE544B18F1:8EB46044FFFDFF8D9656FE6154F71DF2" box="[539,682,434,456]" country="Iran" latitude="37.516666" longLatPrecision="1181" longitude="49.316666" municipality="Galugah" name="Bandar Pahlevi" pageId="23" pageNumber="390" stateProvince="Gilan">Bandar Pahlevi</location>
,
<geoCoordinate id="EE5F5058FFFDFF8D96F9FE6155581DFC" box="[692,773,434,455]" degrees="37" direction="north" minutes="31" orientation="latitude" pageId="23" pageNumber="390" precision="925" value="37.516666">37°31 'N</geoCoordinate>
,
<geoCoordinate id="EE5F5058FFFDFF8D975DFE6055031DF2" box="[784,862,435,456]" degrees="49" direction="east" minutes="19" orientation="longitude" pageId="23" pageNumber="390" precision="925" value="49.316666">49°19 'E</geoCoordinate>
,
<date id="FFD5105FFFFDFF8D9725FE61558E1DFD" box="[872,979,434,455]" pageId="23" pageNumber="390" value="1973-07-04">
<collectingDate id="EF91E9B7FFFDFF8D9725FE61558E1DFD" box="[872,979,434,455]" pageId="23" pageNumber="390" value="1973-07-04">4.VII.1973</collectingDate>
</date>
, leg.
<collectorName id="269E5349FFFDFF8D9048FE6152151DF2" box="[1029,1096,434,456]" pageId="23" pageNumber="390">Senglet</collectorName>
(
<collectionCode id="ED7AAE5AFFFDFF8D943EFE1C56991DD9" box="[115,196,463,483]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="23" pageNumber="390" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3716514FFFDFF8D9426FE28572E1F82" pageId="23" pageNumber="390" type="diagnosis">
<paragraph id="8BD4369FFFFDFF8D9426FE2856841E29" blockId="23.[107,217,507,531]" box="[107,217,507,531]" pageId="23" pageNumber="390">
<emphasis id="B91FEA8DFFFDFF8D9426FE2856841E29" bold="true" box="[107,217,507,531]" pageId="23" pageNumber="390">Diagnosis:</emphasis>
</paragraph>
<paragraph id="8BD4369FFFFDFF8D9426FDF255AC1E03" blockId="23.[107,1009,545,569]" box="[107,1009,545,569]" pageId="23" pageNumber="390">
Extremely similar to
<taxonomicName id="4C6B4D1CFFFDFF8D950CFDF157EF1E0E" authorityName="Fagel" authorityYear="1969" box="[321,434,546,564]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="390" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B91FEA8DFFFDFF8D950CFDF157EF1E0E" box="[321,434,546,564]" italics="true" pageId="23" pageNumber="390">P. anatolica</emphasis>
</taxonomicName>
; distinguished only by the morphology of the aedeagus.
</paragraph>
<paragraph id="8BD4369FFFFDFF8D9423FD9954701E44" blockId="23.[107,1096,583,638]" pageId="23" pageNumber="390">
<specimenCount id="9D6DFD16FFFDFF8D9423FD9956261E60" box="[110,123,586,602]" pageId="23" pageNumber="390" type="male">♂</specimenCount>
: aedeagus of similar morphology as that of
<taxonomicName id="4C6B4D1CFFFDFF8D967AFD9B54F41E60" authorityName="Fagel" authorityYear="1969" box="[567,681,584,602]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="390" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B91FEA8DFFFDFF8D967AFD9B54F41E60" box="[567,681,584,602]" italics="true" pageId="23" pageNumber="390">P. anatolica</emphasis>
</taxonomicName>
, but apical internal structure of different shape, in lateral view very thin (
<figureCitation id="13502A1AFFFDFF8D95FDFDB5547F1E44" box="[432,546,614,638]" captionStart="Figs 44-51" captionStartId="21.[112,149,1338,1360]" captionTargetBox="[116,1075,184,1323]" captionTargetId="figure-76@21.[111,1092,174,1336]" captionTargetPageId="21" captionText="Figs 44-51: Pronomaea anatolica FAGEL (44-47), P. khnzoriani SEMENOV (48-50), and P. flavirostris SEMENOV (51): median lobe of aedeagus in lateral view (44-46, 48-51) of holotype (44) and of males from Muğla (45-46), Iran (48-49), Amasya (50), and Tajikistan (51); spermatheca (47). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364182" httpUri="https://zenodo.org/record/5364182/files/figure.png" pageId="23" pageNumber="390">Figs 48-50</figureCitation>
).
</paragraph>
<paragraph id="8BD4369FFFFDFF8D9422FD5E56EB1EFB" blockId="23.[108,1096,652,705]" pageId="23" pageNumber="390">
<specimenCount id="9D6DFD16FFFDFF8D9422FD5E56241EA5" box="[111,121,653,671]" pageId="23" pageNumber="390" type="female">♀</specimenCount>
: spermatheca shaped like that of
<taxonomicName id="4C6B4D1CFFFDFF8D95A1FD5E54001EA5" authorityName="Fagel" authorityYear="1969" box="[492,605,653,671]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="390" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B91FEA8DFFFDFF8D95A1FD5E54001EA5" box="[492,605,653,671]" italics="true" pageId="23" pageNumber="390">P. anatolica</emphasis>
</taxonomicName>
; for an illustration see figure
<quantity id="4C939B7AFFFDFF8D97EFFD5F55811E9E" box="[930,988,652,676]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="23" pageNumber="390" unit="in" value="13.0">13 in</quantity>
<bibRefCitation id="EFFA4B6EFFFDFF8D97AAFD5E56EC1EFB" author="SEMENOV, V. B." pageId="23" pageNumber="390" pagination="199 - 202" refId="ref17954" refString="SEMENOV, V. B. 2003: Die kaukasischen und mittelasiatischen Arten der Gattung Pronomaea ERICHSON, 1837 (Coleoptera: Staphylinidae: Aleocharinae). - Russian Entomological Journal 12: 199 - 202." type="journal article" year="2003">SEMENOV (2003)</bibRefCitation>
.
</paragraph>
<paragraph id="8BD4369FFFFDFF8D9426FD3357651EC2" blockId="23.[107,312,736,760]" box="[107,312,736,760]" pageId="23" pageNumber="390">
<emphasis id="B91FEA8DFFFDFF8D9426FD3357651EC2" bold="true" box="[107,312,736,760]" pageId="23" pageNumber="390">Comparative notes:</emphasis>
</paragraph>
<paragraph id="8BD4369FFFFDFF8D9426FCD4572E1F82" blockId="23.[107,1096,774,952]" pageId="23" pageNumber="390">
From the highly similar
<taxonomicName id="4C6B4D1CFFFDFF8D9520FCD457831F23" box="[365,478,775,793]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="390" phylum="Arthropoda" rank="species" species="anatolicaanatolica">
<emphasis id="B91FEA8DFFFDFF8D9520FCD457831F23" box="[365,478,775,793]" italics="true" pageId="23" pageNumber="390">P. anatolicaanatolica</emphasis>
</taxonomicName>
- doubtlessly its sister species, as can be inferred from the similarly derived morphology of the male primary sexual characters -, this species is reliably separated only by the shape of the apical internal structure of the aedeagus. From all other congeners it is at once distinguished also by the conspicuous shape of the median lobe of the aedeagus, and from all the species, except
<taxonomicName id="4C6B4D1CFFFDFF8D9652FC5154CD1FAE" authorityName="Fagel" authorityYear="1969" box="[543,656,898,916]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="390" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B91FEA8DFFFDFF8D9652FC5154CD1FAE" box="[543,656,898,916]" italics="true" pageId="23" pageNumber="390">P. anatolica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6B4D1CFFFDFF8D96D2FC5155411FA3" box="[671,796,897,921]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="390" phylum="Arthropoda" rank="species" species="flavirostris">
<emphasis id="B91FEA8DFFFDFF8D96D2FC5155411FA3" box="[671,796,897,921]" italics="true" pageId="23" pageNumber="390">P. flavirostris</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C6B4D1CFFFDFF8D9715FC5155971FA3" box="[856,970,897,921]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="390" phylum="Arthropoda" rank="species" species="procerula">
<emphasis id="B91FEA8DFFFDFF8D9715FC5155971FA3" box="[856,970,897,921]" italics="true" pageId="23" pageNumber="390">P. procerula</emphasis>
</taxonomicName>
, also by the shape of the spermatheca.
</paragraph>
</subSubSection>
<subSubSection id="C3716514FFFDFF8D9426FC05544B18F1" pageId="23" pageNumber="390" type="distribution">
<paragraph id="8BD4369FFFFDFF8D9426FC0557CB1FD2" blockId="23.[107,406,982,1000]" box="[107,406,982,1000]" pageId="23" pageNumber="390">
<emphasis id="B91FEA8DFFFDFF8D9426FC0557CB1FD2" bold="true" box="[107,406,982,1000]" pageId="23" pageNumber="390">Distribution and bionomics:</emphasis>
</paragraph>
<paragraph id="8BD4369FFFFDFF8D9427FC2F544B18F1" blockId="23.[106,1097,1019,1227]" pageId="23" pageNumber="390">
<taxonomicName id="4C6B4D1CFFFDFF8D9427FC2F57111834" box="[106,332,1020,1038]" class="Insecta" family="Staphylinidae" genus="Pronomaea" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="390" phylum="Arthropoda" rank="species" species="khnzorianikhnzoriani">
<emphasis id="B91FEA8DFFFDFF8D9427FC2F57111834" box="[106,332,1020,1038]" italics="true" pageId="23" pageNumber="390">Pronomaea khnzorianikhnzoriani</emphasis>
</taxonomicName>
was previously known from
<collectingCountry id="F37C760FFFFDFF8D963FFC2854931829" box="[626,718,1019,1043]" name="Armenia" pageId="23" pageNumber="390">Armenia</collectingCountry>
,
<collectingCountry id="F37C760FFFFDFF8D969AFC2855711829" box="[727,812,1019,1043]" name="Georgia" pageId="23" pageNumber="390">Georgia</collectingCountry>
, and
<collectingCountry id="F37C760FFFFDFF8D972EFC2F55D01829" box="[867,909,1020,1043]" name="Iran" pageId="23" pageNumber="390">Iran</collectingCountry>
((ASSING in press). It is here recorded from
<collectingCountry id="F37C760FFFFDFF8D952FFBC957F71808" box="[354,426,1050,1074]" name="Turkey" pageId="23" pageNumber="390">Turkey</collectingCountry>
, Dagestan, and
<collectingCountry id="F37C760FFFFDFF8D9602FBC954831808" box="[591,734,1050,1074]" name="Turkmenistan" pageId="23" pageNumber="390">Turkmenistan</collectingCountry>
for the first time (
<figureCitation id="13502A1AFFFDFF8D97D0FBC955BB1808" box="[925,998,1050,1074]" captionStart="Map 4" captionStartId="22.[195,239,844,865]" captionTargetBox="[195,1178,174,836]" captionTargetId="figure-325@22.[195,1178,174,836]" captionTargetPageId="22" captionText="Map 4: Distributions of Pronomaea anatolica FAGEL (filled circles) and P. khnzoriani SEMENOV (open circles), based on examined and literature records." figureDoi="http://doi.org/10.5281/zenodo.5364184" httpUri="https://zenodo.org/record/5364184/files/figure.png" pageId="23" pageNumber="390">Map 4</figureCitation>
). In
<collectingCountry id="F37C760FFFFDFF8D905AFBC852191808" box="[1047,1092,1051,1074]" name="Iran" pageId="23" pageNumber="390">Iran</collectingCountry>
, the species is rather common on the banks of rivers and streams; for numerous additional records see ASSING (in press). The altitudes specified on the labes range from
<quantity id="4C939B7AFFFDFF8D9761FB8455971855" box="[812,970,1111,1135]" metricMagnitude="3" metricUnit="m" metricValue="1.555" metricValueMax="2.71" metricValueMin="0.4" pageId="23" pageNumber="390" unit="m" value="1555.0" valueMax="2710.0" valueMin="400.0">400 to 2710 m</quantity>
. The species has been collected in April (
<bibRefCitation id="EFFA4B6EFFFDFF8D95C2FBA4547318B4" author="SEMENOV, V. B." box="[399,558,1142,1166]" pageId="23" pageNumber="390" pagination="199 - 202" refId="ref17954" refString="SEMENOV, V. B. 2003: Die kaukasischen und mittelasiatischen Arten der Gattung Pronomaea ERICHSON, 1837 (Coleoptera: Staphylinidae: Aleocharinae). - Russian Entomological Journal 12: 199 - 202." type="journal article" year="2003">SEMENOV 2003</bibRefCitation>
) and in large numbers during the period from June through August ((ASSING in press; material examined). Three specimens were taken in September. Some beetles collected in June are teneral.
</paragraph>
</subSubSection>
</treatment>
</document>
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