354 lines
48 KiB
XML
354 lines
48 KiB
XML
<document id="A9FFA2D3220651AE9C1B3C18643FA45C" ID="10.11646/zootaxa.4250.5.5SLASH10746" ID-CLB-Dataset="33274" ID-DOI="10.11646/zootaxa.4250.5.5" ID-GBIF-Dataset="3e3123a6-d497-4ef4-afa0-7cf0ef7f9afa" ID-ISSN="1175-5326" ID-Zenodo-Dep="495550" ID-ZooBank="5448198D-7CF2-47CE-A3CF-E7760EC5DDB6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1491551433022" checkinUser="plazi" docAuthor="Ng, Peter K. L. & Lemaitre, Rafael" docDate="2017" docId="03B88780FF92EC4560A13414FDA9B7A1" docLanguage="en" docName="zootaxa.4250.5.5SLASH10746.pdf" docOrigin="Zootaxa 4250 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geosesarma rathbunae Serene 1968" docType="treatment" docVersion="7" lastPageNumber="482" masterDocId="FF81FFF8FF90EC426036332FFFF0B23C" masterDocTitle="On the taxonomy of two poorly known species of Geosesarma De Man, 1892 (Crustacea: Brachyura: Sesarmidae) from the Philippines" masterLastPageNumber="483" masterPageNumber="475" pageNumber="477" updateTime="1698438789559" updateUser="ExternalLinkService">
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<mods:titleInfo id="50F683B4166D0A85B3AE290CE8E5710D">
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<mods:title id="259B57BD2CB192CA5B56048FB458BFA2">On the taxonomy of two poorly known species of Geosesarma De Man, 1892 (Crustacea: Brachyura: Sesarmidae) from the Philippines</mods:title>
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<mods:name id="4B85E839443B7B209401BC57FD143565" type="personal">
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<mods:namePart id="CF4DBF0DC525ECE7ED1F545AEDA29838">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="5A2DCB6FE6AB6CE8B11B5B17795E3877">Lemaitre, Rafael</mods:namePart>
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<mods:title id="6B3AA1D3141B7485B55BDFAB678EEDC8">Zootaxa</mods:title>
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<mods:date id="18A2E3EDACFC21C2A7148D512996C42A">2017</mods:date>
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<mods:number id="C6EC975F1A21D6BD1BF360B4B0672C2F">4250</mods:number>
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<treatment id="03B88780FF92EC4560A13414FDA9B7A1" ID-DOI="http://doi.org/10.5281/zenodo.6028325" ID-GBIF-Taxon="128344777" ID-Zenodo-Dep="6028325" LSID="urn:lsid:plazi:treatment:03B88780FF92EC4560A13414FDA9B7A1" httpUri="http://treatment.plazi.org/id/03B88780FF92EC4560A13414FDA9B7A1" lastPageId="7" lastPageNumber="482" pageId="2" pageNumber="477">
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<subSubSection id="C30B651DFF92EC4060A13414FE91B54B" pageId="2" pageNumber="477" type="nomenclature">
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<paragraph id="8BAE3696FF92EC4060A13414FD87B56A" blockId="2.[151,631,1851,1911]" box="[151,631,1851,1878]" pageId="2" pageNumber="477">
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<heading id="D0E681FAFF92EC4060A13414FD87B56A" bold="true" box="[151,631,1851,1878]" fontSize="11" level="1" pageId="2" pageNumber="477" reason="1">
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<taxonomicName id="4C114D15FF92EC4060A13414FD87B56A" ID-CoL="3FS9Y" authority="Serene, 1968a" authorityName="Serene" authorityYear="1968" box="[151,631,1851,1878]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="rathbunae">
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<emphasis id="B965EA84FF92EC4060A13414FD87B56A" bold="true" box="[151,631,1851,1878]" pageId="2" pageNumber="477">
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<emphasis id="B965EA84FF92EC4060A13414FE5FB569" bold="true" box="[151,431,1851,1877]" italics="true" pageId="2" pageNumber="477">Geosesarma rathbunae</emphasis>
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(
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<bibRefCitation id="EF804B67FF92EC4061893413FD9FB56A" author="Serene" box="[447,623,1852,1878]" pageId="2" pageNumber="477" refString="Serene, R. (1968 a) Note preliminaire sur de nouvelles especes de Sesarma (Decapoda Brachyura). Bulletin du Museum national d'Histoire naturelle, 39 (5), 1084 - 1095." type="journal article" year="1968" yearSuffix="a">Serène, 1968a</bibRefCitation>
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)
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||
</emphasis>
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</taxonomicName>
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||
</heading>
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</paragraph>
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<paragraph id="8BAE3696FF92EC4060A13471FE91B54B" blockId="2.[151,631,1851,1911]" box="[151,353,1886,1911]" pageId="2" pageNumber="477">
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(
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<figureCitation id="132A2A13FF92EC4060A93471FF1DB54B" box="[159,237,1886,1911]" captionStart="FIGURE 3" captionStartId="4.[151,250,1893,1915]" captionTargetBox="[151,1436,193,1872]" captionTargetId="figure@4.[151,1436,193,1872]" captionTargetPageId="4" captionText="FIGURE 3. Geosesarma rathbunae (Serène, 1968 a). A, holotype male (13.0 × 12.2 mm) (USNM 54315) (after Serène, 1968 a: pl. 1 fig. 4); B – I, non-type male (13.1 × 12.3 mm) (USNM 123208). A, B, overall habitus; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of right chelae at different angles; H, I, dorsal views of right chelipeds at different angles." httpUri="https://zenodo.org/record/495553/files/figure.png" pageId="2" pageNumber="477">Figs. 3</figureCitation>
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,
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<figureCitation id="132A2A13FF92EC4060CC3470FEFAB54B" box="[250,266,1887,1911]" captionStart="FIGURE 4" captionStartId="5.[151,250,1836,1858]" captionTargetBox="[151,1436,193,1814]" captionTargetId="figure@5.[151,1436,193,1814]" captionTargetPageId="5" captionText="FIGURE 4. Geosesarma rathbunae (Serène, 1968 a). A, paratype female (13.5 × 12.1 mm) (USNM 1277697, ex USNM 123208); B – G, non-type male (14.9 × 14.5 mm) (USNM 123208); H, non-type female (14.1 × 12.6 mm) (USNM ex 54315); I, non-type male (10.4 × 9.4 mm) (USNM 123208). A, B, H, I, overall habitus; C, frontal view of cephalothorax; D, E, ventral view of cephalothorax showing thoracic sternum and pleon; F, G, outer views of left chelae at different angles." httpUri="https://zenodo.org/record/495554/files/figure.png" pageId="2" pageNumber="477">4</figureCitation>
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,
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<figureCitation id="132A2A13FF92EC4061203471FED2B54B" box="[278,290,1886,1911]" captionStart="FIGURE 5" captionStartId="6.[151,250,1840,1862]" captionTargetBox="[173,1416,210,1804]" captionTargetId="figure@6.[151,1436,193,1819]" captionTargetPageId="6" captionText="FIGURE 5. A – D, Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758); E – H, Geosesarma rathbunae (Serène, 1968 a), male (13.1 × 12.3 mm) (USNM 123208). A, E, left third maxilliped; B, F, ventral view of left G 1; C, G, mesial view of left G 1; D, H, dorsal view of left G 1. Scales = 1.0 mm." httpUri="https://zenodo.org/record/495555/files/figure.png" pageId="2" pageNumber="477">5</figureCitation>
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E–H)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30B651DFF92EC4060A1348DFB82B5C8" pageId="2" pageNumber="477" type="reference_group">
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||
<paragraph id="8BAE3696FF92EC4060A1348DFD20B5EA" blockId="2.[151,1138,1953,2037]" pageId="2" pageNumber="477">
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||
<treatmentCitationGroup id="AB0111B8FF92EC4060A1348DFD20B5EA" pageId="2" pageNumber="477">
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||
<taxonomicName id="4C114D15FF92EC4060A1348DFD3AB58B" ID-CoL="4WYV5" authority="Serene, 1968a: 1086" authorityName="Serene" authorityPageNumber="1086" authorityYear="1968" box="[151,714,1953,1975]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="rathbunae" subGenus="Geosesarma">
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<emphasis id="B965EA84FF92EC4060A1348DFF1EB58B" box="[151,238,1954,1975]" italics="true" pageId="2" pageNumber="477">Sesarma</emphasis>
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||
(
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<emphasis id="B965EA84FF92EC4060CB348DFE8BB58B" box="[253,379,1954,1975]" italics="true" pageId="2" pageNumber="477">Geosesarma</emphasis>
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)
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<emphasis id="B965EA84FF92EC4061BC348EFE02B58B" box="[394,498,1953,1975]" italics="true" pageId="2" pageNumber="477">rathbunae</emphasis>
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<treatmentCitation id="0AB01087FF92EC4061CE348EFD3AB58B" author="Serene" box="[504,714,1953,1975]" page="1086" pageId="2" pageNumber="477" year="1968">
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||
<bibRefCitation id="EF804B67FF92EC4061CE348EFD7CB58B" author="Serene" box="[504,652,1953,1975]" pageId="2" pageNumber="477" refString="Serene, R. (1968 a) Note preliminaire sur de nouvelles especes de Sesarma (Decapoda Brachyura). Bulletin du Museum national d'Histoire naturelle, 39 (5), 1084 - 1095." type="journal article" year="1968" yearSuffix="a">Serène, 1968a</bibRefCitation>
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: 1086
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||
</treatmentCitation>
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||
</taxonomicName>
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||
, figs. 5, 6; pl. 1 fig. 4.
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<taxonomicName id="4C114D15FF92EC4060A134EEFE02B5EA" ID-CoL="4WYV5" box="[151,498,1984,2006]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="rathbunae" subGenus="Geosesarma">
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||
<emphasis id="B965EA84FF92EC4060A134EEFF1EB5EA" box="[151,238,1985,2006]" italics="true" pageId="2" pageNumber="477">Sesarma</emphasis>
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||
(
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||
<emphasis id="B965EA84FF92EC4060CB34EEFE8BB5EA" box="[253,379,1985,2006]" italics="true" pageId="2" pageNumber="477">Geosesarma</emphasis>
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||
)
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<emphasis id="B965EA84FF92EC4061BC34EFFE02B5EA" box="[394,498,1984,2006]" italics="true" pageId="2" pageNumber="477">rathbunae</emphasis>
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||
</taxonomicName>
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||
—
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||
<treatmentCitation id="0AB01087FF92EC40623A34EFFD3CB5EA" author="Serene" box="[524,716,1984,2006]" page="106" pageId="2" pageNumber="477" year="1968">
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||
<bibRefCitation id="EF804B67FF92EC40623A34EFFD6AB5EA" author="Serene" box="[524,666,1984,2006]" pageId="2" pageNumber="477" refString="Serene, R. (1968 b) The Brachyura of the Indo-West Pacific Region. In: Prodromus for a Check List of the (non-planctonic) Marine Fauna of South East Asia. Singapore National Academy of Sciences, Special Publication No. 1, Fauna IIICc 3, pp. 33 - 112." type="journal article" year="1968" yearSuffix="b">Serène 1968b</bibRefCitation>
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||
: 106
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
|
||
</paragraph>
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||
<paragraph id="8BAE3696FF92EC4060A134F0FB82B5C8" blockId="2.[151,1138,1953,2037]" box="[151,1138,2014,2037]" pageId="2" pageNumber="477">
|
||
<treatmentCitationGroup id="AB0111B8FF92EC4060A134F0FB82B5C8" box="[151,1138,2014,2037]" pageId="2" pageNumber="477">
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||
<taxonomicName id="4C114D15FF92EC4060A134F0FE74B5C8" ID-CoL="3FS9Y" box="[151,388,2014,2036]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="rathbunae">
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||
<emphasis id="B965EA84FF92EC4060A134F0FE74B5C8" box="[151,388,2014,2036]" italics="true" pageId="2" pageNumber="477">Geosesarma rathbunae</emphasis>
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</taxonomicName>
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—
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<treatmentCitation id="0AB01087FF92EC4061AB34F1FD69B5C8" author="Serene" box="[413,665,2014,2036]" page="403" pageId="2" pageNumber="477" year="1970">
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||
<bibRefCitation id="EF804B67FF92EC4061AB34F1FD96B5C8" author="Serene" box="[413,614,2014,2036]" pageId="2" pageNumber="477" refString="Serene, R. & Soh, C. L. (1970) New Indo-Pacific genera allied to Sesarma Say, 1877 (Brachyura, Decapoda, Crustacea). Treubia, 27 (4), 387 - 416." type="journal article" year="1970">Serène & Soh 1970</bibRefCitation>
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||
: 403
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</treatmentCitation>
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; Ng 1988: 119 (list); Ng
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<emphasis id="B965EA84FF92EC4063A034CFFC39B5C8" box="[918,969,2014,2037]" italics="true" pageId="2" pageNumber="477">et al.</emphasis>
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2008: 221 (list).
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</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<caption id="DF6E661EFF93EC4160A137D2FB20B70D" httpUri="https://zenodo.org/record/495552/files/figure.png" pageId="3" pageNumber="478" targetBox="[180,1408,187,1255]" targetPageId="3">
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||
<paragraph id="8BAE3696FF93EC4160A137D2FB20B70D" blockId="3.[151,1436,1276,1329]" pageId="3" pageNumber="478">
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<emphasis id="B965EA84FF93EC4160A137D2FEE1B72E" bold="true" box="[151,273,1277,1299]" pageId="3" pageNumber="478">FIGURE 2</emphasis>
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.
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<taxonomicName id="4C114D15FF93EC41611437D2FD24B72E" authority="De Man, 1892" authorityName="De Man" authorityYear="1892" box="[290,724,1276,1298]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="478" phylum="Arthropoda" rank="species" species="maculatum">
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||
<emphasis id="B965EA84FF93EC41611437D2FDECB72E" box="[290,540,1276,1298]" italics="true" pageId="3" pageNumber="478">Geosesarma maculatum</emphasis>
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(De Man, 1892)
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</taxonomicName>
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, male (12.4 × 11.9 mm) (ZRC 2017.0113), Halmahera. A, overall habitus; B, dorsal view of carapace; C, ventral view of cephalothorax showing thoracic sternum and pleon.
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||
</paragraph>
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||
</caption>
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||
<subSubSection id="C30B651DFF93EC4160A13677FAF8B7C3" pageId="3" pageNumber="478" type="materials_examined">
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<paragraph id="8BAE3696FF93EC4160A13677FC86B784" blockId="3.[151,1437,1368,2032]" pageId="3" pageNumber="478">
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||
<materialsCitation id="3B793CCBFF93EC4160A13677FE7EB7A9" ID-GBIF-Occurrence="1455709147" country="Iceland" elevation="800" location="Culasi" pageId="3" pageNumber="478" specimenCount="2" typeStatus="holotype">
|
||
<emphasis id="B965EA84FF93EC4160A13677FECEB74D" bold="true" box="[151,318,1368,1393]" pageId="3" pageNumber="478">
|
||
<typeStatus id="54AA8834FF93EC4160A13677FF22B74D" box="[151,210,1368,1393]" pageId="3" pageNumber="478">Type</typeStatus>
|
||
material
|
||
</emphasis>
|
||
.
|
||
<typeStatus id="54AA8834FF93EC41617A3677FE44B74D" box="[332,436,1368,1393]" pageId="3" pageNumber="478" type="holotype">Holotype</typeStatus>
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||
(missing, see Remarks): male (13.0 ×
|
||
<quantity id="4CE99B73FF93EC41635C3676FC23B74D" box="[874,979,1369,1393]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="3" pageNumber="478" unit="mm" value="12.2">12.2 mm</quantity>
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), on trunks of forest trees,
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||
<location id="8ECE604DFF93EC4165373677FABBB74D" LSID="urn:lsid:plazi:treatment:03B88780FF92EC4560A13414FDA9B7A1:8ECE604DFF93EC4165373677FABBB74D" box="[1281,1355,1368,1393]" country="Iceland" name="Culasi" pageId="3" pageNumber="478">Culasi</location>
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||
, Panay
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||
<collectingCountry id="F3067606FF93EC4160A13653FF2EB7A9" box="[151,222,1404,1429]" name="Iceland" pageId="3" pageNumber="478">Island</collectingCountry>
|
||
,
|
||
<elevation id="003CD1A5FF93EC4160DF3652FEC0B7A8" box="[233,304,1405,1429]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="478" unit="m" value="800.0">
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<quantity id="4CE99B73FF93EC4160DF3652FEC0B7A8" box="[233,304,1405,1429]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="478" unit="m" value="800.0">800 m</quantity>
|
||
</elevation>
|
||
altitude
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||
</materialsCitation>
|
||
,
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||
<materialsCitation id="3B793CCBFF93EC4161AF3653FBDFB7A9" ID-GBIF-Occurrence="1455709143" box="[409,1071,1404,1429]" collectingDate="1918-05-18" collectionCode="USNM" collectorName="McGregor" country="Philippines" location="Philippines" pageId="3" pageNumber="478" specimenCode="USNM 54315" specimenCount="1" typeStatus="holotype">
|
||
<collectingCountry id="F3067606FF93EC4161AF3653FDEAB7A9" box="[409,538,1404,1429]" name="Philippines" pageId="3" pageNumber="478">Philippines</collectingCountry>
|
||
, coll.
|
||
<collectorName id="26E45340FF93EC41626B3652FD24B7A9" box="[605,724,1405,1429]" pageId="3" pageNumber="478">McGregor</collectorName>
|
||
,
|
||
<collectingDate id="EFEBE9BEFF93EC4162E93652FC84B7A9" box="[735,884,1405,1429]" pageId="3" pageNumber="478" value="1918-05-18">
|
||
<date id="FFAF1056FF93EC4162E93652FC84B7A9" box="[735,884,1405,1429]" pageId="3" pageNumber="478" value="1918-05-18">18 May 1918</date>
|
||
</collectingDate>
|
||
(
|
||
<specimenCode id="DBB79EEDFF93EC4163B53652FBD8B7A9" box="[899,1064,1404,1429]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="478">
|
||
<collectionCode id="ED00AE53FF93EC4163B53652FC26B7A9" box="[899,982,1405,1429]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="478">USNM</collectionCode>
|
||
54315
|
||
</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B793CCBFF93EC41640C3652FC83B784" ID-GBIF-Occurrence="1455709144" collectingDate="1918-05-18" collectionCode="USNM" collectorName="McGregor" country="Philippines" location="Philippines" pageId="3" pageNumber="478" specimenCode="USNM 1277697" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<typeStatus id="54AA8834FF93EC41640C3652FB51B7A9" box="[1082,1185,1405,1429]" pageId="3" pageNumber="478" type="paratype">Paratype</typeStatus>
|
||
:
|
||
<specimenCount id="9D17FD1FFF93EC41649D3652FAFCB7A9" box="[1195,1292,1404,1429]" pageId="3" pageNumber="478" type="female">1 female</specimenCount>
|
||
(13.3 × 12.0 mm), same locality and other data as
|
||
<typeStatus id="54AA8834FF93EC41620F36B0FD6BB784" box="[569,667,1439,1464]" pageId="3" pageNumber="478" type="holotype">holotype</typeStatus>
|
||
(
|
||
<specimenCode id="DBB79EEDFF93EC41629C368FFC9AB784" box="[682,874,1439,1464]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="478">
|
||
<collectionCode id="ED00AE53FF93EC41629C368FFD0CB784" box="[682,764,1440,1464]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="478">USNM</collectionCode>
|
||
1277697
|
||
</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAE3696FF93EC4160F136ECFAF8B7C3" blockId="3.[151,1437,1368,2032]" pageId="3" pageNumber="478">
|
||
<emphasis id="B965EA84FF93EC4160F136ECFE51B7E0" bold="true" box="[199,417,1475,1500]" pageId="3" pageNumber="478">Non-type material</emphasis>
|
||
. 4 males (14.9 ×
|
||
<quantity id="4CE99B73FF93EC41625436ECFD37B7E7" box="[610,711,1475,1500]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="3" pageNumber="478" unit="mm" value="14.5">14.5 mm</quantity>
|
||
, 13.1 ×
|
||
<quantity id="4CE99B73FF93EC41631636ECFC76B7E7" box="[800,902,1475,1499]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="3" pageNumber="478" unit="mm" value="12.3">12.3 mm</quantity>
|
||
, 11.1 ×
|
||
<quantity id="4CE99B73FF93EC4163E836ECFBC5B7E7" box="[990,1077,1475,1499]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="3" pageNumber="478" unit="mm" value="9.7">9.7 mm</quantity>
|
||
, 10.4 ×
|
||
<quantity id="4CE99B73FF93EC4164B836ECFB18B7E7" box="[1166,1256,1475,1499]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="3" pageNumber="478" unit="mm" value="9.4">9.4 mm</quantity>
|
||
),
|
||
<materialsCitation id="3B793CCBFF93EC4164CC36ECFAF4B7C3" ID-GBIF-Occurrence="1455709145" collectionCode="USNM" country="Iceland" elevation="800" location="Culasi" pageId="3" pageNumber="478" specimenCode="USNM 123208" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="9D17FD1FFF93EC4164CC36ECFAABB7E0" box="[1274,1371,1475,1500]" pageId="3" pageNumber="478" type="female">1 female</specimenCount>
|
||
(14.1 ×
|
||
<quantity id="4CE99B73FF93EC41609B36C9FEE5B7C3" box="[173,277,1510,1535]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="3" pageNumber="478" unit="mm" value="12.6">12.6 mm</quantity>
|
||
), 4 immature juveniles, locality and data same as holotype and
|
||
<typeStatus id="54AA8834FF93EC4163ED36C8FBCBB7C3" box="[987,1083,1511,1535]" pageId="3" pageNumber="478" type="paratype">paratype</typeStatus>
|
||
(
|
||
<specimenCode id="DBB79EEDFF93EC41647F36C8FB0CB7C3" box="[1097,1276,1511,1535]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="478">
|
||
<collectionCode id="ED00AE53FF93EC41647F36C8FB6CB7C3" box="[1097,1180,1511,1535]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="478">USNM</collectionCode>
|
||
123208
|
||
</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B651DFF93EC4160F13525FD14B5CC" pageId="3" pageNumber="478" type="diagnosis">
|
||
<paragraph id="8BAE3696FF93EC4160F13525FD14B5CC" blockId="3.[151,1437,1368,2032]" pageId="3" pageNumber="478">
|
||
<emphasis id="B965EA84FF93EC4160F13525FECAB41F" bold="true" box="[199,314,1546,1571]" pageId="3" pageNumber="478">Diagnosis</emphasis>
|
||
. Carapace quadrate to subquadrate, just wider than long, width to length ratio 1.0–1.1, lateral margins parallel (
|
||
<figureCitation id="132A2A13FF93EC4161523502FE45B47A" box="[356,437,1581,1606]" captionStart="FIGURE 3" captionStartId="4.[151,250,1893,1915]" captionTargetBox="[151,1436,193,1872]" captionTargetId="figure@4.[151,1436,193,1872]" captionTargetPageId="4" captionText="FIGURE 3. Geosesarma rathbunae (Serène, 1968 a). A, holotype male (13.0 × 12.2 mm) (USNM 54315) (after Serène, 1968 a: pl. 1 fig. 4); B – I, non-type male (13.1 × 12.3 mm) (USNM 123208). A, B, overall habitus; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of right chelae at different angles; H, I, dorsal views of right chelipeds at different angles." httpUri="https://zenodo.org/record/495553/files/figure.png" pageId="3" pageNumber="478">Figs. 3</figureCitation>
|
||
A, B, 4A, B, H, I); dorsal surface with regions just visible, anterior regions densely covered with low, small, rounded, flattened granules and rugosities (
|
||
<figureCitation id="132A2A13FF93EC416383357EFBF5B455" box="[949,1029,1617,1642]" captionStart="FIGURE 3" captionStartId="4.[151,250,1893,1915]" captionTargetBox="[151,1436,193,1872]" captionTargetId="figure@4.[151,1436,193,1872]" captionTargetPageId="4" captionText="FIGURE 3. Geosesarma rathbunae (Serène, 1968 a). A, holotype male (13.0 × 12.2 mm) (USNM 54315) (after Serène, 1968 a: pl. 1 fig. 4); B – I, non-type male (13.1 × 12.3 mm) (USNM 123208). A, B, overall habitus; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of right chelae at different angles; H, I, dorsal views of right chelipeds at different angles." httpUri="https://zenodo.org/record/495553/files/figure.png" pageId="3" pageNumber="478">Figs. 3</figureCitation>
|
||
A, B, 4A, B, H, I); front deflexed, frontal lobes broad with subtruncate margins, separated by shallow concavity; postfrontal, postorbital cristae low but relatively sharp (
|
||
<figureCitation id="132A2A13FF93EC4161BE35B8FE28B48C" box="[392,472,1687,1712]" captionStart="FIGURE 3" captionStartId="4.[151,250,1893,1915]" captionTargetBox="[151,1436,193,1872]" captionTargetId="figure@4.[151,1436,193,1872]" captionTargetPageId="4" captionText="FIGURE 3. Geosesarma rathbunae (Serène, 1968 a). A, holotype male (13.0 × 12.2 mm) (USNM 54315) (after Serène, 1968 a: pl. 1 fig. 4); B – I, non-type male (13.1 × 12.3 mm) (USNM 123208). A, B, overall habitus; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of right chelae at different angles; H, I, dorsal views of right chelipeds at different angles." httpUri="https://zenodo.org/record/495553/files/figure.png" pageId="3" pageNumber="478">Figs. 3</figureCitation>
|
||
A, B, E, 4A, B, C, H, I); external orbital tooth triangular, directed obliquely, outer margin convex, tip not distinctly extending beyond lateral margin (
|
||
<figureCitation id="132A2A13FF93EC41639B3594FC0EB4E8" box="[941,1022,1723,1748]" captionStart="FIGURE 3" captionStartId="4.[151,250,1893,1915]" captionTargetBox="[151,1436,193,1872]" captionTargetId="figure@4.[151,1436,193,1872]" captionTargetPageId="4" captionText="FIGURE 3. Geosesarma rathbunae (Serène, 1968 a). A, holotype male (13.0 × 12.2 mm) (USNM 54315) (after Serène, 1968 a: pl. 1 fig. 4); B – I, non-type male (13.1 × 12.3 mm) (USNM 123208). A, B, overall habitus; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of right chelae at different angles; H, I, dorsal views of right chelipeds at different angles." httpUri="https://zenodo.org/record/495553/files/figure.png" pageId="3" pageNumber="478">Figs. 3</figureCitation>
|
||
A, B, 4A, B, H, I). Merus of third maxilliped subovate, subequal in length to ischium; exopod slender without trace of flagellum (
|
||
<figureCitation id="132A2A13FF93EC4164DD35F1FAC1B4CB" box="[1259,1329,1758,1783]" captionStart="FIGURE 5" captionStartId="6.[151,250,1840,1862]" captionTargetBox="[173,1416,210,1804]" captionTargetId="figure@6.[151,1436,193,1819]" captionTargetPageId="6" captionText="FIGURE 5. A – D, Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758); E – H, Geosesarma rathbunae (Serène, 1968 a), male (13.1 × 12.3 mm) (USNM 123208). A, E, left third maxilliped; B, F, ventral view of left G 1; C, G, mesial view of left G 1; D, H, dorsal view of left G 1. Scales = 1.0 mm." httpUri="https://zenodo.org/record/495555/files/figure.png" pageId="3" pageNumber="478">Fig. 5</figureCitation>
|
||
E). Outer surface of palm of cheliped of adult male covered with numerous low granules; inner surface granulated but without transverse ridge; dorsal margin of dactylus with 6–8 low granules on proximal half or two-thirds, granules not chitinous, unarmed on distal half (
|
||
<figureCitation id="132A2A13FF93EC41626F3466FD59B55E" box="[601,681,1865,1890]" captionStart="FIGURE 3" captionStartId="4.[151,250,1893,1915]" captionTargetBox="[151,1436,193,1872]" captionTargetId="figure@4.[151,1436,193,1872]" captionTargetPageId="4" captionText="FIGURE 3. Geosesarma rathbunae (Serène, 1968 a). A, holotype male (13.0 × 12.2 mm) (USNM 54315) (after Serène, 1968 a: pl. 1 fig. 4); B – I, non-type male (13.1 × 12.3 mm) (USNM 123208). A, B, overall habitus; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of right chelae at different angles; H, I, dorsal views of right chelipeds at different angles." httpUri="https://zenodo.org/record/495553/files/figure.png" pageId="3" pageNumber="478">Figs. 3</figureCitation>
|
||
F–I, 4F, G). Ambulatory legs each with relatively slender merus having sharp subdistal spine on dorsal margin and lateral, mesial surfaces gently rugose (
|
||
<figureCitation id="132A2A13FF93EC4164A33443FB14B5B9" box="[1173,1252,1900,1925]" captionStart="FIGURE 3" captionStartId="4.[151,250,1893,1915]" captionTargetBox="[151,1436,193,1872]" captionTargetId="figure@4.[151,1436,193,1872]" captionTargetPageId="4" captionText="FIGURE 3. Geosesarma rathbunae (Serène, 1968 a). A, holotype male (13.0 × 12.2 mm) (USNM 54315) (after Serène, 1968 a: pl. 1 fig. 4); B – I, non-type male (13.1 × 12.3 mm) (USNM 123208). A, B, overall habitus; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of right chelae at different angles; H, I, dorsal views of right chelipeds at different angles." httpUri="https://zenodo.org/record/495553/files/figure.png" pageId="3" pageNumber="478">Figs. 3</figureCitation>
|
||
A, 4A, B, H, I); propodus without brush-like setae on ventral margin. Male pleon triangular, relatively broad; somite 6 wide, with convex lateral margins; telson semicircular (
|
||
<figureCitation id="132A2A13FF93EC4162A4349CFD11B5F0" box="[658,737,1971,1996]" captionStart="FIGURE 3" captionStartId="4.[151,250,1893,1915]" captionTargetBox="[151,1436,193,1872]" captionTargetId="figure@4.[151,1436,193,1872]" captionTargetPageId="4" captionText="FIGURE 3. Geosesarma rathbunae (Serène, 1968 a). A, holotype male (13.0 × 12.2 mm) (USNM 54315) (after Serène, 1968 a: pl. 1 fig. 4); B – I, non-type male (13.1 × 12.3 mm) (USNM 123208). A, B, overall habitus; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of right chelae at different angles; H, I, dorsal views of right chelipeds at different angles." httpUri="https://zenodo.org/record/495553/files/figure.png" pageId="3" pageNumber="478">Figs. 3</figureCitation>
|
||
C, D, 4D, E). G1 relatively slender, distal chitinous part bent obliquely laterally, tapering to sharp tip (
|
||
<figureCitation id="132A2A13FF93EC41625634F8FD51B5CC" box="[608,673,2007,2032]" captionStart="FIGURE 5" captionStartId="6.[151,250,1840,1862]" captionTargetBox="[173,1416,210,1804]" captionTargetId="figure@6.[151,1436,193,1819]" captionTargetPageId="6" captionText="FIGURE 5. A – D, Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758); E – H, Geosesarma rathbunae (Serène, 1968 a), male (13.1 × 12.3 mm) (USNM 123208). A, E, left third maxilliped; B, F, ventral view of left G 1; C, G, mesial view of left G 1; D, H, dorsal view of left G 1. Scales = 1.0 mm." httpUri="https://zenodo.org/record/495555/files/figure.png" pageId="3" pageNumber="478">Fig. 5</figureCitation>
|
||
F–H).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6E661EFF94EC4660A1344AFD61B5EB" httpUri="https://zenodo.org/record/495553/files/figure.png" pageId="4" pageNumber="479" targetBox="[151,1436,193,1872]" targetPageId="4">
|
||
<paragraph id="8BAE3696FF94EC4660A1344AFD61B5EB" blockId="4.[151,1436,1893,2007]" pageId="4" pageNumber="479">
|
||
<emphasis id="B965EA84FF94EC4660A1344AFEFCB546" bold="true" box="[151,268,1893,1915]" pageId="4" pageNumber="479">FIGURE 3</emphasis>
|
||
.
|
||
<taxonomicName id="4C114D15FF94EC46612E3449FD5DB547" authority="Serene, 1968a" authorityName="Serene" authorityYear="1968" box="[280,685,1893,1915]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="479" phylum="Arthropoda" rank="species" species="rathbunae">
|
||
<emphasis id="B965EA84FF94EC46612E3449FDF5B547" box="[280,517,1893,1915]" italics="true" pageId="4" pageNumber="479">Geosesarma rathbunae</emphasis>
|
||
(Serène, 1968a)
|
||
</taxonomicName>
|
||
. A, holotype male (13.0 × 12.2 mm) (USNM 54315) (after Serène, 1968a: pl. 1 fig. 4); B–I, non-type male (13.1 × 12.3 mm) (USNM 123208). A, B, overall habitus; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of right chelae at different angles; H, I, dorsal views of right chelipeds at different angles.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6E661EFF95EC4760A13403FB04B5A1" httpUri="https://zenodo.org/record/495554/files/figure.png" pageId="5" pageNumber="480" targetBox="[151,1436,193,1814]" targetPageId="5">
|
||
<paragraph id="8BAE3696FF95EC4760A13403FB04B5A1" blockId="5.[151,1436,1836,1949]" pageId="5" pageNumber="480">
|
||
<emphasis id="B965EA84FF95EC4760A13403FEE0B57D" bold="true" box="[151,272,1836,1858]" pageId="5" pageNumber="480">FIGURE 4</emphasis>
|
||
.
|
||
<taxonomicName id="4C114D15FF95EC4761173403FD32B57E" authority="Serene, 1968a" authorityName="Serene" authorityYear="1968" box="[289,706,1836,1858]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="480" phylum="Arthropoda" rank="species" species="rathbunae">
|
||
<emphasis id="B965EA84FF95EC4761173403FDE1B57E" box="[289,529,1836,1858]" italics="true" pageId="5" pageNumber="480">Geosesarma rathbunae</emphasis>
|
||
(Serène, 1968a)
|
||
</taxonomicName>
|
||
. A, paratype female (13.5 × 12.1 mm) (USNM 1277697, ex USNM 123208); B–G, non-type male (14.9 × 14.5 mm) (USNM 123208); H, non-type female (14.1 × 12.6 mm) (USNM ex 54315); I, non-type male (10.4 × 9.4 mm) (USNM 123208). A, B, H, I, overall habitus; C, frontal view of cephalothorax; D, E, ventral view of cephalothorax showing thoracic sternum and pleon; F, G, outer views of left chelae at different angles.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6E661EFF96EC4460A1341FFC68B5BF" httpUri="https://zenodo.org/record/495555/files/figure.png" pageId="6" pageNumber="481" targetBox="[173,1416,210,1804]" targetPageId="6">
|
||
<paragraph id="8BAE3696FF96EC4460A1341FFC68B5BF" blockId="6.[151,1436,1840,1923]" pageId="6" pageNumber="481">
|
||
<emphasis id="B965EA84FF96EC4460A1341FFEFDB579" bold="true" box="[151,269,1840,1862]" pageId="6" pageNumber="481">FIGURE 5</emphasis>
|
||
. A–D,
|
||
<taxonomicName id="4C114D15FF96EC446161341EFD04B57A" authority="Rathbun, 1914" authorityName="Rathbun" authorityYear="1914" box="[343,756,1840,1862]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="481" phylum="Arthropoda" rank="species" species="vicentense">
|
||
<emphasis id="B965EA84FF96EC446161341EFE26B57A" box="[343,470,1841,1862]" italics="true" pageId="6" pageNumber="481">Geosesarma</emphasis>
|
||
<emphasis id="B965EA84FF96EC4461EA341EFDB5B57A" box="[476,581,1841,1862]" italics="true" pageId="6" pageNumber="481">vicentense</emphasis>
|
||
(Rathbun, 1914)
|
||
</taxonomicName>
|
||
, holotype male (9.2 × 8.5 mm) (USNM 45758); E–H,
|
||
<taxonomicName id="4C114D15FF96EC44652B341EFE58B559" authority="Serene, 1968a" authorityName="Serene" authorityYear="1968" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="481" phylum="Arthropoda" rank="species" species="rathbunae">
|
||
<emphasis id="B965EA84FF96EC44652B341EFF0FB559" italics="true" pageId="6" pageNumber="481">Geosesarma rathbunae</emphasis>
|
||
(Serène, 1968a)
|
||
</taxonomicName>
|
||
, male (13.1 × 12.3 mm) (USNM 123208). A, E, left third maxilliped; B, F, ventral view of left G1; C, G, mesial view of left G1; D, H, dorsal view of left G1. Scales = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30B651DFF97EC4560F133B8FDA9B7A1" pageId="7" pageNumber="482" type="discussion">
|
||
<paragraph id="8BAE3696FF97EC4560F133B8FB7DB024" blockId="7.[151,1437,151,1437]" pageId="7" pageNumber="482">
|
||
<emphasis id="B965EA84FF97EC4560F133B8FECBB28C" bold="true" box="[199,315,151,176]" pageId="7" pageNumber="482">Remarks.</emphasis>
|
||
While examining the type material of
|
||
<taxonomicName id="4C114D15FF97EC4562D933B7FC05B28C" box="[751,1013,152,176]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="482" phylum="Arthropoda" rank="species" species="rathbunae">
|
||
<emphasis id="B965EA84FF97EC4562D933B7FC05B28C" box="[751,1013,152,176]" italics="true" pageId="7" pageNumber="482">Geosesarma rathbunae</emphasis>
|
||
</taxonomicName>
|
||
, some discrepancies were discovered in the contents of the jars that were supposed to contain the holotype and paratype specimens of this species.
|
||
<bibRefCitation id="EF804B67FF97EC4560A133CFFECCB2C4" author="Serene" box="[151,316,223,248]" pageId="7" pageNumber="482" refString="Serene, R. (1968 a) Note preliminaire sur de nouvelles especes de Sesarma (Decapoda Brachyura). Bulletin du Museum national d'Histoire naturelle, 39 (5), 1084 - 1095." type="journal article" year="1968" yearSuffix="a">Serène (1968a</bibRefCitation>
|
||
: 1086, 1087) based his description on only two specimens, one male and one female, loaned to him by the USNM. His two specimens were part of a lot (USNM 54315), containing as per the labels and USNM records, a total of five males and two females collected in Culasi, on the west coast of Panay Island, Philippines. The specimens in this lot had been identified by M. J. Rathbun as “
|
||
<taxonomicName id="4C114D15FF97EC4563B43263FAC6B358" authority="de Man" authorityName="de Man" box="[898,1334,332,357]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="482" phylum="Arthropoda" rank="species" species="sylvicola" subGenus="Sesarma">
|
||
<emphasis id="B965EA84FF97EC4563B43263FC12B35F" box="[898,994,332,355]" italics="true" pageId="7" pageNumber="482">Sesarma</emphasis>
|
||
(
|
||
<emphasis id="B965EA84FF97EC4563C43263FBA2B35F" box="[1010,1106,332,355]" italics="true" pageId="7" pageNumber="482">Sesarma</emphasis>
|
||
)
|
||
<emphasis id="B965EA84FF97EC4564553263FB37B358" box="[1123,1223,332,356]" italics="true" pageId="7" pageNumber="482">sylvicola</emphasis>
|
||
(de Man)
|
||
</taxonomicName>
|
||
, var” but never reported in the literature. Of the two specimens that Serène borrowed, he designated as holotype the male measuring 13.0 ×
|
||
<quantity id="4CE99B73FF97EC45615032BBFE3EB390" box="[358,462,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="7" pageNumber="482" unit="mm" value="12.2">12.2 mm</quantity>
|
||
, which he illustrated and photographed (
|
||
<bibRefCitation id="EF804B67FF97EC45639632BBFACCB390" author="Serene" box="[928,1340,404,429]" pageId="7" pageNumber="482" refString="Serene, R. (1968 a) Note preliminaire sur de nouvelles especes de Sesarma (Decapoda Brachyura). Bulletin du Museum national d'Histoire naturelle, 39 (5), 1084 - 1095." type="journal article" year="1968" yearSuffix="a">Serène 1968a: figs 5, 6, pl. 1, fig. 4</bibRefCitation>
|
||
), and as paratype the female measuring 13.3 × 12.0 mm. The holotype and paratype were then returned to the USNM where the holotype was assigned for itself the same original catalogue number used for all the Culasi material, USNM 54315. A new catalogue number, USNM 123208, was then issued for the female paratype.
|
||
</paragraph>
|
||
<paragraph id="8BAE3696FF97EC4560F1310BFB11B129" blockId="7.[151,1437,151,1437]" pageId="7" pageNumber="482">
|
||
When the two USNM jars with Serène’s (1968a)
|
||
<typeStatus id="54AA8834FF97EC456338310AFCCEB001" box="[782,830,549,573]" pageId="7" pageNumber="482">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="4C114D15FF97EC4563E5310BFAAAB000" box="[979,1370,548,572]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="482" phylum="Arthropoda" rank="species" species="rathbunae" subGenus="Geosesarma">
|
||
<emphasis id="B965EA84FF97EC4563E5310BFBC3B007" box="[979,1075,548,571]" italics="true" pageId="7" pageNumber="482">Sesarma</emphasis>
|
||
(
|
||
<emphasis id="B965EA84FF97EC456471310BFB23B007" box="[1095,1235,548,571]" italics="true" pageId="7" pageNumber="482">Geosesarma</emphasis>
|
||
)
|
||
<emphasis id="B965EA84FF97EC4564D1310BFAAAB000" box="[1255,1370,548,572]" italics="true" pageId="7" pageNumber="482">rathbunae</emphasis>
|
||
</taxonomicName>
|
||
were pulled from the shelves for this study, we discovered that the lot that should contain the male
|
||
<typeStatus id="54AA8834FF97EC4564EE3168FACDB05C" box="[1240,1341,583,608]" pageId="7" pageNumber="482" type="holotype">holotype</typeStatus>
|
||
, USNM 54315, actually contained a female specimen (
|
||
<figureCitation id="132A2A13FF97EC4562953143FD17B0B8" box="[675,743,620,645]" captionStart="FIGURE 4" captionStartId="5.[151,250,1836,1858]" captionTargetBox="[151,1436,193,1814]" captionTargetId="figure@5.[151,1436,193,1814]" captionTargetPageId="5" captionText="FIGURE 4. Geosesarma rathbunae (Serène, 1968 a). A, paratype female (13.5 × 12.1 mm) (USNM 1277697, ex USNM 123208); B – G, non-type male (14.9 × 14.5 mm) (USNM 123208); H, non-type female (14.1 × 12.6 mm) (USNM ex 54315); I, non-type male (10.4 × 9.4 mm) (USNM 123208). A, B, H, I, overall habitus; C, frontal view of cephalothorax; D, E, ventral view of cephalothorax showing thoracic sternum and pleon; F, G, outer views of left chelae at different angles." httpUri="https://zenodo.org/record/495554/files/figure.png" pageId="7" pageNumber="482">Fig. 4</figureCitation>
|
||
A). Furthermore, the lot that should contain the single female
|
||
<typeStatus id="54AA8834FF97EC4560A131BFFF0AB094" box="[151,250,656,680]" pageId="7" pageNumber="482" type="paratype">paratype</typeStatus>
|
||
, USNM 123208, actually contained four males, one female, and a vial with four small and dismembered immature juveniles unaccounted for in the labels and USNM records. It should be noted that
|
||
<bibRefCitation id="EF804B67FF97EC456481319BFA92B0F1" author="Serene" box="[1207,1378,692,717]" pageId="7" pageNumber="482" refString="Serene, R. (1968 a) Note preliminaire sur de nouvelles especes de Sesarma (Decapoda Brachyura). Bulletin du Museum national d'Histoire naturelle, 39 (5), 1084 - 1095." type="journal article" year="1968" yearSuffix="a">Serène (1968a)</bibRefCitation>
|
||
only examined and reported in his description the male and female
|
||
<typeStatus id="54AA8834FF97EC45635931F7FC5EB0CC" box="[879,942,728,752]" pageId="7" pageNumber="482">types</typeStatus>
|
||
; thus, the rest of the Culasi material, even though it has been found to be conspecific with Serène’s species, cannot be part of the
|
||
<typeStatus id="54AA8834FF97EC45645431D2FB62B129" box="[1122,1170,765,789]" pageId="7" pageNumber="482">type</typeStatus>
|
||
series.
|
||
</paragraph>
|
||
<paragraph id="8BAE3696FF97EC4560F13030FC69B69D" blockId="7.[151,1437,151,1437]" pageId="7" pageNumber="482">
|
||
After measuring and comparing all the specimens from Culasi with Serène’s (1968a) description, we determined that the female mistakenly found in the holotype jar matches precisely Serène’s female paratype, and this female specimen has now been assigned a new catalogue number, USNM 1277697. However, none of the males found still in USNM 54315 could be matched with confidence to the size and features of the holotype photographed and illustrated by
|
||
<bibRefCitation id="EF804B67FF97EC456234309FFD5CB1F4" author="Serene" box="[514,684,943,968]" pageId="7" pageNumber="482" refString="Serene, R. (1968 a) Note preliminaire sur de nouvelles especes de Sesarma (Decapoda Brachyura). Bulletin du Museum national d'Histoire naturelle, 39 (5), 1084 - 1095." type="journal article" year="1968" yearSuffix="a">Serène (1968a)</bibRefCitation>
|
||
(
|
||
<figureCitation id="132A2A13FF97EC45628D3080FD0DB1F4" box="[699,765,943,968]" captionStart="FIGURE 3" captionStartId="4.[151,250,1893,1915]" captionTargetBox="[151,1436,193,1872]" captionTargetId="figure@4.[151,1436,193,1872]" captionTargetPageId="4" captionText="FIGURE 3. Geosesarma rathbunae (Serène, 1968 a). A, holotype male (13.0 × 12.2 mm) (USNM 54315) (after Serène, 1968 a: pl. 1 fig. 4); B – I, non-type male (13.1 × 12.3 mm) (USNM 123208). A, B, overall habitus; C, D, ventral view of cephalothorax showing thoracic sternum and pleon; E, frontal view of cephalothorax; F, G, outer views of right chelae at different angles; H, I, dorsal views of right chelipeds at different angles." httpUri="https://zenodo.org/record/495553/files/figure.png" pageId="7" pageNumber="482">Fig. 3</figureCitation>
|
||
A), and moreover, none was found to have the G1 dissected and Serène had a well-known practice of dissecting out the G1 for illustration. More conclusive, one of the five males in the original jar with all the Culasi material, is missing and we have concluded it must be the holotype. Given the possibility, even if remote, that Serène’s holotype may be found in the future, we prefer to be cautious and make a note in the records of the missing holotype in USNM 54315. Finally, the status of lot USNM 123208 that was originally to contain the female paratype but found during this study to contain non-paratypes from Culasi, has been adjusted to reflect that the specimens therein are non-types.
|
||
</paragraph>
|
||
<paragraph id="8BAE3696FF97EC4560F13783FCE7B6D4" blockId="7.[151,1437,151,1437]" pageId="7" pageNumber="482">
|
||
The identity of the species, however, is not in doubt as the series of remaining specimens all agree very well with the
|
||
<typeStatus id="54AA8834FF97EC4560CF37FEFED9B6D5" box="[249,297,1233,1257]" pageId="7" pageNumber="482">type</typeStatus>
|
||
descriptions and figures by
|
||
<bibRefCitation id="EF804B67FF97EC45625337FFFCE3B6D4" author="Serene" box="[613,787,1231,1256]" pageId="7" pageNumber="482" refString="Serene, R. (1968 a) Note preliminaire sur de nouvelles especes de Sesarma (Decapoda Brachyura). Bulletin du Museum national d'Histoire naturelle, 39 (5), 1084 - 1095." type="journal article" year="1968" yearSuffix="a">Serène (1968a)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAE3696FF97EC4560F137DBFDA9B7A1" blockId="7.[151,1437,151,1437]" pageId="7" pageNumber="482">
|
||
<taxonomicName id="4C114D15FF97EC4560F137DBFE23B730" box="[199,467,1268,1292]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="482" phylum="Arthropoda" rank="species" species="rathbunae">
|
||
<emphasis id="B965EA84FF97EC4560F137DBFE23B730" box="[199,467,1268,1292]" italics="true" pageId="7" pageNumber="482">Geosesarma rathbunae</emphasis>
|
||
</taxonomicName>
|
||
is unique among known Philippine
|
||
<taxonomicName id="4C114D15FF97EC4563B937DBFBEBB737" box="[911,1051,1268,1291]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="482" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B965EA84FF97EC4563B937DBFBEBB737" box="[911,1051,1268,1291]" italics="true" pageId="7" pageNumber="482">Geosesarma</emphasis>
|
||
</taxonomicName>
|
||
in that the exopod of the third maxilliped does not have a flagellum (
|
||
<figureCitation id="132A2A13FF97EC4562773638FD73B70C" box="[577,643,1303,1328]" captionStart="FIGURE 5" captionStartId="6.[151,250,1840,1862]" captionTargetBox="[173,1416,210,1804]" captionTargetId="figure@6.[151,1436,193,1819]" captionTargetPageId="6" captionText="FIGURE 5. A – D, Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758); E – H, Geosesarma rathbunae (Serène, 1968 a), male (13.1 × 12.3 mm) (USNM 123208). A, E, left third maxilliped; B, F, ventral view of left G 1; C, G, mesial view of left G 1; D, H, dorsal view of left G 1. Scales = 1.0 mm." httpUri="https://zenodo.org/record/495555/files/figure.png" pageId="7" pageNumber="482">Fig. 5</figureCitation>
|
||
E). All the other species have a long flagellum on the exopod (e.g.,
|
||
<figureCitation id="132A2A13FF97EC4565463638FF4DB769" captionStart="FIGURE 5" captionStartId="6.[151,250,1840,1862]" captionTargetBox="[173,1416,210,1804]" captionTargetId="figure@6.[151,1436,193,1819]" captionTargetPageId="6" captionText="FIGURE 5. A – D, Geosesarma vicentense (Rathbun, 1914), holotype male (9.2 × 8.5 mm) (USNM 45758); E – H, Geosesarma rathbunae (Serène, 1968 a), male (13.1 × 12.3 mm) (USNM 123208). A, E, left third maxilliped; B, F, ventral view of left G 1; C, G, mesial view of left G 1; D, H, dorsal view of left G 1. Scales = 1.0 mm." httpUri="https://zenodo.org/record/495555/files/figure.png" pageId="7" pageNumber="482">Fig. 5A</figureCitation>
|
||
). This is a character, however, present in many highland species from Peninsular
|
||
<collectingCountry id="F3067606FF97EC4564733613FB5BB769" box="[1093,1195,1340,1365]" name="Malaysia" pageId="7" pageNumber="482">Malaysia</collectingCountry>
|
||
and Borneo (e.g., see Ng, 1988, 2015). These species also have G1 structures similar to that of
|
||
<taxonomicName id="4C114D15FF97EC4563D0364FFB89B744" box="[998,1145,1376,1400]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="482" phylum="Arthropoda" rank="species" species="rathbunae">
|
||
<emphasis id="B965EA84FF97EC4563D0364FFB89B744" box="[998,1145,1376,1400]" italics="true" pageId="7" pageNumber="482">G. rathbunae</emphasis>
|
||
</taxonomicName>
|
||
, with the distal chitinous part relatively shorter and laterally bent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |