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<document id="A6561391A6F86AE396645FFCA3D6B7FA" ID-DOI="10.5962/bhl.part.117802" ID-ISSN="0035-418X" ID-Zenodo-Dep="6311706" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645021050174" checkinUser="carolina" docAuthor="Cuccodoro, Giulio" docDate="2011" docId="03F7D871AF735D58FF0FABFE7248D812" docLanguage="en" docName="RevSuisseZool.118.107-148.pdf.imf" docOrigin="Revue suisse de Zoologie 118 (1)" docStyle="DocumentStyle:0846B485C0C87F6EB7648DEEBF34B77F.2:RevSuisseZool.2011-2014.journal_article" docStyleId="0846B485C0C87F6EB7648DEEBF34B77F" docStyleName="RevSuisseZool.2011-2014.journal_article" docStyleVersion="2" docTitle="Megarthrus zunilensis Sharp. These 1887" docType="treatment" docVersion="4" lastPageNumber="122" masterDocId="FFCEA009AF7E5D57FF98AE43766CDA37" masterDocTitle="Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae)" masterLastPageNumber="147" masterPageNumber="107" pageNumber="120" updateTime="1699296264638" updateUser="plazi">
<mods:mods id="020B2FDFEDB4CA377467DE0DCA13DD6B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0DCB7941A50A8159FECC778FF7DC31CD">
<mods:title id="1AFE46F1F170741F47B6230B4744B89B">Revision of the Neotropical types of Megarthrus Curtis, 1829 and description of two new species from Costa Rica and Peru (Coleoptera, Staphylinidae, Proteininae)</mods:title>
</mods:titleInfo>
<mods:name id="266FE9EE257F3F95907D4F5AEFF091D6" type="personal">
<mods:role id="A3FEB7F747D5231109D406E1BC986117">
<mods:roleTerm id="7840D8BFF07A9E8E1F9E75D7D1B443AB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E5083186DD427DB7F3ACCDE77E493E64">Cuccodoro, Giulio</mods:namePart>
</mods:name>
<mods:typeOfResource id="075CB8745489E98ACBF9CC43115631B9">text</mods:typeOfResource>
<mods:relatedItem id="13968A4545D3ADC06B510C4DCA7DE9E9" type="host">
<mods:titleInfo id="92B21634B8229A5685769E5C60EF1045">
<mods:title id="820E95277E22CCF9C23B0F5547C199CB">Revue suisse de Zoologie</mods:title>
</mods:titleInfo>
<mods:part id="5D1ECA8E5863B9290D242B09CB1FCEBF">
<mods:date id="C6D1BE7BA650F94C416E48A35E0E2DF0">2011</mods:date>
<mods:detail id="BEB2B9D8869F4E0C9CB6DD8EF4B33DC3" type="pubDate">
<mods:number id="0BF94ED3D054B4BEC47E689C134CD445">2011-03-31</mods:number>
</mods:detail>
<mods:detail id="CF1EA0D99DAE28A69558845C4E4E7ED5" type="volume">
<mods:number id="F8A870BB3A887658647A14E0545D792B">118</mods:number>
</mods:detail>
<mods:detail id="BA37AAD328018388D6EE126E3F1012CA" type="issue">
<mods:number id="E50F564A8F81910CF3BD67CED3AA7920">1</mods:number>
</mods:detail>
<mods:extent id="8FACCDD3EF7147E6A0B3B83C914B9BE9" unit="page">
<mods:start id="9ED82A5EEFE3F6C61C3D6146AA32A8C0">107</mods:start>
<mods:end id="9D7442686D099402AD10D7BB082E0703">147</mods:end>
</mods:extent>
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<mods:classification id="AFBCEB7265A27CA6F3EDE32E55951CAE">journal article</mods:classification>
<mods:identifier id="0BF5E654A40FE2F0B50B3035F20E322A" type="DOI">10.5962/bhl.part.117802</mods:identifier>
<mods:identifier id="AA1863D6707E749195B36B672E980B1C" type="ISSN">0035-418X</mods:identifier>
<mods:identifier id="C1AB0281D7504FC0A5FF74EE6730EE84" type="Zenodo-Dep">6311706</mods:identifier>
</mods:mods>
<treatment id="03F7D871AF735D58FF0FABFE7248D812" ID-DOI="http://doi.org/10.5281/zenodo.6311375" ID-Zenodo-Dep="6311375" LSID="urn:lsid:plazi:treatment:03F7D871AF735D58FF0FABFE7248D812" httpUri="http://treatment.plazi.org/id/03F7D871AF735D58FF0FABFE7248D812" lastPageId="15" lastPageNumber="122" pageId="13" pageNumber="120">
<subSubSection id="C3443AECAF735D5AFF0FABFE725DDFE2" box="[151,1073,1469,1493]" pageId="13" pageNumber="120" type="nomenclature">
<paragraph id="8BE16967AF735D5AFF0FABFE725DDFE2" blockId="13.[151,1073,1469,1493]" box="[151,1073,1469,1493]" pageId="13" pageNumber="120">
<heading id="D0A9DE0BAF735D5AFF0FABFE725DDFE2" box="[151,1073,1469,1493]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="120" reason="2">
<taxonomicName id="4C5E12E4AF735D5AFF0FABFE747BDFE2" ID-CoL="3YXDC" authority="Sharp, 1887" authorityName="Sharp. These" authorityYear="1887" box="[151,535,1469,1493]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="120" phylum="Arthropoda" rank="species" species="zunilensis">
<emphasis id="B92AB575AF735D5AFF0FABFE77E3DFE2" bold="true" box="[151,399,1469,1493]" italics="true" pageId="13" pageNumber="120">Megarthrus zunilensis</emphasis>
<bibRefCitation id="EFCF1496AF735D5AFE0EABFD747BDFE2" author="SHARP, D." box="[406,535,1470,1493]" pageId="13" pageNumber="120" pagination="673 - 824" refId="ref12839" refString="SHARP, D. 1887. Staphylinidae (pp. 673 - 824). In: Biologia Centrali-Americana. Insecta, Coleoptera 1 (2). Taylor &amp; Francis, London, XVI + 824 pp. + 18 pl." type="book chapter" year="1887">Sharp, 1887</bibRefCitation>
</taxonomicName>
Figs 6, 171-187
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3443AECAF735D5AFF0FABA2744DDFC1" box="[151,545,1505,1526]" pageId="13" pageNumber="120" type="reference_group">
<paragraph id="8BE16967AF735D5AFF0FABA2744DDFC1" blockId="13.[151,545,1505,1526]" box="[151,545,1505,1526]" pageId="13" pageNumber="120">
<treatmentCitationGroup id="AB4E4E49AF735D5AFF0FABA2744DDFC1" box="[151,545,1505,1526]" pageId="13" pageNumber="120">
<treatmentCitation id="0AFF4F76AF735D5AFF0FABA27471DFC1" author="SHARP, D." box="[151,541,1505,1526]" page="743" pageId="13" pageNumber="120" year="1887">
<taxonomicName id="4C5E12E4AF735D5AFF0FABA27471DFC1" ID-CoL="3YXDC" authority="Sharp, 1887: 743" authorityName="Sharp. These" authorityPageNumber="743" authorityYear="1887" box="[151,541,1505,1526]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="120" phylum="Arthropoda" rank="species" species="zunilensis">
<emphasis id="B92AB575AF735D5AFF0FABA27702DFC1" box="[151,366,1505,1526]" italics="true" pageId="13" pageNumber="120">Megarthrus zunilensis</emphasis>
<bibRefCitation id="EFCF1496AF735D5AFEECABA27471DFC1" author="SHARP, D." box="[372,541,1505,1526]" pageId="13" pageNumber="120" pagination="673 - 824" refId="ref12839" refString="SHARP, D. 1887. Staphylinidae (pp. 673 - 824). In: Biologia Centrali-Americana. Insecta, Coleoptera 1 (2). Taylor &amp; Francis, London, XVI + 824 pp. + 18 pl." type="book chapter" year="1887">Sharp, 1887: 743</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3443AECAF735D59FF7BA84177FEDBC4" lastPageId="14" lastPageNumber="121" pageId="13" pageNumber="120" type="materials_examined">
<paragraph id="8BE16967AF735D5AFF7BA84177E9DC4A" blockId="13.[151,1074,1537,1661]" pageId="13" pageNumber="120">
<materialsCitation id="3B36633AAF735D5AFF7BA8417446DC7D" collectionCode="TYPE, MATERIAL, BMNH" collectorName="B. C. A. Col." country="Guatemala" location="Zunil" municipality="Zunil" pageId="13" pageNumber="120" specimenCount="1" typeStatus="lectotype">
<collectionCode id="ED4FF1A2AF735D5AFF7BA841771BDC21" box="[227,375,1538,1559]" pageId="13" pageNumber="120">TYPE MATERIAL</collectionCode>
:
<typeStatus id="54E5D7C5AF735D5AFE1DA8417789DC20" box="[389,485,1538,1559]" pageId="13" pageNumber="120" type="lectotype">Lectotype</typeStatus>
(Ƌ, in
<collectionCode id="ED4FF1A2AF735D5AFDABA84174ECDC20" box="[563,640,1538,1559]" pageId="13" pageNumber="120">BMNH</collectionCode>
): “
<taxonomicName id="4C5E12E4AF735D5AFD04A8407518DC2F" authorityName="Sharp. These" authorityYear="1887" box="[668,884,1539,1560]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="120" phylum="Arthropoda" rank="species" species="zunilensis">
<emphasis id="B92AB575AF735D5AFD04A8407518DC2F" box="[668,884,1539,1560]" italics="true" pageId="13" pageNumber="120">Megarthrus zunilensis</emphasis>
</taxonomicName>
. Type D. S., Cerro
<location id="8E813FBCAF735D5AFF0FA85F76A2DC06" LSID="urn:lsid:plazi:treatment:03F7D871AF735D58FF0FABFE7248D812:8E813FBCAF735D5AFF0FA85F76A2DC06" box="[151,206,1564,1585]" country="Guatemala" municipality="Zunil" name="Zunil" pageId="13" pageNumber="120">Zunil</location>
. Guatem., Champion [Sharp’s handwritting] | Cerro
<collectingMunicipality id="6B85F31DAF735D5AFD5DA85F7497DC06" box="[709,763,1564,1585]" pageId="13" pageNumber="120">Zunil</collectingMunicipality>
,
<collectingCountry id="F34929F7AF735D5AFC9DA85F7503DC06" box="[773,879,1564,1585]" name="Guatemala" pageId="13" pageNumber="120">Guatemala</collectingCountry>
, Champion |
<collectorName id="26AB0CB1AF735D5AFC76A85F76ADDC7D" pageId="13" pageNumber="120">B.C.A. Col.</collectorName>
1. 2.
<taxonomicName id="4C5E12E4AF735D5AFE9AA876744CDC7D" authority="Sharp" authorityName="Sharp. These" authorityYear="1887" box="[258,544,1589,1610]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="120" phylum="Arthropoda" rank="species" species="zunilensis">Megarthrus zunilensis Sharp</taxonomicName>
”
</materialsCitation>
. –
<materialsCitation id="3B36633AAF735D5AFDD5A8767712DC4B" collectionCode="BMNH" collectorName="B. C. A. Col." country="Guatemala" elevation="763" location="Cerro Zunil" municipality="Zunil" pageId="13" pageNumber="120" specimenCount="2" specimenCount-female="2" typeStatus="paralectotype">
<typeStatus id="54E5D7C5AF735D5AFDD5A87674B5DC7D" box="[589,729,1589,1610]" pageId="13" pageNumber="120" type="paralectotype">Paralectotypes</typeStatus>
(
<specimenCount id="9D58A2EEAF735D5AFD72A8767545DC7D" box="[746,809,1588,1610]" pageId="13" pageNumber="120" type="female">2 ♀♀</specimenCount>
in
<collectionCode id="ED4FF1A2AF735D5AFCD7A87675F0DC7D" box="[847,924,1589,1610]" pageId="13" pageNumber="120">BMNH</collectionCode>
): same data as lectotype, but “Sp. figured”; same data as
<typeStatus id="54E5D7C5AF735D5AFDA0A80D74FEDC54" box="[568,658,1614,1635]" pageId="13" pageNumber="120" type="lectotype">lectotype</typeStatus>
, but “
<location id="8E813FBCAF735D5AFD55A80D752FDC54" LSID="urn:lsid:plazi:treatment:03F7D871AF735D58FF0FABFE7248D812:8E813FBCAF735D5AFD55A80D752FDC54" box="[717,835,1614,1635]" municipality="Zunil" name="Cerro Zunil" pageId="13" pageNumber="120">
Cerro
<collectingMunicipality id="6B85F31DAF735D5AFC95A80D752FDC54" box="[781,835,1614,1635]" pageId="13" pageNumber="120">Zunil</collectingMunicipality>
</location>
,
<quantity id="4CA6C482AF735D5AFCD7A80D75DDDC54" box="[847,945,1614,1635]" metricMagnitude="2" metricUnit="m" metricValue="7.6260959999999995" metricValueMax="15.24" metricValueMin="0.012192" pageId="13" pageNumber="120" unit="ft" value="2502.0" valueMax="5000.0" valueMin="4.0">
<elevation id="00738E54AF735D5AFCD7A80D75DDDC54" box="[847,945,1614,1635]" metricMagnitude="2" metricUnit="m" metricValue="7.6260959999999995" metricValueMax="15.24" metricValueMin="0.012192" pageId="13" pageNumber="120" unit="ft" value="2502.0" valueMax="5000.0" valueMin="4.0">4-5000 ft.</elevation>
</quantity>
, Champion”,
<emphasis id="B92AB575AF735D5AFF0FA8247712DC4B" bold="true" box="[151,382,1639,1660]" pageId="13" pageNumber="120">by present designation</emphasis>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE16967AF705D59FF4CAEA077FEDBC4" blockId="14.[136,1059,227,499]" pageId="14" pageNumber="121">
ADDITIONAL
<collectionCode id="ED4FF1A2AF705D59FED7AEA477C6DAC0" box="[335,426,231,247]" pageId="14" pageNumber="121">MATERIAL</collectionCode>
:
<materialsCitation id="3B36633AAF705D59FE21AEA07464DB26" collectingDate="1979-05-23" collectingDateMax="1979-05-24" collectingDateMin="1979-05-23" collectionCode="CNCI" collectorName="Campbell" country="Costa Rica" elevation="1500" location="Monteverde Biol. Res." pageId="14" pageNumber="121" specimenCount="1" stateProvince="Puntarenas">
<collectingCountry id="F34929F7AF705D59FE21AEA07424DACF" box="[441,584,227,248]" name="Costa Rica" pageId="14" pageNumber="121">COSTA RICA</collectingCountry>
,
<collectingRegion id="499AA785AF705D59FDCCAEA074ACDACF" box="[596,704,227,248]" country="Costa Rica" name="Puntarenas" pageId="14" pageNumber="121">Puntarenas</collectingRegion>
,
<location id="8E813FBCAF705D59FD53AEA075C7DACF" LSID="urn:lsid:plazi:treatment:03F7D871AF735D58FF0FABFE7248D812:8E813FBCAF705D59FD53AEA075C7DACF" box="[715,939,227,248]" country="Costa Rica" name="Monteverde Biol. Res." pageId="14" pageNumber="121" stateProvince="Puntarenas">Monteverde Biol. Res.</location>
, ca.
<quantity id="4CA6C482AF705D59FC42AEA07273DACF" box="[986,1055,227,248]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="121" unit="m" value="1500.0">
<elevation id="00738E54AF705D59FC42AEA07273DACF" box="[986,1055,227,248]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="121" unit="m" value="1500.0">1500m</elevation>
</quantity>
,
<date id="FFE04FA7AF705D59FF10AEBF776BDB26" box="[136,263,252,273]" pageId="14" pageNumber="121" value="1979-05-23" valueMax="1979-05-24" valueMin="1979-05-23">
<collectingDate id="EFA4B64FAF705D59FF10AEBF776BDB26" box="[136,263,252,273]" pageId="14" pageNumber="121" value="1979-05-23" valueMax="1979-05-24" valueMin="1979-05-23">23-24.v.1979</collectingDate>
</date>
(
<collectorName id="26AB0CB1AF705D59FE8CAEBF7718DB26" box="[276,372,252,273]" pageId="14" pageNumber="121">Campbell</collectorName>
) 1 Ƌ in
<collectionCode id="ED4FF1A2AF705D59FE54AEBF7464DB26" box="[460,520,252,273]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="14" pageNumber="121">CNCI</collectionCode>
</materialsCitation>
. –
<materialsCitation id="3B36633AAF705D59FDBEAEBF756ADB1D" collectingDate="1993-06-20" collectionCode="SEMC" collectorName="Genier" country="Guatemala" elevation="1340" location="14.2 km SW Zunil" pageId="14" pageNumber="121" specimenCount="1" specimenCount-female="1" stateProvince="Quetzaltenango">
<collectingCountry id="F34929F7AF705D59FDBEAEBF74D2DB26" box="[550,702,252,273]" name="Guatemala" pageId="14" pageNumber="121">GUATEMALA</collectingCountry>
,
<collectingRegion id="499AA785AF705D59FD51AEBF750EDB26" box="[713,866,252,273]" country="Guatemala" name="Quetzaltenango" pageId="14" pageNumber="121">Quetzaltenango</collectingRegion>
,
<location id="8E813FBCAF705D59FCF5AEBF7272DB26" LSID="urn:lsid:plazi:treatment:03F7D871AF735D58FF0FABFE7248D812:8E813FBCAF705D59FCF5AEBF7272DB26" box="[877,1054,252,273]" country="Guatemala" name="14.2 km SW Zunil" pageId="14" pageNumber="121" stateProvince="Quetzaltenango">
<quantity id="4CA6C482AF705D59FCF5AEBF75DADB26" box="[877,950,252,273]" metricMagnitude="4" metricUnit="m" metricValue="1.42" pageId="14" pageNumber="121" unit="km" value="14.2">14.2km</quantity>
SW Zunil
</location>
,
<quantity id="4CA6C482AF705D59FF10AF5676A1DB1D" box="[136,205,277,298]" metricMagnitude="3" metricUnit="m" metricValue="1.34" pageId="14" pageNumber="121" unit="m" value="1340.0">
<elevation id="00738E54AF705D59FF10AF5676A1DB1D" box="[136,205,277,298]" metricMagnitude="3" metricUnit="m" metricValue="1.34" pageId="14" pageNumber="121" unit="m" value="1340.0">1340m</elevation>
</quantity>
,
<date id="FFE04FA7AF705D59FF40AF567752DB1D" box="[216,318,277,298]" pageId="14" pageNumber="121" value="1993-06-20">
<collectingDate id="EFA4B64FAF705D59FF40AF567752DB1D" box="[216,318,277,298]" pageId="14" pageNumber="121" value="1993-06-20">20.vi.1993</collectingDate>
</date>
(
<collectorName id="26AB0CB1AF705D59FED4AF5677FCDB1D" box="[332,400,277,298]" pageId="14" pageNumber="121">Génier</collectorName>
)
<collectingMethod id="521F1170AF705D59FE06AF567403DB1D" box="[414,623,277,298]" pageId="14" pageNumber="121">ex human faeces trap</collectingMethod>
,
<specimenCount id="9D58A2EEAF705D59FDE2AF5674CDDB1D" box="[634,673,276,298]" pageId="14" pageNumber="121" type="female">1 ♀</specimenCount>
in
<collectionCode id="ED4FF1A2AF705D59FD5AAF56756ADB1D" box="[706,774,277,298]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="14" pageNumber="121" type="Museum">SEMC</collectionCode>
</materialsCitation>
. –
<materialsCitation id="3B36633AAF705D59FCB3AF5676A0DB6A" collectingDate="1991-06-12" collectionCode="SEMC" collectorName="Anderson" country="Guatemala" elevation="1760" location="Duenas" municipality="Miguel" pageId="14" pageNumber="121" specimenCount="1" stateProvince="Sacatepequez">
<collectingRegion id="499AA785AF705D59FCB3AF5675DCDB1D" box="[811,944,277,298]" country="Guatemala" name="Sacatepequez" pageId="14" pageNumber="121">Sacatepequez</collectingRegion>
,
<quantity id="4CA6C482AF705D59FC22AF567594DB1D" box="[954,1016,277,298]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="121" unit="km" value="4.5">4.5km</quantity>
SW san
<collectingMunicipality id="6B85F31DAF705D59FF28AF6D7694DB74" box="[176,248,302,323]" pageId="14" pageNumber="121">Miguel</collectingMunicipality>
,
<location id="8E813FBCAF705D59FE9BAF6D7721DB74" LSID="urn:lsid:plazi:treatment:03F7D871AF735D58FF0FABFE7248D812:8E813FBCAF705D59FE9BAF6D7721DB74" box="[259,333,302,323]" country="Guatemala" municipality="Miguel" name="Duenas" pageId="14" pageNumber="121" stateProvince="Sacatepequez">Duenas</location>
,
<quantity id="4CA6C482AF705D59FEC1AF6D77F2DB74" box="[345,414,302,323]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="14" pageNumber="121" unit="m" value="1760.0">
<elevation id="00738E54AF705D59FEC1AF6D77F2DB74" box="[345,414,302,323]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="14" pageNumber="121" unit="m" value="1760.0">1760m</elevation>
</quantity>
,
<date id="FFE04FA7AF705D59FE32AF6D747CDB74" box="[426,528,302,323]" pageId="14" pageNumber="121" value="1991-06-12">
<collectingDate id="EFA4B64FAF705D59FE32AF6D747CDB74" box="[426,528,302,323]" pageId="14" pageNumber="121" value="1991-06-12">12.vi.1991</collectingDate>
</date>
(
<collectorName id="26AB0CB1AF705D59FD87AF6D7413DB74" box="[543,639,302,323]" pageId="14" pageNumber="121">Anderson</collectorName>
) #91-61, ex mesic hardwood litter, 1 Ƌ in
<collectionCode id="ED4FF1A2AF705D59FF10AF0B76A0DB6A" box="[136,204,328,349]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="14" pageNumber="121" type="Museum">SEMC</collectionCode>
</materialsCitation>
. –
<materialsCitation id="3B36633AAF705D59FF74AF0B773EDB41" collectingDate="1991-06-04" collectionCode="SEMC" collectorName="Anderson" country="Guatemala" elevation="1500" location="3.5 km SE La Union" pageId="14" pageNumber="121" specimenCount="1" specimenCount-female="1" stateProvince="Zacapa">
<collectingRegion id="499AA785AF705D59FF74AF0B7758DB6A" box="[236,308,328,349]" country="Guatemala" name="Zacapa" pageId="14" pageNumber="121">Zacapa</collectingRegion>
,
<location id="8E813FBCAF705D59FEA7AF0B7464DB6A" LSID="urn:lsid:plazi:treatment:03F7D871AF735D58FF0FABFE7248D812:8E813FBCAF705D59FEA7AF0B7464DB6A" box="[319,520,328,349]" country="Guatemala" name="3.5 km SE La Union" pageId="14" pageNumber="121" stateProvince="Zacapa">
<quantity id="4CA6C482AF705D59FEA7AF0B7710DB6A" box="[319,380,328,349]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="121" unit="km" value="3.5">3.5km</quantity>
SE La Union
</location>
,
<quantity id="4CA6C482AF705D59FD8CAF0B7435DB6A" box="[532,601,328,349]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="121" unit="m" value="1500.0">
<elevation id="00738E54AF705D59FD8CAF0B7435DB6A" box="[532,601,328,349]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="121" unit="m" value="1500.0">1500m</elevation>
</quantity>
,
<date id="FFE04FA7AF705D59FDFDAF0B74ACDB6A" box="[613,704,328,349]" pageId="14" pageNumber="121" value="1991-06-04">
<collectingDate id="EFA4B64FAF705D59FDFDAF0B74ACDB6A" box="[613,704,328,349]" pageId="14" pageNumber="121" value="1991-06-04">4.vi.1991</collectingDate>
</date>
(
<collectorName id="26AB0CB1AF705D59FD56AF0B7542DB6A" box="[718,814,328,349]" pageId="14" pageNumber="121">Anderson</collectorName>
) #91-50, ex cloud forest litter,
<specimenCount id="9D58A2EEAF705D59FF5BAF227687DB42" box="[195,235,351,374]" pageId="14" pageNumber="121" type="female">1♀</specimenCount>
in
<collectionCode id="ED4FF1A2AF705D59FE96AF22773EDB41" box="[270,338,353,374]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="14" pageNumber="121" type="Museum">SEMC</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B36633AAF705D59FEC6AF227721DBB8" collectingDate="1993-06-23" collectingDateMax="1993-06-25" collectingDateMin="1993-06-23" collectionCode="MHNG" collectorName="Brooks &amp; Ashe" country="Guatemala" elevation="1500" location="3.5 km SE La Union" pageId="14" pageNumber="121" specimenCount="1" specimenCount-female="1" stateProvince="Zacapa">
same data, but
<date id="FFE04FA7AF705D59FE6DAF227410DB41" box="[501,636,353,374]" pageId="14" pageNumber="121" value="1993-06-23" valueMax="1993-06-25" valueMin="1993-06-23">
<collectingDate id="EFA4B64FAF705D59FE6DAF227410DB41" box="[501,636,353,374]" pageId="14" pageNumber="121" value="1993-06-23" valueMax="1993-06-25" valueMin="1993-06-23">23-25.vi.1993</collectingDate>
</date>
(
<collectorName id="26AB0CB1AF705D59FD13AF2274BEDB41" box="[651,722,353,374]" pageId="14" pageNumber="121">Brooks</collectorName>
&amp;
<collectorName id="26AB0CB1AF705D59FD6BAF22754ADB41" box="[755,806,353,374]" pageId="14" pageNumber="121">Ashe</collectorName>
) #103,
<collectingMethod id="521F1170AF705D59FCEBAF2276DCDBB8" pageId="14" pageNumber="121">ex flight intercept trap</collectingMethod>
,
<specimenCount id="9D58A2EEAF705D59FF22AF39768DDBB9" box="[186,225,376,399]" pageId="14" pageNumber="121" type="female">1 ♀</specimenCount>
in
<collectionCode id="ED4FF1A2AF705D59FE99AF397721DBB8" box="[257,333,378,399]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="14" pageNumber="121" type="Museum">MHNG</collectionCode>
</materialsCitation>
. –
<materialsCitation id="3B36633AAF705D59FEF1AF39746ADBF6" collectingDate="1994-06-20" collectionCode="MHNG, SEMC" collectorName="Brooks &amp; Ashe" country="Honduras" elevation="1800" latitude="14.95" location="Blanca" longLatPrecision="1286" longitude="-88.083336" municipality="Mt." pageId="14" pageNumber="121" specimenCount="2" specimenCount-female="2" stateProvince="Santa Barbara">
<collectingCountry id="F34929F7AF705D59FEF1AF39779DDBB8" box="[361,497,378,399]" name="Honduras" pageId="14" pageNumber="121">HONDURAS</collectingCountry>
,
<collectingRegion id="499AA785AF705D59FE63AF3974E9DBB8" box="[507,645,378,399]" country="Honduras" name="Santa Barbara" pageId="14" pageNumber="121">Santa Barbara</collectingRegion>
,
<collectingMunicipality id="6B85F31DAF705D59FD08AF3974DFDBB8" box="[656,691,378,399]" pageId="14" pageNumber="121">Mt.</collectingMunicipality>
<collectingRegion id="499AA785AF705D59FD20AF39752EDBB8" box="[696,834,378,399]" country="Honduras" name="Santa Barbara" pageId="14" pageNumber="121">Santa Barbara</collectingRegion>
,
<quantity id="4CA6C482AF705D59FCD5AF3975F9DBB8" box="[845,917,378,399]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="14" pageNumber="121" unit="km" value="11.5">11.5km</quantity>
S &amp;
<locationDeviation id="9A537045AF705D59FC51AF3976C0DB9F" location="Pen" pageId="14" pageNumber="121">
<quantity id="4CA6C482AF705D59FC51AF397269DBB8" box="[969,1029,378,399]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="14" pageNumber="121" unit="km" value="5.6">5.6km</quantity>
W Peñ
</locationDeviation>
a
<location id="8E813FBCAF705D59FF27AFD0776EDB9F" LSID="urn:lsid:plazi:treatment:03F7D871AF735D58FF0FABFE7248D812:8E813FBCAF705D59FF27AFD0776EDB9F" box="[191,258,403,424]" country="Honduras" latitude="14.95" longLatPrecision="1286" longitude="-88.083336" municipality="Mt." name="Blanca" pageId="14" pageNumber="121" stateProvince="Santa Barbara">Blanca</location>
&lt;
<geoCoordinate id="EE6A0FA0AF705D59FE8DAFD0770ADB9F" box="[277,358,403,424]" degrees="14" direction="north" minutes="57" orientation="latitude" pageId="14" pageNumber="121" precision="925" value="14.95">14°57'N</geoCoordinate>
;
<geoCoordinate id="EE6A0FA0AF705D59FEF7AFD077A8DB9F" box="[367,452,403,424]" degrees="88" direction="west" minutes="05" orientation="longitude" pageId="14" pageNumber="121" precision="925" value="-88.083336">88°05'W</geoCoordinate>
&gt;
<quantity id="4CA6C482AF705D59FE4FAFD07470DB9F" box="[471,540,403,424]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="14" pageNumber="121" unit="m" value="1800.0">
<elevation id="00738E54AF705D59FE4FAFD07470DB9F" box="[471,540,403,424]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="14" pageNumber="121" unit="m" value="1800.0">1800m</elevation>
</quantity>
,
<date id="FFE04FA7AF705D59FDBDAFD074E0DB9F" box="[549,652,403,424]" pageId="14" pageNumber="121" value="1994-06-20">
<collectingDate id="EFA4B64FAF705D59FDBDAFD074E0DB9F" box="[549,652,403,424]" pageId="14" pageNumber="121" value="1994-06-20">20.vi.1994</collectingDate>
</date>
(
<collectorName id="26AB0CB1AF705D59FD0FAFD074B2DB9F" box="[663,734,403,424]" pageId="14" pageNumber="121">Brooks</collectorName>
&amp;
<collectorName id="26AB0CB1AF705D59FD62AFD07541DB9F" box="[762,813,403,424]" pageId="14" pageNumber="121">Ashe</collectorName>
) #163, ex decaying slash, 2 ƋƋ and
<specimenCount id="9D58A2EEAF705D59FF69AFEF7747DBF7" box="[241,299,426,449]" pageId="14" pageNumber="121" type="female">2 ♀♀</specimenCount>
in
<collectionCode id="ED4FF1A2AF705D59FED2AFEF77FFDBF6" box="[330,403,428,449]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="14" pageNumber="121" type="Museum">MHNG</collectionCode>
and
<collectionCode id="ED4FF1A2AF705D59FE5AAFEF746ADBF6" box="[450,518,428,449]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="14" pageNumber="121" type="Museum">SEMC</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B36633AAF705D59FD88AFEF76A0DBED" collectingDate="1994-06-20" collectionCode="SEMC" collectorName="Brooks &amp; Ashe" country="Honduras" elevation="1800" latitude="14.95" location="Blanca" longLatPrecision="1286" longitude="-88.083336" municipality="Mt." pageId="14" pageNumber="121" specimenCount="2" specimenCount-female="2" stateProvince="Santa Barbara">
same data, but #164, ex treefall litter, 1 Ƌ and
<specimenCount id="9D58A2EEAF705D59FC48AFEF7266DBF7" box="[976,1034,426,449]" pageId="14" pageNumber="121" type="female">2 ♀♀</specimenCount>
in
<collectionCode id="ED4FF1A2AF705D59FF10AF8676A0DBED" box="[136,204,453,474]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="14" pageNumber="121" type="Museum">SEMC</collectionCode>
</materialsCitation>
. –
<materialsCitation id="3B36633AAF705D59FF61AF8677E2DBC4" collectingDate="1971-11-20" collectionCode="FMNH" collectorName="Stockwell" country="Nicaragua" elevation="1400" latitude="13.033334" location="El Cerro Cimborazo" longLatPrecision="1291" longitude="-85.933334" pageId="14" pageNumber="121" specimenCount="1">
<collectingCountry id="F34929F7AF705D59FF61AF8677E0DBED" box="[249,396,453,474]" name="Nicaragua" pageId="14" pageNumber="121">NICARAGUA</collectingCountry>
,
<location id="8E813FBCAF705D59FE06AF86741EDBED" LSID="urn:lsid:plazi:treatment:03F7D871AF735D58FF0FABFE7248D812:8E813FBCAF705D59FE06AF86741EDBED" box="[414,626,453,474]" country="Nicaragua" latitude="13.033334" longLatPrecision="1291" longitude="-85.933334" name="El Cerro Cimborazo" pageId="14" pageNumber="121">El Cerro Cimborazo</location>
&lt;
<geoCoordinate id="EE6A0FA0AF705D59FD17AF86748DDBED" box="[655,737,453,474]" degrees="13" direction="north" minutes="02" orientation="latitude" pageId="14" pageNumber="121" precision="925" value="13.033334">13°02'N</geoCoordinate>
;
<geoCoordinate id="EE6A0FA0AF705D59FD6BAF86752BDBED" box="[755,839,453,474]" degrees="85" direction="west" minutes="56" orientation="longitude" pageId="14" pageNumber="121" precision="925" value="-85.933334">85°56'W</geoCoordinate>
&gt;
<quantity id="4CA6C482AF705D59FCFCAF8675C5DBED" box="[868,937,453,474]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="121" unit="m" value="1400.0">
<elevation id="00738E54AF705D59FCFCAF8675C5DBED" box="[868,937,453,474]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="121" unit="m" value="1400.0">1400m</elevation>
</quantity>
,
<date id="FFE04FA7AF705D59FC24AF86724EDBED" box="[956,1058,453,474]" pageId="14" pageNumber="121" value="1971-11-20">
<collectingDate id="EFA4B64FAF705D59FC24AF86724EDBED" box="[956,1058,453,474]" pageId="14" pageNumber="121" value="1971-11-20">20.xi.1971</collectingDate>
</date>
(
<collectorName id="26AB0CB1AF705D59FF17AF9D769EDBC4" box="[143,242,478,499]" pageId="14" pageNumber="121">Stockwell</collectorName>
) 1 Ƌ in
<collectionCode id="ED4FF1A2AF705D59FEDDAF9D77E2DBC4" box="[325,398,478,499]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="14" pageNumber="121" type="Museum">FMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3443AECAF705D59FF4CAC4377D6DC4A" pageId="14" pageNumber="121" type="description">
<paragraph id="8BE16967AF705D59FF4CAC437791D970" blockId="14.[136,1059,512,1661]" pageId="14" pageNumber="121">
DESCRIPTION: Habitus as in Fig. 6. Combined length of pronotum and elytra =
<quantity id="4CA6C482AF705D59FF10AC5D776BD802" box="[136,263,542,565]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="14" pageNumber="121" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4-1.5 mm</quantity>
; maximal pronotal width = 1.0-
<quantity id="4CA6C482AF705D59FDC1AC5D74C1D802" box="[601,685,542,565]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="14" pageNumber="121" unit="mm" value="1.1">1.1 mm</quantity>
. Body dark brown wih appendages slightly paler; antennomeres 10-11 paler than antennomeres 1-9. Dorsal pubescence denser on head and pronotum than on elytral disc, becoming denser along medial groove of pronotum and on anterior portion of elytral disc; frontal setae directed forward; elytral and pronotal setae slightly arcuate, recumbent; metasternal pubescence becoming denser anteriorly, longer than that of prosternum; pubescence on abdominal tergites parallel, uniform; that on sternites IV-VII uniform. Frons, pronotum, lateral portion of elytral disc and anterior portion of prohypomera granulate; frontal granulation conspicuous, with granula about as high as their diameter, or higher; metasternum coarsely granulofossulate.
</paragraph>
<paragraph id="8BE16967AF705D59FF4CAD0D76B8D9E8" blockId="14.[136,1059,512,1661]" pageId="14" pageNumber="121">Frons forming above clypeus a sharp ridge, the latter finely evenly carinate, weakly arcuate in middle and laterally oblique in dorsal view; mesal portion of disc strongly evenly convex in lateral view; U-shaped frontal impression shallow. Temples strongly convex in dorsal view. Antennae (Fig. 182) 2.1-2.3 times longer than pronotum.</paragraph>
<paragraph id="8BE16967AF705D59FF4CADA475C4DEA0" blockId="14.[136,1059,512,1661]" pageId="14" pageNumber="121">Pronotum (Fig. 186) with center strongly convex in frontal view; disc deeply depressed near middle of lateral edges, shallowly depressed along anterior margin, posterior margin and posterior portion of medial groove; the latter slightly arcuate in lateral view, deep, parallel-sided; hypomera ridged from anterior margin to laterobasal angle, disc without pit. Prosternal medial ridge absent. Scutellum with anterior margin rounded and posterior margin slighly arcuate toward acutely angular apex.</paragraph>
<paragraph id="8BE16967AF705D59FF4CAADD7510DEC5" blockId="14.[136,1059,512,1661]" pageId="14" pageNumber="121">Elytra gradually widened (Fig. 6); humeral callus low, moderately convex; disc with low swellings, moderately depressed posteriorly along lateral edge; the latter finely carinate, indistinctly denticulate, slightly arcuate in dorsal view.</paragraph>
<paragraph id="8BE16967AF705D59FF4CAAB97452DF18" blockId="14.[136,1059,512,1661]" pageId="14" pageNumber="121">Abdominal sternites II and III with medial processes as in Fig. 184, posterior portion of process of sternite III straight.</paragraph>
<paragraph id="8BE16967AF705D59FF4CAB7574F1DC15" blockId="14.[136,1059,512,1661]" pageId="14" pageNumber="121">Male: Frontoclypeal area not modified. Protarsomere 1 lacking tenent setae. Mesofemora (Fig. 175) as long as metafemora. Mesotibiae (Fig. 174) shorter than metatibiae (Fig. 173). Metatarsomeres 1 about as long as combined length of metatarsomeres 2-4. Peg-like setae arranged in a single row on mesotrochanters (Fig. 175), grouped in a field on mesotibiae and metatibiae, and absent from protrochanters, profemora, protibiae, mesofemora, metatrochanters and metafemora. Apex of abdominal tergite VIII as in Figs 176, 178. Sternite VIII as in Fig. 177. Sternite IX lacking subbasal protuberance. Aedeagus as in Figs 171-172.</paragraph>
<paragraph id="8BE16967AF705D59FF4CA86977D6DC4A" blockId="14.[136,1059,512,1661]" pageId="14" pageNumber="121">Female: Abdominal tergite VIII (Figs 185, 187) bearing medioapical projection. Sternite VIII as in Fig. 183. Genital segments as in Figs 179-181; gonocoxal plate lacking a mediodorsal ridge.</paragraph>
</subSubSection>
<subSubSection id="C3443AECAF715D58FF7BAEA0771DDB4D" pageId="15" pageNumber="122" type="distribution">
<paragraph id="8BE16967AF715D58FF7BAEA0771DDB4D" blockId="15.[151,1074,227,378]" pageId="15" pageNumber="122">
DISTRIBUTION AND NATURAL HISTORY: The species is known from
<collectingCountry id="F34929F7AF715D58FC2BAEA07242DACD" box="[947,1070,227,250]" name="Costa Rica" pageId="15" pageNumber="122">Costa Rica</collectingCountry>
,
<collectingCountry id="F34929F7AF715D58FF0FAF407761DB2D" box="[151,269,259,282]" name="Guatemala" pageId="15" pageNumber="122">Guatemala</collectingCountry>
,
<collectingCountry id="F34929F7AF715D58FE84AF4077E8DB2D" box="[284,388,259,282]" name="Honduras" pageId="15" pageNumber="122">Honduras</collectingCountry>
and
<collectingCountry id="F34929F7AF715D58FE26AF407443DB2D" box="[446,559,259,282]" name="Nicaragua" pageId="15" pageNumber="122">Nicaragua</collectingCountry>
. It was collected by sifting leaf litter or using flight intercept and faeces traps in mesic hardwood forests and cloud forests at elevations ranging between 1300 and
<quantity id="4CA6C482AF715D58FDB2AF00741BDB6D" box="[554,631,323,346]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="15" pageNumber="122" unit="m" value="3100.0">3100m</quantity>
. It is the most widespread
<taxonomicName id="4C5E12E4AF715D58FC2EAF00725DDB6D" authorityName="Curtis" authorityYear="1829" box="[950,1073,323,346]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="122" phylum="Arthropoda" rank="genus">Megarthrus</taxonomicName>
treated in this study.
</paragraph>
</subSubSection>
<subSubSection id="C3443AECAF715D58FF7BAFCD7248D812" pageId="15" pageNumber="122" type="discussion">
<paragraph id="8BE16967AF715D58FF7BAFCD7248D812" blockId="15.[151,1074,398,549]" pageId="15" pageNumber="122">
COMMENTS: Among Neotropical
<taxonomicName id="4C5E12E4AF715D58FDC3AFCD74B5DB92" authorityName="Curtis" authorityYear="1829" box="[603,729,398,421]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="B92AB575AF715D58FDC3AFCD74B5DB92" box="[603,729,398,421]" italics="true" pageId="15" pageNumber="122">Megarthrus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5E12E4AF715D58FD68AFCD75E9DB92" authorityName="Sharp. These" authorityYear="1887" box="[752,901,398,421]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="122" phylum="Arthropoda" rank="species" species="zunilensis">
<emphasis id="B92AB575AF715D58FD68AFCD75E9DB92" box="[752,901,398,421]" italics="true" pageId="15" pageNumber="122">M. zunilensis</emphasis>
</taxonomicName>
is particularly notable by the gonocoxal plate lacking a mediodorsal ridge in females, in combination with the frontal setae directed forward and the eleventh antennomere ovoid. The latter two characters are shared only with
<taxonomicName id="4C5E12E4AF715D58FD8FAFAD74A4D832" authorityName="Bierig" authorityYear="1940" box="[535,712,494,517]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="122" phylum="Arthropoda" rank="species" species="flavosignatus">
<emphasis id="B92AB575AF715D58FD8FAFAD74A4D832" box="[535,712,494,517]" italics="true" pageId="15" pageNumber="122">M. flavosignatus</emphasis>
</taxonomicName>
, of which only the male is known and which has a differently shaped aedeagus. See comments under
<taxonomicName id="4C5E12E4AF715D58FCF3AC4D7271D812" authorityName="Bierig" authorityYear="1940" box="[875,1053,526,549]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="122" phylum="Arthropoda" rank="species" species="flavosignatus">
<emphasis id="B92AB575AF715D58FCF3AC4D7271D812" box="[875,1053,526,549]" italics="true" pageId="15" pageNumber="122">M. flavosignatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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