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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.403.7145" ID-GBIF-Dataset="a6fda06f-575a-4ae4-8d40-ae927a01f959" ID-PMC="PMC4023238" ID-Pensoft-Pub="1313-2970-403-15" ID-PubMed="24843265" ID-ZBK="328E01EFBF324352AD7DBE989A3D716B" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-403-15" ModsDocOrigin="ZooKeys 403" ModsDocTitle="A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)" checkinTime="1451245966222" checkinUser="pensoft" docAuthor="Yoshida, Takahiro &amp; Hirowatari, Toshiya" docDate="2014" docId="FC02408AD8270DF01FD18D9DAA4E3F3B" docLanguage="en" docName="ZooKeys 403: 15-45" docOrigin="ZooKeys 403" docSource="http://dx.doi.org/10.3897/zookeys.403.7145" docTitle="Psammoecus labyrinthicus Yoshida &amp; Hirowatari, 2014, sp. n." docType="treatment" docUuid="D11F2212-7A76-46A8-951C-B3FCC1543BCC" docUuidSource="ZooBank" docVersion="4" lastPageNumber="29" masterDocId="F8057B36FF80C436875CFFBFFFE6FF8F" masterDocTitle="A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)" masterLastPageNumber="45" masterPageNumber="15" pageNumber="28" updateTime="1668158244412" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yoshida, Takahiro</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hirowatari, Toshiya</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>403</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>15</mods:start>
<mods:end>45</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.403.7145</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.403.7145</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-403-15</mods:identifier>
<mods:identifier type="ZBK">328E01EFBF324352AD7DBE989A3D716B</mods:identifier>
<mods:identifier type="ZooBank">328E01EFBF324352AD7DBE989A3D716B</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152052611" LSID="urn:lsid:zoobank.org:act:D11F2212-7A76-46A8-951C-B3FCC1543BCC" httpUri="http://treatment.plazi.org/id/FC02408AD8270DF01FD18D9DAA4E3F3B" lastPageId="14" lastPageNumber="29" pageId="13" pageNumber="28">
<subSubSection pageId="13" pageNumber="28" type="nomenclature">
<paragraph pageId="13" pageNumber="28">
<taxonomicName LSID="http://zoobank.org/D11F2212-7A76-46A8-951C-B3FCC1543BCC" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus labyrinthicus" order="Coleoptera" pageId="13" pageNumber="28" phylum="Arthropoda" rank="species" species="labyrinthicus">
<pageBreakToken pageId="13" pageNumber="28" start="start">Psammoecus</pageBreakToken>
labyrinthicus
</taxonomicName>
<taxonomicNameLabel pageId="13" pageNumber="28">sp. n.</taxonomicNameLabel>
Figs 1E, 7and 14
<normalizedToken originalValue="AC">A-C</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="28" type="diagnosis">
<paragraph pageId="13" pageNumber="28">Diagnosis.</paragraph>
<paragraph pageId="13" pageNumber="28">
This species is closely similar to
<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus trimaculatus" order="Coleoptera" pageId="13" pageNumber="28" phylum="Arthropoda" rank="species" species="trimaculatus">Psammoecus trimaculatus</taxonomicName>
and
<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus triguttatus" order="Coleoptera" pageId="13" pageNumber="28" phylum="Arthropoda" rank="species" species="triguttatus">Psammoecus triguttatus</taxonomicName>
. However, it can be distinguished by the male genital structure, especially the shape of the parameres, and the comparatively dense punctuation of the dorsal pronotum.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="28" type="description">
<paragraph pageId="13" pageNumber="28">Description.</paragraph>
<paragraph pageId="13" pageNumber="28">Body length: 2.66-3.38 mm (n=25).</paragraph>
<paragraph pageId="13" pageNumber="28">Coloration (Fig. 1E). Head and pronotum reddish-brown. Elytra yellowish-brown with dark maculae; round ones at half, oblong ones on the posterior half to posterior 1/4 along elytral suture, darkened around end of elytra: these maculae sometimes connected each other. Antennae reddish-brown basally, 7th antennomere darkened, from 8th to 10th blackish-brown, 11th (apex) comparatively bright.</paragraph>
<paragraph pageId="13" pageNumber="28">Head (Fig. 7A, B, C). Rounded-triangular, HW/HL 1.25-2.00; IE/HL 0.75-1.27. Temples slightly expanded behind eyes, gradually narrowed. Eyes large, prominent. Dorsal surface with dense and strong punctuation. Antennae 1.35-1.56 mm, moderately long; covered with medium length semi-erect pubescence and some long erect setae on each antennomere; approximate ratio of holotype as follows: 2.4: 1.0: 1.0: 1.2: 1.3: 1.1: 1.1: 1.1: 1.1: 1.1: 1.9 (Fig. 7A).</paragraph>
<paragraph pageId="13" pageNumber="28">Pronotum (Fig. 7B, C). Roundly subquadrate, PW/PL 1.25-1.53. Relatively strong and dense punctuation on dorsal surface. Pubescence composed of short setae, a long seta on each tooth on lateral margins and anterior and posterior angles. Each anterior angle with several small teeth, each lateral margin with four short teeth; tooth I small, tooth II longer than tooth I, teeth III and IV almost the same size, longer than tooth II, teeth II, III and IV broadened at base, each posterior angle with a very small tooth, the shape of these teeth variable.</paragraph>
<paragraph pageId="13" pageNumber="28">Elytra (Fig. 7E). Elongate-oval, EW/BL 0.41-0.47. Rows of punctures narrower than interstices. Pubescence composed of numerous medium length semi-erect setae and some long setae in a row around lateral margins.</paragraph>
<paragraph pageId="13" pageNumber="28">9th abdominal sternite (Fig. 14A). Strut relatively short, cut at anterior 1/3. Lateral sclerites fused with strut, curved inwards.</paragraph>
<paragraph pageId="13" pageNumber="28">Aedeagus (Fig. 14B, C). Parameres stout, club-shaped, punctuated on anterior half of inner margins, anterolateral portions well punctuated, anterior half of inner margins with many setae of variable size, apex with a long seta. Phallobase broadened toward posterior margin, posterior margin widely incised, protuberances around 1/3 of inner margins of branches, especially right protuberance thin. Penis flat, narrowed toward apex, apex pointed, punctuated around anterior 1/8, apex densely punctuated.</paragraph>
</subSubSection>
<subSubSection lastPageId="14" lastPageNumber="29" pageId="13" pageNumber="28" type="type series">
<paragraph pageId="13" pageNumber="28">Type series.</paragraph>
<paragraph lastPageId="14" lastPageNumber="29" pageId="13" pageNumber="28">
Holotype: male,
<normalizedToken originalValue="Hachijô">Hachijo</normalizedToken>
Island,
<normalizedToken originalValue="Hachijô">Hachijo</normalizedToken>
Town, Tokyo Prefecture, Japan, 21
<normalizedToken originalValue="VII">-VII-</normalizedToken>
1957, S. Hisamatsu leg. (EUMJ). Paratypes: [Tokyo Pref.] 3 females, same data as holotype. (EUMJ). [Mie Pref.] 4 males,
<normalizedToken originalValue="Ujitachi-chô">Ujitachi-cho</normalizedToken>
, Ise City, 29
<normalizedToken originalValue="VIII">-VIII-</normalizedToken>
<pageBreakToken pageId="14" pageNumber="29" start="start">2009</pageBreakToken>
, N. Narukawa leg. (ELKU). [Nagasaki Pref.] 1 male and 11 exs., Ta,
<normalizedToken originalValue="Toyotama-chô">Toyotama-cho</normalizedToken>
, Tsushima Island, 7
<normalizedToken originalValue="VII">-VII-</normalizedToken>
2013, T. Yoshida leg. (ELKU). [Kagoshima Pref.] 2 males and 3 exs., Takarajima Island, Toshima Village, 2
<normalizedToken originalValue="VII">-VII-</normalizedToken>
1960, M.
<normalizedToken originalValue="Satô">Sato</normalizedToken>
leg. (EUMJ); 1 ex., same locality, 3
<normalizedToken originalValue="VII">-VII-</normalizedToken>
1960, M.
<normalizedToken originalValue="Satô">Sato</normalizedToken>
leg. (EUMJ); 1 ex., same locality, 1
<normalizedToken originalValue="VI">-VI-</normalizedToken>
1962, M.
<normalizedToken originalValue="Satô">Sato</normalizedToken>
leg. (EUMJ).
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="29" type="distribution">
<paragraph pageId="14" pageNumber="29">Distribution.</paragraph>
<paragraph pageId="14" pageNumber="29">
JAPAN: Honshu,
<normalizedToken originalValue="Hachijô">Hachijo</normalizedToken>
(Izu Islands), Tsushima, Takarajima Islands (Tokara Islands).
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="29" type="biological notes">
<paragraph pageId="14" pageNumber="29">Biological notes.</paragraph>
<paragraph pageId="14" pageNumber="29">The first author (Yoshida) collected many individuals of this new species from dead leaves of evergreen trees in Tsushima Island, Nagasaki Prefecture (Fig. 15A).</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="29" type="etymology">
<paragraph pageId="14" pageNumber="29">Etymology.</paragraph>
<paragraph pageId="14" pageNumber="29">
The specific name means
<normalizedToken originalValue="labyrinthine">'labyrinthine'</normalizedToken>
. Identification of
<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus trimaculatus" order="Coleoptera" pageId="14" pageNumber="29" phylum="Arthropoda" rank="species" species="trimaculatus">Psammoecus trimaculatus</taxonomicName>
and closely similar species is very difficult. The addition of this new species to this group of closely similar species made their identification more difficult like the labyrinth.
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="29" type="remarks">
<paragraph pageId="14" pageNumber="29">Remarks.</paragraph>
<paragraph pageId="14" pageNumber="29">
<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus trimaculatus" order="Coleoptera" pageId="14" pageNumber="29" phylum="Arthropoda" rank="species" species="trimaculatus">Psammoecus trimaculatus</taxonomicName>
is closely related to this species, however the former has been often collected from dead leaves of monocotyledon plant. These two species may have each distinct ecological trait.
</paragraph>
</subSubSection>
</treatment>
</document>