175 lines
18 KiB
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175 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.73.837" ID-GBIF-Dataset="aa5436d3-f5df-447b-a4d1-ca85c4f9eb24" ID-PMC="PMC3088455" ID-Pensoft-Pub="1313-2970-73-1" ID-PubMed="21594049" ModsDocAuthor="" ModsDocDate="2010" ModsDocID="1313-2970-73-1" ModsDocOrigin="ZooKeys 73" ModsDocTitle="A new species of Heser Tuneva, 2005 (Araneae, Gnaphosidae) from the south of India" checkinTime="1451250859721" checkinUser="pensoft" docAuthor="Bosselaers, Jan" docDate="2010" docId="E20013018701335DF06C50B999EA6BEB" docLanguage="en" docName="ZooKeys 73: 1-11" docOrigin="ZooKeys 73" docSource="http://dx.doi.org/10.3897/zookeys.73.837" docTitle="Heser vijayanagara Bosselaers, 2010, sp. n." docType="treatment" docVersion="5" lastPageNumber="7" masterDocId="FFEC2A1FFFA2FFF4FFB0FFE0E5063612" masterDocTitle="A new species of Heser Tuneva, 2005 (Araneae, Gnaphosidae) from the south of India" masterLastPageNumber="11" masterPageNumber="1" pageNumber="3" updateTime="1668165349070" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new species of Heser Tuneva, 2005 (Araneae, Gnaphosidae) from the south of India</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bosselaers, Jan</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="volume">
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<mods:number>73</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>11</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.73.837</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.73.837</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-73-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159360951" LSID="urn:lsid:zoobank.org:act:4CD46D69-8560-49BD-B58F-825F96510C14" httpUri="http://treatment.plazi.org/id/E20013018701335DF06C50B999EA6BEB" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3">
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<subSubSection pageId="2" pageNumber="3" type="nomenclature">
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<paragraph pageId="2" pageNumber="3">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:4CD46D69-8560-49BD-B58F-825F96510C14" class="Arachnida" family="Gnaphosidae" genus="Heser" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heser vijayanagara" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="vijayanagara">Heser vijayanagara</taxonomicName>
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<taxonomicNameLabel pageId="2" pageNumber="3">sp. n.</taxonomicNameLabel>
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Figs 1-8, 10
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<normalizedToken originalValue="–1112–">-1112-</normalizedToken>
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19
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="type material">
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<paragraph pageId="2" pageNumber="3">Type material.</paragraph>
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<paragraph pageId="2" pageNumber="3">Holotype male:India, Karnataka, Hospet, Hampi World Heritage Site, 15°18'27.7"N 76°28'32.8"E, alt. 455 m, under stone close to archaeological office, 25 November 2006, J. Bosselaers leg. (RBINS). Allotype female, same data (RBINS).</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="diagnosis">
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<paragraph pageId="2" pageNumber="3">Diagnosis.</paragraph>
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<paragraph pageId="2" pageNumber="3">
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The species can be distinguished from the other three species of
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Heser" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heser" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Heser</taxonomicName>
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by the abdominal dorsal scutum in males, the pronounced S-shaped curve of the sperm duct, the small, transversally oriented median apophysis and the hook-shaped embolus tip circling the broad membranous conductor in the male palp, as well as the narrow anterior epigynal hood, the large spermathecae and the circular insemination ducts in females.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="entymology">
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<paragraph pageId="2" pageNumber="3">Entymology.</paragraph>
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<paragraph pageId="2" pageNumber="3">The species epithet is a noun in apposition and refers to the imperial city of Vijayanagara (Hampi, Karnataka, India) among whose ruins the type specimens of the new species were found (Fig. 9).</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
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<paragraph pageId="2" pageNumber="3">Description.</paragraph>
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<paragraph lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
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Male (holotype). Total length: 5.00. Carapace length: 2.24; width: 1.63. Carapace orange brown, unicolorous, with a deep and narrow fovea in the posterior half (Fig. 1). Eight eyes in two rows, ringed with black, AER width 0.47, straight from above, slightly procurved from front, PER width 0.56, procurved from above, strongly procurved from front. MOQ depth 0.35, anterior width 0.29, posterior width 0.32. Eyes of AER subequal, AME grey and circular, separated by half their diameter, ALE oval, pearly white, touching AME. PME oval to subtriangular, pearly white, touching, larger than AME (Fig. 1). PLE subquadratic, pearly white, slightly smaller than ALE, separated from PME by half the PLE diameter. Clypeus vertical, slightly larger than diameter of AME. Chilum small, sclerotised, subtriangular and single, orange-brown. Chelicerae brown, with a few scattered thin setae on anterior face, anterior cheliceral rim with two very small teeth close to fang base and three larger teeth further from fang base, posterior cheliceral rim with one very small tooth close to fang base end two medium-sized teeth further from fang base. Sternum smooth, orange-brown, shield-shaped with a thin border, length 1.32; width 1.08. No precoxal triangles (
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<bibRefCitation author="Bosselaers, J" journalOrPublisher="Zoologica Scripta" pageId="6" pageNumber="7" pagination="241 - 270" title="Studies in Corinnidae: cladistic analysis of 38 corinnid and liocranid genera, and transfer of Phrurolithinae." volume="31" year="2002">
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Bosselaers and
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<normalizedToken originalValue="Jocqué">Jocque</normalizedToken>
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2002
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</bibRefCitation>
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: 247, fig. 1K,
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<bibRefCitation author="Penniman, A" journalOrPublisher="Columbus: The Ohio State University" pageId="6" pageNumber="7" title="Revision of the britcheri and pugnata groups of Scotinella (Araneae, Corinnidae, Phrurolithinae) with a reclassification of phrurolithine spiders" year="1985">Penniman 1985</bibRefCitation>
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: 16) or intercoxal sclerites (
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<bibRefCitation author="Bosselaers, J" journalOrPublisher="Zoologica Scripta" pageId="6" pageNumber="7" pagination="241 - 270" title="Studies in Corinnidae: cladistic analysis of 38 corinnid and liocranid genera, and transfer of Phrurolithinae." volume="31" year="2002">
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Bosselaers and
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<normalizedToken originalValue="Jocqué">Jocque</normalizedToken>
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2002
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</bibRefCitation>
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: 247), pleural bars (
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<bibRefCitation author="Bosselaers, J" journalOrPublisher="Zoologica Scripta" pageId="6" pageNumber="7" pagination="241 - 270" title="Studies in Corinnidae: cladistic analysis of 38 corinnid and liocranid genera, and transfer of Phrurolithinae." volume="31" year="2002">
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Bosselaers and
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<normalizedToken originalValue="Jocqué">Jocque</normalizedToken>
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2002
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</bibRefCitation>
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: 247, fig. 1P;
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<bibRefCitation author="Simon, E" journalOrPublisher="Roret, Paris" pageId="7" pageNumber="8" title="Histoire Naturelle des Araignees. Tome 1" year="1892">Simon 1892</bibRefCitation>
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: 11, fig. 29) yellow-brown, weakly sclerotised and isolated, not protruding between coxae. Labium brown, longer than wide, with a slightly thickened anterior rim. Endites longer than labium, dumbbell-shaped with oblique depression, and provided with an apical hair tuft (Fig. 2). Abdomen mottled grey dorsally, with a
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<pageBreakToken pageId="3" pageNumber="4" start="start">frontal</pageBreakToken>
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row of curved hairs and a small, shiny, brown anterior do scutum covering less than 10% of abdominal do surface area. (Fig. 1). Lateral and ventral sides of abdomen creamy white (Figs 1, 2). ALS and PLS large and cylindrical, PMS thin and slender, shorter than PLS. Legs yellow-brown, unicolorous (Figs 1, 2). No trochanter notch, no retrocoxal hymen (
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<bibRefCitation pageId="3" pageNumber="4">Raven 1998</bibRefCitation>
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,
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<bibRefCitation author="Bosselaers, J" journalOrPublisher="Zoologica Scripta" pageId="6" pageNumber="7" pagination="241 - 270" title="Studies in Corinnidae: cladistic analysis of 38 corinnid and liocranid genera, and transfer of Phrurolithinae." volume="31" year="2002">
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Bosselaers and
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<normalizedToken originalValue="Jocqué">Jocque</normalizedToken>
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2002
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</bibRefCitation>
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), patellar indentation
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<pageBreakToken pageId="4" pageNumber="5" start="start">(</pageBreakToken>
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<bibRefCitation author="Simon, E" journalOrPublisher="Roret, Paris" pageId="7" pageNumber="8" title="Histoire Naturelle des Araignees. Tome 1" year="1892">Simon 1892</bibRefCitation>
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: 22,
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<bibRefCitation author="Ledoux, JC" journalOrPublisher="J. - C. Ledoux, imprimeur & editeur, Aramon" pageId="6" pageNumber="7" title="Initiation a l'etude systematique des araignees" year="1991">Ledoux and Canard 1991</bibRefCitation>
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: fig. 15A-15B) long and narrow, 2/3 of pa length. Metatarsi III and IV with ventral terminal preening comb composed of stiff, black setae. Tarsi with two pectinate claws, no claw tufts. Leg formula 4123. Leg spination (Fig. 10): fe: palp do 0-1-2; I pl 0-0-1 do 1-1-0;II pl 0-0-1 do 1-1-0;III do 1-3-3; IV do 1-3-2; pa: palp pl 1-0 do 0-1; III do 0-2-0; ti: palp do 0-0-1; III pl 1-0-1 do 3-0-0 rl 1-0-1 ve 2-2-2; IV pl 1-0-1 do 1-1-0 rl 1-0-1 ve 2-2-2; mt: III pl 0-1-1 do 2-1-2 rl 0-1-1 ve 2-2-1; IV pl 0-1-1 do 2-2-2 rl 0-1-1 ve 2-2-1; ta: palp pl 1-0-0 do 1-0-0 rl 1-0-0.
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</paragraph>
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<caption pageId="4" pageNumber="5">
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<paragraph pageId="4" pageNumber="5">
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Figures 1-11.
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Heser" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heser vijayanagara" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="vijayanagara">Heser vijayanagara</taxonomicName>
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sp. n. 1 Male holotype, dorsal 2 Male holotype, ventral 3 Female allotype, dorsal 4 Female allotype, ventral 5 Male palp, prolateral 6 Male palp, ventral 7 Male palp, retrolateral 8 Epigyne, ventral 9 Pattabhirama temple in close vicinity of the locus typicus, giving a good impression of the type of terrain where the type specimens were found 10 Male leg spination diagram, legend below right 11 Female leg spination diagram. Scale bars: 1-4: 1.0; 5-8: 0.25.
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</paragraph>
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</caption>
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<caption pageId="4" pageNumber="5">
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<paragraph pageId="4" pageNumber="5">
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Figures 12-19.
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Heser" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heser vijayanagara" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="vijayanagara">Heser vijayanagara</taxonomicName>
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sp. n. 12 Female allotype, dorsal view of body 13 Male palp, ventral, with conductor (C), embolus (E), and median apophysis (MA) indicated 14 Male palp, retrolateral 15 Epigyne, ventral 16 Female metatarsi III (left) and IV, with ventral terminal preening comb 17 Female cheliceral teeth 18 Vulva, ventral 19 Vulva, dorsal. Scale bars: 12: 1.0; 13-17: 0.5; 18-19: 0.1.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="leg measurements">
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<paragraph pageId="4" pageNumber="5">Leg measurements:</paragraph>
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<paragraph pageId="4" pageNumber="5">.</paragraph>
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<paragraph pageId="4" pageNumber="5">
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<table pageId="4" pageNumber="5">
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<tr pageId="4" pageNumber="5">
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<th colspan="1" pageId="4" pageNumber="5" rowspan="1">fe</th>
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<th colspan="1" pageId="4" pageNumber="5" rowspan="1">pa</th>
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<th colspan="1" pageId="4" pageNumber="5" rowspan="1">ti</th>
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<th colspan="1" pageId="4" pageNumber="5" rowspan="1">mt</th>
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<th colspan="1" pageId="4" pageNumber="5" rowspan="1">ta</th>
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<th colspan="1" pageId="4" pageNumber="5" rowspan="1">Total</th>
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</tr>
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</table>
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</paragraph>
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<paragraph pageId="5" pageNumber="6">
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<pageBreakToken pageId="5" pageNumber="6" start="start">Male</pageBreakToken>
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palp with a slender, basally-prolaterally inserted embolus circling more than half of the tegulum, having a hook-shaped tip pointing in prolateral direction, which is curling around the broad, membranous conductor. Median apophysis small and subtriangular, oriented transversally. Sperm duct with a pronounced, S-shaped curve in basal half of tegulum. Retrolateral tibial apophysis pointed, subtriangular (Figs 5-7, 13-14).
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</paragraph>
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<paragraph pageId="5" pageNumber="6">Female (allotype). Total length: 6.31. Carapace length: 2.37; width: 1.79. Carapace as in male (Fig. 3). Eyes as in male, AER width 0.53, PER width 0.58, MOQ depth 0.39, anterior width 0.30, posterior width 0.31. PME subtriangular, somewhat smaller than in male, almost touching (Figs 3, 12). Clypeus and chilum as in male. Cheliceral teeth as in male (Fig. 17). Sternum smooth, yellow-brown with a darker margin (Fig. 4), shield-shaped with a thin border, length 1.45; width 1.13. No precoxal triangles or intercoxal sclerites, pleural bars as in male. Labium and endites as in male (Fig. 4). Abdomen pale grey dorsally, with a frontal row of curved hairs and a number of paler chevrons in posterior half, no do scutum (Fig. 3). Ventral side of abdomen pale white (Fig. 4). Legs yellow-brown, unicolorous (Figs 3, 4). No trochanter notch, no retrocoxal hymen, patellar indentation as in male. Metatarsi III and IV with ventral terminal preening comb composed of stiff, black setae (Fig. 16). Tarsi with two pectinate claws, no claw tufts. Leg formula 4123. Leg spination (Fig. 11): fe: palp do 0-1-2; I pl 0-0-1 do 1-1-0;II pl 0-0-1 do 1-1-0;III do 1-3-3; IV do 1-3-2; pa: palp pl 1-0 do 1-1; III do 0-2-0; ti: palp pl 0-1-2 do 0-0-1; III pl 1-0-1 do 3-0-0 rl 1-0-1 ve 2-2-2; IV pl 1-0-1 do 1-1-0 rl 1-0-1 ve 2-2-2; mt: III pl 0-1-1 do 2-1-2 rl 0-1-1 ve 2-2-1; IV pl 0-1-1 do 2-2-2 rl 0-1-1 ve 2-2-1; ta: palp pl 0-2-1 do 1-0-0 rl 0-1-0 ve 0-0-2.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="leg measurements">
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<paragraph pageId="5" pageNumber="6">Leg measurements:</paragraph>
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<paragraph pageId="5" pageNumber="6">.</paragraph>
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<paragraph pageId="5" pageNumber="6">
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<table pageId="5" pageNumber="6">
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<tr pageId="5" pageNumber="6">
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<th colspan="1" pageId="5" pageNumber="6" rowspan="1">fe</th>
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<th colspan="1" pageId="5" pageNumber="6" rowspan="1">pa</th>
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<th colspan="1" pageId="5" pageNumber="6" rowspan="1">ti</th>
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<th colspan="1" pageId="5" pageNumber="6" rowspan="1">mt</th>
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<th colspan="1" pageId="5" pageNumber="6" rowspan="1">ta</th>
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<th colspan="1" pageId="5" pageNumber="6" rowspan="1">Total</th>
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</tr>
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</table>
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</paragraph>
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<paragraph pageId="6" pageNumber="7">
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<pageBreakToken pageId="6" pageNumber="7" start="start">Epigyne</pageBreakToken>
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simple and poorly sclerotised, with a narrow anterior hood and showing large, oval spermathecae, as well as the connected, stout, inward directed part of the looped anterior insemination ducts. Copulatory openings small and medially situated (Figs 8, 15, 18, 19). Vulva (Figs 18, 19) with two large, thick-walled, oval, posterior spermathecae, connected to the anterior, medially situated copulatory openings by an insemination duct looped over 360°. The first, hemicircular stretch of the looped insemination duct passes dorsally behind the anterior part of the spermathecae, while the second, straight, longitudinally directed stretch carries an accessory gland (Fig. 19), and the third, stout, outward directed and ventrally situated stretch connects to the large spermathecae (Fig. 18).
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</paragraph>
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<caption pageId="6" pageNumber="7">
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<paragraph pageId="6" pageNumber="7">
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Figure 20. Distribution map of the four species of the genus
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Heser" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heser" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Heser</taxonomicName>
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.
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Heser" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heser aradensis" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="aradensis">Heser aradensis</taxonomicName>
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: blue square;
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Heser" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heser infumatus" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="infumatus">Heser infumatus</taxonomicName>
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: yellow triangle;
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Heser" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heser malefactor" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="malefactor">Heser malefactor</taxonomicName>
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: brown lozenge;
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Heser" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heser vijayanagara" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="vijayanagara">Heser vijayanagara</taxonomicName>
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sp. n.; red circle.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |