treatments-xml/data/03/93/87/0393878AFFE19B550695FB5A5844F821.xml
2024-06-21 12:22:17 +02:00

208 lines
22 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="10.11646/zootaxa.5258.4.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="7784525" ID-ZooBank="915148C7-872B-4434-8E8B-F326EC568362" checkinTime="1680165208046" checkinUser="plazi" docAuthor="Brock, Paul D. &amp; Hennemann, Frank H." docDate="2023" docId="0393878AFFE19B550695FB5A5844F821" docLanguage="en" docName="zootaxa.5258.4.5.pdf" docOrigin="Zootaxa 5258 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5258.4.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Xeroderus conlei Brock &amp; Hennemann 2023, sp. n." docType="treatment" docVersion="3" lastPageNumber="450" masterDocId="FFAAFFF2FFE49B520602FFDC5D12FF8B" masterDocTitle="A new species of Xeroderus Gray, 1835 (Phasmida: Phasmatidae: Xeroderinae) from Papua New Guinea and notes on the genus" masterLastPageNumber="454" masterPageNumber="443" pageNumber="448" updateTime="1680204935416" updateUser="diego" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Xeroderus Gray, 1835 (Phasmida: Phasmatidae: Xeroderinae) from Papua New Guinea and notes on the genus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Brock, Paul D.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hennemann, Frank H.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5258</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>443</mods:start>
<mods:end>454</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5258.4.5</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5258.4.5</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7784525</mods:identifier>
<mods:identifier type="ZooBank">915148C7-872B-4434-8E8B-F326EC568362</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:0393878AFFE19B550695FB5A5844F821" httpUri="http://treatment.plazi.org/id/0393878AFFE19B550695FB5A5844F821" lastPageId="7" lastPageNumber="450" pageId="5" pageNumber="448">
<subSubSection box="[151,431,1158,1185]" pageId="5" pageNumber="448" type="nomenclature">
<paragraph blockId="5.[151,431,1158,1257]" box="[151,431,1158,1185]" pageId="5" pageNumber="448">
<heading bold="true" box="[151,431,1158,1185]" fontSize="11" level="1" pageId="5" pageNumber="448" reason="1">
<emphasis bold="true" box="[151,431,1158,1185]" pageId="5" pageNumber="448">
<taxonomicName authority="Brock &amp; Hennemann, 2023" authorityName="Brock &amp; Hennemann" authorityYear="2023" box="[151,357,1158,1185]" class="Insecta" family="Phasmatidae" genus="Xeroderus" kingdom="Animalia" order="Phasmida" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="conlei" status="sp. n.">
<emphasis bold="true" box="[151,357,1158,1185]" italics="true" pageId="5" pageNumber="448">Xeroderus conlei</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[365,431,1159,1185]" pageId="5" pageNumber="448" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,379,1195,1221]" pageId="5" pageNumber="448" type="vernacular_names">
<paragraph blockId="5.[151,431,1158,1257]" box="[151,379,1195,1221]" pageId="5" pageNumber="448">
[Conles
<taxonomicName authorityName="Gray" authorityYear="1835" box="[253,373,1195,1221]" class="Insecta" family="Phasmatidae" genus="Xeroderus" kingdom="Animalia" order="Phasmida" pageId="5" pageNumber="448" phylum="Arthropoda" rank="genus">Xeroderus</taxonomicName>
]
</paragraph>
</subSubSection>
<subSubSection box="[151,274,1231,1257]" pageId="5" pageNumber="448" type="description">
<paragraph blockId="5.[151,431,1158,1257]" box="[151,274,1231,1257]" pageId="5" pageNumber="448">
(
<figureCitation box="[159,266,1231,1257]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="4.[151,250,1917,1941]" captionStartId-1="6.[151,250,1903,1927]" captionTargetBox-0="[173,1434,314,1890]" captionTargetBox-1="[151,1436,597,1880]" captionTargetId-0="figure-88@4.[151,1436,309,1894]" captionTargetId-1="figure-212@6.[151,1436,597,1880]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionText-0="FIGURE 3: Xeroderus conlei sp. n. A. ♀ holotype, dorsal view [MNHU]; B. ventral view; C. lateral view; D. Head &amp; thorax, in dorsal view; E. F. G. End of abdomen lateral, dorsal and ventral views. All photos by Oskar Conle." captionText-1="FIGURE 4: Distribution map of Xeroderus species." figureDoi-0="http://doi.org/10.5281/zenodo.7784538" figureDoi-1="http://doi.org/10.5281/zenodo.7784542" httpUri-0="https://zenodo.org/record/7784538/files/figure.png" httpUri-1="https://zenodo.org/record/7784542/files/figure.png" pageId="5" pageNumber="448">Figs. 34</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection box="[151,1425,1303,1329]" pageId="5" pageNumber="448" type="materials_examined">
<paragraph blockId="5.[151,1437,1303,2013]" box="[151,1425,1303,1329]" pageId="5" pageNumber="448">
<materialsCitation box="[151,1425,1303,1329]" collectionCode="MNHU" collectorName="D. Neu-Guinea' &amp; Prof. Neuhauss S." country="Papua New Guinea" location="PAPUA NEW GUINEA" pageId="5" pageNumber="448" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis bold="true" box="[151,260,1303,1329]" pageId="5" pageNumber="448">
<typeStatus box="[151,260,1303,1329]" pageId="5" pageNumber="448">Holotype</typeStatus>
</emphasis>
<specimenCount box="[267,288,1303,1329]" pageId="5" pageNumber="448" type="female"></specimenCount>
,
<collectingCountry box="[302,573,1303,1329]" name="Papua New Guinea" pageId="5" pageNumber="448">PAPUA NEW GUINEA</collectingCountry>
[as
<collectorName box="[625,815,1303,1329]" pageId="5" pageNumber="448">D. Neu-Guinea</collectorName>
], Sattelberg,
<collectorName box="[959,1154,1303,1329]" pageId="5" pageNumber="448">Prof. Neuhauss S.</collectorName>
, 10 iii.[19]09 [
<collectionCode box="[1320,1415,1303,1329]" pageId="5" pageNumber="448">MNHU</collectionCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="449" pageId="5" pageNumber="448" type="description">
<paragraph blockId="5.[151,1437,1303,2013]" pageId="5" pageNumber="448">
<emphasis bold="true" box="[199,345,1338,1364]" pageId="5" pageNumber="448">Description.</emphasis>
Attractive part brownish and greyish medium-sized insect, heavily mottled greyish and whitish dorsally, possibly partly green, including legs, forewings and pre-anal part of hindwings, the latter uniform brown (
<figureCitation box="[159,249,1411,1437]" captionStart="FIGURE 3" captionStartId="4.[151,250,1917,1941]" captionTargetBox="[173,1434,314,1890]" captionTargetId="figure-88@4.[151,1436,309,1894]" captionTargetPageId="4" captionText="FIGURE 3: Xeroderus conlei sp. n. A. ♀ holotype, dorsal view [MNHU]; B. ventral view; C. lateral view; D. Head &amp; thorax, in dorsal view; E. F. G. End of abdomen lateral, dorsal and ventral views. All photos by Oskar Conle." figureDoi="http://doi.org/10.5281/zenodo.7784538" httpUri="https://zenodo.org/record/7784538/files/figure.png" pageId="5" pageNumber="448">Fig. 3A</figureCitation>
). Thorax with lateral lobes, mesonotum with paired broad spines (
<figureCitation box="[991,1126,1411,1437]" captionStart="FIGURE 3" captionStartId="4.[151,250,1917,1941]" captionTargetBox="[173,1434,314,1890]" captionTargetId="figure-88@4.[151,1436,309,1894]" captionTargetPageId="4" captionText="FIGURE 3: Xeroderus conlei sp. n. A. ♀ holotype, dorsal view [MNHU]; B. ventral view; C. lateral view; D. Head &amp; thorax, in dorsal view; E. F. G. End of abdomen lateral, dorsal and ventral views. All photos by Oskar Conle." figureDoi="http://doi.org/10.5281/zenodo.7784538" httpUri="https://zenodo.org/record/7784538/files/figure.png" pageId="5" pageNumber="448">Figs. 3CD</figureCitation>
).
</paragraph>
<paragraph blockId="5.[151,1437,1303,2013]" lastBlockId="6.[151,1437,151,573]" lastPageId="6" lastPageNumber="449" pageId="5" pageNumber="448">
<emphasis bold="true" box="[199,322,1446,1476]" pageId="5" pageNumber="448">
<specimenCount box="[199,220,1448,1476]" pageId="5" pageNumber="448" type="female"></specimenCount>
(
<figureCitation box="[236,308,1446,1472]" captionStart="FIGURE 3" captionStartId="4.[151,250,1917,1941]" captionTargetBox="[173,1434,314,1890]" captionTargetId="figure-88@4.[151,1436,309,1894]" captionTargetPageId="4" captionText="FIGURE 3: Xeroderus conlei sp. n. A. ♀ holotype, dorsal view [MNHU]; B. ventral view; C. lateral view; D. Head &amp; thorax, in dorsal view; E. F. G. End of abdomen lateral, dorsal and ventral views. All photos by Oskar Conle." figureDoi="http://doi.org/10.5281/zenodo.7784538" httpUri="https://zenodo.org/record/7784538/files/figure.png" pageId="5" pageNumber="448">Fig. 3</figureCitation>
).
</emphasis>
Back of head with sparse tubercles/small spines, segment about as long as wide, eyes large; three distinct ocelli present, the hind pair larger. Antennae broken, therefore length not known, nor number of segments. However, basal segment broader than following segments, segment 2 only half length, and shorter than similar sized segments 36. Segments 7 about 1.5 x longer than preceding segments. Pronotum slightly shorter than head, with central impression between swollen upper and lower areas; black longitudinal, central line present along upper half, which has a pair of central spines. Mesonotum broader than head and pronotum, at its widest towards hind part, overall segment 1.7 x length of pronotum, with raised central area, armed with two large, broad paired spines posteriorly, two more large, paired spines and six smaller, stout spines, also swollen areas, in a distinct pattern; small lateral spines also present; on the outside rim there are a further four small spines each side with small lateral spines/ tubercles also present. Lateral margins of meso- and metathorax with series of lobes and few small spines (
<figureCitation box="[1334,1423,1771,1797]" captionStart="FIGURE 3" captionStartId="4.[151,250,1917,1941]" captionTargetBox="[173,1434,314,1890]" captionTargetId="figure-88@4.[151,1436,309,1894]" captionTargetPageId="4" captionText="FIGURE 3: Xeroderus conlei sp. n. A. ♀ holotype, dorsal view [MNHU]; B. ventral view; C. lateral view; D. Head &amp; thorax, in dorsal view; E. F. G. End of abdomen lateral, dorsal and ventral views. All photos by Oskar Conle." figureDoi="http://doi.org/10.5281/zenodo.7784538" httpUri="https://zenodo.org/record/7784538/files/figure.png" pageId="5" pageNumber="448">Fig. 3D</figureCitation>
). Ventral surface of head, thorax and abdomen smooth, lacking tubercles (
<figureCitation box="[966,1054,1807,1833]" captionStart="FIGURE 3" captionStartId="4.[151,250,1917,1941]" captionTargetBox="[173,1434,314,1890]" captionTargetId="figure-88@4.[151,1436,309,1894]" captionTargetPageId="4" captionText="FIGURE 3: Xeroderus conlei sp. n. A. ♀ holotype, dorsal view [MNHU]; B. ventral view; C. lateral view; D. Head &amp; thorax, in dorsal view; E. F. G. End of abdomen lateral, dorsal and ventral views. All photos by Oskar Conle." figureDoi="http://doi.org/10.5281/zenodo.7784538" httpUri="https://zenodo.org/record/7784538/files/figure.png" pageId="5" pageNumber="448">Fig. 3B</figureCitation>
). Metathorax slightly broader and longer than mesothorax and marginally shorter than length of median segment. Tegmina broad, oval; alae long, reaching almost end of 9
<superScript attach="left" box="[439,452,1878,1892]" fontSize="6" pageId="5" pageNumber="448">th</superScript>
abdominal segment. Veins with frequent whitish flecks of irregular length. Abdomen elongate, parallel-sided, with rounded lateral lobes on abdominal segments (hook-tipped on some), smaller on curved anal segment, which is lobed at sides of tip. Subgenital plate broad and rounded at tip, reaching beyond end of 9th abdominal segment (
<figureCitation box="[522,615,1987,2013]" captionStart="FIGURE 3" captionStartId="4.[151,250,1917,1941]" captionTargetBox="[173,1434,314,1890]" captionTargetId="figure-88@4.[151,1436,309,1894]" captionTargetPageId="4" captionText="FIGURE 3: Xeroderus conlei sp. n. A. ♀ holotype, dorsal view [MNHU]; B. ventral view; C. lateral view; D. Head &amp; thorax, in dorsal view; E. F. G. End of abdomen lateral, dorsal and ventral views. All photos by Oskar Conle." figureDoi="http://doi.org/10.5281/zenodo.7784538" httpUri="https://zenodo.org/record/7784538/files/figure.png" pageId="5" pageNumber="448">Fig. 3G</figureCitation>
); epiproct plate reaching end of abdomen, triangular incised at tip. On ventral surface, hind part of abdominal segment 9 has four dark spots (these dark spots or marks appear on former abdominal segments to a lesser extent), whereas base of anal segment has dark triangular side marks and central mark. Epiproct plate has four short dark flecks. Cerci broad, leaf-like, shorter than length of anal segment (
<figureCitation captionStart="FIGURE 3" captionStartId="4.[151,250,1917,1941]" captionTargetBox="[173,1434,314,1890]" captionTargetId="figure-88@4.[151,1436,309,1894]" captionTargetPageId="4" captionText="FIGURE 3: Xeroderus conlei sp. n. A. ♀ holotype, dorsal view [MNHU]; B. ventral view; C. lateral view; D. Head &amp; thorax, in dorsal view; E. F. G. End of abdomen lateral, dorsal and ventral views. All photos by Oskar Conle." figureDoi="http://doi.org/10.5281/zenodo.7784538" httpUri="https://zenodo.org/record/7784538/files/figure.png" pageId="6" pageNumber="449">Figs. 3EG</figureCitation>
). Legs moderately long and slender, reaching end of segment 7. Femora with well spread paired lobes and spines, some similarly shaped to abdominal lobes, whilst tibiae have few short spines and are broadened at tip. Tarsi of modest length, with all tarsi well over half length of tibiae.
</paragraph>
<paragraph blockId="6.[151,1437,151,573]" pageId="6" pageNumber="449">Measurements [mm]: Length of body: 95.6, head 5.8, antennae:&gt;13.0 (tips broken off), pronotum 5.2, mesonotum 8.7, metanotum 9.5, median segment: 9.9, tegmina 16.0, alae 68.5, profemora 18.3, mesofemora 12.1, metafemora 17.5, protibiae 15.3, mesotibiae 9.8, metatibiae 14.3, cerci 1.9.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="449" type="discussion">
<paragraph blockId="6.[151,1437,151,573]" pageId="6" pageNumber="449">
<emphasis bold="true" box="[199,262,475,501]" pageId="6" pageNumber="449">Note.</emphasis>
As is usual in phasmids, there can be variation in colour and spines within a species (
<bibRefCitation author="Brock, P. D. &amp; Buscher, T. H." pageId="6" pageNumber="449" refId="ref4764" refString="Brock, P. D. &amp; Buscher, T. H. (2022) Stick and leaf-insects of the world. Phasmids. N. A. P. Editions, Verrieres-le-Buisson, 612 pp." type="book" year="2022">Brock &amp; Büscher, 2022</bibRefCitation>
). Like
<taxonomicName authorityName="Gray" authorityYear="1837" box="[299,400,511,537]" class="Insecta" family="Phasmatidae" genus="Xeroderus" kingdom="Animalia" order="Phasmida" pageId="6" pageNumber="449" phylum="Arthropoda" rank="species" species="kirbyi">
<emphasis box="[299,400,511,537]" italics="true" pageId="6" pageNumber="449">X. kirbyi</emphasis>
</taxonomicName>
, there is likely to be geographic variation, perhaps dependant on habitat and foodplant selection.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7784542" ID-Zenodo-Dep="7784542" box="[151,681,1903,1927]" httpUri="https://zenodo.org/record/7784542/files/figure.png" pageId="6" pageNumber="449" startId="6.[151,250,1903,1927]" targetBox="[151,1436,597,1880]" targetPageId="6">
<paragraph blockId="6.[151,681,1903,1927]" box="[151,681,1903,1927]" pageId="6" pageNumber="449">
<emphasis bold="true" box="[151,277,1903,1927]" pageId="6" pageNumber="449">FIGURE 4:</emphasis>
Distribution map of
<taxonomicName box="[490,681,1903,1927]" class="Insecta" family="Phasmatidae" genus="Xeroderus" kingdom="Animalia" order="Phasmida" pageId="6" pageNumber="449" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[490,595,1903,1927]" italics="true" pageId="6" pageNumber="449">Xeroderus</emphasis>
species.
</taxonomicName>
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7784544" ID-Zenodo-Dep="7784544" httpUri="https://zenodo.org/record/7784544/files/figure.png" pageId="7" pageNumber="450" startId="7.[151,250,1470,1494]" targetBox="[151,1436,181,1446]" targetPageId="7">
<paragraph blockId="7.[151,1436,1470,1530]" pageId="7" pageNumber="450">
<emphasis bold="true" box="[151,280,1470,1494]" pageId="7" pageNumber="450">FIGURE 5:</emphasis>
Queensland, Golden Hole, Biggs Recreation area, Bartle Frere. Riverine lowland tropical rainforest habitat of
<taxonomicName authorityName="Gray" authorityYear="1837" box="[151,321,1506,1530]" class="Insecta" family="Phasmatidae" genus="Xeroderus" kingdom="Animalia" order="Phasmida" pageId="7" pageNumber="450" phylum="Arthropoda" rank="species" species="kirbyi">
<emphasis box="[151,321,1506,1530]" italics="true" pageId="7" pageNumber="450">Xeroderus kirbyi</emphasis>
</taxonomicName>
, photos by Ross Coupland.
</paragraph>
</caption>
<subSubSection pageId="7" pageNumber="450" type="etymology">
<paragraph blockId="7.[151,1437,1576,1963]" pageId="7" pageNumber="450">
<emphasis bold="true" box="[199,334,1576,1602]" pageId="7" pageNumber="450">Etymology.</emphasis>
Named after Oskar V. Conle (Duisburg,
<collectingCountry box="[809,918,1576,1602]" name="Germany" pageId="7" pageNumber="450">Germany</collectingCountry>
), for his outstanding contribution to phasmid research and efforts in re-organizing the phasmid collection of MNHU, in which the
<typeStatus box="[1117,1215,1612,1638]" pageId="7" pageNumber="450">holotype</typeStatus>
of this species was found.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="450" type="distribution">
<paragraph blockId="7.[151,1437,1576,1963]" pageId="7" pageNumber="450">
<emphasis bold="true" box="[199,443,1684,1710]" pageId="7" pageNumber="450">
Distribution (
<figureCitation box="[359,428,1684,1710]" captionStart="FIGURE 4" captionStartId="6.[151,250,1903,1927]" captionTargetBox="[151,1436,597,1880]" captionTargetId="figure-212@6.[151,1436,597,1880]" captionTargetPageId="6" captionText="FIGURE 4: Distribution map of Xeroderus species." figureDoi="http://doi.org/10.5281/zenodo.7784542" httpUri="https://zenodo.org/record/7784542/files/figure.png" pageId="7" pageNumber="450">Fig. 4</figureCitation>
).
</emphasis>
Only known from the
<typeStatus box="[692,740,1685,1711]" pageId="7" pageNumber="450">type</typeStatus>
locality in
<collectingCountry box="[863,1081,1684,1711]" name="Papua New Guinea" pageId="7" pageNumber="450">Papua New Guinea</collectingCountry>
. Sattelberg (Saddle Mountain) is a village on the Huon Peninsula, in
<collectingRegion box="[577,775,1720,1746]" country="Papua New Guinea" name="Morobe" pageId="7" pageNumber="450">Morobe Province</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="450" type="biology_ecology">
<paragraph blockId="7.[151,1437,1576,1963]" pageId="7" pageNumber="450">
<emphasis bold="true" box="[199,481,1756,1782]" pageId="7" pageNumber="450">Habitat and foodplants.</emphasis>
Presumably forest, probably up to
<quantity box="[867,938,1756,1783]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="450" unit="m" value="900.0">900 m</quantity>
, where the area was formerly part of German New
<collectingCountry box="[211,292,1792,1818]" name="Guinea-Bissau" pageId="7" pageNumber="450">Guinea</collectingCountry>
[D(eutsch) Neu-Guinea] when the specimen would have been collected in 1909. The foodplants are unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="450" type="discussion">
<paragraph blockId="7.[151,1437,1576,1963]" pageId="7" pageNumber="450">
<emphasis bold="true" box="[199,489,1864,1890]" pageId="7" pageNumber="450">Notes on the collector.</emphasis>
Professor Richard Gustav Neuhauss (18551915) was a German doctor and anthropologist, who was in German New
<collectingCountry box="[615,696,1900,1926]" name="Guinea-Bissau" pageId="7" pageNumber="450">Guinea</collectingCountry>
from 19081910. In 1909 he went with several missionaries on an expedition to Laewomba territory, then conducted research in the area of Huon Gulf (
<bibRefCitation author="Baumann, K. &amp; Klein, D. &amp; Apitzsch, W." box="[1107,1354,1936,1963]" pageId="7" pageNumber="450" refId="ref4729" refString="Baumann, K., Klein, D., Apitzsch, W. (2002) Biographisches Handbuch Deutsches-Neuguinea 1882 - 1922. 2 nd Edition. Baumann, Fassberg, 338 pp." type="book" year="2002">
Baumann
<emphasis box="[1223,1279,1936,1962]" italics="true" pageId="7" pageNumber="450">et al.</emphasis>
, 2002
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>