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<document id="88D38D8DE23BB0831F9E74BCD43DD0E2" ID-CLB-Dataset="102843" ID-DOI="10.11646/zootaxa.5168.2.5" ID-GBIF-Dataset="16df2448-1e2c-4410-ac7b-af3623ff401f" ID-ISSN="1175-5326" ID-Zenodo-Dep="6877267" ID-ZooBank="36831FDB-E9BE-4255-82B9-BFFCDCD40098" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658420700313" checkinUser="plazi" docAuthor="Zhao, Ming-Zhi &amp; Zorn, Carsten" docDate="2022" docId="03B887EBDF7BFFB517AC03170A44FE74" docLanguage="en" docName="zootaxa.5168.2.5.pdf" docOrigin="Zootaxa 5168 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anomala kanshireiensis Zorn &amp; Zhao 2022, new species" docType="treatment" docUuid="75B007A5-92AA-4FA5-A599-8B0980C08944" docUuidSource="ZooBank" docVersion="6" lastPageNumber="187" masterDocId="FF81FF93DF72FFB9173B045C0836FF81" masterDocTitle="Contribution to the genus Anomala Samouelle, 1819 (Coleoptera: Scarabaeidae Rutelinae) of China and adjacent regions. Part II: six new species from Taiwan and Hainan" masterLastPageNumber="195" masterPageNumber="175" pageNumber="184" updateTime="1699361553354" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F9E48FFC41A8D910AC746E8B36C5FA83" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C358393E760678DB4B6A853B9BCAB0F7">
<mods:title id="A9E14705C53C39AAD132818745FDCADA">Contribution to the genus Anomala Samouelle, 1819 (Coleoptera: Scarabaeidae Rutelinae) of China and adjacent regions. Part II: six new species from Taiwan and Hainan</mods:title>
</mods:titleInfo>
<mods:name id="6D4DB409F52924D562C30D2268D9F034" type="personal">
<mods:role id="3AB7711167042ADA3AE82C3CF8424EDA">
<mods:roleTerm id="C41008F0FCC7C8599357964D418D4E7D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="03E9F756BC7244D7E66A19976CE4DD6D">Zhao, Ming-Zhi</mods:namePart>
<mods:nameIdentifier id="D2D5DB900D3A6AF980EA6A1A32304667" type="ORCID">0000-0002-3158-6177</mods:nameIdentifier>
<mods:affiliation id="19DEF98F1AF02522257859F1810A7EE9">College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, P. R. China. zhaomzhai @ 126. com; https: // orcid. org / 0000 - 0002 - 3158 - 6177</mods:affiliation>
</mods:name>
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<mods:role id="054989F4B8835489AE4BB1E3067BB1BD">
<mods:roleTerm id="3208ED55F25CBDEEA8F0B95A7382E01B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5BA99D8E8C3CEB9BFA968EB47262E0C2">Zorn, Carsten</mods:namePart>
<mods:affiliation id="34E0FAFEC4976245BF1979E7B9570287">Rostocker Strasse 1 a, Gnoien, 17179, Germany.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="C9FEEDD6676E7A8B96DD2D580D1A6048">text</mods:typeOfResource>
<mods:relatedItem id="5E2565EF7B676EC8E0A89089B33A3F2F" type="host">
<mods:titleInfo id="6293C8AE009FF9914339C103EBFF06DB">
<mods:title id="13D81AA3A7EDDD14CC7E16B9F18D442A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3EF30BAAE947C6B5067FA6543E3FB6B4">
<mods:date id="2E934DBC2873459A847DD134685DA047">2022</mods:date>
<mods:detail id="C704C7B1A74AF61394C3F00DB5B36CE0" type="pubDate">
<mods:number id="659046ADDD2796FCF6260A63A5A350F6">2022-07-21</mods:number>
</mods:detail>
<mods:detail id="2E4049EE6B427994AA4BDA6414BDAB20" type="volume">
<mods:number id="115C66364E269F61401411CF48FCCD59">5168</mods:number>
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<mods:number id="362705B71BC2CCB79561CDB525EC438F">2</mods:number>
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<mods:start id="06B90C51520BC8EEFFED0BA6B9A5FD35">175</mods:start>
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<mods:classification id="510B56E1F8A6D72782887D3BDD502016">journal article</mods:classification>
<mods:identifier id="BEBA625D1F930441E9AD6323B4013F7F" type="CLB-Dataset">102843</mods:identifier>
<mods:identifier id="85894FCF77DE16D05C058DDFDDD84011" type="DOI">10.11646/zootaxa.5168.2.5</mods:identifier>
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<mods:identifier id="49F253AAABFB11C16C267EB162417736" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="03B887EBDF7BFFB517AC03170A44FE74" ID-DOI="http://doi.org/10.5281/zenodo.6877215" ID-GBIF-Taxon="197841319" ID-Zenodo-Dep="6877215" LSID="urn:lsid:zoobank.org:act:75B007A5-92AA-4FA5-A599-8B0980C08944" httpUri="http://treatment.plazi.org/id/03B887EBDF7BFFB517AC03170A44FE74" lastPageId="12" lastPageNumber="187" pageId="9" pageNumber="184">
<subSubSection id="C30B6576DF7BFFB017AC03170C87F8E7" box="[151,1201,1867,1894]" pageId="9" pageNumber="184" type="nomenclature">
<paragraph id="8BAE36FDDF7BFFB017AC03170C87F8E7" blockId="9.[151,1201,1867,1930]" box="[151,1201,1867,1894]" pageId="9" pageNumber="184">
<heading id="D0E68191DF7BFFB017AC03170C87F8E7" bold="true" box="[151,1201,1867,1894]" fontSize="11" level="1" pageId="9" pageNumber="184" reason="1">
<emphasis id="B965EAEFDF7BFFB017AC03170C87F8E7" bold="true" box="[151,1201,1867,1894]" pageId="9" pageNumber="184">
<taxonomicName id="4C114D7EDF7BFFB017AC03170A5BF8E7" authority="Zorn &amp; Zhao" authorityName="Zorn &amp; Zhao" authorityYear="2022" box="[151,621,1867,1894]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="kanshireiensis" status="sp. nov.">
<emphasis id="B965EAEFDF7BFFB017AC0317098DF8E7" bold="true" box="[151,443,1867,1894]" italics="true" pageId="9" pageNumber="184">Anomala kanshireiensis</emphasis>
Zorn &amp; Zhao
</taxonomicName>
,
<taxonomicNameLabel id="A2565794DF7BFFB0154103100B3CF8E7" box="[634,778,1868,1894]" pageId="9" pageNumber="184" rank="species">new species</taxonomicNameLabel>
[关T岭异fflŵoi/ṞTĂ條ŵff]
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30B6576DF7BFFB017AC032C09ECF80B" box="[151,474,1904,1930]" pageId="9" pageNumber="184" type="description">
<paragraph id="8BAE36FDDF7BFFB017AC032C09ECF80B" blockId="9.[151,1201,1867,1930]" box="[151,474,1904,1930]" pageId="9" pageNumber="184">
(
<figureCitation id="132A2A78DF7BFFB017A4032C08D9F80B" box="[159,239,1904,1930]" captionStart="FIGURE 6" captionStartId="10.[151,250,1964,1988]" captionTargetBox="[171,1416,402,1917]" captionTargetId="figure-91@10.[151,1435,381,1941]" captionTargetPageId="10" captionText="FIGURE 6. Habitus of Anomala kanshireiensis Zorn &amp; Zhao, new species. A–C, holotype; D–F, female paratype. A, D, dorsal view; B, E, lateral view; C, F, ventral view. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877279" httpUri="https://zenodo.org/record/6877279/files/figure.png" pageId="9" pageNumber="184">
Figs.
<geoCoordinate id="EE25503ADF7BFFB017E4032C08D9F80B" box="[223,239,1904,1930]" degrees="6, 8" direction="east" orientation="longitude" pageId="9" pageNumber="184" precision="5555" value="6.8">6</geoCoordinate>
</figureCitation>
,
<figureCitation id="132A2A78DF7BFFB017C0032C090EF80B" box="[251,312,1904,1930]" captionStart="FIGURE 8" captionStartId="14.[151,250,1881,1905]" captionTargetBox="[151,1426,181,1857]" captionTargetId="figure-21@14.[151,1435,181,1857]" captionTargetPageId="14" captionText="FIGURE 8. Head of Anomala spp. A, A. linwenhsini Zhao &amp; Zorn, new species, holotype; B, ditto, female paratype; C, A. wutaiensis Zhao &amp; Zorn, new species, holotype; D, ditto, female paratype; E, A. kanshireiensis Zorn &amp; Zhao, new species, holotype; F, ditto, female paratype; G, A. inclinata Zhao &amp; Zorn, new species, holotype; H, ditto, female paratype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877285" httpUri="https://zenodo.org/record/6877285/files/figure.png" pageId="9" pageNumber="184">8E–F</figureCitation>
,
<figureCitation id="132A2A78DF7BFFB01678032C09E4F80B" box="[323,466,1904,1930]" captionStart="FIGURE 9" captionStartId="15.[151,250,1583,1607]" captionTargetBox="[163,1411,190,1552]" captionTargetId="figure-18@15.[151,1435,181,1560]" captionTargetPageId="15" captionText="FIGURE 9. Male genitalia (A–M) and everted endophallus (N–Q) of Anomala spp. A–C, A. linwenhsini Zhao &amp; Zorn, new species, holotype; D–G, A. wutaiensis Zhao &amp; Zorn, new species, holotype; H–J, N–O, A. kanshireiensis Zorn &amp; Zhao, new species, holotype; K–M, P–Q, A. inclinata Zhao &amp; Zorn, new species, holotype. A, D, H, K, dorsal view; B, F, I, L, N, P, right lateral view; E, O, Q, left lateral view; C, G, J, M, ventral view. Sc 1, scale bar for A–M = 2 mm; Sc 2, scale bar for N–Q = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877289" httpUri="https://zenodo.org/record/6877289/files/figure.png" pageId="9" pageNumber="184">9H–J, 9N–O</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30B6576DF7BFFB317AC03E40980FF78" lastPageId="10" lastPageNumber="185" pageId="9" pageNumber="184" type="materials_examined">
<paragraph id="8BAE36FDDF7BFFB317AC03E40980FF78" blockId="9.[151,1437,1976,2039]" lastBlockId="10.[151,1437,151,357]" lastPageId="10" lastPageNumber="185" pageId="9" pageNumber="184">
<emphasis id="B965EAEFDF7BFFB017AC03E40973F853" bold="true" box="[151,325,1976,2002]" pageId="9" pageNumber="184">Type material.</emphasis>
<materialsCitation id="3B793CA0DF7BFFB0167603E408EDF877" ID-GBIF-Occurrence="3864839309" collectionCode="MFNB" county="Formosa" location="Kanshirei" municipality="Sauter" pageId="9" pageNumber="184" specimenCount="1" specimenCount-male="1" stateProvince="Tainan" typeStatus="holotype">
<emphasis id="B965EAEFDF7BFFB0167603E409F5F853" bold="true" box="[333,451,1976,2002]" pageId="9" pageNumber="184">
<typeStatus id="54AA885FDF7BFFB0167603E40988F853" box="[333,446,1976,2002]" pageId="9" pageNumber="184" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D17FD74DF7BFFB016F103E509E9F852" box="[458,479,1977,2003]" pageId="9" pageNumber="184" type="male">♂</specimenCount>
(
<collectionCode id="ED00AE38DF7BFFB016D503E40A75F853" box="[494,579,1976,2002]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="184">MFNB</collectionCode>
),
<collectingCounty id="62CF4E71DF7BFFB0156C03E50A8AF852" box="[599,700,1977,2003]" pageId="9" pageNumber="184">Formosa</collectingCounty>
,
<collectingMunicipality id="6BCAAC87DF7BFFB015FC03E40B26F853" box="[711,784,1976,2002]" pageId="9" pageNumber="184">Sauter</collectingMunicipality>
,
<location id="8ECE6026DF7BFFB0142703E40BBFF853" LSID="urn:lsid:plazi:treatment:03B887EBDF7BFFB517AC03170A44FE74:8ECE6026DF7BFFB0142703E40BBFF853" box="[796,905,1976,2002]" county="Formosa" municipality="Sauter" name="Kanshirei" pageId="9" pageNumber="184" stateProvince="Tainan">Kanshirei</location>
[= Guanziling in
<collectingRegion id="49D5F81FDF7BFFB0136A03E40CABF853" box="[1105,1181,1976,2002]" country="Taiwan" name="Tainan" pageId="9" pageNumber="184">Tainan</collectingRegion>
City,
<emphasis id="B965EAEFDF7BFFB013DA03E50D03F851" bold="true" box="[1249,1333,1977,2000]" pageId="9" pageNumber="184">ṞTĂ</emphasis>
], 908. V. 19-27
</materialsCitation>
.
<materialsCitation id="3B793CA0DF7BFFB017DC03800B79F877" ID-GBIF-Occurrence="3864839329" box="[231,847,2012,2039]" collectionCode="MFNB" county="Formosa" location="Kanshirei" municipality="Sauter" pageId="9" pageNumber="184" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Tainan" typeStatus="paratype">
<emphasis id="B965EAEFDF7BFFB017DC0380095EF877" bold="true" box="[231,360,2012,2038]" pageId="9" pageNumber="184">
<typeStatus id="54AA885FDF7BFFB017DC03800955F877" box="[231,355,2012,2038]" pageId="9" pageNumber="184" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D17FD74DF7BFFB0164B038009A2F877" box="[368,404,2012,2038]" pageId="9" pageNumber="184" type="male">1♂</specimenCount>
,
<specimenCount id="9D17FD74DF7BFFB01699038009EFF877" box="[418,473,2012,2038]" pageId="9" pageNumber="184" type="female">3♀♀</specimenCount>
(
<collectionCode id="ED00AE38DF7BFFB016D203800A08F877" box="[489,574,2012,2038]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="184">MFNB</collectionCode>
), same data as holotype
</materialsCitation>
;
<materialsCitation id="3B793CA0DF7BFFB31461038008E8FF30" ID-GBIF-Occurrence="3864839319" collectionCode="MFNB" county="Kanshirei" lastPageId="10" lastPageNumber="185" location="Formosa" municipality="Sauter" pageId="9" pageNumber="184" specimenCount="3" specimenCount-male="3" stateProvince="Tainan" typeStatus="paratype">
<specimenCount id="9D17FD74DF7BFFB0146103800BA4F877" box="[858,914,2012,2038]" pageId="9" pageNumber="184" type="male">3♂♂</specimenCount>
(
<collectionCode id="ED00AE38DF7BFFB0149903800BC1F877" box="[930,1015,2012,2038]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="184">MFNB</collectionCode>
),
<location id="8ECE6026DF7BFFB0133003810C46F876" LSID="urn:lsid:plazi:treatment:03B887EBDF7BFFB517AC03170A44FE74:8ECE6026DF7BFFB0133003810C46F876" box="[1035,1136,2013,2039]" county="Kanshirei" municipality="Sauter" name="Formosa" pageId="9" pageNumber="184" stateProvince="Tainan">Formosa</location>
,
<collectingMunicipality id="6BCAAC87DF7BFFB0134703800CF3F877" box="[1148,1221,2012,2038]" pageId="9" pageNumber="184">Sauter</collectingMunicipality>
,
<collectingCounty id="62CF4E71DF7BFFB013EA03800D77F877" box="[1233,1345,2012,2038]" pageId="9" pageNumber="184">Kanshirei</collectingCounty>
, 908. V. 17-24
</materialsCitation>
.;
<materialsCitation id="3B793CA0DF78FFB317CA04CB0B6EFF30" ID-GBIF-Occurrence="3864839324" box="[241,856,151,177]" collectionCode="MFNB" county="Kanshirei" location="Formosa" municipality="Sauter" pageId="10" pageNumber="185" specimenCount="1" specimenCount-male="1" stateProvince="Tainan" typeStatus="paratype">
<specimenCount id="9D17FD74DF78FFB317CA04CB0922FF30" box="[241,276,151,177]" pageId="10" pageNumber="185" type="male">1♂</specimenCount>
(
<collectionCode id="ED00AE38DF78FFB3161E04CB094CFF30" box="[293,378,151,177]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="10" pageNumber="185">MFNB</collectionCode>
),
<location id="8ECE6026DF78FFB316B404CB09C2FF30" LSID="urn:lsid:plazi:treatment:03B887EBDF7BFFB517AC03170A44FE74:8ECE6026DF78FFB316B404CB09C2FF30" box="[399,500,151,177]" county="Kanshirei" municipality="Sauter" name="Formosa" pageId="10" pageNumber="185" stateProvince="Tainan">Formosa</location>
,
<collectingMunicipality id="6BCAAC87DF78FFB3153A04CB0A7CFF30" box="[513,586,151,177]" pageId="10" pageNumber="185">Sauter</collectingMunicipality>
,
<collectingCounty id="62CF4E71DF78FFB3156304CB0AFEFF30" box="[600,712,151,177]" pageId="10" pageNumber="185">Kanshirei</collectingCounty>
, 908. V. 1-9
</materialsCitation>
;
<materialsCitation id="3B793CA0DF78FFB3145D04CB08E9FF54" ID-GBIF-Occurrence="3864839332" collectionCode="MFNB" county="Kanshirei" location="Formosa" municipality="Sauter" pageId="10" pageNumber="185" specimenCount="1" specimenCount-male="1" stateProvince="Tainan" typeStatus="paratype">
<specimenCount id="9D17FD74DF78FFB3145D04CB0BBFFF30" box="[870,905,151,177]" pageId="10" pageNumber="185" type="male">1♂</specimenCount>
(
<collectionCode id="ED00AE38DF78FFB314A104CB0BD9FF30" box="[922,1007,151,177]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="10" pageNumber="185">MFNB</collectionCode>
),
<location id="8ECE6026DF78FFB3133E04CB0C5CFF30" LSID="urn:lsid:plazi:treatment:03B887EBDF7BFFB517AC03170A44FE74:8ECE6026DF78FFB3133E04CB0C5CFF30" box="[1029,1130,151,177]" county="Kanshirei" municipality="Sauter" name="Formosa" pageId="10" pageNumber="185" stateProvince="Tainan">Formosa</location>
,
<collectingMunicipality id="6BCAAC87DF78FFB3134C04CB0CF6FF30" box="[1143,1216,151,177]" pageId="10" pageNumber="185">Sauter</collectingMunicipality>
,
<collectingCounty id="62CF4E71DF78FFB313F504CB0D08FF30" box="[1230,1342,151,177]" pageId="10" pageNumber="185">Kanshirei</collectingCounty>
, 908. V. 19-26
</materialsCitation>
.;
<materialsCitation id="3B793CA0DF78FFB317CB04E70C1AFF54" ID-GBIF-Occurrence="3864839322" box="[240,1068,187,213]" collectingDate="1908-06-02" collectingDateMax="1908-06-14" collectingDateMin="1908-06-02" collectionCode="SMNS" collectorName="H. Sauter" country="Taiwan" location="Kanshirei" pageId="10" pageNumber="185" specimenCount="4" specimenCount-female="1" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="9D17FD74DF78FFB317CB04E7091CFF54" box="[240,298,187,213]" pageId="10" pageNumber="185" type="male">3♂♂</specimenCount>
,
<specimenCount id="9D17FD74DF78FFB3160304E7096DFF54" box="[312,347,187,213]" pageId="10" pageNumber="185" type="female">1♀</specimenCount>
(
<collectionCode id="ED00AE38DF78FFB3165704E7098BFF54" box="[364,445,187,213]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="10" pageNumber="185">SMNS</collectionCode>
),
<collectingCountry id="F306766DDF78FFB316E904E70A77FF54" box="[466,577,187,213]" name="Taiwan" pageId="10" pageNumber="185">TAIWAN</collectingCountry>
,
<location id="8ECE6026DF78FFB3157504E70A88FF54" LSID="urn:lsid:plazi:treatment:03B887EBDF7BFFB517AC03170A44FE74:8ECE6026DF78FFB3157504E70A88FF54" box="[590,702,187,213]" country="Taiwan" name="Kanshirei" pageId="10" pageNumber="185">Kanshirei</location>
,
<date id="FFAF103DDF78FFB315F104E70B49FF54" box="[714,895,187,213]" pageId="10" pageNumber="185" value="1908-06-02" valueMax="1908-06-14" valueMin="1908-06-02">
<collectingDate id="EFEBE9D5DF78FFB315F104E70B49FF54" box="[714,895,187,213]" pageId="10" pageNumber="185" value="1908-06-02" valueMax="1908-06-14" valueMin="1908-06-02">2. - 14. 6. 1908</collectingDate>
</date>
,
<collectorName id="26E4532BDF78FFB314B004E70BC1FF54" box="[907,1015,187,213]" pageId="10" pageNumber="185">H. Sauter</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B793CA0DF78FFB3130204E70984FF78" ID-GBIF-Occurrence="3864839315" collectionCode="HNHM" country="Taiwan" county="Kanshirei" location="Formosa" municipality="Sauter" pageId="10" pageNumber="185" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D17FD74DF78FFB3130204E70C6AFF54" box="[1081,1116,187,213]" pageId="10" pageNumber="185" type="male">1♂</specimenCount>
(
<collectionCode id="ED00AE38DF78FFB3135604E70CFEFF54" box="[1133,1224,187,213]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="185">HNHM</collectionCode>
),
<location id="8ECE6026DF78FFB313E604E70D74FF54" LSID="urn:lsid:plazi:treatment:03B887EBDF7BFFB517AC03170A44FE74:8ECE6026DF78FFB313E604E70D74FF54" box="[1245,1346,187,213]" country="Taiwan" county="Kanshirei" municipality="Sauter" name="Formosa" pageId="10" pageNumber="185">Formosa</location>
,
<collectingMunicipality id="6BCAAC87DF78FFB3127404E70DAEFF54" box="[1359,1432,187,213]" pageId="10" pageNumber="185">Sauter</collectingMunicipality>
,
<collectingCounty id="62CF4E71DF78FFB317AC04830931FF78" box="[151,263,223,249]" pageId="10" pageNumber="185">Kanshirei</collectingCounty>
, 908. V-1-2-14
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B6576DF78FFB217FC055E0D7EF838" lastPageId="11" lastPageNumber="186" pageId="10" pageNumber="185" type="description">
<paragraph id="8BAE36FDDF78FFB317FC055E0D7BFE9C" blockId="10.[151,1437,151,357]" box="[199,1357,258,285]" pageId="10" pageNumber="185">
<emphasis id="B965EAEFDF78FFB317FC055E0A38FE9C" bold="true" box="[199,526,258,285]" pageId="10" pageNumber="185">
Description.
<typeStatus id="54AA885FDF78FFB3165B055F09FBFE9C" box="[352,461,259,285]" pageId="10" pageNumber="185" type="holotype">Holotype</typeStatus>
male
</emphasis>
(
<figureCitation id="132A2A78DF78FFB31527055F0A97FE9C" box="[540,673,259,285]" captionStart="FIGURE 6" captionStartId="10.[151,250,1964,1988]" captionTargetBox="[171,1416,402,1917]" captionTargetId="figure-91@10.[151,1435,381,1941]" captionTargetPageId="10" captionText="FIGURE 6. Habitus of Anomala kanshireiensis Zorn &amp; Zhao, new species. A–C, holotype; D–F, female paratype. A, D, dorsal view; B, E, lateral view; C, F, ventral view. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877279" httpUri="https://zenodo.org/record/6877279/files/figure.png" pageId="10" pageNumber="185">Figs. 6A–C</figureCitation>
,
<figureCitation id="132A2A78DF78FFB31597055F0AF8FE9C" box="[684,718,259,285]" captionStart="FIGURE 8" captionStartId="14.[151,250,1881,1905]" captionTargetBox="[151,1426,181,1857]" captionTargetId="figure-21@14.[151,1435,181,1857]" captionTargetPageId="14" captionText="FIGURE 8. Head of Anomala spp. A, A. linwenhsini Zhao &amp; Zorn, new species, holotype; B, ditto, female paratype; C, A. wutaiensis Zhao &amp; Zorn, new species, holotype; D, ditto, female paratype; E, A. kanshireiensis Zorn &amp; Zhao, new species, holotype; F, ditto, female paratype; G, A. inclinata Zhao &amp; Zorn, new species, holotype; H, ditto, female paratype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877285" httpUri="https://zenodo.org/record/6877285/files/figure.png" pageId="10" pageNumber="185">8E</figureCitation>
,
<figureCitation id="132A2A78DF78FFB315E1055F0B13FE9C" box="[730,805,259,285]" captionStart="FIGURE 9" captionStartId="15.[151,250,1583,1607]" captionTargetBox="[163,1411,190,1552]" captionTargetId="figure-18@15.[151,1435,181,1560]" captionTargetPageId="15" captionText="FIGURE 9. Male genitalia (A–M) and everted endophallus (N–Q) of Anomala spp. A–C, A. linwenhsini Zhao &amp; Zorn, new species, holotype; D–G, A. wutaiensis Zhao &amp; Zorn, new species, holotype; H–J, N–O, A. kanshireiensis Zorn &amp; Zhao, new species, holotype; K–M, P–Q, A. inclinata Zhao &amp; Zorn, new species, holotype. A, D, H, K, dorsal view; B, F, I, L, N, P, right lateral view; E, O, Q, left lateral view; C, G, J, M, ventral view. Sc 1, scale bar for A–M = 2 mm; Sc 2, scale bar for N–Q = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877289" httpUri="https://zenodo.org/record/6877289/files/figure.png" pageId="10" pageNumber="185">9K–M</figureCitation>
). Body length:
<quantity id="4CE99B18DF78FFB314E9055F0C01FE9C" box="[978,1079,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="10" pageNumber="185" unit="mm" value="11.8">11.8 mm</quantity>
, greatest width:
<quantity id="4CE99B18DF78FFB313CB055F0D7FFE9C" box="[1264,1353,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="10" pageNumber="185" unit="mm" value="6.7">6.7 mm</quantity>
.
</paragraph>
<paragraph id="8BAE36FDDF78FFB317FC057A09B3FEE4" blockId="10.[151,1437,151,357]" pageId="10" pageNumber="185">
<emphasis id="B965EAEFDF78FFB317FC057A09F4FEC0" bold="true" box="[199,450,294,321]" pageId="10" pageNumber="185">General appearance.</emphasis>
Body shape elongated ovoid, weakly convex. Entirely dark reddish brown, with weak green metallic sheen.
</paragraph>
<caption id="DF6E6675DF78FFB317AC03F00B3BF869" ID-DOI="http://doi.org/10.5281/zenodo.6877279" ID-Zenodo-Dep="6877279" httpUri="https://zenodo.org/record/6877279/files/figure.png" pageId="10" pageNumber="185" startId="10.[151,250,1964,1988]" targetBox="[171,1416,402,1917]" targetPageId="10">
<paragraph id="8BAE36FDDF78FFB317AC03F00B3BF869" blockId="10.[151,1436,1964,2024]" pageId="10" pageNumber="185">
<emphasis id="B965EAEFDF78FFB317AC03F00925F845" bold="true" box="[151,275,1964,1988]" pageId="10" pageNumber="185">FIGURE 6.</emphasis>
Habitus of
<taxonomicName id="4C114D7EDF78FFB316B203F00B3FF845" authority="Zorn &amp; Zhao" authorityName="Zorn &amp; Zhao" authorityYear="2022" box="[393,777,1964,1988]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="kanshireiensis" status="sp. nov.">
<emphasis id="B965EAEFDF78FFB316B203F00A4DF845" box="[393,635,1964,1988]" italics="true" pageId="10" pageNumber="185">Anomala kanshireiensis</emphasis>
Zorn &amp; Zhao
</taxonomicName>
,
<emphasis id="B965EAEFDF78FFB3142803F00BB9F845" bold="true" box="[787,911,1964,1988]" pageId="10" pageNumber="185">
<taxonomicNameLabel id="A2565794DF78FFB3142803F00BB9F845" box="[787,911,1964,1988]" pageId="10" pageNumber="185" rank="species">new species</taxonomicNameLabel>
</emphasis>
. A–C, holotype; D–F, female paratype. A, D, dorsal view; B, E, lateral view; C, F, ventral view. Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="8BAE36FDDF79FFB217FC04CB0A53FEC0" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="186">
<emphasis id="B965EAEFDF79FFB217FC04CB0931FF30" bold="true" box="[199,263,151,177]" pageId="11" pageNumber="186">Head</emphasis>
(
<figureCitation id="132A2A78DF79FFB2162C04CB095AFF30" box="[279,364,151,177]" captionStart="FIGURE 8" captionStartId="14.[151,250,1881,1905]" captionTargetBox="[151,1426,181,1857]" captionTargetId="figure-21@14.[151,1435,181,1857]" captionTargetPageId="14" captionText="FIGURE 8. Head of Anomala spp. A, A. linwenhsini Zhao &amp; Zorn, new species, holotype; B, ditto, female paratype; C, A. wutaiensis Zhao &amp; Zorn, new species, holotype; D, ditto, female paratype; E, A. kanshireiensis Zorn &amp; Zhao, new species, holotype; F, ditto, female paratype; G, A. inclinata Zhao &amp; Zorn, new species, holotype; H, ditto, female paratype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877285" httpUri="https://zenodo.org/record/6877285/files/figure.png" pageId="11" pageNumber="186">Fig. 8E</figureCitation>
). Clypeus subtrapezoidal, anterior corner broadly rounded; anterior margin somewhat rounded and strongly reflexed; surface densely rugopunctate. Frontoclypeal suture distinct. An inverted triangular area at anterior half of frons densely rugopunctate, other portions of frons and vertex with irregularly distributed dense small punctures. Antennal club longer than antennomeres 2–6 combined. Inner margin of eye with several moderately long setae (mostly worn in the
<typeStatus id="54AA885FDF79FFB216C9057B0A6FFEC0" box="[498,601,295,321]" pageId="11" pageNumber="186" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BAE36FDDF79FFB217FC051709F6FE74" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="186">
<emphasis id="B965EAEFDF79FFB217FC05170971FEE4" bold="true" box="[199,327,331,357]" pageId="11" pageNumber="186">Pronotum.</emphasis>
Sides gently arched and convergent anteriad. Anterior angle acute and protruding, posterior angle obtuse, rounded off. Basal marginal line interrupted before scutellum, other marginal lines complete. With dense small punctures, punctures being partly coalescent at marginal portions; with a very weakly indicated longitudinal medial furrow not reaching the base. Lateral margin with several long setae, two moderately long to long semierect setae on disc of pronotum.
</paragraph>
<paragraph id="8BAE36FDDF79FFB217FC05A20BF4FDBC" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="186">
<emphasis id="B965EAEFDF79FFB217FC05A20970FD99" bold="true" box="[199,326,510,536]" pageId="11" pageNumber="186">Scutellum.</emphasis>
Subtriangular, lateral margin arched. Margins with polished appearance, disc with irregularly distributed moderately large punctures. Disc with two semierect long setae.
</paragraph>
<paragraph id="8BAE36FDDF79FFB217FC061B0B0CFCB8" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="186">
<emphasis id="B965EAEFDF79FFB217FC061B092CFDE0" bold="true" box="[199,282,583,609]" pageId="11" pageNumber="186">Elytra.</emphasis>
Intervals strongly convex; strial punctures dense to very dense, annulated and large, punctures of secondary striae unequally spaced; the whole surface with scattered minute punctures. Interstices I to III each with a secondary stria; interstice I (the subsutural interstice) broadest, with secondary stria shortly doubled in basal fourth; the other secondary striae reaching the level of apical protuberance and usually interrupted, secondary stria of interstice II obsolete between level of scutellar apex to midway of elytron. Primary costae without additional striae. Humeral umbone and apical protuberance moderately bulging. Lateral carina indistinct. Epipleura with a row of moderately dense long setae. Marginal membrane complete.
</paragraph>
<paragraph id="8BAE36FDDF79FFB217FC071E0D67FCDC" blockId="11.[151,1437,151,2013]" box="[199,1361,834,861]" pageId="11" pageNumber="186">
<emphasis id="B965EAEFDF79FFB217FC071E095DFCDD" bold="true" box="[199,363,834,860]" pageId="11" pageNumber="186">Propygidium.</emphasis>
With dense transverse punctures, usually coalescent into transverse striolation; glabrous.
</paragraph>
<paragraph id="8BAE36FDDF79FFB217FC073A0B0BFC48" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="186">
<emphasis id="B965EAEFDF79FFB217FC073A0974FC01" bold="true" box="[199,322,870,896]" pageId="11" pageNumber="186">Pygidium.</emphasis>
Distinctly bulging, posterior margin protruding. Disc with dense ovoid large punctures, becoming striate near all margins; punctures encircling greatest tumidity. With several short setae distributed along posterior margin, posterior margin with a row of long setae near apex.
</paragraph>
<paragraph id="8BAE36FDDF79FFB217FC078F0D53FBD8" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="186">
<emphasis id="B965EAEFDF79FFB217FC078F09D8FC6D" bold="true" box="[199,494,978,1005]" pageId="11" pageNumber="186">Ventral thoracic surface.</emphasis>
Hypomeron with dense coalescent striolation and moderately dense long setae. Collar of mesosternite transversely striolate with dense recumbent short setae. Other portions of mesosternite with small punctures and somewhat rugopunctate, with dense short setae. Ventral metathoracic surface with dense annulated moderately large punctures and dense, rather long setae; with minute punctures and almost glabrous medially.
</paragraph>
<paragraph id="8BAE36FDDF79FFB217FC003E08D8FB68" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="186">
<emphasis id="B965EAEFDF79FFB217FC003E09F1FBFD" bold="true" box="[199,455,1122,1148]" pageId="11" pageNumber="186">Abdominal ventrites.</emphasis>
Ventrites 1–2 and anterior half of 3 weakly carinate laterally. Ventrites 2–5 with irregularly distributed small punctures, punctures coalescent into striolation in ventrite 6. Ventrites 2–5 each with a transverse row of sparse and moderately long setae; ventrite 6 with a complete row of long setae along posterior margin.
</paragraph>
<paragraph id="8BAE36FDDF79FFB217FC00AF0992F9AC" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="186">
<emphasis id="B965EAEFDF79FFB217FC00AF0930FA8C" bold="true" box="[199,262,1267,1293]" pageId="11" pageNumber="186">Legs.</emphasis>
Protibia tridentate, apical and second tooth acute at apex, proximal tooth small; apical tooth extending to level of halfway of protarsomere 2. Inner spur inserted at the level of the indention between proximal and second tooth. Inner protarsal claw and outer mesotarsal claw split apically, lower branches longer and wider, the lower branch of inner protarsal claw concave internobasally. Each tarsomere 5 with an internomedial protuberance. Outer metatarsal claw slightly longer than inner one. Mesofemoral surface with transverse rows of long setae, several irregular rows between two regular rows, one at anterior margin and another situated behind middle, the row at posterior margin usually with shorter setae. Metafemoral surface with three transverse rows of sparse long setae, the first row at anterior margin, a second row situated behind middle with robuster setae, and a third row at posterior margin with short setae.
</paragraph>
<paragraph id="8BAE36FDDF79FFB217FC026B0D4BF9D0" blockId="11.[151,1437,151,2013]" box="[199,1405,1590,1617]" pageId="11" pageNumber="186">
<emphasis id="B965EAEFDF79FFB217FC026B094FF9D1" bold="true" box="[199,377,1590,1617]" pageId="11" pageNumber="186">Male genitalia.</emphasis>
See
<figureCitation id="132A2A78DF79FFB21694026B0A16F9D0" box="[431,544,1591,1617]" captionStart="FIGURE 9" captionStartId="15.[151,250,1583,1607]" captionTargetBox="[163,1411,190,1552]" captionTargetId="figure-18@15.[151,1435,181,1560]" captionTargetPageId="15" captionText="FIGURE 9. Male genitalia (A–M) and everted endophallus (N–Q) of Anomala spp. A–C, A. linwenhsini Zhao &amp; Zorn, new species, holotype; D–G, A. wutaiensis Zhao &amp; Zorn, new species, holotype; H–J, N–O, A. kanshireiensis Zorn &amp; Zhao, new species, holotype; K–M, P–Q, A. inclinata Zhao &amp; Zorn, new species, holotype. A, D, H, K, dorsal view; B, F, I, L, N, P, right lateral view; E, O, Q, left lateral view; C, G, J, M, ventral view. Sc 1, scale bar for A–M = 2 mm; Sc 2, scale bar for N–Q = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877289" httpUri="https://zenodo.org/record/6877289/files/figure.png" pageId="11" pageNumber="186">Fig. 9H–J</figureCitation>
. Parameres asymmetric, right paramere longer. Ventral plate flat, apex acute.
</paragraph>
<paragraph id="8BAE36FDDF79FFB217FC02070D7EF838" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="186">
<emphasis id="B965EAEFDF79FFB217FC020709A8F9F4" bold="true" box="[199,414,1627,1653]" pageId="11" pageNumber="186">
<typeStatus id="54AA885FDF79FFB217FC02070977F9F4" box="[199,321,1627,1653]" pageId="11" pageNumber="186" type="paratype">Paratypes</typeStatus>
. Males.
</emphasis>
Body length: 11.5–13.0 mm, greatest width:
<quantity id="4CE99B18DF79FFB2149302060C02F9F4" box="[936,1076,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="7.5" metricValueMin="6.5" pageId="11" pageNumber="186" unit="mm" value="7.0" valueMax="7.5" valueMin="6.5">6.5–7.5 mm</quantity>
. External features and shape of parameres consistent. Semierect longer setae on scutellum and disc of pronotum are only present in few males but might be worn off in other specimens.
<emphasis id="B965EAEFDF79FFB2156402FF0AF7F93C" bold="true" box="[607,705,1699,1725]" pageId="11" pageNumber="186">Females</emphasis>
(
<figureCitation id="132A2A78DF79FFB215EF02FF0B61F93C" box="[724,855,1699,1725]" captionStart="FIGURE 6" captionStartId="10.[151,250,1964,1988]" captionTargetBox="[171,1416,402,1917]" captionTargetId="figure-91@10.[151,1435,381,1941]" captionTargetPageId="10" captionText="FIGURE 6. Habitus of Anomala kanshireiensis Zorn &amp; Zhao, new species. A–C, holotype; D–F, female paratype. A, D, dorsal view; B, E, lateral view; C, F, ventral view. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877279" httpUri="https://zenodo.org/record/6877279/files/figure.png" pageId="11" pageNumber="186">Figs. 6D–F</figureCitation>
,
<figureCitation id="132A2A78DF79FFB2145E02FF0BB2F93C" box="[869,900,1699,1725]" captionStart="FIGURE 8" captionStartId="14.[151,250,1881,1905]" captionTargetBox="[151,1426,181,1857]" captionTargetId="figure-21@14.[151,1435,181,1857]" captionTargetPageId="14" captionText="FIGURE 8. Head of Anomala spp. A, A. linwenhsini Zhao &amp; Zorn, new species, holotype; B, ditto, female paratype; C, A. wutaiensis Zhao &amp; Zorn, new species, holotype; D, ditto, female paratype; E, A. kanshireiensis Zorn &amp; Zhao, new species, holotype; F, ditto, female paratype; G, A. inclinata Zhao &amp; Zorn, new species, holotype; H, ditto, female paratype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877285" httpUri="https://zenodo.org/record/6877285/files/figure.png" pageId="11" pageNumber="186">8F</figureCitation>
). Body length: 12.3–13.0 mm, greatest width: 7.0–
<quantity id="4CE99B18DF79FFB217F2029B0916F960" box="[201,288,1735,1761]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="11" pageNumber="186" unit="mm" value="7.9">7.9 mm</quantity>
. Generally similar to male. Antennal club almost equally as long as antennomeres 2–6 combined. Sides of pronotum more strongly curved. Pygidium more bulging, shorter than in male. Protibia and protarsomeres thinner than in male; apical protibial tooth tongue-shaped, almost extends to level of apex of protarsomere 3; the other two teeth blunt at apex, proximal tooth indistinct; inner spur situated at the level of the proximal tooth; internomedial protuberance of protarsomere 5 small; upper and lower branches of inner protarsal claw and outer mesotarsal claw almost equal in length. Ventral metathoracic surface less setose than in male. Semierect longer setae on scutellum and disc of pronotum are only present in the minority of females but might be worn off in some specimens.
</paragraph>
</subSubSection>
<subSubSection id="C30B6576DF79FFB517FC039E0C96FE2C" lastPageId="12" lastPageNumber="187" pageId="11" pageNumber="186" type="diagnosis">
<paragraph id="8BAE36FDDF79FFB517FC039E0C96FE2C" blockId="11.[151,1437,151,2013]" lastBlockId="12.[151,1437,150,501]" lastPageId="12" lastPageNumber="187" pageId="11" pageNumber="186">
<emphasis id="B965EAEFDF79FFB217FC039E09FBF85D" bold="true" box="[199,461,1986,2012]" pageId="11" pageNumber="186">Differential diagnosis.</emphasis>
<taxonomicName id="4C114D7EDF79FFB216E8039F0B4CF85C" authority="Zorn &amp; Zhao" authorityName="Zorn &amp; Zhao" authorityYear="2022" box="[467,890,1987,2013]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="kanshireiensis" status="sp. nov.">
<emphasis id="B965EAEFDF79FFB216E8039F0AE8F85C" box="[467,734,1987,2013]" italics="true" pageId="11" pageNumber="186">Anomala kanshireiensis</emphasis>
Zorn &amp; Zhao
</taxonomicName>
,
<emphasis id="B965EAEFDF79FFB214BF039F0C3BF85D" bold="true" box="[900,1037,1986,2013]" pageId="11" pageNumber="186">
<taxonomicNameLabel id="A2565794DF79FFB214BF039F0C3BF85D" box="[900,1037,1986,2013]" pageId="11" pageNumber="186" rank="species">new species</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="4C114D7EDF79FFB513E1039F08D8FF30" authority="Zhao &amp; Zorn" authorityName="Zhao &amp; Zorn" authorityYear="2022" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" lastPageId="12" lastPageNumber="187" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="inclinata" status="sp. nov.">
<emphasis id="B965EAEFDF79FFB213E1039F0D6BF85C" box="[1242,1373,1987,2013]" italics="true" pageId="11" pageNumber="186">A. inclinata</emphasis>
Zhao &amp; Zorn
</taxonomicName>
,
<emphasis id="B965EAEFDF7EFFB517C004CB09B3FF31" bold="true" box="[251,389,150,177]" pageId="12" pageNumber="187">
<taxonomicNameLabel id="A2565794DF7EFFB517C004CB09B3FF31" box="[251,389,150,177]" pageId="12" pageNumber="187" rank="species">new species</taxonomicNameLabel>
</emphasis>
. Above all, the general body color is reddish brown in
<taxonomicName id="4C114D7EDF7EFFB514C704CB0C8DFF30" authorityName="Zorn &amp; Zhao" authorityYear="2022" box="[1020,1211,151,177]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="kanshireiensis">
<emphasis id="B965EAEFDF7EFFB514C704CB0C8DFF30" box="[1020,1211,151,177]" italics="true" pageId="12" pageNumber="187">A. kanshireiensis</emphasis>
</taxonomicName>
but blackish brown in
<taxonomicName id="4C114D7EDF7EFFB5178E04E7090CFF54" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[181,314,187,213]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="inclinata">
<emphasis id="B965EAEFDF7EFFB5178E04E7090CFF54" box="[181,314,187,213]" italics="true" pageId="12" pageNumber="187">A. inclinata</emphasis>
</taxonomicName>
. The interrupted secondary stria of interstice II is more complete than in
<taxonomicName id="4C114D7EDF7EFFB513BB04E70D32FF54" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[1152,1284,187,213]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="inclinata">
<emphasis id="B965EAEFDF7EFFB513BB04E70D32FF54" box="[1152,1284,187,213]" italics="true" pageId="12" pageNumber="187">A. inclinata</emphasis>
</taxonomicName>
. The aedeagi of the two species are similar in shape. The left paramere is shorter and has a less curved apex compared to
<taxonomicName id="4C114D7EDF7EFFB51266048308D3FE9C" authorityName="Zhao &amp; Zorn" authorityYear="2022" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="inclinata">
<emphasis id="B965EAEFDF7EFFB51266048308D3FE9C" italics="true" pageId="12" pageNumber="187">A. inclinata</emphasis>
</taxonomicName>
. The left paramere is placed slightly above the right paramere in lateral view (parameres are overlapping in
<taxonomicName id="4C114D7EDF7EFFB517AC057B092CFEC0" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[151,282,295,321]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="inclinata">
<emphasis id="B965EAEFDF7EFFB517AC057B092CFEC0" box="[151,282,295,321]" italics="true" pageId="12" pageNumber="187">A. inclinata</emphasis>
</taxonomicName>
). The apex of the ventral plate is acute in
<taxonomicName id="4C114D7EDF7EFFB515D0057B0B9EFEC0" authorityName="Zorn &amp; Zhao" authorityYear="2022" box="[747,936,295,321]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="kanshireiensis">
<emphasis id="B965EAEFDF7EFFB515D0057B0B9EFEC0" box="[747,936,295,321]" italics="true" pageId="12" pageNumber="187">A. kanshireiensis</emphasis>
</taxonomicName>
but arched in
<taxonomicName id="4C114D7EDF7EFFB5137D057B0CFFFEC0" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[1094,1225,295,321]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="inclinata">
<emphasis id="B965EAEFDF7EFFB5137D057B0CFFFEC0" box="[1094,1225,295,321]" italics="true" pageId="12" pageNumber="187">A. inclinata</emphasis>
</taxonomicName>
. The endophallites of the two species show considerable differences: the two dorsal endophallites are spaced with one being relatively long in
<taxonomicName id="4C114D7EDF7EFFB517D50533099AFE08" authorityName="Zorn &amp; Zhao" authorityYear="2022" box="[238,428,367,393]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="kanshireiensis">
<emphasis id="B965EAEFDF7EFFB517D50533099AFE08" box="[238,428,367,393]" italics="true" pageId="12" pageNumber="187">A. kanshireiensis</emphasis>
</taxonomicName>
(
<figureCitation id="132A2A78DF7EFFB5168705330A0CFE08" box="[444,570,367,393]" captionStart="FIGURE 9" captionStartId="15.[151,250,1583,1607]" captionTargetBox="[163,1411,190,1552]" captionTargetId="figure-18@15.[151,1435,181,1560]" captionTargetPageId="15" captionText="FIGURE 9. Male genitalia (A–M) and everted endophallus (N–Q) of Anomala spp. A–C, A. linwenhsini Zhao &amp; Zorn, new species, holotype; D–G, A. wutaiensis Zhao &amp; Zorn, new species, holotype; H–J, N–O, A. kanshireiensis Zorn &amp; Zhao, new species, holotype; K–M, P–Q, A. inclinata Zhao &amp; Zorn, new species, holotype. A, D, H, K, dorsal view; B, F, I, L, N, P, right lateral view; E, O, Q, left lateral view; C, G, J, M, ventral view. Sc 1, scale bar for A–M = 2 mm; Sc 2, scale bar for N–Q = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877289" httpUri="https://zenodo.org/record/6877289/files/figure.png" pageId="12" pageNumber="187">Fig. 9N–O</figureCitation>
), they are adjacent and both short in
<taxonomicName id="4C114D7EDF7EFFB514E205330C6BFE08" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[985,1117,367,393]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="inclinata">
<emphasis id="B965EAEFDF7EFFB514E205330C6BFE08" box="[985,1117,367,393]" italics="true" pageId="12" pageNumber="187">A. inclinata</emphasis>
</taxonomicName>
(
<figureCitation id="132A2A78DF7EFFB5135705330CD3FE08" box="[1132,1253,367,393]" captionStart="FIGURE 9" captionStartId="15.[151,250,1583,1607]" captionTargetBox="[163,1411,190,1552]" captionTargetId="figure-18@15.[151,1435,181,1560]" captionTargetPageId="15" captionText="FIGURE 9. Male genitalia (A–M) and everted endophallus (N–Q) of Anomala spp. A–C, A. linwenhsini Zhao &amp; Zorn, new species, holotype; D–G, A. wutaiensis Zhao &amp; Zorn, new species, holotype; H–J, N–O, A. kanshireiensis Zorn &amp; Zhao, new species, holotype; K–M, P–Q, A. inclinata Zhao &amp; Zorn, new species, holotype. A, D, H, K, dorsal view; B, F, I, L, N, P, right lateral view; E, O, Q, left lateral view; C, G, J, M, ventral view. Sc 1, scale bar for A–M = 2 mm; Sc 2, scale bar for N–Q = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6877289" httpUri="https://zenodo.org/record/6877289/files/figure.png" pageId="12" pageNumber="187">Fig. 9P–Q</figureCitation>
); the ventral endophallite is a curved spine in
<taxonomicName id="4C114D7EDF7EFFB516D605CF0A9DFE2C" authorityName="Zorn &amp; Zhao" authorityYear="2022" box="[493,683,403,429]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="kanshireiensis">
<emphasis id="B965EAEFDF7EFFB516D605CF0A9DFE2C" box="[493,683,403,429]" italics="true" pageId="12" pageNumber="187">A. kanshireiensis</emphasis>
</taxonomicName>
, while it is a serrated sclerite in
<taxonomicName id="4C114D7EDF7EFFB5132C05CF0CACFE2C" authorityName="Zhao &amp; Zorn" authorityYear="2022" box="[1047,1178,403,429]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="187" phylum="Arthropoda" rank="species" species="inclinata">
<emphasis id="B965EAEFDF7EFFB5132C05CF0CACFE2C" box="[1047,1178,403,429]" italics="true" pageId="12" pageNumber="187">A. inclinata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B6576DF7EFFB517FC05EB0D37FE50" box="[199,1281,439,465]" pageId="12" pageNumber="187" type="etymology">
<paragraph id="8BAE36FDDF7EFFB517FC05EB0D37FE50" blockId="12.[151,1437,150,501]" box="[199,1281,439,465]" pageId="12" pageNumber="187">
<emphasis id="B965EAEFDF7EFFB517FC05EB0978FE50" bold="true" box="[199,334,439,465]" pageId="12" pageNumber="187">Etymology.</emphasis>
The specific epithet is alluding to the historical name of the
<typeStatus id="54AA885FDF7EFFB514CE05EB0C13FE50" box="[1013,1061,439,465]" pageId="12" pageNumber="187">type</typeStatus>
locality, Kanshirei.
</paragraph>
</subSubSection>
<subSubSection id="C30B6576DF7EFFB517FC05860A44FE74" box="[199,626,474,501]" pageId="12" pageNumber="187" type="distribution">
<paragraph id="8BAE36FDDF7EFFB517FC05860A44FE74" blockId="12.[151,1437,150,501]" box="[199,626,474,501]" pageId="12" pageNumber="187">
<emphasis id="B965EAEFDF7EFFB517FC05860957FE75" bold="true" box="[199,353,474,500]" pageId="12" pageNumber="187">Distribution.</emphasis>
<collectingCountry id="F306766DDF7EFFB5165C05870A32FE74" box="[359,516,475,501]" name="Taiwan Island" pageId="12" pageNumber="187">Taiwan Island</collectingCountry>
(Tainan).
</paragraph>
</subSubSection>
</treatment>
</document>
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