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<document id="D7FBE7B4D82724D9B7F8B5B1650151B8" ID-DOI="10.1046/j.1439-0469.2001.00155.x" ID-GBIF-Dataset="95b3d62f-8fcc-4088-873f-b33f4a63d3fc" ID-Zenodo-Dep="2360323" IM.bibliography_requiresApprovalFor="plazi" IM.metadata_requiresApprovalFor="plazi" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1545431078479" checkinUser="PlaziZenodoSync" docAuthor="R. G. Beutel &amp; S. N. Gorb" docDate="2001" docId="0394A3592F668E1E017EFA02FE41FCCE" docLanguage="en" docName="j.zool.syst.evol.res.j.1439-0469.2001.00155.x.pdf" docOrigin="J. Zool. Syst. Evol. Research 39" docStyle="DocumentStyle{}" docTitle="Plecoptera" docType="treatment" docVersion="5" lastPageNumber="179" masterDocId="FFADDB212F648E1D022AFFC1FF81FFBA" masterDocTitle="Ultrastructure of attachment specializations of hexapods (Arthropoda): evolutionary patterns inferred from a revised ordinal phylogeny" masterLastPageNumber="207" masterPageNumber="177" originalUpdateDomain="Boston" originalUpdateTime="1545061171653" originalUpdateUser="PlaziZenodoSync" pageNumber="179" updateTime="1698553195662" updateUser="plazi">
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<mods:title id="FB28B58B6E2B36F4A0059097EE87D6C8">Ultrastructure of attachment specializations of hexapods (Arthropoda): evolutionary patterns inferred from a revised ordinal phylogeny</mods:title>
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<mods:namePart id="74727582304893EE8F0302115DC5E07C">R. G. Beutel</mods:namePart>
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<mods:namePart id="04914BCE082DE1597269DC84E4C4D2FA">S. N. Gorb</mods:namePart>
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8 
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<paragraph id="8B82124F2F668E1E017EFA20FE41FCCE" blockId="2.[852,1489,1475,1556]" lastBlockId="3.[144,782,804,884]" lastPageId="3" lastPageNumber="180" pageId="2" pageNumber="179">
The two proximal tarsomeres of the three­segmented tarsus bear euplantulae in the larger species (
<bibRefCitation id="EFAC6FBE2F668E1F0738FA3FFA16F9AE" author="Zwick, P." box="[1298,1431,1534,1556]" journalOrPublisher="Handb. Zool. IV. Insecta. Inst." pageId="2" pageNumber="179" pagination="1 - 115" part="26" refId="ref18396" refString="Zwick, P., 1980: Plecoptera (Sternfliegen). Handb. Zool. IV. Insecta. Inst. 26, 1 - 115." title="Plecoptera (Sternfliegen)" type="journal article" year="1980">Zwick 1980</bibRefCitation>
). An arolium is present between the strong, paired claws (
<figureCitation id="13060ECA2F678E1E009CFCE5FC80FC80" box="[694,769,804,826]" captionStart="Fig. 3" captionStartId="3.[1132,1165,405,424]" captionTargetBox="[149,976,192,769]" captionTargetId="figure@3.[144,979,190,771]" captionTargetPageId="3" captionText="Fig. 3. Diversity of the leg attachment devices (gray­colored areas) in hexapods. (a) Arolium (smooth). (b) Pulvilli (smooth or hairy). (c) Empodial pulvillus (ep) (hairy). (d) Hairy adhesive soles of tarsomeres. (e) Eversible pretarsal bladder (smooth). (f) Eversible structure between tibia and tarsus (smooth). (g) Fossula spongiosa (hairy). (h) Euplantulae (eu) and claw pad (cp) (both smooth). (i) Tarsal thorns transformed into adhesive structures (th), claw pad (cp) (both smooth). (j) Adhesive claw setae" httpUri="https://zenodo.org/record/2360333/files/figure.png" pageId="3" pageNumber="180">Fig. 3a</figureCitation>
). The arolium cuticle bears chitinous threads oriented at an angle to the surface (
<figureCitation id="13060ECA2F678E1E0340FC9FFE32FCCE" box="[362,435,862,884]" captionStart="Fig. 4" captionStartId="4.[182,215,1663,1682]" captionTargetBox="[220,1453,190,1630]" captionTargetId="figure@4.[220,1453,190,1630]" captionTargetPageId="4" captionText="Fig. 4. Details of hexapod tarsi and pretarsi. (A) Lepisma saccharina (Zygentoma), foreleg, pretarsus, lateral aspect. (B) Paraleptophlebia sp. (Ephemeroptera), midleg, pretarsus, ventral aspect. (C) Blatella germanica (Blattodea), midleg, tarsus, ventro­lateral aspect. (D) Omocestus viridulus (Caelifera), foreleg, tarsus, ventral aspect. (E) Cercopis vulnerata (Auchenorrhyncha), midleg, pretarsus, ventro­lateral aspect. (F) Chrysoperla carnea (Neuroptera), pretarsus, ventral aspect. (G, H) Schistocerca gregaria (Caelifera), foreleg, arolium sagittal section, triple stain after Cason. (I) Brachyptera risi (Plecoptera), midleg, arolium, sagittal section, triple stain after Cason. (J) Cercopis vulnerata (Auchenorrhyncha), midleg, sagittal section, triple stain after Cason. Black arrows indicate ventral directions. AR, arolium; CP, claw pad; EU, euplantulae; RD, chitinous threads; TAR, tarsal segments; TI, tibia; UN, claw, UN1, ephemeropteran hook­like claw, UN2, ephemeropteran pad­bearing claw" httpUri="https://zenodo.org/record/2360341/files/figure.png" pageId="3" pageNumber="180">Fig. 4I</figureCitation>
).
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