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<document id="239A1F596EBD69598005771C36529EDE" ID-DOI="10.5962/bhl.part.117802" ID-ISSN="0035-418X" ID-Zenodo-Dep="6311706" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645021050174" checkinUser="carolina" docAuthor="Cuccodoro, Giulio" docDate="2011" docId="03F7D871AF775D5DFF0FAEA17471D842" docLanguage="en" docName="RevSuisseZool.118.107-148.pdf.imf" docOrigin="Revue suisse de Zoologie 118 (1)" docStyle="DocumentStyle:0846B485C0C87F6EB7648DEEBF34B77F.2:RevSuisseZool.2011-2014.journal_article" docStyleId="0846B485C0C87F6EB7648DEEBF34B77F" docStyleName="RevSuisseZool.2011-2014.journal_article" docStyleVersion="2" docTitle="Megarthrus mammiger Bierig 1940" docType="treatment" docVersion="3" lastPageNumber="117" masterDocId="FFCEA009AF7E5D57FF98AE43766CDA37" masterDocTitle="Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae)" masterLastPageNumber="147" masterPageNumber="107" pageNumber="116" updateTime="1699296264638" updateUser="plazi">
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<mods:title id="6479C4B0BCCBD16F0134CA7930DEA4E2">Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae)</mods:title>
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<mods:namePart id="032576C24BB5F9C698D43F786919D6A3">Cuccodoro, Giulio</mods:namePart>
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<mods:date id="F71DB141DD1DA1A71D2C2DB4AF639040">2011</mods:date>
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<treatment id="03F7D871AF775D5DFF0FAEA17471D842" LSID="urn:lsid:plazi:treatment:03F7D871AF775D5DFF0FAEA17471D842" httpUri="http://treatment.plazi.org/id/03F7D871AF775D5DFF0FAEA17471D842" lastPageId="10" lastPageNumber="117" pageId="9" pageNumber="116">
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<heading id="D0A9DE0BAF775D5EFF0FAEA1725EDACD" box="[151,1074,226,250]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="116" reason="2">
<taxonomicName id="4C5E12E4AF775D5EFF0FAEA17448DACD" ID-CoL="737VD" authority="Bierig, 1940" authorityName="Bierig" authorityYear="1940" box="[151,548,226,250]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="mammiger">
<emphasis id="B92AB575AF775D5EFF0FAEA177F4DACD" bold="true" box="[151,408,226,250]" italics="true" pageId="9" pageNumber="116">Megarthrus mammiger</emphasis>
<bibRefCitation id="EFCF1496AF775D5EFE07AEA07448DACD" author="BIERIG, A." box="[415,548,227,250]" pageId="9" pageNumber="116" pagination="373 - 380" refId="ref12543" refString="BIERIG, A. 1940. Proteinini (Col. Staph.) Costaricenses. Revista de Entomologia 11: 373 - 380." type="journal article" year="1940">Bierig, 1940</bibRefCitation>
</taxonomicName>
Figs 2, 105-123
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3443AECAF775D5EFF0FAF447709DB58" pageId="9" pageNumber="116" type="reference_group">
<paragraph id="8BE16967AF775D5EFF0FAF44725DDB0A" blockId="9.[151,557,262,282]" lastBlockId="9.[151,1075,295,499]" pageId="9" pageNumber="116">
<treatmentCitationGroup id="AB4E4E49AF775D5EFF0FAF44725DDB0A" pageId="9" pageNumber="116">
<treatmentCitation id="0AFF4F76AF775D5EFF0FAF447445DB2C" author="BIERIG, A." box="[151,553,262,284]" page="375" pageId="9" pageNumber="116" year="1940">
<taxonomicName id="4C5E12E4AF775D5EFF0FAF447445DB2C" ID-CoL="737VD" authority="Bierig, 1940: 375" authorityName="Bierig" authorityPageNumber="375" authorityYear="1940" box="[151,553,262,284]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="mammiger">
<emphasis id="B92AB575AF775D5EFF0FAF44771ADB2B" box="[151,374,263,284]" italics="true" pageId="9" pageNumber="116">Megarthrus mammiger</emphasis>
<materialsCitation id="3B36633AAF775D5EFEE4AF457445DB2C" box="[380,553,262,283]" collectionCode="TYPE, MATERIAL, FMNH" country="Costa Rica" location="Vara Blanca" pageId="9" pageNumber="116" specimenCount="1" typeStatus="lectotype">
<bibRefCitation id="EFCF1496AF775D5EFEE4AF457445DB2C" author="BIERIG, A." box="[380,553,262,283]" pageId="9" pageNumber="116" pagination="373 - 380" refId="ref12543" refString="BIERIG, A. 1940. Proteinini (Col. Staph.) Costaricenses. Revista de Entomologia 11: 373 - 380." type="journal article" year="1940">Bierig, 1940: 375</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
. TYPE MATERIAL:
<typeStatus id="54E5D7C5AF775D5EFE11AF6B7785DB0A" box="[393,489,296,317]" pageId="9" pageNumber="116" type="lectotype">Lectotype</typeStatus>
(Ƌ, in FMNH): “
<location id="8E813FBCAF775D5EFD3BAF6B754CDB0A" LSID="urn:lsid:plazi:treatment:03F7D871AF775D5DFF0FAEA17471D842:8E813FBCAF775D5EFD3BAF6B754CDB0A" box="[675,800,296,317]" country="Costa Rica" name="Vara Blanca" pageId="9" pageNumber="116">Vara Blanca</location>
, viii.38,
<collectingCountry id="F34929F7AF775D5EFCE4AF6B7585DB0A" box="[892,1001,296,317]" name="Costa Rica" pageId="9" pageNumber="116">Costa Rica</collectingCountry>
[hand-
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE16967AF775D5EFF0FAF027709DB58" blockId="9.[151,1075,295,499]" pageId="9" pageNumber="116">
<materialsCitation id="3B36633AAF775D5EFF0FAF027732DB58" collectionCode="FMNH" country="Costa Rica" location="Vara Blanca" pageId="9" pageNumber="116" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
written] / Typus”.
<typeStatus id="54E5D7C5AF775D5EFEC1AF0277B0DB61" box="[345,476,321,342]" pageId="9" pageNumber="116" type="paralectotype">Paralectotype</typeStatus>
(
<specimenCount id="9D58A2EEAF775D5EFE73AF02747CDB61" box="[491,528,320,342]" pageId="9" pageNumber="116" type="female">1 ♀</specimenCount>
, in
<collectionCode id="ED4FF1A2AF775D5EFDADAF027413DB61" box="[565,639,321,342]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="116" type="Museum">FMNH</collectionCode>
): same data as lectotype, but “
<typeStatus id="54E5D7C5AF775D5EFC39AF027269DB61" box="[929,1029,321,342]" pageId="9" pageNumber="116" type="paratype">Paratypus</typeStatus>
”,
<emphasis id="B92AB575AF775D5EFB80AF027732DB58" bold="true" pageId="9" pageNumber="116">by present designation</emphasis>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3443AECAF775D5DFF7BAF3A7401DB6D" lastPageId="10" lastPageNumber="117" pageId="9" pageNumber="116" type="description">
<paragraph id="8BE16967AF775D5EFF7BAF3A75B2DBC4" blockId="9.[151,1075,295,499]" pageId="9" pageNumber="116">
ADDITIONAL MATERIAL:
<materialsCitation id="3B36633AAF775D5EFE55AF3A745FDB90" collectingDate="1985-04" collectionCode="CNCI" collectorName="Goulet &amp; Masner" country="Costa Rica" elevation="3200" latitude="9.6" location="Hwy" longLatPrecision="1299" longitude="-83.75" pageId="9" pageNumber="116" specimenCount="1">
<collectingCountry id="F34929F7AF775D5EFE55AF3A7433DBB9" box="[461,607,377,398]" name="Costa Rica" pageId="9" pageNumber="116">COSTA RICA</collectingCountry>
,
<location id="8E813FBCAF775D5EFDF5AF3A74F0DBB9" LSID="urn:lsid:plazi:treatment:03F7D871AF775D5DFF0FAEA17471D842:8E813FBCAF775D5EFDF5AF3A74F0DBB9" box="[621,668,377,398]" country="Costa Rica" latitude="9.6" longLatPrecision="1299" longitude="-83.75" name="Hwy" pageId="9" pageNumber="116">Hwy</location>
#2, km 93 &lt;
<geoCoordinate id="EE6A0FA0AF775D5EFCB1AF3A7513DBB9" box="[809,895,377,398]" degrees="83" direction="west" minutes="45" orientation="longitude" pageId="9" pageNumber="116" precision="925" value="-83.75">83°45'W</geoCoordinate>
;
<geoCoordinate id="EE6A0FA0AF775D5EFC15AF3A75BCDBB9" box="[909,976,377,398]" degrees="9" direction="north" minutes="36" orientation="latitude" pageId="9" pageNumber="116" precision="925" value="9.6">9°36'N</geoCoordinate>
&gt;
<quantity id="4CA6C482AF775D5EFC71AF3A7241DBB9" box="[1001,1069,377,398]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="9" pageNumber="116" unit="m" value="3200.0">
<elevation id="00738E54AF775D5EFC71AF3A7241DBB9" box="[1001,1069,377,398]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="9" pageNumber="116" unit="m" value="3200.0">3200m</elevation>
</quantity>
,
<date id="FFE04FA7AF775D5EFF0FAFD176B3DB90" box="[151,223,402,423]" pageId="9" pageNumber="116" value="1985-04">
<collectingDate id="EFA4B64FAF775D5EFF0FAFD176B3DB90" box="[151,223,402,423]" pageId="9" pageNumber="116" value="1985-04">iv.1985</collectingDate>
</date>
(
<collectorName id="26AB0CB1AF775D5EFF74AFD17743DB90" box="[236,303,402,423]" pageId="9" pageNumber="116">Goulet</collectorName>
&amp;
<collectorName id="26AB0CB1AF775D5EFEC8AFD177F0DB90" box="[336,412,402,423]" pageId="9" pageNumber="116">Masner</collectorName>
), 1 Ƌ in
<collectionCode id="ED4FF1A2AF775D5EFE6FAFD1745FDB90" box="[503,563,402,423]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="9" pageNumber="116">CNCI</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B36633AAF775D5EFDC9AFD17588DBF6" collectingDate="1989-05-21" collectionCode="SEMC" collectorName="Ashe &amp; Brooks &amp; Leschen" country="Costa Rica" elevation="1780" location="Cerro Amigos" municipality="Monte Verde" pageId="9" pageNumber="116" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas">
<collectingRegion id="499AA785AF775D5EFDC9AFD174D1DB90" box="[593,701,402,423]" country="Costa Rica" name="Puntarenas" pageId="9" pageNumber="116">Puntarenas</collectingRegion>
,
<collectingMunicipality id="6B85F31DAF775D5EFD50AFD17524DB90" box="[712,840,402,423]" pageId="9" pageNumber="116">Monte Verde</collectingMunicipality>
,
<location id="8E813FBCAF775D5EFCCBAFD175B1DB90" LSID="urn:lsid:plazi:treatment:03F7D871AF775D5DFF0FAEA17471D842:8E813FBCAF775D5EFCCBAFD175B1DB90" box="[851,989,402,423]" country="Costa Rica" municipality="Monte Verde" name="Cerro Amigos" pageId="9" pageNumber="116" stateProvince="Puntarenas">Cerro Amigos</location>
,
<quantity id="4CA6C482AF775D5EFC71AFD17242DB90" box="[1001,1070,402,423]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="9" pageNumber="116" unit="m" value="1780.0">
<elevation id="00738E54AF775D5EFC71AFD17242DB90" box="[1001,1070,402,423]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="9" pageNumber="116" unit="m" value="1780.0">1780m</elevation>
</quantity>
,
<date id="FFE04FA7AF775D5EFF0FAFEF7699DBF6" box="[151,245,428,449]" pageId="9" pageNumber="116" value="1989-05-21">
<collectingDate id="EFA4B64FAF775D5EFF0FAFEF7699DBF6" box="[151,245,428,449]" pageId="9" pageNumber="116" value="1989-05-21">21.v.1989</collectingDate>
</date>
(
<collectorName id="26AB0CB1AF775D5EFE9BAFEF7754DBF6" box="[259,312,428,449]" pageId="9" pageNumber="116">Ashe</collectorName>
,
<collectorName id="26AB0CB1AF775D5EFED9AFEF77EADBF6" box="[321,390,428,449]" pageId="9" pageNumber="116">Brooks</collectorName>
&amp;
<collectorName id="26AB0CB1AF775D5EFE3EAFEF7794DBF6" box="[422,504,428,449]" pageId="9" pageNumber="116">Leschen</collectorName>
)
<collectingMethod id="521F1170AF775D5EFD9CAFEF74ACDBF6" box="[516,704,428,449]" pageId="9" pageNumber="116">flight intercept trap</collectingMethod>
, #315, 1 Ƌ and
<specimenCount id="9D58A2EEAF775D5EFCC3AFEF75EDDBF7" box="[859,897,426,449]" pageId="9" pageNumber="116" type="female">1 ♀</specimenCount>
in
<collectionCode id="ED4FF1A2AF775D5EFC38AFEF7588DBF6" box="[928,996,428,449]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="116" type="Museum">SEMC</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B36633AAF775D5EFB98AFEF7411DBED" collectingDate="1940-01-18" collectionCode="MHNG" collectorName="Bierig" country="Costa Rica" elevation="2800" location="Volc. Irazu" pageId="9" pageNumber="116" specimenCount="1" stateProvince="Puntarenas">
<location id="8E813FBCAF775D5EFB98AFEF76A7DBED" LSID="urn:lsid:plazi:treatment:03F7D871AF775D5DFF0FAEA17471D842:8E813FBCAF775D5EFB98AFEF76A7DBED" country="Costa Rica" name="Volc. Irazu" pageId="9" pageNumber="116" stateProvince="Puntarenas">Volc. Irazú</location>
,
<quantity id="4CA6C482AF775D5EFF4FAF867770DBED" box="[215,284,453,474]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="9" pageNumber="116" unit="m" value="2800.0">
<elevation id="00738E54AF775D5EFF4FAF867770DBED" box="[215,284,453,474]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="9" pageNumber="116" unit="m" value="2800.0">2800m</elevation>
</quantity>
,
<date id="FFE04FA7AF775D5EFEB0AF8677EFDBED" box="[296,387,453,474]" pageId="9" pageNumber="116" value="1940-01-18">
<collectingDate id="EFA4B64FAF775D5EFEB0AF8677EFDBED" box="[296,387,453,474]" pageId="9" pageNumber="116" value="1940-01-18">18.i.1940</collectingDate>
</date>
(
<collectorName id="26AB0CB1AF775D5EFE0AAF8677BFDBED" box="[402,467,453,474]" pageId="9" pageNumber="116">Bierig</collectorName>
), 1 Ƌ in
<collectionCode id="ED4FF1A2AF775D5EFDA9AF867411DBED" box="[561,637,453,474]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="116" type="Museum">MHNG</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B36633AAF775D5EFD05AF86774BDBC4" collectingDate="1944-05-19" collectionCode="FMNH" collectorName="El Jardiu &amp; Bierig" country="Costa Rica" elevation="3000" pageId="9" pageNumber="116" specimenCount="1" stateProvince="Puntarenas">
<collectorName id="26AB0CB1AF775D5EFD05AF867494DBED" box="[669,760,453,474]" pageId="9" pageNumber="116">El Jardiú</collectorName>
, ca.
<quantity id="4CA6C482AF775D5EFCB0AF867500DBED" box="[808,876,453,474]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="116" unit="m" value="3000.0">
<elevation id="00738E54AF775D5EFCB0AF867500DBED" box="[808,876,453,474]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="116" unit="m" value="3000.0">3000m</elevation>
</quantity>
,
<date id="FFE04FA7AF775D5EFCE1AF8675BBDBED" box="[889,983,453,474]" pageId="9" pageNumber="116" value="1944-05-19">
<collectingDate id="EFA4B64FAF775D5EFCE1AF8675BBDBED" box="[889,983,453,474]" pageId="9" pageNumber="116" value="1944-05-19">19.v.1944</collectingDate>
</date>
(
<collectorName id="26AB0CB1AF775D5EFC7EAF86724BDBED" box="[998,1063,453,474]" pageId="9" pageNumber="116">Bierig</collectorName>
), 1 Ƌ in
<collectionCode id="ED4FF1A2AF775D5EFF46AF9D774BDBC4" box="[222,295,478,499]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="116" type="Museum">FMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B36633AAF775D5EFEDAAF9D75B7DBC4" box="[322,987,477,499]" collectingDate="1944-05-19" collectionCode="MHNG" collectorName="il Tajar &amp; Bierig" country="Costa Rica" elevation="1500" location="San Isidro" pageId="9" pageNumber="116" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas">
<location id="8E813FBCAF775D5EFEDAAF9D77C9DBC4" LSID="urn:lsid:plazi:treatment:03F7D871AF775D5DFF0FAEA17471D842:8E813FBCAF775D5EFEDAAF9D77C9DBC4" box="[322,421,478,499]" country="Costa Rica" name="San Isidro" pageId="9" pageNumber="116" stateProvince="Puntarenas">San Isidro</location>
de
<collectorName id="26AB0CB1AF775D5EFE51AF9D7463DBC4" box="[457,527,478,499]" pageId="9" pageNumber="116">il Tajar</collectorName>
, ca.
<quantity id="4CA6C482AF775D5EFDA3AF9D74ECDBC4" box="[571,640,478,499]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="116" unit="m" value="1500.0">
<elevation id="00738E54AF775D5EFDA3AF9D74ECDBC4" box="[571,640,478,499]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="116" unit="m" value="1500.0">1500m</elevation>
</quantity>
,
<date id="FFE04FA7AF775D5EFD12AF9D7485DBC4" box="[650,745,478,499]" pageId="9" pageNumber="116" value="1944-05-19">
<collectingDate id="EFA4B64FAF775D5EFD12AF9D7485DBC4" box="[650,745,478,499]" pageId="9" pageNumber="116" value="1944-05-19">19.v.1944</collectingDate>
</date>
(
<collectorName id="26AB0CB1AF775D5EFD6EAF9D755BDBC4" box="[758,823,478,499]" pageId="9" pageNumber="116">Bierig</collectorName>
),
<specimenCount id="9D58A2EEAF775D5EFCD0AF9D7502DBC4" box="[840,878,477,499]" pageId="9" pageNumber="116" type="female">1 ♀</specimenCount>
in
<collectionCode id="ED4FF1A2AF775D5EFC16AF9D75B7DBC4" box="[910,987,478,499]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="116" type="Museum">MHNG</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE16967AF775D5EFF7BAC477695D964" blockId="9.[151,1074,516,1660]" pageId="9" pageNumber="116">
DESCRIPTION: Habitus as in Fig. 2. Combined length of pronotum and elytra =
<quantity id="4CA6C482AF775D5EFF0FAC607779D80D" box="[151,277,547,570]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="9" pageNumber="116" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.8-1.9 mm</quantity>
; maximal pronotal width =
<quantity id="4CA6C482AF775D5EFDA0AC6074E0D80D" box="[568,652,547,570]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="116" unit="mm" value="1.3">1.3 mm</quantity>
. Body dark brown with appendages and elytra slightly paler; antennomeres 10-11 paler than antennomeres 1-9. Dorsal pubescence fairly uniform, denser on head and pronotum than on elytral disc; frontal setae directed forward; elytral and pronotal setae fairly straight, recumbent; metasternal pubescence fairly uniform, shorter than prosternal; pubescence on abdominal tergites parallel, uniform. Frons, anterior portion of prohypomera, humeral callus and lateral area of elytral disc granulate; frontal granulation inconspicuous, with granula about as high as half of their diameter; pronotum granulofossulate; metasternum with granulofossulation becoming finer posteriomedially; adsutural half of elytral disc punctate.
</paragraph>
<paragraph id="8BE16967AF775D5EFF7BAD1877A4D9D8" blockId="9.[151,1074,516,1660]" pageId="9" pageNumber="116">Frons forming above clypeus a sharp ridge, the latter finely evenly carinate, weakly arcuate in middle and laterally oblique in dorsal view; mesal portion of disc strongly evenly convex in lateral view; U-shaped frontal impression deep in middle, shallow laterally. Temples strongly convex in dorsal view. Antennae (Fig. 115) 2.3-2.6 times longer than pronotum.</paragraph>
<paragraph id="8BE16967AF775D5EFF7BADB475E4DE9D" blockId="9.[151,1074,516,1660]" pageId="9" pageNumber="116">Pronotum (Fig. 123) with center strongly convex in frontal view; disc deeply depressed near middle of lateral edges, shallowly depressed along anterior and posterior margins; medial groove nearly straight in lateral view, deep, somewhat narrowed in middle; hypomera ridged from anterior margin to laterobasal angle, disc without pit. Prosternal medial ridge absent. Scutellum with anterior margin angulate in middle and posterior margin slighly arcuate toward right-angled apex.</paragraph>
<paragraph id="8BE16967AF775D5EFF7BAAF177BBDF10" blockId="9.[151,1074,516,1660]" pageId="9" pageNumber="116">Elytra gradually widened (Fig. 2); humeral callus raised, forming a blunt longitudinal ridge; disc with low anterior and posterior adsutural humps, moderately depressed posteriorly along lateral edge; the latter finely carinate, finely denticulate, nearly straight in dorsal view.</paragraph>
<paragraph id="8BE16967AF775D5EFF7BAB6C745FDF52" blockId="9.[151,1074,516,1660]" pageId="9" pageNumber="116">Abdominal sternites II and III with medial processes as in Fig. 120, posterior portion of process of sternite III trifid.</paragraph>
<paragraph id="8BE16967AF775D5EFF7BAB2E726FDC4B" blockId="9.[151,1074,516,1660]" pageId="9" pageNumber="116">Male: Frontoclypeal area not modified. Protarsomeres 1 lacking tenent setae. Mesofemora (Fig. 111) as long as metafemora (Fig. 110). Mesotibiae (Fig. 107) shorter than metatibiae (Fig. 108). Metatarsomeres 1 about as long as combined length of metatarsomeres 2-4. Peg-like setae arranged in a single row on mesotrochanters (Fig. 111), mesotibiae and metatibiae, absent from protrochanters, profemora, protibiae, mesofemora, metatrochanters metatrochanter (Figs 109-110) and metafemora; metatrochanters with a projecting process. Pubescence on sternites IV-VII becoming denser posteriomedially. Apex of abdominal tergite VIII as in Figs 113, 114. Sternite VIII as in Fig. 112. Sternite IX lacking subbasal protuberance. Aedeagus as in Figs 105-106.</paragraph>
<paragraph id="8BE16967AF745D5DFF4CAEA07401DB6D" blockId="10.[136,1059,227,346]" pageId="10" pageNumber="117">Female: Pubescence on sternites IV-VII uniform, except for a pair of subapical macrosetae on each sternite. Abdominal tergite VIII (Figs 121-122) without medioapical projection. Sternite VIII as in Fig. 119. Genital segments as in Figs 116-118; gonocoxal plate bearing a mediodorsal ridge.</paragraph>
</subSubSection>
<subSubSection id="C3443AECAF745D5DFF4CAF32749FDBDF" pageId="10" pageNumber="117" type="distribution">
<paragraph id="8BE16967AF745D5DFF4CAF32749FDBDF" blockId="10.[136,1059,369,488]" pageId="10" pageNumber="117">
<materialsCitation id="3B36633AAF745D5DFF4CAF327483DBDF" collectorName="A. Bierig" country="Costa Rica" elevation="2350" location="The" pageId="10" pageNumber="117" specimenCount="1">
DISTRIBUTION AND NATURAL HISTORY:
<location id="8E813FBCAF745D5DFDEAAF3274F7DBBF" LSID="urn:lsid:plazi:treatment:03F7D871AF775D5DFF0FAEA17471D842:8E813FBCAF745D5DFDEAAF3274F7DBBF" box="[626,667,369,392]" country="Costa Rica" name="The" pageId="10" pageNumber="117">The</location>
species is known only from
<collectingCountry id="F34929F7AF745D5DFC7EAF3276D0DB9F" name="Costa Rica" pageId="10" pageNumber="117">Costa Rica</collectingCountry>
, where it was found at elevations ranging from
<quantity id="4CA6C482AF745D5DFD54AFD2750CDB9F" box="[716,864,401,424]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="3.2" metricValueMin="1.5" pageId="10" pageNumber="117" unit="m" value="2350.0" valueMax="3200.0" valueMin="1500.0">
<elevation id="00738E54AF745D5DFD54AFD2750CDB9F" box="[716,864,401,424]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="3.2" metricValueMin="1.5" pageId="10" pageNumber="117" unit="m" value="2350.0" valueMax="3200.0" valueMin="1500.0">1500-3200 m</elevation>
</quantity>
. According to the original description,
<collectorName id="26AB0CB1AF745D5DFEF2AFF277BCDBFF" box="[362,464,433,456]" pageId="10" pageNumber="117">A. Bierig</collectorName>
collected the
<typeStatus id="54E5D7C5AF745D5DFDF6AFF274CADBFF" box="[622,678,433,456]" pageId="10" pageNumber="117">types</typeStatus>
between the Volcanoes Barba and Poás, at
<quantity id="4CA6C482AF745D5DFF7BAF927759DBDF" box="[227,309,465,488]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="10" pageNumber="117" unit="m" value="1800.0">
<elevation id="00738E54AF745D5DFF7BAF927759DBDF" box="[227,309,465,488]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="10" pageNumber="117" unit="m" value="1800.0">1800 m</elevation>
</quantity>
in cow dung and in dead bamboo leaves
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3443AECAF745D5DFF4CAFBD7471D842" pageId="10" pageNumber="117" type="discussion">
<paragraph id="8BE16967AF745D5DFF4CAFBD7471D842" blockId="10.[136,1059,510,629]" pageId="10" pageNumber="117">
COMMENTS: Among the Neotropical
<taxonomicName id="4C5E12E4AF745D5DFDEFAFBD7499D822" authorityName="Curtis" authorityYear="1829" box="[631,757,510,533]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B92AB575AF745D5DFDEFAFBD7499D822" box="[631,757,510,533]" italics="true" pageId="10" pageNumber="117">Megarthrus</emphasis>
</taxonomicName>
possessing a conspicuous humeral callus (see comments under
<taxonomicName id="4C5E12E4AF745D5DFDDBAC5D74DBD802" authorityName="Cuccodoro &amp; C. &amp; P. &amp; Ch-" authorityYear="2011" box="[579,695,542,565]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="117" phylum="Arthropoda" rank="species" species="bierigi">
<emphasis id="B92AB575AF745D5DFDDBAC5D74DBD802" box="[579,695,542,565]" italics="true" pageId="10" pageNumber="117">M. bierigi</emphasis>
</taxonomicName>
),
<taxonomicName id="4C5E12E4AF745D5DFD4FAC5D751AD802" authorityName="Bierig" authorityYear="1940" box="[727,886,542,565]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="117" phylum="Arthropoda" rank="species" species="mammiger">
<emphasis id="B92AB575AF745D5DFD4FAC5D751AD802" box="[727,886,542,565]" italics="true" pageId="10" pageNumber="117">M. mammiger</emphasis>
</taxonomicName>
can be easily recognised by its laterobasally shallowly emarginated pronotum. See also comments under
<taxonomicName id="4C5E12E4AF745D5DFF54AC1D7733D842" authorityName="Bierig" authorityYear="1940" box="[204,351,606,629]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="117" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B92AB575AF745D5DFF54AC1D7733D842" box="[204,351,606,629]" italics="true" pageId="10" pageNumber="117">M. inaequalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5E12E4AF745D5DFE0DAC1D747AD842" authorityName="Bierig" authorityYear="1940" box="[405,534,606,629]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="117" phylum="Arthropoda" rank="species" species="mastiger">
<emphasis id="B92AB575AF745D5DFE0DAC1D747AD842" box="[405,534,606,629]" italics="true" pageId="10" pageNumber="117">M. mastiger</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>