147 lines
17 KiB
XML
147 lines
17 KiB
XML
<document id="3BB28A298542F2B3669746408EFC12EE" ID-DOI="10.11646/zootaxa.5047.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5540714" ID-ZooBank="072B07D8-324A-412E-A76E-C39067AC77AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632985111669" checkinUser="plazi" docAuthor="Saucier, Esprit Heestand, France, Scott C. & Watling, Les" docDate="2021" docId="03B887C4FF9DFFDDD3CD672F0A04F9CF" docLanguage="en" docName="zootaxa.5047.3.2.pdf" docOrigin="Zootaxa 5047 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Isididae Lamouroux 1812" docType="treatment" docVersion="3" lastPageNumber="266" masterDocId="FF81FFBCFF8EFFCED35A62150F55FE12" masterDocTitle="Toward a revision of the bamboo corals: Part 3, deconstructing the Family Isididae" masterLastPageNumber="272" masterPageNumber="247" pageNumber="266" updateTime="1699416498060" updateUser="plazi" zenodo-license-document="CLOSED">
|
||
<mods:mods id="ABB6009E2D1AFEDB9D83E0D4E9DBB8E0" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="CD47614D34E7D8AE8B207E5289B93F28">
|
||
<mods:title id="79F35F931FC3F58FDC296632E7E823C7">Toward a revision of the bamboo corals: Part 3, deconstructing the Family Isididae</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="D1CA85910917B308DB24B5425D19DE6F" type="personal">
|
||
<mods:role id="1A5A87CEEF8B4B8C6CD1416E47AE6AB0">
|
||
<mods:roleTerm id="59327D136A43A3D96A60AA035FB7010B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="38F3AA528434956C969127EA9F804006">Saucier, Esprit Heestand</mods:namePart>
|
||
<mods:affiliation id="E632F3C061A4BDA02D1BE69453F95D7E">Faculty of Science, Brigham Young University - Hawaii, 55 - 220 Kulanui St, Laie, Hawaii, 96762, USA</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="37B24D7A04256476D6014B985C4DF7CC" type="personal">
|
||
<mods:role id="A3A8B741F9A4C12598AE982D5C73C576">
|
||
<mods:roleTerm id="3A1E68390352BDD9450DBD10AA9FB18A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1AB0D0BEC98A5F57AAE60E7E0BC6E9E4">France, Scott C.</mods:namePart>
|
||
<mods:affiliation id="A21901A381EF2061E233C33EE7A903E1">Department of Biology, University of Louisiana at Lafayette, Lafayette, Louisiana, USA</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="DA66D97D056EE414D951CF077914A356" type="personal">
|
||
<mods:role id="8EC711D333DE9D9C581CB59EA34EFED5">
|
||
<mods:roleTerm id="C23BE749AB686EF0A970698381E23377">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2EA5A122C9EACB7A269E26137C3299D3">Watling, Les</mods:namePart>
|
||
<mods:affiliation id="DFFFADB6A70D1EE4F2C57A64A29334CD">School of Life Sciences, University of Hawaii at Manoa, Honolulu, Hawaii 96822</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="F5B962B035BC93574E23674E5ECEE262">text</mods:typeOfResource>
|
||
<mods:relatedItem id="1281B1350429D5FC0BCF88630E572DA5" type="host">
|
||
<mods:titleInfo id="08EC474421A02383DE70BCC8F087808A">
|
||
<mods:title id="E1DCA8FA89C379F562E1AA2CEFAAFB37">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="6F83C57E1502883409D2C1562FC865F4">
|
||
<mods:date id="3E656CC0565BD6C943E2210FFA4E89FB">2021</mods:date>
|
||
<mods:detail id="DBC3D460FEB8B0F79DB1D1499225BCB7" type="pubDate">
|
||
<mods:number id="173E0B52567B8839CBF355D7B7CC9035">2021-09-30</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="1BB2D5251E0CE78D01838525297C6ADE" type="volume">
|
||
<mods:number id="F988E6F4AE7C8F454BF2E4A4377C3FB3">5047</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="8300E9738388FD3B1D2689F2991EBFA2" type="issue">
|
||
<mods:number id="2941F49B709B78989AE9A4E70B0F23BC">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="DF1672DEA28EB397C79C6417E3F91F50" unit="page">
|
||
<mods:start id="E4C17AF9384D38ED1959DA68B899A03B">247</mods:start>
|
||
<mods:end id="C63FB8BCACA725C19F30E9D05CC9C2F2">272</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="C3DCEE6183D6245F666FFC630B26C716">journal article</mods:classification>
|
||
<mods:identifier id="5305F152AFA5370F6BF58235414AE09B" type="DOI">10.11646/zootaxa.5047.3.2</mods:identifier>
|
||
<mods:identifier id="63A8C675E528A5B6081CBFBFEACC78C0" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="9CA02B53AF7C4C366DB475D5C972D176" type="Zenodo-Dep">5540714</mods:identifier>
|
||
<mods:identifier id="CFE76AB862FF9AA58710514EA54B04F5" type="ZooBank">072B07D8-324A-412E-A76E-C39067AC77AE</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03B887C4FF9DFFDDD3CD672F0A04F9CF" ID-DOI="http://doi.org/10.5281/zenodo.5540720" ID-Zenodo-Dep="5540720" LSID="urn:lsid:plazi:treatment:03B887C4FF9DFFDDD3CD672F0A04F9CF" httpUri="http://treatment.plazi.org/id/03B887C4FF9DFFDDD3CD672F0A04F9CF" lastPageNumber="266" pageId="19" pageNumber="266">
|
||
<subSubSection id="C30B6559FF9DFFDDD3CD672F0D6AFB46" box="[151,575,1338,1365]" pageId="19" pageNumber="266" type="nomenclature">
|
||
<paragraph id="8BAE36D2FF9DFFDDD3CD672F0D6AFB46" blockId="19.[151,575,1338,1365]" box="[151,575,1338,1365]" pageId="19" pageNumber="266">
|
||
<heading id="D0E681BEFF9DFFDDD3CD672F0D6AFB46" bold="true" box="[151,575,1338,1365]" fontSize="11" level="1" pageId="19" pageNumber="266" reason="1">
|
||
<emphasis id="B965EAC0FF9DFFDDD3CD672F0D6AFB46" bold="true" box="[151,575,1338,1365]" pageId="19" pageNumber="266">
|
||
Family
|
||
<taxonomicName id="4C114D51FF9DFFDDD3AD672F0D6AFB46" ID-CoL="84KBH" authority="Lamouroux, 1812" authorityName="Lamouroux" authorityYear="1812" box="[247,575,1338,1365]" class="Anthozoa" family="Isididae" kingdom="Animalia" order="Alcyonacea" pageId="19" pageNumber="266" phylum="Cnidaria" rank="family">
|
||
Isididae
|
||
<bibRefCitation id="EF804B23FF9DFFDDD23B672E0D6AFB46" author="Lamouroux, J. V. F." box="[353,575,1338,1365]" pageId="19" pageNumber="266" pagination="181 - 188" refId="ref16281" refString="Lamouroux, J. V. F. (1812) Extrait d'un memoire sur la classification des Polypiers coralligenes non entierement pierreux. Nouveau Bulletin des Sciences, par la Societe Philomatique, Paris, 3, 181 - 188." type="journal article" year="1812">Lamouroux, 1812</bibRefCitation>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B6559FF9DFFDDD3CD67960D5DFB8F" box="[151,520,1410,1437]" pageId="19" pageNumber="266" type="type_taxon">
|
||
<paragraph id="8BAE36D2FF9DFFDDD3CD67960D5DFB8F" blockId="19.[151,1437,1410,2013]" box="[151,520,1410,1437]" pageId="19" pageNumber="266">
|
||
<emphasis id="B965EAC0FF9DFFDDD3CD67960E73FB8F" bold="true" box="[151,294,1411,1437]" pageId="19" pageNumber="266">
|
||
<typeStatus id="54AA8870FF9DFFDDD3CD67960F87FB8F" box="[151,210,1411,1437]" pageId="19" pageNumber="266">Type</typeStatus>
|
||
genus:
|
||
</emphasis>
|
||
<taxonomicName id="4C114D51FF9DFFDDD27467960D5DFB8F" ID-CoL="84QRG" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[302,520,1410,1437]" class="Anthozoa" family="Isididae" genus="Isis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="19" pageNumber="266" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B965EAC0FF9DFFDDD27467960E00FB8E" box="[302,341,1411,1436]" italics="true" pageId="19" pageNumber="266">Isis</emphasis>
|
||
Linnaeus, 1758
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B6559FF9DFFDDD39D67B30D21F88B" pageId="19" pageNumber="266" type="diagnosis">
|
||
<paragraph id="8BAE36D2FF9DFFDDD39D67B30B55F867" blockId="19.[151,1437,1410,2013]" pageId="19" pageNumber="266">
|
||
<emphasis id="B965EAC0FF9DFFDDD39D67B30E17FBD2" bold="true" box="[199,322,1446,1472]" pageId="19" pageNumber="266">Diagnosis.</emphasis>
|
||
Colony with articulated skeleton of gorgonin nodes alternating with calcareous internodes that are solid and composed of radiating bundles of sclerite-like units. Branches originate from the internodes, always beginning with a node at the base of the branch. Coenenchyme thick; polyps completely retractile into small pockets in the coenenchyme, leaving only small pores visible on the surface. Polyps mostly devoid of sclerites, with small rods only in the tentacles. Coenenchyme sclerites mostly small clubs, but may include 6-, 7- or 8-radiates, warty rods, spindles and crosses, heavily sculpted capstans, dumbbells, and double cones.
|
||
</paragraph>
|
||
<paragraph id="8BAE36D2FF9DFFDDD39D646A0D21F88B" blockId="19.[151,1437,1410,2013]" box="[199,628,1662,1689]" pageId="19" pageNumber="266">
|
||
<emphasis id="B965EAC0FF9DFFDDD39D646A0EC7F88B" bold="true" box="[199,402,1663,1689]" pageId="19" pageNumber="266">Included genera:</emphasis>
|
||
<taxonomicName id="4C114D51FF9DFFDDD2C3646A0D21F88B" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[409,628,1662,1689]" class="Anthozoa" family="Isididae" genus="Isis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="19" pageNumber="266" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B965EAC0FF9DFFDDD2C3646A0E95F88A" box="[409,448,1663,1688]" italics="true" pageId="19" pageNumber="266">Isis</emphasis>
|
||
Linnaeus, 1758
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B6559FF9DFFDDD39D64B60A04F9CF" pageId="19" pageNumber="266" type="discussion">
|
||
<paragraph id="8BAE36D2FF9DFFDDD39D64B60A04F9CF" blockId="19.[151,1437,1410,2013]" pageId="19" pageNumber="266">
|
||
<emphasis id="B965EAC0FF9DFFDDD39D64B60E6EF8AF" bold="true" box="[199,315,1699,1725]" pageId="19" pageNumber="266">Remarks.</emphasis>
|
||
The genus
|
||
<taxonomicName id="4C114D51FF9DFFDDD29D64B60D06F8AF" box="[455,595,1699,1725]" class="Anthozoa" family="Isididae" genus="Chelidonisis" kingdom="Animalia" order="Alcyonacea" pageId="19" pageNumber="266" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B965EAC0FF9DFFDDD29D64B60D06F8AF" box="[455,595,1699,1725]" italics="true" pageId="19" pageNumber="266">Chelidonisis</emphasis>
|
||
</taxonomicName>
|
||
has routinely been included in the
|
||
<taxonomicName id="4C114D51FF9DFFDDD0A864B60B1EF8AF" authorityName="Lamouroux" authorityYear="1812" box="[1010,1099,1699,1725]" class="Anthozoa" family="Isididae" kingdom="Animalia" order="Alcyonacea" pageId="19" pageNumber="266" phylum="Cnidaria" rank="family">Isididae</taxonomicName>
|
||
subfamily
|
||
<taxonomicName id="4C114D51FF9DFFDDD79564B60A7DF8AF" box="[1231,1320,1699,1725]" class="Anthozoa" family="Gorgoniidae" kingdom="Animalia" order="Alcyonacea" pageId="19" pageNumber="266" phylum="Cnidaria" rank="subFamily" subFamily="Isidinae">Isidinae</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C114D51FF9DFFDDD63464B60AC0F8AE" authorityName="Linnaeus" authorityYear="1758" box="[1390,1429,1699,1724]" class="Anthozoa" family="Isididae" genus="Isis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="19" pageNumber="266" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B965EAC0FF9DFFDDD63464B60AC0F8AE" box="[1390,1429,1699,1724]" italics="true" pageId="19" pageNumber="266">Isis</emphasis>
|
||
</taxonomicName>
|
||
, chiefly on the basis of the presence of 6-radiate sclerites in the coenenchyme. However, there are several morphological differences that distinguish the two genera. The axis of
|
||
<taxonomicName id="4C114D51FF9DFFDDD0C564FE0C93F916" authorityName="Linnaeus" authorityYear="1758" box="[927,966,1771,1796]" class="Anthozoa" family="Isididae" genus="Isis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="19" pageNumber="266" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B965EAC0FF9DFFDDD0C564FE0C93F916" box="[927,966,1771,1796]" italics="true" pageId="19" pageNumber="266">Isis</emphasis>
|
||
</taxonomicName>
|
||
is thick with smooth longitudinal ridges on the internodes, whereas in
|
||
<taxonomicName id="4C114D51FF9DFFDDD2B7651A0D2CF93B" box="[493,633,1807,1833]" class="Anthozoa" family="Isididae" genus="Chelidonisis" kingdom="Animalia" order="Alcyonacea" pageId="19" pageNumber="266" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B965EAC0FF9DFFDDD2B7651A0D2CF93B" box="[493,633,1807,1833]" italics="true" pageId="19" pageNumber="266">Chelidonisis</emphasis>
|
||
</taxonomicName>
|
||
the axis is slender and the ridges on the internodes are festooned with small spines. The coenenchyme of
|
||
<taxonomicName id="4C114D51FF9DFFDDD17565260D03F95E" authorityName="Linnaeus" authorityYear="1758" box="[559,598,1843,1868]" class="Anthozoa" family="Isididae" genus="Isis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="19" pageNumber="266" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B965EAC0FF9DFFDDD17565260D03F95E" box="[559,598,1843,1868]" italics="true" pageId="19" pageNumber="266">Isis</emphasis>
|
||
</taxonomicName>
|
||
is very thick and encompasses retractile polyps as compared to the thin coenenchyme of
|
||
<taxonomicName id="4C114D51FF9DFFDDD20165420EB2F963" box="[347,487,1879,1905]" class="Anthozoa" family="Isididae" genus="Chelidonisis" kingdom="Animalia" order="Alcyonacea" pageId="19" pageNumber="266" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B965EAC0FF9DFFDDD20165420EB2F963" box="[347,487,1879,1905]" italics="true" pageId="19" pageNumber="266">Chelidonisis</emphasis>
|
||
</taxonomicName>
|
||
, wherein the polyps contract into armored verrucae on the branch. In addition, the coenenchyme sclerites of
|
||
<taxonomicName id="4C114D51FF9DFFDDD2ED656E0E8BF986" authorityName="Linnaeus" authorityYear="1758" box="[439,478,1915,1940]" class="Anthozoa" family="Isididae" genus="Isis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="19" pageNumber="266" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B965EAC0FF9DFFDDD2ED656E0E8BF986" box="[439,478,1915,1940]" italics="true" pageId="19" pageNumber="266">Isis</emphasis>
|
||
</taxonomicName>
|
||
are very diverse capstans, tuberculate spindles, and clubs, whereas in
|
||
<taxonomicName id="4C114D51FF9DFFDDD7B2656E0A21F987" box="[1256,1396,1915,1941]" class="Anthozoa" family="Isididae" genus="Chelidonisis" kingdom="Animalia" order="Alcyonacea" pageId="19" pageNumber="266" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B965EAC0FF9DFFDDD7B2656E0A21F987" box="[1256,1396,1915,1941]" italics="true" pageId="19" pageNumber="266">Chelidonisis</emphasis>
|
||
</taxonomicName>
|
||
the sclerites are predominantly 6-radiates and spindles, and capstans are never present. Finally, the two genera occur in widely separated clades on the molecular phylogeny (
|
||
<figureCitation id="132A2A57FF9DFFDDD1AA65D60C08F9CF" box="[752,861,1987,2013]" captionStart="FIGURE 2" captionStartId="13.[151,250,1709,1733]" captionTargetBox="[153,1434,858,1682]" captionTargetId="figure-370@13.[151,1436,848,1685]" captionTargetPageId="13" captionText="FIGURE 2. Maximum likelihood tree showing the divergence among the subfamilies of the family Isididae sensu lato. The tree is based on a concatenated alignment of mtMutS-5’, mtMutS-3’, and 18S and includes 61 total taxa (26 isidids). The clades that include taxa currently classified as Isididae are indicated in bold font." figureDoi="http://doi.org/10.5281/zenodo.5540722" httpUri="https://zenodo.org/record/5540722/files/figure.png" pageId="19" pageNumber="266">Figures 2</figureCitation>
|
||
,
|
||
<figureCitation id="132A2A57FF9DFFDDD03365D60C2DF9CF" box="[873,888,1987,2013]" captionStart="FIGURE 3" captionStartId="14.[151,250,1576,1600]" captionTargetBox="[160,1423,605,1545]" captionTargetId="figure-213@14.[151,1436,597,1553]" captionTargetPageId="14" captionText="FIGURE 3. Midpoint rooted RAxML cladogram. Bootstrap values> 50 are shown at the nodes. Baysian posterior probability values>0.5 are shown below the node. The tree is based on a concatenated alignment of mtMutS-5’, mtMutS-3’, and 18S, and includes 61 total taxa (26 isidids). Clades with an articulated skeleton are marked with a large asterisk (*) on the branch. The type genus of the Isididae is indicated by the black rectangle and Isididae sensu lato are indicated by the purple rectangles. Light yellow circles indicate clades that have the “ancestral” mt genome arrangement (as defined by Brockman & McFadden, 2012). The dark blue circle indicates the clade that has the Keratoisidinae mt genome arrangement mapped by Brugler and France (2008). Only isidid mt genomes are mapped on the tree. Small asterisks (*) following taxon names indicate specimens used to test mt gene boundaries. The “S1 clade” of Keratoisidinae is the one with taxa Cladarisis sp. 1 and sp. 2." figureDoi="http://doi.org/10.5281/zenodo.5540724" httpUri="https://zenodo.org/record/5540724/files/figure.png" pageId="19" pageNumber="266">3</figureCitation>
|
||
), as also shown by
|
||
<bibRefCitation id="EF804B23FF9DFFDDD70E65D60A18F9CF" author="Quattrini, A. M. & Rodriguez, E. & Faircloth, B. C. & Cowman, P. F. & Brugler, M. R. & Farfan, G. A. & Hellberg, M. E. & Kitahara, M. V. & Morrison, C. L. & Paz-Garcia, D. A. & Reimer, J. D." box="[1108,1357,1987,2013]" pageId="19" pageNumber="266" pagination="1531 - 1538" refId="ref17038" refString="Quattrini, A. M., Rodriguez, E., Faircloth, B. C., Cowman, P. F., Brugler, M. R., Farfan, G. A., Hellberg, M. E., Kitahara, M. V., Morrison, C. L., Paz-Garcia, D. A. & Reimer, J. D. (2020) Palaeoclimate ocean conditions shaped the evolution of corals and their skeletons through deep time. Nature Ecology & Evolution, 4, 1531 - 1538. https: // doi. org / 10.1038 / s 41559 - 020 - 01291 - 1" type="journal article" year="2020">
|
||
Quattrini
|
||
<emphasis id="B965EAC0FF9DFFDDD79A65D60BADF9CF" box="[1216,1272,1987,2013]" italics="true" pageId="19" pageNumber="266">et al.</emphasis>
|
||
(2020)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |