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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.328.5439" ID-GBIF-Dataset="dae4ece1-a5a6-4e1d-8121-53672e946970" ID-PMC="PMC3800822" ID-Pensoft-Pub="1313-2970-328-59" ID-PubMed="24146548" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-328-59" ModsDocOrigin="ZooKeys 328" ModsDocTitle="Further revision of the genus Megalopsalis (Opiliones, Neopilionidae), with the description of seven new species" checkinTime="1451246982575" checkinUser="pensoft" docAuthor="Taylor, Christopher K." docDate="2013" docId="5633AA9AABADB892A7DDE0AA1DA67631" docLanguage="en" docName="ZooKeys 328: 59-117" docOrigin="ZooKeys 328" docSource="http://dx.doi.org/10.3897/zookeys.328.5439" docTitle="Megalopsalis Roewer 1923" docType="treatment" docVersion="4" lastPageNumber="64" masterDocId="DB53CC7BFF8FA05DFFA3FFDBFFF4FFC0" masterDocTitle="Further revision of the genus Megalopsalis (Opiliones, Neopilionidae), with the description of seven new species" masterLastPageNumber="117" masterPageNumber="59" pageNumber="62" updateTime="1668156393602" updateUser="ExternalLinkService">
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<mods:title>Further revision of the genus Megalopsalis (Opiliones, Neopilionidae), with the description of seven new species</mods:title>
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<mods:namePart>Taylor, Christopher K.</mods:namePart>
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<mods:date>2013</mods:date>
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<treatment ID-GBIF-Taxon="152047436" LSID="urn:lsid:plazi:treatment:5633AA9AABADB892A7DDE0AA1DA67631" httpUri="http://treatment.plazi.org/id/5633AA9AABADB892A7DDE0AA1DA67631" lastPageId="6" lastPageNumber="64" pageId="3" pageNumber="62">
<subSubSection pageId="3" pageNumber="62" type="nomenclature">
<paragraph pageId="3" pageNumber="62">
<taxonomicName LSID="http://species-id.net/wiki/Megalopsalis" authority="Roewer, 1923" authorityName="Roewer" authorityYear="1923" class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Megalopsalis Roewer, 1923</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="62" type="reference_group">
<paragraph pageId="3" pageNumber="62">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Macropsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macropsalis" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Macropsalis</taxonomicName>
<normalizedToken originalValue="Sørensen">Sorensen</normalizedToken>
, 1886: 54-55 non Sclater 1866 -
<bibRefCitation author="Pocock, RI" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="43" pageNumber="102" pagination="392 - 413" title="On some new harvest-spiders of the order Opiliones from the southern continents." volume="1902" year="1903">Pocock 1903</bibRefCitation>
: 398;
<bibRefCitation author="Hogg, HR" journalOrPublisher="Transactions and Proceedings of the New Zealand Institute" pageId="43" pageNumber="102" pagination="273 - 283" title="Some New Zealand and Tasmanian Arachnidae." volume="42" year="1910">Hogg 1910</bibRefCitation>
: 277;
<bibRefCitation author="Roewer, CF" journalOrPublisher="Archiv fuer Naturgeschichte" pageId="43" pageNumber="102" pagination="1 - 106" title="Uebersicht der Genera der Subfamilie der Phalangiini der Opiliones Palpatores nebst Beschreibung einiger neuer Gattungen und Arten." volume="77" year="1911">Roewer 1911</bibRefCitation>
: 102,
<bibRefCitation author="Roewer, CF" journalOrPublisher="Abhandlungen aus dem Gebiete der Naturwissenschaften, herausgegeben von Naturwissenschaftlichen Verein in Hamburg" pageId="43" pageNumber="102" pagination="1 - 295" title="Revision der Opiliones Palpatores (= Opiliones Plagiostethi). II. Teil: Familie der Phalangiidae. (Subfamilien: Sclerosomini, Oligolophini, Phalangiini)." volume="20" year="1912">1912</bibRefCitation>
: 278.
</paragraph>
<paragraph pageId="3" pageNumber="62">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Megalopsalis</taxonomicName>
Roewer, 1923: 866 (replacement name for
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Macropsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macropsalis" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Macropsalis</taxonomicName>
<normalizedToken originalValue="Sørensen">Sorensen</normalizedToken>
) -
<bibRefCitation author="Forster, RR" journalOrPublisher="Records of the Dominion Museum" pageId="42" pageNumber="101" pagination="183 - 192" title="The genus Megalopsalis Roewer in New Zealand with keys to the New Zealand genera of Opiliones." volume="1" year="1944">Forster 1944</bibRefCitation>
: 184-185 (referring to material of
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Forsteropsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Forsteropsalis" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Forsteropsalis</taxonomicName>
Taylor, 2011);
<bibRefCitation author="Crawford, RL" journalOrPublisher="Burke Museum Contributions in Anthropology and Natural History" pageId="42" pageNumber="101" pagination="1 - 60" title="Catalogue of the genera and type species of the harvestman superfamily Phalangioidea (Arachnida)." volume="8" year="1992">Crawford 1992</bibRefCitation>
: 28, 29;
<bibRefCitation author="Taylor, CK" journalOrPublisher="Zootaxa" pageId="44" pageNumber="103" pagination="1 - 65" title="Revision of the genus Megalopsalis (Arachnida: Opiliones: Phalangioidea) in Australia and New Zealand and implications for phalangioid classification." volume="2773" year="2011">Taylor 2011</bibRefCitation>
: 31.
</paragraph>
<paragraph pageId="3" pageNumber="62">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Spinicrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinicrus" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Spinicrus</taxonomicName>
Forster, 1949: 63 syn. n.;
<bibRefCitation author="Hickman, VV" journalOrPublisher="Papers and Proceedings of the Royal Society of Tasmania" pageId="42" pageNumber="101" pagination="65 - 79" title="Some Tasmanian harvestmen of the sub-order Palpatores." volume="91" year="1957">Hickman 1957</bibRefCitation>
: 73;
<bibRefCitation author="Crawford, RL" journalOrPublisher="Burke Museum Contributions in Anthropology and Natural History" pageId="42" pageNumber="101" pagination="1 - 60" title="Catalogue of the genera and type species of the harvestman superfamily Phalangioidea (Arachnida)." volume="8" year="1992">Crawford 1992</bibRefCitation>
: 43.
</paragraph>
<paragraph pageId="3" pageNumber="62">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Hypomegalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypomegalopsalis" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Hypomegalopsalis</taxonomicName>
Taylor, 2011: 45 syn. n.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="62" type="type species">
<paragraph pageId="3" pageNumber="62">Type species.</paragraph>
<paragraph pageId="3" pageNumber="62">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Macropsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macropsalis serritarsus" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="serritarsus">Macropsalis serritarsus</taxonomicName>
<normalizedToken originalValue="Sørensen">Sorensen</normalizedToken>
, 1886 by monotypy.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="62" type="other included species">
<paragraph pageId="3" pageNumber="62">Other included species.</paragraph>
<paragraph pageId="3" pageNumber="62">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis serritarsus" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="serritarsus">Megalopsalis serritarsus</taxonomicName>
-species group:
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis epizephyros" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="epizephyros">Megalopsalis epizephyros</taxonomicName>
Taylor, 2011,
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis eremiotis" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="eremiotis">Megalopsalis eremiotis</taxonomicName>
Taylor, 2011,
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis hoggi" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="hoggi">Megalopsalis hoggi</taxonomicName>
Pocock, 1903,
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis pilliga" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="pilliga">Megalopsalis pilliga</taxonomicName>
Taylor, 2011.
</paragraph>
<paragraph pageId="3" pageNumber="62">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis leptekes" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="leptekes">Megalopsalis leptekes</taxonomicName>
-species group:
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis leptekes" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="leptekes">Megalopsalis leptekes</taxonomicName>
, 2011,
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis tanisphyros" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="tanisphyros">Megalopsalis tanisphyros</taxonomicName>
(Taylor, 2011), comb. n. (=
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Hypomegalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypomegalopsalis tanisphyros" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="tanisphyros">Hypomegalopsalis tanisphyros</taxonomicName>
).
</paragraph>
<paragraph pageId="3" pageNumber="62">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis minima" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="minima">Megalopsalis minima</taxonomicName>
-species group:
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis minima" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="minima">Megalopsalis minima</taxonomicName>
(Kauri, 1954), comb. n. (=
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Spinicrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinicrus minimum" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="minimum">Spinicrus minimum</taxonomicName>
),
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis porongorupensis" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="porongorupensis">Megalopsalis porongorupensis</taxonomicName>
(Kauri, 1954), comb. n. (=
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Spinicrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinicrus porongorupense" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="porongorupense">Spinicrus porongorupense</taxonomicName>
),
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis suffugiens" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="suffugiens">Megalopsalis suffugiens</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis walpolensis" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="walpolensis">Megalopsalis walpolensis</taxonomicName>
sp. n..
</paragraph>
<paragraph pageId="3" pageNumber="62">
Species not placed in species groups:
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis atrocidiana" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="atrocidiana">Megalopsalis atrocidiana</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis caeruleomontium" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="caeruleomontium">Megalopsalis caeruleomontium</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis coronata" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="coronata">Megalopsalis coronata</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis puerilis" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="puerilis">Megalopsalis puerilis</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis stewarti" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="stewarti">Megalopsalis stewarti</taxonomicName>
(Forster, 1949), comb. n. (=
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Spinicrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinicrus stewarti" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="stewarti">Spinicrus stewarti</taxonomicName>
),
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis sublucens" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="sublucens">Megalopsalis sublucens</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis tasmanica" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="tasmanica">Megalopsalis tasmanica</taxonomicName>
(Hogg, 1910), comb. n. (=
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis tasmanica" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="tasmanica">Pantopsalis tasmanica</taxonomicName>
),
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis thryptica" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="thryptica">Megalopsalis thryptica</taxonomicName>
(Hickman, 1957), comb. n. (=
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Spinicrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinicrus thrypticum" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="thrypticum">Spinicrus thrypticum</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="62" type="diagnosis">
<paragraph pageId="3" pageNumber="62">Diagnosis.</paragraph>
<paragraph pageId="3" pageNumber="62">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Megalopsalis</taxonomicName>
can be distinguished from all other genera of
<taxonomicName lsidName="" pageId="3" pageNumber="62" rank="subFamily" subFamily="Enantiobuninae">Enantiobuninae</taxonomicName>
by its male genital morphology, with the glans being relatively short, broad, distally flattened, and more or less subtriangular in ventral view (e.g. Fig. 3d). It can be further distinguished from
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Monoscutum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monoscutum" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Monoscutum</taxonomicName>
,
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Acihasta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acihasta" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Acihasta</taxonomicName>
,
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Templar" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Templar" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Templar</taxonomicName>
and
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Australiscutum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Australiscutum" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Australiscutum</taxonomicName>
by having the legs relatively long and thin, and the dorsum of the opisthosoma weakly sclerotised and unarmed (except
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis atrocidiana" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="atrocidiana">Megalopsalis atrocidiana</taxonomicName>
;
<bibRefCitation author="Forster, RR" journalOrPublisher="Records of the Auckland Institute and Museum" pageId="42" pageNumber="101" pagination="313 - 318" title="A new sub-family and species of New Zealand Opiliones." volume="3" year="1948">Forster 1948</bibRefCitation>
;
<bibRefCitation author="Taylor, CK" journalOrPublisher="Journal of Arachnology" pageId="44" pageNumber="103" pagination="176 - 179" title="A new species of Monoscutinae (Arachnida, Opiliones, Monoscutidae) from New Zealand, with a redescription of Monoscutum titirangiense." url="10.1636/H07-10SC.1" volume="36" year="2008 a">Taylor 2008a</bibRefCitation>
,
<bibRefCitation author="Taylor, CK" journalOrPublisher="Insect Systematics and Evolution" pageId="44" pageNumber="103" pagination="319 - 332" title="Australiscutum, a new genus of Monoscutidae (Arachnida: Opiliones) from eastern Australia, with the first record of asymmetrical chelicerae in Opiliones." url="10.1163/187631209X458367" volume="40" year="2009">2009</bibRefCitation>
).
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Pantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pantopsalis" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Pantopsalis</taxonomicName>
,
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Forsteropsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Forsteropsalis" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Forsteropsalis</taxonomicName>
,
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Neopantopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neopantopsalis" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Neopantopsalis</taxonomicName>
and
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Mangatangi" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mangatangi" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Mangatangi</taxonomicName>
differ from all
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Megalopsalis</taxonomicName>
species except
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis caeruleomontium" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="caeruleomontium">Megalopsalis caeruleomontium</taxonomicName>
by the presence of setae on the mobile finger of the chelicera (
<bibRefCitation author="Taylor, CK" journalOrPublisher="ZooKeys" pageId="44" pageNumber="103" pagination="59 - 73" title="Further notes on New Zealand Enantiobuninae (Opiliones, Neopilionidae), with the description of a new genus and two new species." url="10.3897/zookeys.263.4158" volume="263" year="2013">Taylor 2013</bibRefCitation>
: figs 1d, 2c).
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Tercentenarium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tercentenarium" order="Opiliones" pageId="3" pageNumber="62" phylum="Arthropoda" rank="genus">Tercentenarium</taxonomicName>
has males with a distinct frontodistal bulge on the chelicerae (
<bibRefCitation author="Taylor, CK" journalOrPublisher="Records of the Western Australian Museum" pageId="44" pageNumber="103" pagination="375 - 380" title="A new species of Monoscutidae (Arachnida, Opiliones) from the wheatbelt of Western Australia." volume="24" year="2008 b">Taylor 2008b</bibRefCitation>
: fig. 3), and females with a
<normalizedToken originalValue="keyhole">'keyhole'</normalizedToken>
-like emargination at the front of the genital operculum (
<bibRefCitation author="Taylor, CK" journalOrPublisher="Records of the Western Australian Museum" pageId="44" pageNumber="103" pagination="375 - 380" title="A new species of Monoscutidae (Arachnida, Opiliones) from the wheatbelt of Western Australia." volume="24" year="2008 b">Taylor 2008b</bibRefCitation>
: fig. 10).
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62" type="description">
<paragraph pageId="3" pageNumber="62">Description.</paragraph>
<paragraph lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62">
Ozopores usually large, oblong (small, round in
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis nigricans" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="nigricans">
<pageBreakToken pageId="4" pageNumber="63" start="start">Megalopsalis</pageBreakToken>
nigricans
</taxonomicName>
). Dorsum of opisthosoma unarmed (except with transverse rows of spines in
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis atrocidiana" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="atrocidiana">Megalopsalis atrocidiana</taxonomicName>
). Chelicera segment II denticulate or not; mobile finger usually closing tightly against finger of segment II, fingers bowed apart in larger males of
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis caeruleomontium" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="caeruleomontium">Megalopsalis caeruleomontium</taxonomicName>
. Pedipalp usually with patella shorter than tibia (slightly longer in
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis nigricans" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="nigricans">Megalopsalis nigricans</taxonomicName>
); apophysis present or absent on patella; claw with ventral tooth-row. Glans relatively short, broad, more or less subtriangular in ventral view, proximal section usually somewhat inflated dorsally (except in
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis nigricans" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="nigricans">Megalopsalis nigricans</taxonomicName>
); distal section more or less dorsoventrally flattened. Spiracle with reticulate or partially reticulate covering spines (reduced or absent in
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis minima" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="minima">Megalopsalis minima</taxonomicName>
-species group); lace tubercles present or absent.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="63" type="distribution">
<paragraph pageId="4" pageNumber="63">Distribution</paragraph>
<paragraph pageId="4" pageNumber="63">(Figs 1, 7). Southern and eastern Australia.</paragraph>
<caption pageId="4" pageNumber="63">
<paragraph pageId="4" pageNumber="63">
Figure 1. Locality map for
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis minima" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="minima">Megalopsalis minima</taxonomicName>
species-group in southern Western Australia: open square =
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis minima" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="minima">Megalopsalis minima</taxonomicName>
; solid circle =
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis porongorupensis" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="porongorupensis">Megalopsalis porongorupensis</taxonomicName>
; circle with dot =
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis walpolensis" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="walpolensis">Megalopsalis walpolensis</taxonomicName>
; solid triangle =
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis suffugiens" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="suffugiens">Megalopsalis suffugiens</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="64" pageId="4" pageNumber="63" type="comments">
<paragraph pageId="4" pageNumber="63">Comments.</paragraph>
<paragraph pageId="4" pageNumber="63">
The genus
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Spinicrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spinicrus" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">Spinicrus</taxonomicName>
as previously defined (
<bibRefCitation author="Forster, RR" journalOrPublisher="Memoirs of the National Museum of Victoria" pageId="42" pageNumber="101" pagination="59 - 89" title="Australian Opiliones." volume="16" year="1949">Forster 1949</bibRefCitation>
) is likely to be non-monophyletic with regard to both
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">Megalopsalis</taxonomicName>
and
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Hypomegalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypomegalopsalis" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">Hypomegalopsalis</taxonomicName>
, and lacks clear synapomorphies (see phylogenetic analysis below). In contrast, the clade uniting these three genera is characterised by a distinct penis morphology, and they are hence united into a single genus
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">Megalopsalis</taxonomicName>
. The species groups listed above are clades that were consistently recovered in the phylogenetic analysis under varying analytical conditions; those species not placed in groups did not form consistent subgeneric clades across all analyses. Members of the
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis serritarsus" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="serritarsus">Megalopsalis serritarsus</taxonomicName>
- and
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis leptekes" order="Opiliones" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="leptekes">Megalopsalis leptekes</taxonomicName>
-groups were described by
<bibRefCitation author="Taylor, CK" journalOrPublisher="Zootaxa" pageId="44" pageNumber="103" pagination="1 - 65" title="Revision of the genus Megalopsalis (Arachnida: Opiliones: Phalangioidea) in Australia and New Zealand and implications for phalangioid classification." volume="2773" year="2011">Taylor (2011)</bibRefCitation>
.
</paragraph>
<paragraph pageId="5" pageNumber="64">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis tasmanica" order="Opiliones" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="tasmanica">
<pageBreakToken pageId="5" pageNumber="64" start="start">Megalopsalis</pageBreakToken>
tasmanica
</taxonomicName>
and
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis thryptica" order="Opiliones" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="thryptica">Megalopsalis thryptica</taxonomicName>
were described in detail by
<bibRefCitation author="Hickman, VV" journalOrPublisher="Papers and Proceedings of the Royal Society of Tasmania" pageId="42" pageNumber="101" pagination="65 - 79" title="Some Tasmanian harvestmen of the sub-order Palpatores." volume="91" year="1957">Hickman (1957)</bibRefCitation>
, and so are not redescribed here. Both sexes of
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis tasmanica" order="Opiliones" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="tasmanica">Megalopsalis tasmanica</taxonomicName>
can be distinguished from other
<taxonomicName family="Neopilionidae" lsidName="" pageId="5" pageNumber="64" rank="family">Neopilionidae</taxonomicName>
by their distinctive elongate opisthosoma as illustrated by
<bibRefCitation author="Hickman, VV" journalOrPublisher="Papers and Proceedings of the Royal Society of Tasmania" pageId="42" pageNumber="101" pagination="65 - 79" title="Some Tasmanian harvestmen of the sub-order Palpatores." volume="91" year="1957">Hickman (1957</bibRefCitation>
: fig. 29); this distinction is even more pronounced in the female. See below under
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis stewarti" order="Opiliones" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="stewarti">Megalopsalis stewarti</taxonomicName>
for discussion of the distinction between this species and
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis thryptica" order="Opiliones" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="thryptica">Megalopsalis thryptica</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="65" pageId="5" pageNumber="64" type="key to males of species of megalopsalis">
<paragraph pageId="5" pageNumber="64">
Key to males of species of
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis" order="Opiliones" pageId="5" pageNumber="64" phylum="Arthropoda" rank="genus">Megalopsalis</taxonomicName>
</paragraph>
<paragraph lastPageId="6" lastPageNumber="65" pageId="5" pageNumber="64">
<table lastPageId="6" lastPageNumber="65" pageId="5" pageNumber="64">
<tr pageId="5" pageNumber="64">
<td colspan="1" pageId="5" pageNumber="64" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis hoggi" order="Opiliones" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="hoggi">Megalopsalis hoggi</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="64">
<td colspan="1" pageId="5" pageNumber="64" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis pilliga" order="Opiliones" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="pilliga">Megalopsalis pilliga</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="64">
<td colspan="1" pageId="5" pageNumber="64" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis epizephyros" order="Opiliones" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="epizephyros">Megalopsalis epizephyros</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="64">
<td colspan="1" pageId="5" pageNumber="64" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis serritarsus" order="Opiliones" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="serritarsus">Megalopsalis serritarsus</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="64">
<td colspan="1" pageId="5" pageNumber="64" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis eremiotis" order="Opiliones" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="eremiotis">Megalopsalis eremiotis</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="64">
<td colspan="1" pageId="5" pageNumber="64" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis leptekes" order="Opiliones" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="leptekes">Megalopsalis leptekes</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="64">
<td colspan="1" pageId="5" pageNumber="64" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis tanisphyros" order="Opiliones" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="tanisphyros">Megalopsalis tanisphyros</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="64">
<td colspan="1" pageId="5" pageNumber="64" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis atrocidiana" order="Opiliones" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="atrocidiana">Megalopsalis atrocidiana</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="64">
<td colspan="1" pageId="5" pageNumber="64" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis nigricans" order="Opiliones" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="nigricans">Megalopsalis nigricans</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="64">
<td colspan="1" pageId="5" pageNumber="64" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis caeruleomontium" order="Opiliones" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="caeruleomontium">Megalopsalis caeruleomontium</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="64">
<td colspan="1" pageId="5" pageNumber="64" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis tasmanica" order="Opiliones" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="tasmanica">Megalopsalis tasmanica</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="64">
<td colspan="1" pageId="5" pageNumber="64" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis sublucens" order="Opiliones" pageId="5" pageNumber="64" phylum="Arthropoda" rank="species" species="sublucens">Megalopsalis sublucens</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="65">
<td colspan="1" pageId="6" pageNumber="65" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis thryptica" order="Opiliones" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="thryptica">
<pageBreakToken pageId="6" pageNumber="65" start="start">Megalopsalis</pageBreakToken>
thryptica
</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="65">
<td colspan="1" pageId="6" pageNumber="65" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis stewarti" order="Opiliones" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="stewarti">Megalopsalis stewarti</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="65">
<td colspan="1" pageId="6" pageNumber="65" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis minima" order="Opiliones" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="minima">Megalopsalis minima</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="65">
<td colspan="1" pageId="6" pageNumber="65" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis suffugiens" order="Opiliones" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="suffugiens">Megalopsalis suffugiens</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="65">
<td colspan="1" pageId="6" pageNumber="65" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis porongorupensis" order="Opiliones" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="porongorupensis">Megalopsalis porongorupensis</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="65">
<td colspan="1" pageId="6" pageNumber="65" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis walpolensis" order="Opiliones" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="walpolensis">Megalopsalis walpolensis</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="65">
<td colspan="1" pageId="6" pageNumber="65" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis coronata" order="Opiliones" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="coronata">Megalopsalis coronata</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="65">
<td colspan="1" pageId="6" pageNumber="65" rowspan="1">
<taxonomicName class="Arachnida" family="Neopilionidae" genus="Megalopsalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megalopsalis puerilis" order="Opiliones" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="puerilis">Megalopsalis puerilis</taxonomicName>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>