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<document id="1FDA6A4C2460C1ADEDB8FF20373A9E44" ID-DOI="10.5281/zenodo.5401374" ID-ISSN="1638-9387" ID-Zenodo-Dep="5401374" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630334492992" checkinUser="marcus" docAuthor="Ivanenko, Viatcheslav N. &amp; Defaye, Danielle" docDate="2004" docId="03B887D70357FF94E62B05C2CB165AA3" docLanguage="en" docName="Zoosystema.2004.26.1.49-64.pdf" docOrigin="Zoosystema 26 (1)" docStyle="DocumentStyle:64A3174DE33E974D4EB6573A0A793B12.4:Zoosystema.2000-2008.journal_article.type2" docStyleId="64A3174DE33E974D4EB6573A0A793B12" docStyleName="Zoosystema.2000-2008.journal_article.type2" docStyleVersion="4" docTitle="Heptnerina confusa Ivanenko &amp; Defaye 2004, n. sp." docType="treatment" docVersion="4" lastPageNumber="57" masterDocId="FF81FFAF0355FF9CE562056BCA1E5C26" masterDocTitle="A new genus and species of deep-sea cyclopoid (Crustacea, Copepoda, Cyclopinidae) from the Mid-Atlantic Ridge (Azores Triple Junction, Lucky Strike)" masterLastPageNumber="64" masterPageNumber="49" pageNumber="51" updateTime="1699232485843" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="EBF4B40124C280CDB2433DCA1FD2705D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8475431565D398053128C376842F55D3">
<mods:title id="43BB45B4D4398D23FAB7C8850B45ADE4">A new genus and species of deep-sea cyclopoid (Crustacea, Copepoda, Cyclopinidae) from the Mid-Atlantic Ridge (Azores Triple Junction, Lucky Strike)</mods:title>
</mods:titleInfo>
<mods:name id="22DF2D9D1E399F90F713778CC91F6A38" type="personal">
<mods:role id="E4E5293F8486052FA6A5A6E0901A0920">
<mods:roleTerm id="7CE3FD347BCB3DD98C6A9ADF83E4A9CC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0162902B70092E0AB666EABCFE710BD3">Ivanenko, Viatcheslav N.</mods:namePart>
<mods:affiliation id="C578E4A84B1C5CBFC2F65EB1EB526A8E">Department of Invertebrate Zoology, Biological Faculty, Moscow State University, Moscow 119899 (Russia) Ivanenko @ soil. msu. ru</mods:affiliation>
<mods:nameIdentifier id="FE5D8D2877F983A533CDC3ED1A29EC12" type="email">vanenko@soil.msu.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="B286E089FC6B9BFB59D23FA5AD21CAD2" type="personal">
<mods:role id="8D7755D4EB7CB5EAA732A003B4AC7832">
<mods:roleTerm id="99DD77B9E2B8331ABF4E86F93DDFB59D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D1ADD15F7D0070D02E6B1026954308DC">Defaye, Danielle</mods:namePart>
<mods:affiliation id="EA460F5FA5A088EB4D37C04943E1FFBF">Muséum national d’Histoire naturelle, Département Milieux et Peuplements aquatiques, USM 0403, 61 rue Buffon, F- 75231 Paris cedex 05 (France) ddefaye @ mnhn. fr.</mods:affiliation>
<mods:nameIdentifier id="C0DF51027ED59467769F220758E86E62" type="email">ddefaye@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="51BC4F211EFDC567607A70EBE63A3296">text</mods:typeOfResource>
<mods:relatedItem id="32F58F41C012C1D11A0318EA32A60E3A" type="host">
<mods:titleInfo id="21C652B91D43E304ED359DE108BBD98D">
<mods:title id="4FB35310A2B262372277B3BDB56DA5BD">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="C2FAA5DAC3AF04827029ED5BBDEE674C">
<mods:date id="4C2553BC44058CACB20B6D44107E6708">2004</mods:date>
<mods:detail id="48A0476A812A142C5E8A70D0781B079E" type="volume">
<mods:number id="A8BA3AB7D05870091265635F67486521">26</mods:number>
</mods:detail>
<mods:detail id="75773117D6FBF186753E64AAC251F303" type="issue">
<mods:number id="60FEE02B5AE525B8AC0CD25D2DD68447">1</mods:number>
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<mods:start id="BDF21F6BD30A1275A69A5167B2BD6378">49</mods:start>
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</mods:extent>
</mods:part>
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<mods:classification id="BC16E030EFC03A0532C83CFD3F8AF42D">journal article</mods:classification>
<mods:identifier id="33307CD0D47ABDEB12286C29755CAAD3" type="DOI">10.5281/zenodo.5401374</mods:identifier>
<mods:identifier id="360B31B22A582AE0BF6FFC8878BCB544" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="49165DC9587AC2E44B7E097D5EA1A754" type="Zenodo-Dep">5401374</mods:identifier>
<mods:identifier id="44DC948D217B66CA9A591000E63AC629" type="ZooBank">E70B6263-B5B2-4AB2-9184-07ED438AAE32</mods:identifier>
</mods:mods>
<treatment id="03B887D70357FF94E62B05C2CB165AA3" LSID="urn:lsid:plazi:treatment:03B887D70357FF94E62B05C2CB165AA3" httpUri="http://treatment.plazi.org/id/03B887D70357FF94E62B05C2CB165AA3" lastPageId="8" lastPageNumber="57" pageId="2" pageNumber="51">
<subSubSection id="C30B654A0357FF9EE62B05C2CE7E5CE5" box="[841,1120,169,196]" pageId="2" pageNumber="51" type="nomenclature">
<paragraph id="8BAE36C10357FF9EE62B05C2CE7E5CE5" blockId="2.[841,1120,169,227]" box="[841,1120,169,196]" pageId="2" pageNumber="51">
<heading id="D0E681AD0357FF9EE62B05C2CE7E5CE5" box="[841,1120,169,196]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="51" reason="2">
<taxonomicName id="4C114D420357FF9EE62B05C2CE005CE2" authority="Ivanenko &amp; Defaye, 2004" authorityName="Ivanenko &amp; Defaye" authorityYear="2004" box="[841,1054,169,196]" class="Hexanauplia" family="Cyclopinidae" genus="Heptnerina" kingdom="Animalia" order="Cyclopoida" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="confusa" status="sp. nov.">
<emphasis id="B965EAD30357FF9EE62B05C2CE005CE2" bold="true" box="[841,1054,169,196]" italics="true" pageId="2" pageNumber="51">Heptnerina confusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25657A80357FF9EE14705C2CE7E5CE5" box="[1061,1120,169,195]" pageId="2" pageNumber="51" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30B654A0357FF9EE6C205A2CE175CC5" box="[928,1033,201,227]" pageId="2" pageNumber="51" type="description">
<paragraph id="8BAE36C10357FF9EE6C205A2CE175CC5" blockId="2.[841,1120,169,227]" box="[928,1033,201,227]" pageId="2" pageNumber="51">
(
<figureCitation id="132A2A440357FF9EE6C505A2C9F85CC5" box="[935,998,201,227]" captionStart="FIG" captionStartId="3.[151,162,1262,1279]" captionTargetBox="[189,1200,169,1227]" captionTargetId="figure-135@3.[186,1178,157,1232]" captionTargetPageId="3" captionText="FIG. 1. — Heptnerina confusa n. gen., n. sp., holotype; A, habitus, dorsal; B, habitus, lateral; C, genital double somite, dorsal; D, genital double somite, ventral; E, gonoporal area and P6, lateral. Scale bars: A, B, 0.2 mm; C-E, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401376" httpUri="https://zenodo.org/record/5401376/files/figure.png" pageId="2" pageNumber="51">Figs 1</figureCitation>
-8)
</paragraph>
</subSubSection>
<subSubSection id="C30B654A0357FF9EE7A8047ACE015EA4" pageId="2" pageNumber="51" type="materials_examined">
<paragraph id="8BAE36C10357FF9EE7A8047AC97D5DD2" blockId="2.[714,1248,273,733]" pageId="2" pageNumber="51">
<materialsCitation id="3B793C9C0357FF9EE7A8047ACEC25D0F" box="[714,1244,273,297]" collectionCode="TYPE, MATERIAL" pageId="2" pageNumber="51" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED00AE040357FF9EE7A8047AC97D5D01" box="[714,867,273,296]" pageId="2" pageNumber="51">TYPE MATERIAL</collectionCode>
. —
<typeStatus id="54AA88630357FF9EE6F4047AC9ED5D0F" box="[918,1011,273,297]" pageId="2" pageNumber="51" type="holotype">Holotype</typeStatus>
(MNHN-Cp2155)
</materialsCitation>
.
<materialsCitation id="3B793C9C0357FF9EE7A80441C9415DD2" collectorName="O. Kindler" country="Germany" location="Germany" pageId="2" pageNumber="51" specimenCount="4" typeStatus="paratype">
<typeStatus id="54AA88630357FF9EE7A80441C9365D64" box="[714,808,298,322]" pageId="2" pageNumber="51" type="paratype">Paratypes</typeStatus>
:
<typeStatus id="54AA88630357FF9EE64D0440C9645D65" box="[815,890,299,323]" pageId="2" pageNumber="51" type="allotype">allotype</typeStatus>
(MNHN-Cp2156); 6, 5, 2 copepodids of stage 5, 4 copepodids of stage 4 (MNHN-Cp2157). The description is based mainly on the
<typeStatus id="54AA88630357FF9EE78D041CC95B5DA9" box="[751,837,375,399]" pageId="2" pageNumber="51" type="holotype">holotype</typeStatus>
female and
<typeStatus id="54AA88630357FF9EE6D9041CCE165DA9" box="[955,1032,375,399]" pageId="2" pageNumber="51" type="allotype">allotype</typeStatus>
male. The type material is mounted on slides in glycerol and sealed with Eukitt (
<collectorName id="26E453170357FF9EE67404C2C9615DE7" box="[790,895,425,449]" pageId="2" pageNumber="51">O. Kindler</collectorName>
GmbH &amp; Co., Freiburg,
<collectingCountry id="F30676510357FF9EE11904C2CEC65DE7" box="[1147,1240,425,449]" name="Germany" pageId="2" pageNumber="51">Germany</collectingCountry>
) and deposited in the Muséum national d’Histoire naturelle, Paris
</materialsCitation>
.
</paragraph>
<paragraph id="8BAE36C10357FF9EE7A8076FCE015EA4" blockId="2.[714,1248,273,733]" pageId="2" pageNumber="51">
<collectionCode id="ED00AE040357FF9EE7A8076FC9755E3C" box="[714,875,516,539]" pageId="2" pageNumber="51">TYPE LOCALITY</collectionCode>
. — Atlantic Ocean, Mid-Atlantic Ridge, Azores Triple Junction, Lucky Strike, cruise
<collectionCode id="ED00AE040357FF9EE7A8075CC9175E69" box="[714,777,567,591]" pageId="2" pageNumber="51">PICO</collectionCode>
, vessel
<emphasis id="B965EAD30357FF9EE637075CC9915E69" box="[853,911,567,591]" italics="true" pageId="2" pageNumber="51">Nadir</emphasis>
, submersible
<emphasis id="B965EAD30357FF9EE17E075CCE7B5E69" box="[1052,1125,567,591]" italics="true" pageId="2" pageNumber="51">Nautile</emphasis>
, dive 1270, sample number
<collectionCode id="ED00AE040357FF9EE6C0073ACE365E4F" box="[930,1064,593,617]" pageId="2" pageNumber="51">SMAC A-</collectionCode>
1,
<geoCoordinate id="EE2550060357FF9EE133073ACEC45E4F" box="[1105,1242,593,617]" degrees="37" direction="north" minutes="17.29" orientation="latitude" pageId="2" pageNumber="51" precision="9" value="37.288166">37°17.29’N</geoCoordinate>
,
<geoCoordinate id="EE2550060357FF9EE7A80701C9585EA4" box="[714,838,618,642]" degrees="32" direction="west" minutes="16.45" orientation="longitude" pageId="2" pageNumber="51" precision="9" value="-32.274166">32°16.45’W</geoCoordinate>
,
<quantity id="4CE99B240357FF9EE6320701C9815EA6" box="[848,927,617,642]" metricMagnitude="3" metricUnit="m" metricValue="1.698" pageId="2" pageNumber="51" unit="m" value="1698.0">1698 m</quantity>
,
<date id="FFAF10010357FF9EE6C80701CE055EA4" box="[938,1051,618,642]" pageId="2" pageNumber="51" value="1998-07-07">7.VII.1998</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B654A0357FF9EE7A807FAC9A95EFB" pageId="2" pageNumber="51" type="etymology">
<paragraph id="8BAE36C10357FF9EE7A807FAC9A95EFB" blockId="2.[714,1248,273,733]" pageId="2" pageNumber="51">
ETYMOLOGY. — From the Latin
<emphasis id="B965EAD30357FF9EE17607F9CE7F5E8C" box="[1044,1121,658,682]" italics="true" pageId="2" pageNumber="51">confusus</emphasis>
, referring to the difficulty in interpreting the morphological features of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C30B654A0357FF94E7A80661CB165AA3" lastPageId="8" lastPageNumber="57" pageId="2" pageNumber="51" type="description">
<paragraph id="8BAE36C10357FF9EE7A80661C9455F04" blockId="2.[714,1247,778,1669]" box="[714,859,778,804]" pageId="2" pageNumber="51">DESCRIPTION</paragraph>
<paragraph id="8BAE36C10357FF9EE7A80640C9705F63" blockId="2.[714,1247,778,1669]" box="[714,878,811,837]" pageId="2" pageNumber="51">
<emphasis id="B965EAD30357FF9EE7A80640C9705F63" box="[714,878,811,837]" italics="true" pageId="2" pageNumber="51">
<typeStatus id="54AA88630357FF9EE7A80640C93B5F63" box="[714,805,811,837]" pageId="2" pageNumber="51" type="holotype">Holotype</typeStatus>
female
</emphasis>
</paragraph>
<paragraph id="8BAE36C10357FF9FE7A80620CB4D5983" blockId="2.[714,1247,778,1669]" lastBlockId="3.[151,684,1386,1669]" lastPageId="3" lastPageNumber="52" pageId="2" pageNumber="51">
Body cyclopiform (
<figureCitation id="132A2A440357FF9EE6CC0620CE335F42" box="[942,1069,842,869]" captionStart="FIG" captionStartId="3.[151,162,1262,1279]" captionTargetBox="[189,1200,169,1227]" captionTargetId="figure-135@3.[186,1178,157,1232]" captionTargetPageId="3" captionText="FIG. 1. — Heptnerina confusa n. gen., n. sp., holotype; A, habitus, dorsal; B, habitus, lateral; C, genital double somite, dorsal; D, genital double somite, ventral; E, gonoporal area and P6, lateral. Scale bars: A, B, 0.2 mm; C-E, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401376" httpUri="https://zenodo.org/record/5401376/files/figure.png" pageId="2" pageNumber="51">Fig. 1A, B</figureCitation>
). Total length, excluding caudal setae: 0.82 mm, greatest width 0.32 mm. First pedigerous somite free, concealed laterally by extension of cephalosome. Tergites of second to fourth pedigerous somites sclerotized laterally and pointed posteriorly. Urosome five-segmented. Genital and first abdominal somites fused to form genital double somite (
<figureCitation id="132A2A440357FF9EE7970120C9965842" box="[757,904,1098,1125]" captionStart="FIG" captionStartId="3.[151,162,1262,1279]" captionTargetBox="[189,1200,169,1227]" captionTargetId="figure-135@3.[186,1178,157,1232]" captionTargetPageId="3" captionText="FIG. 1. — Heptnerina confusa n. gen., n. sp., holotype; A, habitus, dorsal; B, habitus, lateral; C, genital double somite, dorsal; D, genital double somite, ventral; E, gonoporal area and P6, lateral. Scale bars: A, B, 0.2 mm; C-E, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401376" httpUri="https://zenodo.org/record/5401376/files/figure.png" pageId="2" pageNumber="51">Fig. 1C, D</figureCitation>
). Genital field with single medioventral copulatory pore, connecting internally into a wide copulatory tube leading to the seminal receptacle organized into two arms, each extending to dorsolateral gonopore associated with leg 6; accessory rounded structure of unknown function (possibly glandular), present on each side, proximal to seminal receptacle (
<figureCitation id="132A2A440357FF9EE7B00020C94A5942" box="[722,852,1354,1381]" captionStart="FIG" captionStartId="3.[151,162,1262,1279]" captionTargetBox="[189,1200,169,1227]" captionTargetId="figure-135@3.[186,1178,157,1232]" captionTargetPageId="3" captionText="FIG. 1. — Heptnerina confusa n. gen., n. sp., holotype; A, habitus, dorsal; B, habitus, lateral; C, genital double somite, dorsal; D, genital double somite, ventral; E, gonoporal area and P6, lateral. Scale bars: A, B, 0.2 mm; C-E, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401376" httpUri="https://zenodo.org/record/5401376/files/figure.png" pageId="2" pageNumber="51">Fig. 1D, E</figureCitation>
). Terminal margin of anal somite with ventral row of small spinules. Caudal rami (
<figureCitation id="132A2A440357FF9EE7B000E0C9365983" box="[722,808,1419,1445]" captionStart="FIG" captionStartId="4.[151,162,1296,1313]" captionTargetBox="[234,1156,176,1253]" captionTargetId="figure-49@4.[232,1176,170,1257]" captionTargetPageId="4" captionText="FIG. 2. — Heptnerina confusa n. gen., n. sp., holotype; A, caudal ramus, ventral; B, rostral and oral area showing labrum and mandibles; C, paragnath; D, paragnath and mandible (part); E, antennule; F, setation of third segment of antennule; G, distal segment of antennule. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401378" httpUri="https://zenodo.org/record/5401378/files/figure.png" pageId="2" pageNumber="51">Fig. 2A</figureCitation>
) parallel, three times longer than wide, ventral side subdistally lined by row of flattened spinules; bearing seven setae. Accessory lateral seta (seta I) short, difficult to see; inner terminal seta 1.47 times longer than caudal ramus and longer than dorsal and distal outer setae; median inner terminal seta long, 5.3 times longer than caudal ramus and 1.7 times longer than median external seta. Median caudal setae with heteronomous setation.
</paragraph>
<caption id="DF6E66490356FF9FE5F50185CE015932" ID-DOI="http://doi.org/10.5281/zenodo.5401376" ID-Zenodo-Dep="5401376" httpUri="https://zenodo.org/record/5401376/files/figure.png" pageId="3" pageNumber="52" startId="3.[151,162,1262,1279]" targetBox="[189,1200,169,1227]" targetPageId="3">
<paragraph id="8BAE36C10356FF9FE5F50185CE015932" blockId="3.[151,1248,1262,1300]" pageId="3" pageNumber="52">
FIG. 1. —
<taxonomicName id="4C114D420356FF9FE58E0185CB8E58D9" authority="Ivanenko &amp; Defaye, 2004" authorityName="Ivanenko &amp; Defaye" authorityYear="2004" box="[236,400,1262,1279]" class="Hexanauplia" family="Cyclopinidae" genus="Heptnerina" kingdom="Animalia" order="Cyclopoida" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="confusa" status="n. gen., n. sp.">
<emphasis id="B965EAD30356FF9FE58E0185CB8E58D9" box="[236,400,1262,1279]" italics="true" pageId="3" pageNumber="52">Heptnerina confusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25657A80356FF9FE4F50185C80C58D9" box="[407,530,1262,1279]" pageId="3" pageNumber="52" rank="species">n. gen., n. sp.</taxonomicNameLabel>
, holotype;
<emphasis id="B965EAD30356FF9FE7E70185C88C58D9" bold="true" box="[645,658,1262,1279]" pageId="3" pageNumber="52">A</emphasis>
, habitus, dorsal;
<emphasis id="B965EAD30356FF9FE6450185C92A58D9" bold="true" box="[807,820,1262,1279]" pageId="3" pageNumber="52">B</emphasis>
, habitus, lateral;
<emphasis id="B965EAD30356FF9FE6AB0185C9C958D9" bold="true" box="[969,983,1262,1279]" pageId="3" pageNumber="52">C</emphasis>
, genital double somite, dorsal;
<emphasis id="B965EAD30356FF9FE5F50068CABB5932" bold="true" box="[151,165,1283,1300]" pageId="3" pageNumber="52">D</emphasis>
, genital double somite, ventral;
<emphasis id="B965EAD30356FF9FE4CD0068CBA55932" bold="true" box="[431,443,1283,1300]" pageId="3" pageNumber="52">E</emphasis>
, gonoporal area and P6, lateral. Scale bars: A, B, 0.2 mm; C-E, 0.05 mm.
</paragraph>
</caption>
<paragraph id="8BAE36C10356FF9FE5F500C1CB9759E3" blockId="3.[151,684,1386,1669]" box="[151,393,1450,1477]" pageId="3" pageNumber="52">Egg sacs not observed.</paragraph>
<paragraph id="8BAE36C10356FF9FE5F500A1C83459C3" blockId="3.[151,684,1386,1669]" box="[151,554,1482,1509]" pageId="3" pageNumber="52">
Rostrum (
<figureCitation id="132A2A440356FF9FE46700A0CB4859C3" box="[261,342,1483,1509]" captionStart="FIG" captionStartId="4.[151,162,1296,1313]" captionTargetBox="[234,1156,176,1253]" captionTargetId="figure-49@4.[232,1176,170,1257]" captionTargetPageId="4" captionText="FIG. 2. — Heptnerina confusa n. gen., n. sp., holotype; A, caudal ramus, ventral; B, rostral and oral area showing labrum and mandibles; C, paragnath; D, paragnath and mandible (part); E, antennule; F, setation of third segment of antennule; G, distal segment of antennule. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401378" httpUri="https://zenodo.org/record/5401378/files/figure.png" pageId="3" pageNumber="52">Fig. 2B</figureCitation>
) weakly developed.
</paragraph>
<paragraph id="8BAE36C10356FF9FE5F50080CBFC5A03" blockId="3.[151,684,1386,1669]" pageId="3" pageNumber="52">
Labrum with anterior outgrowth and patches of setules as shown on
<figureCitation id="132A2A440356FF9FE4120360CBC05A03" box="[368,478,1547,1573]" captionStart="FIG" captionStartId="4.[151,162,1296,1313]" captionTargetBox="[234,1156,176,1253]" captionTargetId="figure-49@4.[232,1176,170,1257]" captionTargetPageId="4" captionText="FIG. 2. — Heptnerina confusa n. gen., n. sp., holotype; A, caudal ramus, ventral; B, rostral and oral area showing labrum and mandibles; C, paragnath; D, paragnath and mandible (part); E, antennule; F, setation of third segment of antennule; G, distal segment of antennule. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401378" httpUri="https://zenodo.org/record/5401378/files/figure.png" pageId="3" pageNumber="52">Figure 2B</figureCitation>
.
</paragraph>
<paragraph id="8BAE36C10356FF9FE5F50340CB3F5AA3" blockId="3.[151,684,1386,1669]" pageId="3" pageNumber="52">
Paragnath (
<figureCitation id="132A2A440356FF9FE4700340CB965A62" box="[274,392,1578,1605]" captionStart="FIG" captionStartId="4.[151,162,1296,1313]" captionTargetBox="[234,1156,176,1253]" captionTargetId="figure-49@4.[232,1176,170,1257]" captionTargetPageId="4" captionText="FIG. 2. — Heptnerina confusa n. gen., n. sp., holotype; A, caudal ramus, ventral; B, rostral and oral area showing labrum and mandibles; C, paragnath; D, paragnath and mandible (part); E, antennule; F, setation of third segment of antennule; G, distal segment of antennule. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401378" httpUri="https://zenodo.org/record/5401378/files/figure.png" pageId="3" pageNumber="52">Fig. 2C, D</figureCitation>
) lobate with four spinulate setae, normally covered ventrally by gnathobase of mandible.
</paragraph>
<paragraph id="8BAE36C10356FF9FE7A80000CEB45AA3" blockId="3.[714,1247,1387,1669]" pageId="3" pageNumber="52">
Antennule (
<figureCitation id="132A2A440356FF9FE63A0000C98159A3" box="[856,927,1387,1413]" captionStart="FIG" captionStartId="4.[151,162,1296,1313]" captionTargetBox="[234,1156,176,1253]" captionTargetId="figure-49@4.[232,1176,170,1257]" captionTargetPageId="4" captionText="FIG. 2. — Heptnerina confusa n. gen., n. sp., holotype; A, caudal ramus, ventral; B, rostral and oral area showing labrum and mandibles; C, paragnath; D, paragnath and mandible (part); E, antennule; F, setation of third segment of antennule; G, distal segment of antennule. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401378" httpUri="https://zenodo.org/record/5401378/files/figure.png" pageId="3" pageNumber="52">Fig. 2</figureCitation>
E-G) 10-segmented; third, fourth and fifth segments subdivided, expressing partially, intersegmental articulations on their medial margin. Setation formula as follows: 3, 5, 8, 4, 6+s, 6+a, 3+a, 2, 2+a, 8+a (s: spine, a: aesthetasc), corresponding to ancestral segments: I- II, III-V, VI-IX, X-XI, XII-XIV, XV-XX, XXI-XXIII, XXIV, XXV, XXVI-XXVIII respectively. First segment ornamented with setules.
</paragraph>
<caption id="DF6E66490351FF98E5F5007BC816596A" ID-DOI="http://doi.org/10.5281/zenodo.5401378" ID-Zenodo-Dep="5401378" httpUri="https://zenodo.org/record/5401378/files/figure.png" pageId="4" pageNumber="53" startId="4.[151,162,1296,1313]" targetBox="[234,1156,176,1253]" targetPageId="4">
<paragraph id="8BAE36C10351FF98E5F5007BC816596A" blockId="4.[151,1247,1296,1356]" pageId="4" pageNumber="53">
FIG. 2. —
<taxonomicName id="4C114D420351FF98E58E007BCB8F5907" authority="Ivanenko &amp; Defaye, 2004" authorityName="Ivanenko &amp; Defaye" authorityYear="2004" box="[236,401,1296,1313]" class="Hexanauplia" family="Cyclopinidae" genus="Heptnerina" kingdom="Animalia" order="Cyclopoida" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="confusa" status="n. gen., n. sp.">
<emphasis id="B965EAD30351FF98E58E007BCB8F5907" box="[236,401,1296,1313]" italics="true" pageId="4" pageNumber="53">Heptnerina confusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25657A80351FF98E4FA007BC80D5907" box="[408,531,1296,1313]" pageId="4" pageNumber="53" rank="species">n. gen., n. sp.</taxonomicNameLabel>
, holotype;
<emphasis id="B965EAD30351FF98E7E5007BC88A5907" bold="true" box="[647,660,1296,1313]" pageId="4" pageNumber="53">A</emphasis>
, caudal ramus, ventral;
<emphasis id="B965EAD30351FF98E606007BC96F5907" bold="true" box="[868,881,1296,1313]" pageId="4" pageNumber="53">B</emphasis>
, rostral and oral area showing labrum and mandibles;
<emphasis id="B965EAD30351FF98E59E004DCB145911" bold="true" box="[252,266,1318,1335]" pageId="4" pageNumber="53">C</emphasis>
, paragnath;
<emphasis id="B965EAD30351FF98E418004DCB965911" bold="true" box="[378,392,1318,1335]" pageId="4" pageNumber="53">D</emphasis>
, paragnath and mandible (part);
<emphasis id="B965EAD30351FF98E7C8004DC8A85911" bold="true" box="[682,694,1318,1335]" pageId="4" pageNumber="53">E</emphasis>
, antennule;
<emphasis id="B965EAD30351FF98E641004DC9305911" bold="true" box="[803,814,1318,1335]" pageId="4" pageNumber="53">F</emphasis>
, setation of third segment of antennule;
<emphasis id="B965EAD30351FF98E1F4004DCEBA5911" bold="true" box="[1174,1188,1318,1335]" pageId="4" pageNumber="53">G</emphasis>
, distal segment of antennule. Scale bars: 0.05 mm.
</paragraph>
</caption>
<paragraph id="8BAE36C10351FF98E5F500C0CEB95A63" blockId="4.[151,685,1450,1669]" lastBlockId="4.[714,1247,1450,1669]" pageId="4" pageNumber="53">
Antenna (
<figureCitation id="132A2A440351FF98E46000C0CB6C59E2" box="[258,370,1450,1477]" captionStart="FIG" captionStartId="5.[151,162,1382,1399]" captionTargetBox="[151,1248,169,1420]" captionTargetId="figure-118@5.[141,1230,143,1364]" captionTargetPageId="5" captionText="FIG. 3. — Heptnerina confusa n. gen., n. sp.," figureDoi="http://doi.org/10.5281/zenodo.5401380" httpUri="https://zenodo.org/record/5401380/files/figure.png" pageId="4" pageNumber="53">Fig. 3A, B</figureCitation>
) consisting of four segments: basis with one inner seta and two external setae representing exopod; endopod three-segmented with setal formula 1, 5, 7. Basis with basal patch of spinules on frontal surface; first endopodal segment with lateral outer distal prominence bearing group of tiny spinules, and with inner margin with two rows of small spinules, one in each half of segment; last endopodal segment with outer spiny prominence distally as on first endopodal segment, a proximal row of spinules on same margin, and terminal patch of spinules on lateral surface.
</paragraph>
<paragraph id="8BAE36C10351FF99E7A80320CB925A63" blockId="4.[714,1247,1450,1669]" lastBlockId="5.[151,684,1482,1669]" lastPageId="5" lastPageNumber="54" pageId="4" pageNumber="53">
Mandible (
<figureCitation id="132A2A440351FF98E6200320C9A55A42" box="[834,955,1610,1637]" captionStart="FIG" captionStartId="5.[151,162,1382,1399]" captionTargetBox="[151,1248,169,1420]" captionTargetId="figure-118@5.[141,1230,143,1364]" captionTargetPageId="5" captionText="FIG. 3. — Heptnerina confusa n. gen., n. sp.," figureDoi="http://doi.org/10.5281/zenodo.5401380" httpUri="https://zenodo.org/record/5401380/files/figure.png" pageId="4" pageNumber="53">Fig. 3C, D</figureCitation>
) with toothed gnathobase; palp biramous; basis with one inner seta and two rows of setules; endopod two-segmented, setal formula 3, 6; exopod four-segmented, setal formula 1, 1, 1, 2; one seta on distal segment with patch of setules on tip.
</paragraph>
<caption id="DF6E66490350FF99E5F5000DC8345951" ID-DOI="http://doi.org/10.5281/zenodo.5401380" ID-Zenodo-Dep="5401380" box="[151,554,1382,1399]" httpUri="https://zenodo.org/record/5401380/files/figure.png" pageId="5" pageNumber="54" startId="5.[151,162,1382,1399]" targetBox="[151,1248,169,1420]" targetPageId="5">
<paragraph id="8BAE36C10350FF99E5F5000DC8345951" blockId="5.[151,566,1382,1420]" box="[151,554,1382,1399]" pageId="5" pageNumber="54">
FIG. 3. —
<taxonomicName id="4C114D420350FF99E592000DCB815951" authority="Ivanenko &amp; Defaye, 2004" authorityName="Ivanenko &amp; Defaye" authorityYear="2004" box="[240,415,1382,1399]" class="Hexanauplia" family="Cyclopinidae" genus="Heptnerina" kingdom="Animalia" order="Cyclopoida" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="confusa" status="n. gen., n. sp.">
<emphasis id="B965EAD30350FF99E592000DCB815951" box="[240,415,1382,1399]" italics="true" pageId="5" pageNumber="54">Heptnerina confusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25657A80350FF99E4C5000DC83B5951" box="[423,549,1382,1399]" pageId="5" pageNumber="54" rank="species">n. gen., n. sp.</taxonomicNameLabel>
,
</paragraph>
</caption>
<paragraph id="8BAE36C10350FF99E5F50320CE395AA3" blockId="5.[151,684,1482,1669]" lastBlockId="5.[714,1247,1482,1669]" pageId="5" pageNumber="54">
Maxillule (
<figureCitation id="132A2A440350FF99E4760320CB8F5A42" box="[276,401,1610,1637]" captionStart="FIG" captionStartId="6.[151,162,1544,1561]" captionTargetBox="[190,1247,169,1505]" captionTargetId="figure-30@6.[177,1266,158,1511]" captionTargetPageId="6" captionText="FIG. 4. — Heptnerina confusa n. gen., n. sp., holotype; A, maxillule; B, inner lobe of maxillule, ventral; C, maxilla; D, maxilliped. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401382" httpUri="https://zenodo.org/record/5401382/files/figure.png" pageId="5" pageNumber="54">Fig. 4A, B</figureCitation>
): praecoxal arthrite with 12 setae; coxal epipodite with two unequal setae, coxal endite with one seta; basis with two endites, proximal endite with three setae, distal endite with two setae; endopod one-segmented with seven setae; exopod one-segmented, with four setae; one seta shorter than others and ornament- ed with patch of spinules on tip.
</paragraph>
<caption id="DF6E66490353FF9AE5F50363CB545A08" ID-DOI="http://doi.org/10.5281/zenodo.5401382" ID-Zenodo-Dep="5401382" httpUri="https://zenodo.org/record/5401382/files/figure.png" pageId="6" pageNumber="55" startId="6.[151,162,1544,1561]" targetBox="[190,1247,169,1505]" targetPageId="6">
<paragraph id="8BAE36C10353FF9AE5F50363CB545A08" blockId="6.[151,1249,1544,1582]" pageId="6" pageNumber="55">
FIG. 4. —
<taxonomicName id="4C114D420353FF9AE5890363CB905A3F" authority="Ivanenko &amp; Defaye, 2004" authorityName="Ivanenko &amp; Defaye" authorityYear="2004" box="[235,398,1544,1561]" class="Hexanauplia" family="Cyclopinidae" genus="Heptnerina" kingdom="Animalia" order="Cyclopoida" pageId="6" pageNumber="55" phylum="Arthropoda" rank="species" species="confusa" status="n. gen., n. sp.">
<emphasis id="B965EAD30353FF9AE5890363CB905A3F" box="[235,398,1544,1561]" italics="true" pageId="6" pageNumber="55">Heptnerina confusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25657A80353FF9AE4F60363C8105A3F" box="[404,526,1544,1561]" pageId="6" pageNumber="55" rank="species">n. gen., n. sp.</taxonomicNameLabel>
, holotype;
<emphasis id="B965EAD30353FF9AE71D0363C8925A3F" bold="true" box="[639,652,1544,1561]" pageId="6" pageNumber="55">A</emphasis>
, maxillule;
<emphasis id="B965EAD30353FF9AE78E0363C8E75A3F" bold="true" box="[748,761,1544,1561]" pageId="6" pageNumber="55">B</emphasis>
, inner lobe of maxillule, ventral;
<emphasis id="B965EAD30353FF9AE16F0363CE055A3F" bold="true" box="[1037,1051,1544,1561]" pageId="6" pageNumber="55">C</emphasis>
, maxilla;
<emphasis id="B965EAD30353FF9AE10E0363CE645A3F" bold="true" box="[1132,1146,1544,1561]" pageId="6" pageNumber="55">D</emphasis>
, maxilliped. Scale bars: 0.05 mm.
</paragraph>
</caption>
<caption id="DF6E66490352FF9BE5F505C2CBDA5CE9" pageId="7" pageNumber="56" startId="7.[151,162,169,186]">
<paragraph id="8BAE36C10352FF9BE5F505C2CBDA5CE9" blockId="7.[151,684,169,207]" pageId="7" pageNumber="56">
TABLE 1. — Spine and setal formula of swimming legs in
<taxonomicName id="4C114D420352FF9BE5F505D5CB315CE9" authority="Ivanenko &amp; Defaye, 2004" authorityName="Ivanenko &amp; Defaye" authorityYear="2004" box="[151,303,190,207]" class="Hexanauplia" family="Cyclopinidae" genus="Hepterina" kingdom="Animalia" order="Cyclopoida" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="confusa" status="n. gen., n. sp.">
<emphasis id="B965EAD30352FF9BE5F505D5CB315CE9" box="[151,303,190,207]" italics="true" pageId="7" pageNumber="56">Hepterina confusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25657A80352FF9BE45605D5CBB75CE9" box="[308,425,190,207]" pageId="7" pageNumber="56" rank="species">n. gen., n. sp.</taxonomicNameLabel>
,.
</paragraph>
</caption>
<paragraph id="8BAE36C10352FF9BE46B0591C8B95DBD" pageId="7" pageNumber="56">
<table id="F911C46103520063E5C90591C8B95DBD" box="[171,679,250,411]" gridcols="5" gridrows="5" pageId="7" pageNumber="56">
<tr id="3521348303520063E5C90591C8B95D28" box="[171,679,250,270]" gridrow="0" pageId="7" pageNumber="56" rowspan-0="1">
<th id="76F05DFF03520063E46B0591CB265D28" box="[265,312,250,270]" gridcol="1" gridrow="0" pageId="7" pageNumber="56">
<emphasis id="B965EAD30352FF9BE46B0591CB265D28" bold="true" box="[265,312,250,270]" pageId="7" pageNumber="56">Coxa</emphasis>
</th>
<th id="76F05DFF03520063E4000591CB8A5D28" box="[354,404,250,270]" gridcol="2" gridrow="0" pageId="7" pageNumber="56">
<emphasis id="B965EAD30352FF9BE4000591CB8A5D28" bold="true" box="[354,404,250,270]" pageId="7" pageNumber="56">Basis</emphasis>
</th>
<th id="76F05DFF03520063E4D30591C8005D28" box="[433,542,250,270]" gridcol="3" gridrow="0" pageId="7" pageNumber="56">
<emphasis id="B965EAD30352FF9BE4AA0591C8105D28" bold="true" box="[456,526,250,270]" pageId="7" pageNumber="56">Exopod</emphasis>
</th>
<th id="76F05DFF03520063E74C0591C8B95D28" box="[558,679,250,270]" gridcol="4" gridrow="0" pageId="7" pageNumber="56">
<emphasis id="B965EAD30352FF9BE7230591C88D5D28" bold="true" box="[577,659,250,270]" pageId="7" pageNumber="56">Endopod</emphasis>
</th>
</tr>
<tr id="3521348303520063E5C90443C8B95D1A" box="[171,679,296,316]" gridrow="1" pageId="7" pageNumber="56">
<th id="76F05DFF03520063E5C90443CAC35D1A" box="[171,221,296,316]" gridcol="0" gridrow="1" pageId="7" pageNumber="56">
<emphasis id="B965EAD30352FF9BE5C90443CAC35D1A" bold="true" box="[171,221,296,316]" pageId="7" pageNumber="56">Leg 1</emphasis>
</th>
<td id="76F05DFF03520063E46B0443CB265D1A" box="[265,312,296,316]" gridcol="1" gridrow="1" pageId="7" pageNumber="56">0-1</td>
<td id="76F05DFF03520063E4000443CB8A5D1A" box="[354,404,296,316]" gridcol="2" gridrow="1" pageId="7" pageNumber="56">1-I</td>
<td id="76F05DFF03520063E4D30443C8005D1A" box="[433,542,296,316]" gridcol="3" gridrow="1" pageId="7" pageNumber="56">I-1; I-1; III,I,4</td>
<td id="76F05DFF03520063E74C0443C8B95D1A" box="[558,679,296,316]" gridcol="4" gridrow="1" pageId="7" pageNumber="56">0-1; 0-1; 1,2,3</td>
</tr>
<tr id="3521348303520063E5C90423C8B95D7A" box="[171,679,328,348]" gridrow="2" pageId="7" pageNumber="56">
<th id="76F05DFF03520063E5C90423CAC35D7A" box="[171,221,328,348]" gridcol="0" gridrow="2" pageId="7" pageNumber="56">
<emphasis id="B965EAD30352FF9BE5C90423CAC35D7A" bold="true" box="[171,221,328,348]" pageId="7" pageNumber="56">Leg 2</emphasis>
</th>
<td id="76F05DFF03520063E46B0423CB265D7A" box="[265,312,328,348]" gridcol="1" gridrow="2" pageId="7" pageNumber="56">0-1</td>
<td id="76F05DFF03520063E4000423CB8A5D7A" box="[354,404,328,348]" gridcol="2" gridrow="2" pageId="7" pageNumber="56">1-0</td>
<td id="76F05DFF03520063E4D30423C8005D7A" box="[433,542,328,348]" gridcol="3" gridrow="2" pageId="7" pageNumber="56">I-1; I-1; III,I,5</td>
<td id="76F05DFF03520063E74C0423C8B95D7A" box="[558,679,328,348]" gridcol="4" gridrow="2" pageId="7" pageNumber="56">0-1; 0-2; 1,2,3</td>
</tr>
<tr id="3521348303520063E5C9040CC8B95D5D" box="[171,679,359,379]" gridrow="3" pageId="7" pageNumber="56">
<th id="76F05DFF03520063E5C9040CCAC35D5D" box="[171,221,359,379]" gridcol="0" gridrow="3" pageId="7" pageNumber="56">
<emphasis id="B965EAD30352FF9BE5C9040CCAC35D5D" bold="true" box="[171,221,359,379]" pageId="7" pageNumber="56">Leg 3</emphasis>
</th>
<td id="76F05DFF03520063E46B040CCB265D5D" box="[265,312,359,379]" gridcol="1" gridrow="3" pageId="7" pageNumber="56">0-1</td>
<td id="76F05DFF03520063E400040CCB8A5D5D" box="[354,404,359,379]" gridcol="2" gridrow="3" pageId="7" pageNumber="56">1-0</td>
<td id="76F05DFF03520063E4D3040CC8005D5D" box="[433,542,359,379]" gridcol="3" gridrow="3" pageId="7" pageNumber="56">I-1; I-1; III,I,5</td>
<td id="76F05DFF03520063E74C040CC8B95D5D" box="[558,679,359,379]" gridcol="4" gridrow="3" pageId="7" pageNumber="56">0-1; 0-2; 1,2,3</td>
</tr>
<tr id="3521348303520063E5C904ECC8B95DBD" box="[171,679,391,411]" gridrow="4" pageId="7" pageNumber="56">
<th id="76F05DFF03520063E5C904ECCAC35DBD" box="[171,221,391,411]" gridcol="0" gridrow="4" pageId="7" pageNumber="56">
<emphasis id="B965EAD30352FF9BE5C904ECCAC35DBD" bold="true" box="[171,221,391,411]" pageId="7" pageNumber="56">Leg 4</emphasis>
</th>
<td id="76F05DFF03520063E46B04ECCB265DBD" box="[265,312,391,411]" gridcol="1" gridrow="4" pageId="7" pageNumber="56">0-1</td>
<td id="76F05DFF03520063E40004ECCB8A5DBD" box="[354,404,391,411]" gridcol="2" gridrow="4" pageId="7" pageNumber="56">1-0</td>
<td id="76F05DFF03520063E4D304ECC8005DBD" box="[433,542,391,411]" gridcol="3" gridrow="4" pageId="7" pageNumber="56">I-1; I-1; II,I,5</td>
<td id="76F05DFF03520063E74C04ECC8B95DBD" box="[558,679,391,411]" gridcol="4" gridrow="4" pageId="7" pageNumber="56">0-1; 0-2; 1,2,3</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAE36C10352FF9BE5F50480CB545EC2" blockId="7.[151,684,490,1669]" pageId="7" pageNumber="56">
Maxilla (
<figureCitation id="132A2A440352FF9BE5980480CB4C5E23" box="[250,338,491,517]" captionStart="FIG" captionStartId="6.[151,162,1544,1561]" captionTargetBox="[190,1247,169,1505]" captionTargetId="figure-30@6.[177,1266,158,1511]" captionTargetPageId="6" captionText="FIG. 4. — Heptnerina confusa n. gen., n. sp., holotype; A, maxillule; B, inner lobe of maxillule, ventral; C, maxilla; D, maxilliped. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401382" httpUri="https://zenodo.org/record/5401382/files/figure.png" pageId="7" pageNumber="56">Fig. 4C</figureCitation>
) with praecoxa separated from coxa by arthrodial membrane posteriorly only and with two endites, proximal with four setae, distal with one seta; coxa with two endites, each with three setae; basis produced into claw, with one seta; endopod three-segmented, first segment with four setae; second and third segments with total of six setae.
</paragraph>
<paragraph id="8BAE36C10352FF9BE5F50780C80D5823" blockId="7.[151,684,490,1669]" pageId="7" pageNumber="56">
Maxilliped (
<figureCitation id="132A2A440352FF9BE44A0780CB9D5F23" box="[296,387,747,773]" captionStart="FIG" captionStartId="6.[151,162,1544,1561]" captionTargetBox="[190,1247,169,1505]" captionTargetId="figure-30@6.[177,1266,158,1511]" captionTargetPageId="6" captionText="FIG. 4. — Heptnerina confusa n. gen., n. sp., holotype; A, maxillule; B, inner lobe of maxillule, ventral; C, maxilla; D, maxilliped. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401382" httpUri="https://zenodo.org/record/5401382/files/figure.png" pageId="7" pageNumber="56">Fig. 4E</figureCitation>
) with praecoxa and coxa separated by arthrodial membrane on both sides. Praecoxa with two endites armed with one and three setae; coxa and basis with two setae each; coxa with row of long setules along outer margin, basis with small distal row of spinules along outer and two others on surface of segment; row of long setules on internal margin; endopod foursegmented, setal formula 0, 0, 1, 4.
</paragraph>
<paragraph id="8BAE36C10352FF9BE5F50160CB455843" blockId="7.[151,684,490,1669]" pageId="7" pageNumber="56">
Swimming legs 1-4 biramous (
<figureCitation id="132A2A440352FF9BE49F0160C8595803" box="[509,583,1035,1061]" captionStart="FIG" captionStartId="8.[151,162,1454,1471]" captionTargetBox="[173,1227,178,1419]" captionTargetId="figure-28@8.[151,1240,162,1432]" captionTargetPageId="8" captionText="FIG. 5. — Heptnerina confusa n. gen., n. sp., holotype; A, leg 1, anterior; B, leg 2, anterior; C, exopod of leg 2, anterior; D, leg 3, anterior; E, endopod of leg 3, anterior. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401384" httpUri="https://zenodo.org/record/5401384/files/figure.png" pageId="7" pageNumber="56">Figs 5</figureCitation>
;
<figureCitation id="132A2A440352FF9BE7350160C8835802" box="[599,669,1034,1061]" captionStart="FIG" captionStartId="9.[151,162,1524,1541]" captionTargetBox="[163,1180,177,1481]" captionTargetId="figure-25@9.[151,1184,170,1486]" captionTargetPageId="9" captionText="FIG. 6. — Heptnerina confusa n. gen., n. sp.,; A, holotype, leg 4; B, holotype, endopod of leg 4; C, holotype, leg 5 with endopod arrowed; D, paratype, leg 5. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401386" httpUri="https://zenodo.org/record/5401386/files/figure.png" pageId="7" pageNumber="56">6A, B</figureCitation>
), with three-segmented rami; spine and setal formula: see
<tableCitation id="C693037A0352FF9BE4600120CB4B5843" box="[258,341,1099,1125]" captionStart="TABLE" captionStartId="7.[151,162,169,186]" captionText="TABLE 1. — Spine and setal formula of swimming legs in Hepterina confusa n. gen., n. sp.,." pageId="7" pageNumber="56">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BAE36C10352FF9BE5F50100CBC45883" blockId="7.[151,684,490,1669]" pageId="7" pageNumber="56">Formula of spines of exopods: 4, 4, 4, 3. Formula of setae of exopods: 4, 5, 5, 5.</paragraph>
<paragraph id="8BAE36C10352FF9BE5F501C1CAD658C3" blockId="7.[151,684,490,1669]" pageId="7" pageNumber="56">Leg 1 with one seta on second segment of endopod.</paragraph>
<paragraph id="8BAE36C10352FF9BE5F50181CB2F5962" blockId="7.[151,684,490,1669]" pageId="7" pageNumber="56">
Leg 4 with setae on second endopodal segment and proximal inner seta on third (
<figureCitation id="132A2A440352FF9BE76F0060C87E5903" box="[525,608,1291,1317]" captionStart="FIG" captionStartId="9.[151,162,1524,1541]" captionTargetBox="[163,1180,177,1481]" captionTargetId="figure-25@9.[151,1184,170,1486]" captionTargetPageId="9" captionText="FIG. 6. — Heptnerina confusa n. gen., n. sp.,; A, holotype, leg 4; B, holotype, endopod of leg 4; C, holotype, leg 5 with endopod arrowed; D, paratype, leg 5. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401386" httpUri="https://zenodo.org/record/5401386/files/figure.png" pageId="7" pageNumber="56">Fig. 6B</figureCitation>
) particularly strong.
</paragraph>
<paragraph id="8BAE36C10352FF9BE5F50021C83A5A63" blockId="7.[151,684,490,1669]" pageId="7" pageNumber="56">
Leg 5 (
<figureCitation id="132A2A440352FF9BE5840020CB235943" box="[230,317,1355,1381]" captionStart="FIG" captionStartId="9.[151,162,1524,1541]" captionTargetBox="[163,1180,177,1481]" captionTargetId="figure-25@9.[151,1184,170,1486]" captionTargetPageId="9" captionText="FIG. 6. — Heptnerina confusa n. gen., n. sp.,; A, holotype, leg 4; B, holotype, endopod of leg 4; C, holotype, leg 5 with endopod arrowed; D, paratype, leg 5. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401386" httpUri="https://zenodo.org/record/5401386/files/figure.png" pageId="7" pageNumber="56">Fig. 6C</figureCitation>
) connected by narrow intercoxal plate; undivided protopod with outer seta and setules along inner margin; exopod one-segmented, armed distally with two spines and one long median seta; inner margin of protopod bearing short, stout, digitiform segment-like structure, with or without ornamenatation of spinules, interpreted as representing endopod.
</paragraph>
<paragraph id="8BAE36C10352FF9BE5F50321C9D85CC3" blockId="7.[151,684,490,1669]" lastBlockId="7.[714,1246,170,229]" pageId="7" pageNumber="56">
Leg 6 (
<figureCitation id="132A2A440352FF9BE58C0320CB555A43" box="[238,331,1611,1637]" captionStart="FIG" captionStartId="3.[151,162,1262,1279]" captionTargetBox="[189,1200,169,1227]" captionTargetId="figure-135@3.[186,1178,157,1232]" captionTargetPageId="3" captionText="FIG. 1. — Heptnerina confusa n. gen., n. sp., holotype; A, habitus, dorsal; B, habitus, lateral; C, genital double somite, dorsal; D, genital double somite, ventral; E, gonoporal area and P6, lateral. Scale bars: A, B, 0.2 mm; C-E, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401376" httpUri="https://zenodo.org/record/5401376/files/figure.png" pageId="7" pageNumber="56">Fig. 1E</figureCitation>
) located laterally on genital double-somite, with three unequal setae, one small; longest seta, located more posteriorly, strong, curved dorsally.
</paragraph>
<paragraph id="8BAE36C10352FF9BE7A80460C94A5D03" blockId="7.[714,1247,267,1061]" box="[714,852,267,293]" pageId="7" pageNumber="56">
<emphasis id="B965EAD30352FF9BE7A80460C94A5D03" box="[714,852,267,293]" italics="true" pageId="7" pageNumber="56">
<typeStatus id="54AA88630352FF9BE7A80460C9025D03" box="[714,796,267,293]" pageId="7" pageNumber="56" type="allotype">Allotype</typeStatus>
male
</emphasis>
</paragraph>
<paragraph id="8BAE36C10352FF9BE7A80440CE305D63" blockId="7.[714,1247,267,1061]" box="[714,1070,298,325]" pageId="7" pageNumber="56">Differing from female as follows:</paragraph>
<paragraph id="8BAE36C10352FF9BE7A80420C9565E63" blockId="7.[714,1247,267,1061]" pageId="7" pageNumber="56">
Body (
<figureCitation id="132A2A440352FF9BE6710420C9965D42" box="[787,904,330,357]" captionStart="FIG" captionStartId="10.[151,162,1547,1564]" captionTargetBox="[157,1194,173,1524]" captionTargetId="figure-28@10.[151,1209,168,1528]" captionTargetPageId="10" captionText="FIG. 7. — Heptnerina confusa n. gen., n. sp.,; A, habitus, dorsal; B, habitus, lateral; C, rostrum and labrum, lateral; D, leg 5 and genital somite, ventral; E, caudal ramus, ventral; F, leg 5. Scale bars: A, B, 0.2 mm; C, 0.1 mm; D-F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401388" httpUri="https://zenodo.org/record/5401388/files/figure.png" pageId="7" pageNumber="56">Fig. 7A, B</figureCitation>
): total length, excluding caudal setae 0.65 mm, greatest width 0.23 mm. Urosome six-segmented; genital and first abdominal somites separate (
<figureCitation id="132A2A440352FF9BE6B504C0CE235DE2" box="[983,1085,426,453]" captionStart="FIG" captionStartId="10.[151,162,1547,1564]" captionTargetBox="[157,1194,173,1524]" captionTargetId="figure-28@10.[151,1209,168,1528]" captionTargetPageId="10" captionText="FIG. 7. — Heptnerina confusa n. gen., n. sp.,; A, habitus, dorsal; B, habitus, lateral; C, rostrum and labrum, lateral; D, leg 5 and genital somite, ventral; E, caudal ramus, ventral; F, leg 5. Scale bars: A, B, 0.2 mm; C, 0.1 mm; D-F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401388" httpUri="https://zenodo.org/record/5401388/files/figure.png" pageId="7" pageNumber="56">Fig. 7 D</figureCitation>
). Caudal rami (
<figureCitation id="132A2A440352FF9BE7B004A0C9535DC2" box="[722,845,458,485]" captionStart="FIG" captionStartId="10.[151,162,1547,1564]" captionTargetBox="[157,1194,173,1524]" captionTargetId="figure-28@10.[151,1209,168,1528]" captionTargetPageId="10" captionText="FIG. 7. — Heptnerina confusa n. gen., n. sp.,; A, habitus, dorsal; B, habitus, lateral; C, rostrum and labrum, lateral; D, leg 5 and genital somite, ventral; E, caudal ramus, ventral; F, leg 5. Scale bars: A, B, 0.2 mm; C, 0.1 mm; D-F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401388" httpUri="https://zenodo.org/record/5401388/files/figure.png" pageId="7" pageNumber="56">Fig. 7A, E</figureCitation>
) with few spinules on each side of ventral distal margin. Rostrum and labrum with anterior outgrowth (shown in lateral view in
<figureCitation id="132A2A440352FF9BE7A80740C9245E63" box="[714,826,555,581]" captionStart="FIG" captionStartId="10.[151,162,1547,1564]" captionTargetBox="[157,1194,173,1524]" captionTargetId="figure-28@10.[151,1209,168,1528]" captionTargetPageId="10" captionText="FIG. 7. — Heptnerina confusa n. gen., n. sp.,; A, habitus, dorsal; B, habitus, lateral; C, rostrum and labrum, lateral; D, leg 5 and genital somite, ventral; E, caudal ramus, ventral; F, leg 5. Scale bars: A, B, 0.2 mm; C, 0.1 mm; D-F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401388" httpUri="https://zenodo.org/record/5401388/files/figure.png" pageId="7" pageNumber="56">Figure 7C</figureCitation>
).
</paragraph>
<paragraph id="8BAE36C10352FF9BE7A80720CE3B5F02" blockId="7.[714,1247,267,1061]" pageId="7" pageNumber="56">Antennule (Fig. 8A, B) indistinctly 17-segment- ed, fifth, sixth, seventh segments incompletely divided. Setation formula as follows: 3, 5, 3, 2, 2, 2, 2, 1, 3, 2, 2, 2+a, 1, 2, 4, 2+a, 11+a; segmental homologies as: I-II, III-V, VI-VII, VIII, IX, X, XI, XII, XIII, XIV, XV, XVI, XVII, XVIII, XIX- XX, XXI-XXIII, XXIV-XXVIII.</paragraph>
<paragraph id="8BAE36C10352FF9BE7A80641C9745F43" blockId="7.[714,1247,267,1061]" pageId="7" pageNumber="56">Leg 1 (Fig. 8C): distal segments of right endopod fused partially.</paragraph>
<paragraph id="8BAE36C10352FF9BE7A80601C9065FE3" blockId="7.[714,1247,267,1061]" pageId="7" pageNumber="56">
Leg 5 (
<figureCitation id="132A2A440352FF9BE6750600C9755FA2" box="[791,875,874,901]" captionStart="FIG" captionStartId="10.[151,162,1547,1564]" captionTargetBox="[157,1194,173,1524]" captionTargetId="figure-28@10.[151,1209,168,1528]" captionTargetPageId="10" captionText="FIG. 7. — Heptnerina confusa n. gen., n. sp.,; A, habitus, dorsal; B, habitus, lateral; C, rostrum and labrum, lateral; D, leg 5 and genital somite, ventral; E, caudal ramus, ventral; F, leg 5. Scale bars: A, B, 0.2 mm; C, 0.1 mm; D-F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401388" httpUri="https://zenodo.org/record/5401388/files/figure.png" pageId="7" pageNumber="56">Fig. 7F</figureCitation>
): without endopod, one-segment- ed exopod with two internal setae, not present in female.
</paragraph>
<paragraph id="8BAE36C10352FF9BE7A806A1CE825803" blockId="7.[714,1247,267,1061]" pageId="7" pageNumber="56">
Leg 6 (
<figureCitation id="132A2A440352FF9BE67706A0C9755FC2" box="[789,875,970,997]" captionStart="FIG" captionStartId="10.[151,162,1547,1564]" captionTargetBox="[157,1194,173,1524]" captionTargetId="figure-28@10.[151,1209,168,1528]" captionTargetPageId="10" captionText="FIG. 7. — Heptnerina confusa n. gen., n. sp.,; A, habitus, dorsal; B, habitus, lateral; C, rostrum and labrum, lateral; D, leg 5 and genital somite, ventral; E, caudal ramus, ventral; F, leg 5. Scale bars: A, B, 0.2 mm; C, 0.1 mm; D-F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401388" httpUri="https://zenodo.org/record/5401388/files/figure.png" pageId="7" pageNumber="56">Fig. 7D</figureCitation>
) with three setae, outermost longest, innermost very short, as small spine, median seta bearing a few (one on
<typeStatus id="54AA88630352FF9BE6880160CE5F5803" box="[1002,1089,1035,1061]" pageId="7" pageNumber="56" type="allotype">allotype</typeStatus>
) setules.
</paragraph>
<paragraph id="8BAE36C10352FF9BE7A80121C9C158A2" blockId="7.[714,1247,1098,1669]" pageId="7" pageNumber="56">
<emphasis id="B965EAD30352FF9BE7A80121C9C158A2" italics="true" pageId="7" pageNumber="56">
Variation: dissected
<typeStatus id="54AA88630352FF9BE6CC0121CE0A5845" box="[942,1044,1098,1123]" pageId="7" pageNumber="56" type="paratype">paratypes</typeStatus>
(
<specimenCount id="9D17FD480352FF9BE1440121CEBF5843" box="[1062,1185,1098,1125]" pageId="7" pageNumber="56" type="female">six females</specimenCount>
,
<specimenCount id="9D17FD480352FF9BE1CD0121C91858A3" pageId="7" pageNumber="56" type="male">four males</specimenCount>
, six copepodid stages)
</emphasis>
</paragraph>
<paragraph id="8BAE36C10352FF9BE7A801E1CE4659E2" blockId="7.[714,1247,1098,1669]" pageId="7" pageNumber="56">
From the
<specimenCount id="9D17FD480352FF9BE65401E0C9815883" box="[822,927,1163,1189]" pageId="7" pageNumber="56" type="female">six female</specimenCount>
<typeStatus id="54AA88630352FF9BE6C401E1CE105882" box="[934,1038,1162,1188]" pageId="7" pageNumber="56" type="paratype">paratypes</typeStatus>
, three have leg 5 of general shape (as in
<figureCitation id="132A2A440352FF9BE6D201C0CE3658E3" box="[944,1064,1195,1221]" captionStart="FIG" captionStartId="9.[151,162,1524,1541]" captionTargetBox="[163,1180,177,1481]" captionTargetId="figure-25@9.[151,1184,170,1486]" captionTargetPageId="9" captionText="FIG. 6. — Heptnerina confusa n. gen., n. sp.,; A, holotype, leg 4; B, holotype, endopod of leg 4; C, holotype, leg 5 with endopod arrowed; D, paratype, leg 5. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5401386" httpUri="https://zenodo.org/record/5401386/files/figure.png" pageId="7" pageNumber="56">Figure 6D</figureCitation>
); in these specimens, the endopodal structure varies in size and proportion; the endopodal structure, when present, is not always symmetrical on right and left legs. In addition, it can be present on one side only. No muscle connected to the presumed endopod has been found within protopod. The three other
<typeStatus id="54AA88630352FF9BE63600E1C9DE5982" box="[852,960,1418,1444]" pageId="7" pageNumber="56" type="paratype">paratypes</typeStatus>
have a leg 5 lacking the endopod-like structure (Fig. 8G, H).
</paragraph>
<paragraph id="8BAE36C10352FF9BE7A800A1C9B95AA3" blockId="7.[714,1247,1098,1669]" pageId="7" pageNumber="56">
In
<specimenCount id="9D17FD480352FF9BE78B00A1C95059C3" box="[745,846,1482,1509]" pageId="7" pageNumber="56" type="adult">one adult</specimenCount>
male, leg 1 does not show any fusion of the distal endopodal segments, as it is the condition in the
<typeStatus id="54AA88630352FF9BE6EA0360C9FC5A03" box="[904,994,1547,1573]" pageId="7" pageNumber="56" type="allotype">allotype</typeStatus>
. Leg 5 lacks the endopod, as in the
<typeStatus id="54AA88630352FF9BE60F0340C9D95A63" box="[877,967,1579,1605]" pageId="7" pageNumber="56" type="allotype">allotype</typeStatus>
. Leg 5 of the three other
<typeStatus id="54AA88630352FF9BE7A80321C9395A42" box="[714,807,1610,1636]" pageId="7" pageNumber="56" type="paratype">paratype</typeStatus>
males lacks the endopodal structure, as in the
<typeStatus id="54AA88630352FF9BE6720300C97B5AA3" box="[784,869,1643,1669]" pageId="7" pageNumber="56" type="allotype">allotype</typeStatus>
male.
</paragraph>
<caption id="DF6E6649035DFF94E5F500C5C89259F2" ID-DOI="http://doi.org/10.5281/zenodo.5401384" ID-Zenodo-Dep="5401384" httpUri="https://zenodo.org/record/5401384/files/figure.png" pageId="8" pageNumber="57" startId="8.[151,162,1454,1471]" targetBox="[173,1227,178,1419]" targetPageId="8">
<paragraph id="8BAE36C1035DFF94E5F500C5C89259F2" blockId="8.[151,1248,1454,1492]" pageId="8" pageNumber="57">
FIG. 5. —
<taxonomicName id="4C114D42035DFF94E58B00C5CB925999" authority="Ivanenko &amp; Defaye, 2004" authorityName="Ivanenko &amp; Defaye" authorityYear="2004" box="[233,396,1454,1471]" class="Hexanauplia" family="Cyclopinidae" genus="Heptnerina" kingdom="Animalia" order="Cyclopoida" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="confusa" status="n. gen., n. sp.">
<emphasis id="B965EAD3035DFF94E58B00C5CB925999" box="[233,396,1454,1471]" italics="true" pageId="8" pageNumber="57">Heptnerina confusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25657A8035DFF94E4F300C5C8175999" box="[401,521,1454,1471]" pageId="8" pageNumber="57" rank="species">n. gen., n. sp.</taxonomicNameLabel>
, holotype;
<emphasis id="B965EAD3035DFF94E71A00C5C89B5999" bold="true" box="[632,645,1454,1471]" pageId="8" pageNumber="57">A</emphasis>
, leg 1, anterior;
<emphasis id="B965EAD3035DFF94E66D00C5C9025999" bold="true" box="[783,796,1454,1471]" pageId="8" pageNumber="57">B</emphasis>
, leg 2, anterior;
<emphasis id="B965EAD3035DFF94E6C700C5C9AD5999" bold="true" box="[933,947,1454,1471]" pageId="8" pageNumber="57">C</emphasis>
, exopod of leg 2, anterior;
<emphasis id="B965EAD3035DFF94E1FA00C5CEB85999" bold="true" box="[1176,1190,1454,1471]" pageId="8" pageNumber="57">D</emphasis>
, leg 3, anterior;
<emphasis id="B965EAD3035DFF94E58300A8CAF359F2" bold="true" box="[225,237,1475,1492]" pageId="8" pageNumber="57">E</emphasis>
, endopod of leg 3, anterior. Scale bars: 0.05 mm.
</paragraph>
</caption>
<paragraph id="8BAE36C1035DFF94E5F50360CB165AA3" blockId="8.[151,684,1546,1669]" pageId="8" pageNumber="57">
Female copepodid stage V: leg 5 as in Figure 8E. Male copepodid stage V: leg 5 as in Figure 8D. Copepodid stage IV (
<specimenCount id="9D17FD48035DFF94E4E30320C8355A43" box="[385,555,1611,1637]" pageId="8" pageNumber="57" type="generic">four specimens</specimenCount>
): leg 5 as in Figure
<specimenCount id="9D17FD48035DFF94E5810300CB165AA3" box="[227,264,1643,1669]" pageId="8" pageNumber="57" type="generic">8F.</specimenCount>
</paragraph>
</subSubSection>
</treatment>
</document>
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