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<document id="801B26911B8FC290D1B581CDAEE4111A" ID-CLB-Dataset="53812" ID-DOI="10.11646/zootaxa.5250.1.1" ID-GBIF-Dataset="da8acfed-df61-403e-9a49-7f856ce56196" ID-ISSN="1175­5334" ID-Zenodo-Dep="7705657" ID-ZooBank="4729EAF3-8B78-4AAD-B373-9179D008F239" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" checkinTime="1678178160677" checkinUser="plazi" docAuthor="WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI &amp; HUANG, JIA" docDate="2023" docId="0393914C2A0C8757C282FD076B96F9EA" docLanguage="en" docName="zootaxa.5250.1.1.pdf" docOrigin="Zootaxa 5250 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5250.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov." docType="treatment" docVersion="10" lastPageNumber="48" masterDocId="FFAAE9342A228778C215FF896958FB43" masterDocTitle="Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae)" masterLastPageNumber="109" masterPageNumber="1" pageNumber="47" updateTime="1699476226253" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F0F4C25977F5783FE3B65B10AE91B134">Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae)</mods:title>
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<mods:namePart id="0996F620F671882D8816A88134829B35">CUI, MENG-DI</mods:namePart>
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<mods:namePart id="F925980B490F1DE1EDCDE2A59CE59DDA">ZHANG, YUAN</mods:namePart>
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<mods:namePart id="47E7BA331B88CE6E2F52BC726DA6C8D9">WANG, LE</mods:namePart>
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<treatment id="0393914C2A0C8757C282FD076B96F9EA" ID-DOI="http://doi.org/10.5281/zenodo.7708847" ID-GBIF-Taxon="209579528" ID-Zenodo-Dep="7708847" LSID="urn:lsid:plazi:treatment:0393914C2A0C8757C282FD076B96F9EA" httpUri="http://treatment.plazi.org/id/0393914C2A0C8757C282FD076B96F9EA" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<subSubSection id="C32073D12A0C8756C282FD076A75F9EB" box="[151,813,654,681]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8B85205A2A0C8756C282FD076A75F9EB" blockId="46.[151,813,654,717]" box="[151,813,654,681]" pageId="46" pageNumber="47">
<heading id="D0CD97362A0C8756C282FD076A75F9EB" bold="true" box="[151,813,654,681]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
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<taxonomicName id="4C3A5BD92A0C8756C282FD076B9AF9EB" ID-CoL="BDMQN" authority="Cui &amp; Chen" authorityName="Cui &amp; Chen" box="[151,706,654,681]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="marenubila" status="sp. nov." subGenus="Steganina">
<emphasis id="B94EFC482A0C8756C282FD0769A1F9EB" bold="true" box="[151,249,654,680]" italics="true" pageId="46" pageNumber="47">Stegana</emphasis>
(
<emphasis id="B94EFC482A0C8756C31FFD0768DCF9EB" bold="true" box="[266,388,654,680]" italics="true" pageId="46" pageNumber="47">Steganina</emphasis>
)
<emphasis id="B94EFC482A0C8756C380FD076B79F9EA" bold="true" box="[405,545,654,681]" italics="true" pageId="46" pageNumber="47">marenubila</emphasis>
Cui &amp; Chen
</taxonomicName>
,
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</emphasis>
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<subSubSection id="C32073D12A0C8756C282FD3B683BF98F" box="[151,355,690,717]" pageId="46" pageNumber="47" type="description">
<paragraph id="8B85205A2A0C8756C282FD3B683BF98F" blockId="46.[151,813,654,717]" box="[151,355,690,717]" pageId="46" pageNumber="47">
(
<figureCitation id="13013CDF2A0C8756C28AFD3B6869F98E" box="[159,305,690,717]" captionStart="FIGURE 5" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-30@28.[173,1415,181,1919]" captionTargetPageId="28" captionText="FIGURE 5. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov.; C, D. S. (S.) S. (S.) nigripalpula Cui &amp; Chen, sp. nov.; E, F. S. (S.) glabra Chen &amp; Chen, 2012; G, H. S. (S.) masanoritodai Okada &amp; Sidorenko, 1992; I, J. S. (S.) multidentata Chen, Gao, &amp; Chen, 2009." figureDoi="http://doi.org/10.5281/zenodo.7705675" httpUri="https://zenodo.org/record/7705675/files/figure.png" pageId="46" pageNumber="47">Figs. 5A, 5B</figureCitation>
,
<figureCitation id="13013CDF2A0C8756C32BFD3B6803F98F" box="[318,347,690,716]" captionStart="FIGURE 21" captionStartId="46.[151,250,1693,1717]" captionTargetBox="[163,1426,901,1662]" captionTargetId="figure-300@46.[151,1436,884,1669]" captionTargetPageId="46" captionText="FIGURE 21. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; D. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705715" httpUri="https://zenodo.org/record/7705715/files/figure.png" pageId="46" pageNumber="47">21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A0C8756C282FD736CCEF81F" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="8B85205A2A0C8756C282FD736CCEF81F" blockId="46.[151,1437,762,861]" pageId="46" pageNumber="47">
<emphasis id="B94EFC482A0C8756C282FD73684AF857" bold="true" box="[151,274,762,788]" pageId="46" pageNumber="47">Diagnosis.</emphasis>
This species resembles
<emphasis id="B94EFC482A0C8756C033FD726B6CF856" box="[550,564,763,789]" italics="true" pageId="46" pageNumber="47">S</emphasis>
. (
<emphasis id="B94EFC482A0C8756C058FD726B03F856" box="[589,603,763,789]" italics="true" pageId="46" pageNumber="47">S</emphasis>
.)
<emphasis id="B94EFC482A0C8756C061FD726B9DF857" box="[628,709,763,788]" italics="true" pageId="46" pageNumber="47">crinata</emphasis>
<bibRefCitation id="EFAB5DAB2A0C8756C0DBFD736AE7F856" author="Zhang, Y. &amp; Chen, H. W." box="[718,959,762,789]" pageId="46" pageNumber="47" pagination="131 - 137" refId="ref75057" refString="Zhang, Y. &amp; Chen, H. W. (2015) Four new species of the Stegana ornatipes species group (Diptera: Drosophilidae) from Yunnan, China, with DNA barcoding information. Zootaxa, 3905 (1), 131 - 137. https: // doi. org / 10.11646 / zootaxa. 3905.1.8" type="journal article" year="2015">
Zhang &amp; Chen,
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in the shape of epandrium, surstylus, and cercus (as fig.
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<bibRefCitation id="EFAB5DAB2A0C8756C379FC976B0CF87A" author="Zhang, Y. &amp; Chen, H. W." box="[364,596,798,825]" pageId="46" pageNumber="47" pagination="131 - 137" refId="ref75057" refString="Zhang, Y. &amp; Chen, H. W. (2015) Four new species of the Stegana ornatipes species group (Diptera: Drosophilidae) from Yunnan, China, with DNA barcoding information. Zootaxa, 3905 (1), 131 - 137. https: // doi. org / 10.11646 / zootaxa. 3905.1.8" type="journal article" year="2015">Zhang &amp; Chen 2015</bibRefCitation>
,
<figureCitation id="13013CDF2A0C8756C04AFC976B9EF87B" box="[607,710,798,824]" captionStart="FIGURE 21" captionStartId="46.[151,250,1693,1717]" captionTargetBox="[163,1426,901,1662]" captionTargetId="figure-300@46.[151,1436,884,1669]" captionTargetPageId="46" captionText="FIGURE 21. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; D. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705715" httpUri="https://zenodo.org/record/7705715/files/figure.png" pageId="46" pageNumber="47">Fig. 21A</figureCitation>
), but can be distinguished from the latter by having the aedeagus with 3 pair of basally fused processes: ventral pair small, medial pair slender, and dorsal pair strong (
<figureCitation id="13013CDF2A0C8756C717FCCB6CD1F81F" box="[1282,1417,834,860]" captionStart="FIGURE 21" captionStartId="46.[151,250,1693,1717]" captionTargetBox="[163,1426,901,1662]" captionTargetId="figure-300@46.[151,1436,884,1669]" captionTargetPageId="46" captionText="FIGURE 21. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; D. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705715" httpUri="https://zenodo.org/record/7705715/files/figure.png" pageId="46" pageNumber="47">Fig. 21B, C</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF4570D22A0C8756C282F91468DCFDBE" ID-DOI="http://doi.org/10.5281/zenodo.7705715" ID-Zenodo-Dep="7705715" httpUri="https://zenodo.org/record/7705715/files/figure.png" pageId="46" pageNumber="47" startId="46.[151,250,1693,1717]" targetBox="[163,1426,901,1662]" targetPageId="46">
<paragraph id="8B85205A2A0C8756C282F91468DCFDBE" blockId="46.[151,1437,1693,1789]" pageId="46" pageNumber="47">
<emphasis id="B94EFC482A0C8756C282F9146879FDF6" bold="true" box="[151,289,1693,1717]" pageId="46" pageNumber="47">FIGURE 21.</emphasis>
<taxonomicName id="4C3A5BD92A0C8756C33DF9176BA1FDF6" authority="Cui &amp; Chen" authorityName="Cui &amp; Chen" box="[296,761,1693,1717]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="marenubila" status="sp. nov." subGenus="Steganina">
<emphasis id="B94EFC482A0C8756C33DF9176822FDF6" box="[296,378,1694,1717]" italics="true" pageId="46" pageNumber="47">Stegana</emphasis>
(
<emphasis id="B94EFC482A0C8756C39CF91768B6FDF6" box="[393,494,1694,1717]" italics="true" pageId="46" pageNumber="47">Steganina</emphasis>
)
<emphasis id="B94EFC482A0C8756C3EBF9146B2AFDF6" box="[510,626,1693,1717]" italics="true" pageId="46" pageNumber="47">marenubila</emphasis>
Cui &amp; Chen
</taxonomicName>
,
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<taxonomicNameLabel id="A27D41332A0C8756C111F9146A0DFDF6" box="[772,853,1693,1717]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; D. Subepandrial sclerite in ventral view.
</paragraph>
</caption>
<subSubSection id="C32073D12A0C8757C2D2F8A26830FA92" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="description">
<paragraph id="8B85205A2A0C8756C2D2F8A26CC5FCCE" blockId="46.[151,1437,1835,1934]" pageId="46" pageNumber="47">
<emphasis id="B94EFC482A0C8756C2D2F8A26801FC06" bold="true" box="[199,345,1835,1861]" pageId="46" pageNumber="47">Description.</emphasis>
Male and female: Frons yellow in male, brown in female. First flagellomere yellow in male, brown in female. Face black, with 1 yellow band medially; facial carina absent. Clypeus black. Palpus mostly yellow. Mesonotum and scutellum yellow in male (
<figureCitation id="13013CDF2A0C8756C0D7F8FA6A67FCCD" box="[706,831,1907,1934]" captionStart="FIGURE 5" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-30@28.[173,1415,181,1919]" captionTargetPageId="28" captionText="FIGURE 5. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov.; C, D. S. (S.) S. (S.) nigripalpula Cui &amp; Chen, sp. nov.; E, F. S. (S.) glabra Chen &amp; Chen, 2012; G, H. S. (S.) masanoritodai Okada &amp; Sidorenko, 1992; I, J. S. (S.) multidentata Chen, Gao, &amp; Chen, 2009." figureDoi="http://doi.org/10.5281/zenodo.7705675" httpUri="https://zenodo.org/record/7705675/files/figure.png" pageId="46" pageNumber="47">Fig. 5A, B</figureCitation>
), brownish in female. Katepisternum yellowish (
<figureCitation id="13013CDF2A0C8756C77AF8FA6CC5FCCE" box="[1391,1437,1907,1933]" captionStart="FIGURE 5" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-30@28.[173,1415,181,1919]" captionTargetPageId="28" captionText="FIGURE 5. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov.; C, D. S. (S.) S. (S.) nigripalpula Cui &amp; Chen, sp. nov.; E, F. S. (S.) glabra Chen &amp; Chen, 2012; G, H. S. (S.) masanoritodai Okada &amp; Sidorenko, 1992; I, J. S. (S.) multidentata Chen, Gao, &amp; Chen, 2009." figureDoi="http://doi.org/10.5281/zenodo.7705675" httpUri="https://zenodo.org/record/7705675/files/figure.png" pageId="46" pageNumber="47">Fig.</figureCitation>
</paragraph>
<paragraph id="8B85205A2A0C8757C282F81E6BF1FBBA" blockId="46.[151,1437,1943,2042]" lastBlockId="47.[151,1436,151,681]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
5B). Halter mostly grey (
<figureCitation id="13013CDF2A0C8756C3A0F81E6B56FCF1" box="[437,526,1943,1970]" captionStart="FIGURE 5" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-30@28.[173,1415,181,1919]" captionTargetPageId="28" captionText="FIGURE 5. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov.; C, D. S. (S.) S. (S.) nigripalpula Cui &amp; Chen, sp. nov.; E, F. S. (S.) glabra Chen &amp; Chen, 2012; G, H. S. (S.) masanoritodai Okada &amp; Sidorenko, 1992; I, J. S. (S.) multidentata Chen, Gao, &amp; Chen, 2009." figureDoi="http://doi.org/10.5281/zenodo.7705675" httpUri="https://zenodo.org/record/7705675/files/figure.png" pageId="46" pageNumber="47">Fig. 5B</figureCitation>
). Fore femur with 3 thin, long setae on distal part of posteroventral surface. Abdominal tergites yellow (
<figureCitation id="13013CDF2A0C8756C3B9F8326B7CFC95" box="[428,548,1979,2006]" captionStart="FIGURE 5" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-30@28.[173,1415,181,1919]" captionTargetPageId="28" captionText="FIGURE 5. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov.; C, D. S. (S.) S. (S.) nigripalpula Cui &amp; Chen, sp. nov.; E, F. S. (S.) glabra Chen &amp; Chen, 2012; G, H. S. (S.) masanoritodai Okada &amp; Sidorenko, 1992; I, J. S. (S.) multidentata Chen, Gao, &amp; Chen, 2009." figureDoi="http://doi.org/10.5281/zenodo.7705675" httpUri="https://zenodo.org/record/7705675/files/figure.png" pageId="46" pageNumber="47">Fig. 5A, B</figureCitation>
), 4th to 6th tergites brownish medially in female; sternites yellow, about 2 times as wide as long. Male terminalia (
<figureCitation id="13013CDF2A0C8756C004F8566B3DFCB9" box="[529,613,2015,2042]" captionStart="FIGURE 21" captionStartId="46.[151,250,1693,1717]" captionTargetBox="[163,1426,901,1662]" captionTargetId="figure-300@46.[151,1436,884,1669]" captionTargetPageId="46" captionText="FIGURE 21. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; D. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705715" httpUri="https://zenodo.org/record/7705715/files/figure.png" pageId="46" pageNumber="47">Fig. 21</figureCitation>
): Epandrium with
<emphasis id="B94EFC482A0C8756C120F8696A17FCBA" box="[821,847,2016,2041]" italics="true" pageId="46" pageNumber="47">ca</emphasis>
. 8 setae on dorsal to posterior portions per side (
<figureCitation id="13013CDF2A0C8757C77AF8566993FBF2" captionStart="FIGURE 21" captionStartId="46.[151,250,1693,1717]" captionTargetBox="[163,1426,901,1662]" captionTargetId="figure-300@46.[151,1436,884,1669]" captionTargetPageId="46" captionText="FIGURE 21. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; D. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705715" httpUri="https://zenodo.org/record/7705715/files/figure.png" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">Fig. 21A</figureCitation>
). Surstylus lacking pubescence (
<figureCitation id="13013CDF2A0D8757C057FF1E6BF5FBF2" box="[578,685,151,177]" captionStart="FIGURE 21" captionStartId="46.[151,250,1693,1717]" captionTargetBox="[163,1426,901,1662]" captionTargetId="figure-300@46.[151,1436,884,1669]" captionTargetPageId="46" captionText="FIGURE 21. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; D. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705715" httpUri="https://zenodo.org/record/7705715/files/figure.png" pageId="47" pageNumber="48">Fig. 21A</figureCitation>
). Cercus with much pubescence (
<figureCitation id="13013CDF2A0D8757C625FF1E6DC3FBF2" box="[1072,1179,151,177]" captionStart="FIGURE 21" captionStartId="46.[151,250,1693,1717]" captionTargetBox="[163,1426,901,1662]" captionTargetId="figure-300@46.[151,1436,884,1669]" captionTargetPageId="46" captionText="FIGURE 21. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; D. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705715" httpUri="https://zenodo.org/record/7705715/files/figure.png" pageId="47" pageNumber="48">Fig. 21A</figureCitation>
). Hypandrium slightly expanded anteromedially (
<figureCitation id="13013CDF2A0D8757C3AAFF326B1CFB96" box="[447,580,187,213]" captionStart="FIGURE 21" captionStartId="46.[151,250,1693,1717]" captionTargetBox="[163,1426,901,1662]" captionTargetId="figure-300@46.[151,1436,884,1669]" captionTargetPageId="46" captionText="FIGURE 21. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; D. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705715" httpUri="https://zenodo.org/record/7705715/files/figure.png" pageId="47" pageNumber="48">Fig. 21B, C</figureCitation>
). Pregonite blurry or absent (
<figureCitation id="13013CDF2A0D8757C193FF326D53FB96" box="[902,1035,187,213]" captionStart="FIGURE 21" captionStartId="46.[151,250,1693,1717]" captionTargetBox="[163,1426,901,1662]" captionTargetId="figure-300@46.[151,1436,884,1669]" captionTargetPageId="46" captionText="FIGURE 21. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; D. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705715" httpUri="https://zenodo.org/record/7705715/files/figure.png" pageId="47" pageNumber="48">Fig. 21B, C</figureCitation>
). Junction between hypandrium and aedeagus with 3 sensilla per side (
<figureCitation id="13013CDF2A0D8757C000FF566BC4FBBA" box="[533,668,223,249]" captionStart="FIGURE 21" captionStartId="46.[151,250,1693,1717]" captionTargetBox="[163,1426,901,1662]" captionTargetId="figure-300@46.[151,1436,884,1669]" captionTargetPageId="46" captionText="FIGURE 21. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov., male terminalia. A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, aedeagus, phallapodeme, and aedeagal sheath in ventral and lateral views; D. Subepandrial sclerite in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705715" httpUri="https://zenodo.org/record/7705715/files/figure.png" pageId="47" pageNumber="48">Fig. 21B, C</figureCitation>
).
</paragraph>
<paragraph id="8B85205A2A0D8757C2D2FE8A6830FA92" blockId="47.[151,1436,151,681]" pageId="47" pageNumber="48">
<emphasis id="B94EFC482A0D8757C2D2FE8A6B55FA5F" bold="true" box="[199,525,258,285]" pageId="47" pageNumber="48">Measurements and indices.</emphasis>
BL =
<quantity id="4CC28DBF2A0D8757C042FE8B6BE4FA5E" box="[599,700,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" pageId="47" pageNumber="48" unit="mm" value="2.45">2.45 mm</quantity>
in
<typeStatus id="54819EF82A0D8757C0F4FE8A6A1BFA5E" box="[737,835,259,285]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
(
<quantity id="4CC28DBF2A0D8757C147FE8A6AFBFA5E" box="[850,931,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="6.9342" pageId="47" pageNumber="48" unit="in" value="2.73">2.73 in</quantity>
<specimenCount id="9D3CEBD32A0D8757C1BFFE8A6A95FA5E" box="[938,973,259,285]" pageId="47" pageNumber="48" type="female">1♀</specimenCount>
<typeStatus id="54819EF82A0D8757C1C0FE8A6D62FA5E" box="[981,1082,259,285]" pageId="47" pageNumber="48" type="paratype">paratype</typeStatus>
), ThL =
<quantity id="4CC28DBF2A0D8757C689FE8A6C59FA5E" box="[1180,1281,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" pageId="47" pageNumber="48" unit="mm" value="1.21">1.21 mm</quantity>
(1.19), WL =
<quantity id="4CC28DBF2A0D8757C282FEAF69A3FA02" box="[151,251,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="2.56" pageId="47" pageNumber="48" unit="mm" value="2.56">2.56 mm</quantity>
(2.75), WW =
<quantity id="4CC28DBF2A0D8757C3B7FEAE6B5DFA02" box="[418,517,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="47" pageNumber="48" unit="mm" value="0.93">0.93 mm</quantity>
(1.12), arb = 7/4 (7/5), avd = 0.73 (0.67), adf = 1.07 (1.21), flw = 1.42 (1.69), FW/ HW = 0.32 (0.37), ch/o = 0.24 (0.24), prorb = 0.96 (0.90), rcorb = 1.20 (0.54), vb = 0.57 (0.46), dcl = 0.28 (1.22), presctl = 0.38 (0.38), sctl = damaged (2.63), sterno = 0.41 (0.46), orbito = 2.22 (1.89), dcp = 0.53 (0.30), sctlp = 1.50 (1.39), C = 2.20 (2.16), 4c = 1.62 (1.02), 4v = 2.49 (1.75), 5x = 1.29 (1.42), ac = 10.75 (8.46), M = 0.93 (0.56), C3F = 0.64 (0.60).
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A0D8757C2D2FE526B6BF97E" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="8B85205A2A0D8757C2D2FE526B6BF97E" blockId="47.[151,1436,151,681]" pageId="47" pageNumber="48">
<emphasis id="B94EFC482A0D8757C2D2FE5268DFFAB7" bold="true" box="[199,391,474,501]" pageId="47" pageNumber="48">Type materials.</emphasis>
<materialsCitation id="3B522A072A0D8757C380FE526A93F95A" ID-GBIF-Occurrence="4097216303" collectingDate="2017-11-08" collectionCode="SCAU" collectorName="H. W. Chen" country="China" elevation="1720" latitude="24.1175" location="Mangshi" longLatPrecision="21" longitude="98.31778" municipality="Manghai" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54819EF82A0D8757C380FE5268A5FAB6" box="[405,509,475,501]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
♂ (
<collectionCode id="ED2BB89F2A0D8757C020FE526BDCFAB6" box="[565,644,475,501]" pageId="47" pageNumber="48">SCAU</collectionCode>
, No. 125411, DNA #2347),
<collectingCountry id="F32D60CA2A0D8757C1F6FE526D18FAB6" box="[995,1088,475,501]" name="China" pageId="47" pageNumber="48">CHINA</collectingCountry>
:
<collectingMunicipality id="6BE1BA202A0D8757C644FE526DEFFAB6" box="[1105,1207,475,501]" pageId="47" pageNumber="48">Manghai</collectingMunicipality>
,
<location id="8EE576812A0D8757C6DFFE526C77FAB6" LSID="urn:lsid:plazi:treatment:0393914C2A0C8757C282FD076B96F9EA:8EE576812A0D8757C6DFFE526C77FAB6" box="[1226,1327,475,501]" country="China" latitude="24.1175" longLatPrecision="21" longitude="98.31778" municipality="Manghai" name="Mangshi" pageId="47" pageNumber="48" stateProvince="Yunnan">Mangshi</location>
,
<collectingRegion id="49FEEEB82A0D8757C755FE526CC1FAB6" box="[1344,1433,475,501]" country="China" name="Yunnan" pageId="47" pageNumber="48">Yunnan</collectingRegion>
,
<geoCoordinate id="EE0E469D2A0D8757C282FE766844F95A" box="[151,284,511,537]" degrees="24" direction="north" minutes="07" orientation="latitude" pageId="47" pageNumber="48" precision="15" seconds="03" value="24.1175">24°07'03''N</geoCoordinate>
,
<geoCoordinate id="EE0E469D2A0D8757C332FE7668F1F95A" box="[295,425,511,537]" degrees="98" direction="east" minutes="19" orientation="longitude" pageId="47" pageNumber="48" precision="15" seconds="04" value="98.31778">98°19'04''E</geoCoordinate>
, alt.
<quantity id="4CC28DBF2A0D8757C3CBFE766B76F95A" box="[478,558,511,537]" metricMagnitude="3" metricUnit="m" metricValue="1.72" pageId="47" pageNumber="48" unit="m" value="1720.0">
<elevation id="0017C7692A0D8757C3CBFE766B76F95A" box="[478,558,511,537]" metricMagnitude="3" metricUnit="m" metricValue="1.72" pageId="47" pageNumber="48" unit="m" value="1720.0">1720m</elevation>
</quantity>
,
<date id="FF84069A2A0D8757C02CFE766BFEF95A" box="[569,678,511,537]" pageId="47" pageNumber="48" value="2017-11-08">
<collectingDate id="EFC0FF722A0D8757C02CFE766BFEF95A" box="[569,678,511,537]" pageId="47" pageNumber="48" value="2017-11-08">8.xi.2017</collectingDate>
</date>
, ex tree trunk,
<collectorName id="26CF458C2A0D8757C15EFE766A93F95A" box="[843,971,511,537]" pageId="47" pageNumber="48">H.W. Chen</collectorName>
</materialsCitation>
.
<materialsCitation id="3B522A072A0D8757C1C1FE766B6BF97E" ID-GBIF-Occurrence="4097216435" collectingDate="2017-11-08" collectionCode="SCAU" collectorName="H. W. Chen" country="China" elevation="1720" latitude="24.1175" location="Mangshi" longLatPrecision="21" longitude="98.31778" municipality="Manghai" pageId="47" pageNumber="48" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54819EF82A0D8757C1C1FE766D62F95A" box="[980,1082,511,537]" pageId="47" pageNumber="48" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D3CEBD32A0D8757C650FE766D3FF95A" box="[1093,1127,511,537]" pageId="47" pageNumber="48" type="female">1♀</specimenCount>
(
<collectionCode id="ED2BB89F2A0D8757C663FE766D9DF95A" box="[1142,1221,511,537]" pageId="47" pageNumber="48">SCAU</collectionCode>
, No. 125412, DNA #2747), other data same as holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A0D8757C2D2FDCF6B79F922" box="[199,545,582,609]" pageId="47" pageNumber="48" type="distribution">
<paragraph id="8B85205A2A0D8757C2D2FDCF6B79F922" blockId="47.[151,1436,151,681]" box="[199,545,582,609]" pageId="47" pageNumber="48">
<emphasis id="B94EFC482A0D8757C2D2FDCF6839F923" bold="true" box="[199,353,582,608]" pageId="47" pageNumber="48">Distribution.</emphasis>
<collectingCountry id="F32D60CA2A0D8757C372FDCE68F2F922" box="[359,426,583,609]" name="China" pageId="47" pageNumber="48">China</collectingCountry>
(
<collectingRegion id="49FEEEB82A0D8757C3ACFDCE6B4DF922" box="[441,533,583,609]" country="China" name="Yunnan" pageId="47" pageNumber="48">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A0D8757C2D2FDE26B96F9EA" pageId="47" pageNumber="48" type="etymology">
<paragraph id="8B85205A2A0D8757C2D2FDE26B96F9EA" blockId="47.[151,1436,151,681]" pageId="47" pageNumber="48">
<emphasis id="B94EFC482A0D8757C2D2FDE26816F9C6" bold="true" box="[199,334,619,645]" pageId="47" pageNumber="48">Etymology.</emphasis>
A combination of the Latin words &quot;
<emphasis id="B94EFC482A0D8757C0CFFDE26A4BF9C7" box="[730,787,619,644]" italics="true" pageId="47" pageNumber="48">mare</emphasis>
&quot; (= sea) and &quot;
<emphasis id="B94EFC482A0D8757C1A5FDE26D5AF9C6" box="[944,1026,619,645]" italics="true" pageId="47" pageNumber="48">nubilus</emphasis>
&quot; (= nubilous), referring to the meaning of the
<typeStatus id="54819EF82A0D8757C31FFD066862F9EA" box="[266,314,655,681]" pageId="47" pageNumber="48">type</typeStatus>
locality (Manghai: a sea of clouds).
</paragraph>
</subSubSection>
</treatment>
</document>
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