304 lines
33 KiB
XML
304 lines
33 KiB
XML
<document id="FEE65780A668CC922D2003DFEEF8E20B" ID-DOI="10.1649/0010-065X-72.1.129" ID-ISSN="1938-4394" ID-Zenodo-Dep="5380372" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630289587697" checkinUser="diego" docAuthor="Schawaller, Wolfgang & Steiner, Warren E." docDate="2018" docId="F709F305FFC63D34FD77FAC435BB60C1" docLanguage="en" docName="0010-065X-72.1.129.pdf" docOrigin="The Coleopterists Bulletin 72 (1)" docSource="http://dx.doi.org/10.1649/0010-065x-72.1.129" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Philhammus ambouli SCHAWALLER AND STEINER 2018, new species" docType="treatment" docUuid="12EC9BC5-B9AB-4392-B457-497DC71E2D18" docUuidSource="ZooBank" docVersion="5" lastPageNumber="132" masterDocId="0B308B7DFFC73D30FFB9FFCA354F620E" masterDocTitle="Philhammus ambouli Schawaller and Steiner (Coleoptera: Tenebrionidae: Cnemeplatiini), New Species from Djibouti" masterLastPageNumber="133" masterPageNumber="129" pageNumber="129" updateTime="1699216185810" updateUser="plazi">
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<mods:title id="28DA6C99626B3CC9CD79B6169F40094D">Philhammus ambouli Schawaller and Steiner (Coleoptera: Tenebrionidae: Cnemeplatiini), New Species from Djibouti</mods:title>
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<mods:namePart id="A14A5501F484C13D86B50B4080D2D574">Schawaller, Wolfgang</mods:namePart>
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<mods:namePart id="09E6D2101637E24237708AEACBBE80CE">Steiner, Warren E.</mods:namePart>
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<mods:title id="F3CE61D4D35B2F4240EF5721C9127F04">The Coleopterists Bulletin</mods:title>
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<mods:part id="AF4696457EE4E9BD40DBC5933FA39DEF">
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<mods:date id="7D519392510736C4AA55D070FBC70558">2018</mods:date>
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<mods:number id="CDE228BA2EF188F18D665BC8DCB7D3DB">2018-03-23</mods:number>
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<mods:number id="2527F4237A69CF09B51668691D3FA9A2">72</mods:number>
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<mods:identifier id="1472CFE7A4FEB73FB74B0A5B09ED7CDB" type="ISSN">1938-4394</mods:identifier>
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<taxonomicName id="B8A03990FFC63D31FD77FAC431D5672D" authority="Schawaller and Steiner" authorityName="SCHAWALLER AND STEINER" authorityYear="2018" box="[718,1178,1294,1315]" class="Insecta" family="Tenebrionidae" genus="Philhammus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="129" phylum="Arthropoda" rank="species" species="ambouli" status="sp. nov.">
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<emphasis id="4DD49E01FFC63D31FD77FAC436EA672D" bold="true" box="[718,933,1294,1315]" italics="true" pageId="1" pageNumber="129">Philhammus ambouli</emphasis>
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Schawaller and Steiner
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</taxonomicName>
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,
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<taxonomicNameLabel id="56E7237AFFC63D31FCC2FAE336BF6730" box="[891,1008,1321,1342]" pageId="1" pageNumber="129" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="7F1F4213FFC63D31FCB8FA8936A46780" blockId="1.[718,1181,1294,1422]" pageId="1" pageNumber="129">
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zoobank.org/
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urn:lsid:zoobank.org:act:
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(
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<figureCitation id="E79B5E96FFC63D31FC3EFAB336856780" box="[903,970,1401,1422]" captionStart="Fig" captionStartId="2.[175,206,1073,1092]" captionTargetBox="[156,1167,196,1055]" captionTargetId="figure-253@2.[154,1169,194,1057]" captionTargetPageId="2" captionText="Fig. 1. Philhammus ambouli, paratype, dorsal and ventral views. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5380374" httpUri="https://zenodo.org/record/5380374/files/figure.png" pageId="1" pageNumber="129">Figs. 1</figureCitation>
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,
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<figureCitation id="E79B5E96FFC63D31FC61FAB336AA6780" box="[984,997,1401,1422]" captionStart="Fig" captionStartId="2.[175,206,1687,1706]" captionTargetBox="[156,632,1477,1669]" captionTargetId="figure-270@2.[154,634,1475,1671]" captionTargetPageId="2" captionText="Fig. 2. Philhammus ambouli, aedeagus of holotype, lateral and dorsal views. Length 0.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5380376" httpUri="https://zenodo.org/record/5380376/files/figure.png" pageId="1" pageNumber="129">2</figureCitation>
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)
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</paragraph>
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<subSubSection id="37BA1198FFC63D32FD62FA693763669A" lastPageId="2" lastPageNumber="130" pageId="1" pageNumber="129" type="materials_examined">
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<paragraph id="7F1F4213FFC63D32FD62FA693763669A" blockId="1.[707,1194,1443,1730]" lastBlockId="2.[151,638,1124,1411]" lastPageId="2" lastPageNumber="130" pageId="1" pageNumber="129">
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<emphasis id="4DD49E01FFC63D31FD62FA6936CC67B6" bold="true" box="[731,899,1443,1464]" pageId="1" pageNumber="129">Type Material.</emphasis>
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<materialsCitation id="CFC8484EFFC63D32FC2FFA693763669A" collectingDate="2014-05-07" collectingDateMax="2014-05-13" collectingDateMin="2014-05-07" collectionCode="USNM, SMNS" collectorName="S. W. Gotte & W. E. Steiner" country="Djibouti" elevation="2" lastPageId="2" lastPageNumber="130" latitude="11.543889" location="Chabelley Airfield" longLatPrecision="21" longitude="43.16889" municipality="Ambouli River valley" pageId="1" pageNumber="129" specimenCount="73" stateProvince="Arta" typeStatus="holotype">
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<typeStatus id="A01BFCB1FFC63D31FC2FFA6936BA67B6" box="[918,1013,1443,1464]" pageId="1" pageNumber="129" type="holotype">Holotype</typeStatus>
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, dissected male (
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), and
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<specimenCount id="69A6899AFFC63D31FCDFFA7436A267DD" box="[870,1005,1470,1491]" pageId="1" pageNumber="129" type="generic" typeStatus="paratype">one paratype</specimenCount>
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(
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<collectionCode id="19B1DAD6FFC63D31FBB1FA74310467DD" box="[1032,1099,1470,1491]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="129">SMNS</collectionCode>
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) labeled “
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<collectingCountry id="07B70283FFC63D31FD74FA12367467E3" box="[717,827,1496,1517]" name="Djibouti" pageId="1" pageNumber="129">DJIBOUTI</collectingCountry>
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: Djb. Region; Camp Lemonnier, E. end of base in scrub flats,
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<quantity id="B858EFF6FFC63D31FC75FA3931476406" box="[972,1032,1523,1544]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="129" unit="km" value="5.0">5 km</quantity>
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SSE Djibouti,
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11°32
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<emphasis id="4DD49E01FFC63D31FD42F9C1364F6416" bold="true" box="[763,768,1547,1560]" italics="true" pageId="1" pageNumber="129">
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<superScript id="88D5EF5BFFC63D31FD42F9C1364F6416" attach="none" box="[763,768,1547,1560]" fontSize="6" pageId="1" pageNumber="129">′</superScript>
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</emphasis>
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38″N
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,
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43°10
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||
<emphasis id="4DD49E01FFC63D31FCC3F9C136306416" bold="true" box="[890,895,1547,1560]" italics="true" pageId="1" pageNumber="129">
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<superScript id="88D5EF5BFFC63D31FCC3F9C136306416" attach="none" box="[890,895,1547,1560]" fontSize="6" pageId="1" pageNumber="129">′</superScript>
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</emphasis>
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08″E
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,
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<elevation id="F48DA520FFC63D31FC05F9C736A8642C" box="[956,999,1549,1570]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="129" unit="m" value="2.0">2 m</elevation>
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</quantity>
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||
.
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</specimenCount>
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;
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<date id="0B1E64D3FFC63D31FC4EF9C73134642C" box="[1015,1147,1549,1570]" pageId="1" pageNumber="129" value="2014-05-13">
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<collectingDate id="1B5A9D3BFFC63D31FC4EF9C73134642C" box="[1015,1147,1549,1570]" pageId="1" pageNumber="129" value="2014-05-13">13 May 2014</collectingDate>
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||
</date>
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||
/ At black [ultraviolet] light in open
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<taxonomicName id="B8A03990FFC63D31FC4DF9E231076433" box="[1012,1096,1576,1597]" class="Magnoliopsida" family="Fabaceae" genus="Prosopis" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="129" phylum="Tracheophyta" rank="genus">
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<emphasis id="4DD49E01FFC63D31FC4DF9E231076433" box="[1012,1096,1576,1597]" italics="true" pageId="1" pageNumber="129">Prosopis</emphasis>
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</taxonomicName>
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&
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<taxonomicName id="B8A03990FFC63D31FBDEF9ED31E66433" box="[1127,1193,1575,1597]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="129" phylum="Tracheophyta" rank="genus">
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<emphasis id="4DD49E01FFC63D31FBDEF9ED31E66433" box="[1127,1193,1575,1597]" italics="true" pageId="1" pageNumber="129">Acacia</emphasis>
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</taxonomicName>
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scrub on dry sandy soil; Colls. W. E. Steiner,
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<collectorName id="D25527C5FFC63D31FBCCF98837B5647C" pageId="1" pageNumber="129">S. W. Gotte</collectorName>
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, et al.” One
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<typeStatus id="A01BFCB1FFC63D31FC25F99736A2647C" box="[924,1005,1629,1650]" pageId="1" pageNumber="129" type="paratype">paratype</typeStatus>
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(
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<collectionCode id="19B1DAD6FFC63D31FBBFF9973102647C" box="[1030,1101,1629,1650]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="129" type="Museum">USNM</collectionCode>
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) labeled “
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<collectingCountry id="07B70283FFC63D31FD74F9B236746483" box="[717,827,1656,1677]" name="Djibouti" pageId="1" pageNumber="129">DJIBOUTI</collectingCountry>
|
||
:
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||
<collectingRegion id="BD648CF1FFC63D31FCF1F9B2368B6483" box="[840,964,1656,1677]" country="Djibouti" name="Arta" pageId="1" pageNumber="129">Arta Region</collectingRegion>
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;
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||
<collectingMunicipality id="9F7BD869FFC63D31FC6BF9B231E66483" box="[978,1193,1656,1677]" pageId="1" pageNumber="129">Ambouli River valley</collectingMunicipality>
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<locationDeviation id="6EAD5B31FFC63D31FD7AF95836D264A9" box="[707,925,1682,1703]" location="Balbala" pageId="1" pageNumber="129">
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|
||
SW Balbala
|
||
</locationDeviation>
|
||
near
|
||
<location id="7A7F14C8FFC63D31FC5EF95831EA64A9" LSID="urn:lsid:plazi:treatment:F709F305FFC63D34FD77FAC435BB60C1:7A7F14C8FFC63D31FC5EF95831EA64A9" box="[999,1189,1682,1703]" country="Djibouti" latitude="11.543889" longLatPrecision="21" longitude="43.16889" municipality="Ambouli River valley" name="Chabelley Airfield" pageId="1" pageNumber="129" stateProvince="Arta">Chabelley Airfield</location>
|
||
,
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||
<geoCoordinate id="1A9424D4FFC63D31FD7AF967367C64CC" box="[707,819,1706,1730]" degrees="11" direction="north" minutes="31" orientation="latitude" pageId="1" pageNumber="129" precision="15" seconds="10" value="11.519444">
|
||
11°31
|
||
<emphasis id="4DD49E01FFC63D31FD42F960364F64B9" bold="true" box="[763,768,1706,1719]" italics="true" pageId="1" pageNumber="129">
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<superScript id="88D5EF5BFFC63D31FD42F960364F64B9" attach="none" box="[763,768,1706,1719]" fontSize="6" pageId="1" pageNumber="129">′</superScript>
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||
</emphasis>
|
||
10″N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1A9424D4FFC63D31FCF9F96736FF64CC" box="[832,944,1706,1730]" degrees="43" direction="east" minutes="04" orientation="longitude" pageId="1" pageNumber="129" precision="15" seconds="52" value="43.081112">
|
||
43°04
|
||
<emphasis id="4DD49E01FFC63D31FCC0F960363164B9" bold="true" box="[889,894,1706,1719]" italics="true" pageId="1" pageNumber="129">
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<superScript id="88D5EF5BFFC63D31FCC0F960363164B9" attach="none" box="[889,894,1706,1719]" fontSize="6" pageId="1" pageNumber="129">′</superScript>
|
||
</emphasis>
|
||
52″E
|
||
</geoCoordinate>
|
||
,
|
||
<specimenCount id="69A6899AFFC63D31FC02F96736B864CC" box="[955,1015,1709,1730]" pageId="1" pageNumber="129" type="generic">
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<quantity id="B858EFF6FFC63D31FC02F96736BD64CC" box="[955,1010,1709,1730]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="129" unit="m" value="70.0">
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||
<elevation id="F48DA520FFC63D31FC02F96736BD64CC" box="[955,1010,1709,1730]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="129" unit="m" value="70.0">70 m</elevation>
|
||
</quantity>
|
||
.
|
||
</specimenCount>
|
||
;
|
||
<date id="0B1E64D3FFC63D31FBBAF967313564CC" box="[1027,1146,1709,1730]" pageId="1" pageNumber="129" value="2014-05-07">
|
||
<collectingDate id="1B5A9D3BFFC63D31FBBAF967313564CC" box="[1027,1146,1709,1730]" pageId="1" pageNumber="129" value="2014-05-07">7 May 2014</collectingDate>
|
||
</date>
|
||
/ At black [ultraviolet] light, rocky edge of drying wadi, sparse
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||
<taxonomicName id="B8A03990FFC53D32FF63FBB43453669A" box="[218,284,1150,1172]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="4DD49E01FFC53D32FF63FBB43453669A" box="[218,284,1150,1172]" italics="true" pageId="2" pageNumber="130">Acacia</emphasis>
|
||
</taxonomicName>
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||
scrub;
|
||
<collectorName id="D25527C5FFC53D32FEDCFBB534A8669A" box="[357,487,1151,1172]" pageId="2" pageNumber="130">W. E. Steiner</collectorName>
|
||
, et al.”
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="2BDF129BFFC53D32FF16FBFB36D3664A" ID-DOI="http://doi.org/10.5281/zenodo.5380374" ID-Zenodo-Dep="5380374" box="[175,924,1072,1092]" httpUri="https://zenodo.org/record/5380374/files/figure.png" pageId="2" pageNumber="130" startId="2.[175,206,1073,1092]" targetBox="[156,1167,196,1055]" targetPageId="2">
|
||
<paragraph id="7F1F4213FFC53D32FF16FBFB36D3664A" blockId="2.[175,924,1072,1092]" box="[175,924,1072,1092]" pageId="2" pageNumber="130">
|
||
<emphasis id="4DD49E01FFC53D32FF16FBFB35A6664A" bold="true" box="[175,233,1073,1092]" pageId="2" pageNumber="130">Fig. 1.</emphasis>
|
||
<taxonomicName id="B8A03990FFC53D32FF47FBFA34FC664D" authorityName="SCHAWALLER AND STEINER" authorityYear="2018" box="[254,435,1072,1091]" class="Insecta" family="Tenebrionidae" genus="Philhammus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="130" phylum="Arthropoda" rank="species" species="ambouli">
|
||
<emphasis id="4DD49E01FFC53D32FF47FBFA34FC664D" box="[254,435,1072,1091]" italics="true" pageId="2" pageNumber="130">Philhammus ambouli</emphasis>
|
||
</taxonomicName>
|
||
, paratype, dorsal and ventral views. Scale line = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="37BA1198FFC53D33FF16FB5037DC6433" lastPageId="3" lastPageNumber="131" pageId="2" pageNumber="130" type="description">
|
||
<paragraph id="7F1F4213FFC53D33FF16FB5037DC6433" blockId="2.[151,638,1124,1411]" lastBlockId="3.[173,660,1550,1730]" lastPageId="3" lastPageNumber="131" pageId="2" pageNumber="130">
|
||
<emphasis id="4DD49E01FFC53D32FF16FB50346566A1" bold="true" box="[175,298,1178,1199]" pageId="2" pageNumber="130">Description.</emphasis>
|
||
Body length
|
||
<quantity id="B858EFF6FFC53D32FE07FB50377566A1" box="[446,570,1177,1199]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="2" pageNumber="130" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.3–2.5 mm</quantity>
|
||
. Body and all appendages unicolorous yellow-brown, all surfaces dull (
|
||
<figureCitation id="E79B5E96FFC53D32FE9CFB05342F66EA" box="[293,352,1231,1252]" captionStart="Fig" captionStartId="2.[175,206,1073,1092]" captionTargetBox="[156,1167,196,1055]" captionTargetId="figure-253@2.[154,1169,194,1057]" captionTargetPageId="2" captionText="Fig. 1. Philhammus ambouli, paratype, dorsal and ventral views. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5380374" httpUri="https://zenodo.org/record/5380374/files/figure.png" pageId="2" pageNumber="130">Fig. 1</figureCitation>
|
||
). Head evenly covered with dense but nonconfluent granulation; margin of clypeus and genae with equally upturned margins and dentate (altogether about 20 teeth); clypeus with a pair of distinct impressions; genae not surpassing eyes; eyes large, with diameter of about length of last 2 antennomeres combined. Pronotum subquadrate, lateral margins parallel, anterolateral angles rounded and not protruding, posterolateral angles distinctly dentate; surface with same dense granulation as on head; disc with deep and nearly complete, medial, longitudinal furrow. Elytra 1.4 times longer than wide, with nonstriate rows of punctures; internal row of punctures somewhat impressed, so 1
|
||
<superScript id="88D5EF5BFFC53D32FCFAFAD336006729" attach="left" box="[835,847,1305,1319]" fontSize="6" pageId="2" pageNumber="130">st</superScript>
|
||
interval slightly convex, all other intervals completely flat; punctures of rows separated from each other by more than their diameter; surface of elytra smooth, with only traces of granules; microgranules visible along suture. Prosternum not prominent. Epipleura, metasternum, and ventrites with traces of fine granulation. Anterior tibiae very short, nearly as wide as long, broadly triangular with rounded inner apical corner and outer apical angle (burrowing tooth), this polished dorsally; outer margin sinuate; inner apical spur large, as long as tarsus, curved and polished (similarly described and illustrated by
|
||
<bibRefCitation id="1B313FE2FFC53D32FCF6F99736BA647C" author="Medvedev, G. S." box="[847,1013,1629,1650]" pageId="2" pageNumber="130" pagination="561 - 563" refId="ref2415" refString="Medvedev, G. S. 2005. New species of tenebrionid beetles (Coleoptera: Tenebrionidae) from Botswana and Afghanistan. Annales Zoologici 55: 561 - 563." type="journal article" year="2005">Medvedev (2005)</bibRefCitation>
|
||
for
|
||
<taxonomicName id="B8A03990FFC53D32FBA1F996367B6483" authority="Ferrer" authorityName="Ferrer" class="Insecta" family="Tenebrionidae" genus="Philhammus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="130" phylum="Arthropoda" rank="species" species="brincki">
|
||
<emphasis id="4DD49E01FFC53D32FBA1F99637BC6483" italics="true" pageId="2" pageNumber="130">Philhammus brincki</emphasis>
|
||
Ferrer
|
||
</taxonomicName>
|
||
[as
|
||
<taxonomicName id="B8A03990FFC53D32FCE3F9B236926483" box="[858,989,1655,1677]" class="Insecta" family="Tenebrionidae" genus="Philhammus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="130" phylum="Arthropoda" rank="species" species="triplehorni">
|
||
<emphasis id="4DD49E01FFC53D32FCE3F9B236926483" box="[858,989,1655,1677]" italics="true" pageId="2" pageNumber="130">P. triplehorni</emphasis>
|
||
</taxonomicName>
|
||
]). Aedeagus (
|
||
<figureCitation id="E79B5E96FFC53D32FBE3F9B231C26483" box="[1114,1165,1656,1677]" captionStart="Fig" captionStartId="2.[175,206,1687,1706]" captionTargetBox="[156,632,1477,1669]" captionTargetId="figure-270@2.[154,634,1475,1671]" captionTargetPageId="2" captionText="Fig. 2. Philhammus ambouli, aedeagus of holotype, lateral and dorsal views. Length 0.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5380376" httpUri="https://zenodo.org/record/5380376/files/figure.png" pageId="2" pageNumber="130">Fig.2</figureCitation>
|
||
)
|
||
<quantity id="B858EFF6FFC53D32FD14F95837BA64A9" box="[685,757,1682,1703]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="130" unit="mm" value="0.6">0.6 mm</quantity>
|
||
long, slender, gently curved dorsally; fused parameres parallel-sided along base to middle in dorsal view, gradually narrowing to a rounded apex; basal piece short, half length of parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="2BDF129BFFC53D32FF16F95D34B564CC" ID-DOI="http://doi.org/10.5281/zenodo.5380376" ID-Zenodo-Dep="5380376" httpUri="https://zenodo.org/record/5380376/files/figure.png" pageId="2" pageNumber="130" startId="2.[175,206,1687,1706]" targetBox="[156,632,1477,1669]" targetPageId="2">
|
||
<paragraph id="7F1F4213FFC53D32FF16F95D34B564CC" blockId="2.[151,638,1686,1730]" pageId="2" pageNumber="130">
|
||
<emphasis id="4DD49E01FFC53D32FF16F95D35A664A4" bold="true" box="[175,233,1687,1706]" pageId="2" pageNumber="130">Fig. 2.</emphasis>
|
||
<taxonomicName id="B8A03990FFC53D32FF47F95C34FB64A7" authorityName="SCHAWALLER AND STEINER" authorityYear="2018" box="[254,436,1686,1705]" class="Insecta" family="Tenebrionidae" genus="Philhammus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="130" phylum="Arthropoda" rank="species" species="ambouli">
|
||
<emphasis id="4DD49E01FFC53D32FF47F95C34FB64A7" box="[254,436,1686,1705]" italics="true" pageId="2" pageNumber="130">Philhammus ambouli</emphasis>
|
||
</taxonomicName>
|
||
, aedeagus of holotype, lateral and dorsal views. Length 0.6 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2BDF129BFFC43D33FF7CFA72315C67EC" ID-DOI="http://doi.org/10.5281/zenodo.5380378" ID-Zenodo-Dep="5380378" httpUri="https://zenodo.org/record/5380378/files/figure.png" pageId="3" pageNumber="131" startId="3.[197,236,1464,1483]" targetBox="[177,1188,196,1446]" targetPageId="3">
|
||
<paragraph id="7F1F4213FFC43D33FF7CFA7231E667C5" blockId="3.[173,1193,1463,1506]" box="[197,1193,1463,1483]" pageId="3" pageNumber="131">
|
||
<emphasis id="4DD49E01FFC43D33FF7CFA72346F67C5" bold="true" box="[197,288,1463,1483]" pageId="3" pageNumber="131">Figs. 3–4.</emphasis>
|
||
Habitat of
|
||
<taxonomicName id="B8A03990FFC43D33FE24FA7D371767C4" authorityName="SCHAWALLER AND STEINER" authorityYear="2018" box="[413,600,1463,1482]" class="Insecta" family="Tenebrionidae" genus="Philhammus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="131" phylum="Arthropoda" rank="species" species="ambouli">
|
||
<emphasis id="4DD49E01FFC43D33FE24FA7D371767C4" box="[413,600,1463,1482]" italics="true" pageId="3" pageNumber="131">Philhammus ambouli</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="4DD49E01FFC43D33FDD0FA72373567C5" bold="true" box="[617,634,1464,1483]" pageId="3" pageNumber="131">3)</emphasis>
|
||
Type locality at Camp Lemonnier, Djibouti. Photograph by
|
||
</paragraph>
|
||
<paragraph id="7F1F4213FFC43D33FF14FA05315C67EC" blockId="3.[173,1193,1463,1506]" box="[173,1043,1487,1506]" pageId="3" pageNumber="131">
|
||
B. K. Schmidt;
|
||
<emphasis id="4DD49E01FFC43D33FE8CFA05340967EC" bold="true" box="[309,326,1487,1506]" pageId="3" pageNumber="131">4)</emphasis>
|
||
Collecting site at the Ambouli River crossing, Djibouti. Photograph by R. Russell.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="37BA1198FFC43D34FF7CF98937306044" lastPageId="4" lastPageNumber="132" pageId="3" pageNumber="131" type="diagnosis">
|
||
<paragraph id="7F1F4213FFC43D34FF7CF98937306044" blockId="3.[173,660,1550,1730]" lastBlockId="4.[151,639,194,719]" lastPageId="4" lastPageNumber="132" pageId="3" pageNumber="131">
|
||
<emphasis id="4DD49E01FFC43D33FF7CF98934636456" bold="true" box="[197,300,1603,1624]" pageId="3" pageNumber="131">Diagnosis.</emphasis>
|
||
<taxonomicName id="B8A03990FFC43D33FE8CF988374B6456" authorityName="SCHAWALLER AND STEINER" authorityYear="2018" box="[309,516,1602,1624]" class="Insecta" family="Tenebrionidae" genus="Philhammus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="131" phylum="Arthropoda" rank="species" species="ambouli">
|
||
<emphasis id="4DD49E01FFC43D33FE8CF988374B6456" box="[309,516,1602,1624]" italics="true" pageId="3" pageNumber="131">Philhammus ambouli</emphasis>
|
||
</taxonomicName>
|
||
can be recognized by the shape of the pronotum with parallel sides, distinctly marked posterolateral angles, and nearly complete, medial, longitudinal impression.
|
||
<taxonomicName id="B8A03990FFC43D33FF14F966374264CC" authority="Kaszab, 1967" authorityName="Kaszab" authorityYear="1967" box="[173,525,1708,1730]" class="Insecta" family="Tenebrionidae" genus="Philhammus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="131" phylum="Arthropoda" rank="species" species="ferenczi">
|
||
<emphasis id="4DD49E01FFC43D33FF14F966343664CC" box="[173,377,1708,1730]" italics="true" pageId="3" pageNumber="131">Philhammus ferenczi</emphasis>
|
||
<bibRefCitation id="1B313FE2FFC43D33FE3CF967374264CC" author="Kaszab, Z." box="[389,525,1709,1730]" pageId="3" pageNumber="131" pagination="291 - 296" refId="ref2322" refString="Kaszab, Z. 1967. Die Arten der Gattung Philhammus Fairm. (Coleoptera, Tenebrionidae). Annales Historiconaturalis Musei Nationalis Hungarici 59: 291 - 296." type="journal article" year="1967">Kaszab, 1967</bibRefCitation>
|
||
</taxonomicName>
|
||
from central Africa is the only other known species of the genus with distinct posterolateral angles of the pronotum, but to a lesser extent. Additionally, in
|
||
<taxonomicName id="B8A03990FFC43D33FB9FF98931C86456" authorityName="Kaszab" authorityYear="1967" box="[1062,1159,1602,1624]" class="Insecta" family="Tenebrionidae" genus="Philhammus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="131" phylum="Arthropoda" rank="species" species="ferenczi">
|
||
<emphasis id="4DD49E01FFC43D33FB9FF98931C86456" box="[1062,1159,1602,1624]" italics="true" pageId="3" pageNumber="131">P. ferenczi</emphasis>
|
||
</taxonomicName>
|
||
the punctures of the elytral rows are closely spaced, the elytral intervals are slightly convex, and the pronotal medial impression is diminished.
|
||
<taxonomicName id="B8A03990FFC43D33FB8BF958368F64CC" authority="(Reitter, 1910)" baseAuthorityName="Reitter" baseAuthorityYear="1910" class="Insecta" family="Tenebrionidae" genus="Philhammus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="131" phylum="Arthropoda" rank="species" species="aharonii">
|
||
<emphasis id="4DD49E01FFC43D33FB8BF958365264CC" italics="true" pageId="3" pageNumber="131">Philhammus aharonii</emphasis>
|
||
(Reitter, 1910)
|
||
</taxonomicName>
|
||
from adjacent
|
||
<collectingCountry id="07B70283FFC43D33FBD3F96731EA64CC" box="[1130,1189,1709,1730]" name="Egypt" pageId="3" pageNumber="131">Egypt</collectingCountry>
|
||
,
|
||
<collectingCountry id="07B70283FFC33D34FF2EFF08359A62D9" box="[151,213,194,215]" name="Sudan" pageId="4" pageNumber="132">Sudan</collectingCountry>
|
||
,
|
||
<collectingCountry id="07B70283FFC33D34FF5CFF08342562D9" box="[229,362,194,215]" name="Saudi Arabia" pageId="4" pageNumber="132">Saudi Arabia</collectingCountry>
|
||
, and
|
||
<collectingCountry id="07B70283FFC33D34FE11FF08349362D9" box="[424,476,194,215]" name="Israel" pageId="4" pageNumber="132">Israel</collectingCountry>
|
||
has completely rounded posterior corners of the pronotum, the median furrow is limited to the basal third of the pronotum, the eyes are smaller, with the genae surpassing the eyes, and the elytra have denser rows of punctures as in
|
||
<taxonomicName id="B8A03990FFC33D34FEFFFE8D34E36352" authorityName="SCHAWALLER AND STEINER" authorityYear="2018" box="[326,428,326,348]" class="Insecta" family="Tenebrionidae" genus="Philhammus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="132" phylum="Arthropoda" rank="species" species="ambouli">
|
||
<emphasis id="4DD49E01FFC33D34FEFFFE8D34E36352" box="[326,428,326,348]" italics="true" pageId="4" pageNumber="132">P. ambouli</emphasis>
|
||
</taxonomicName>
|
||
. For other differences in general habitus, compare figures of other species (
|
||
<bibRefCitation id="1B313FE2FFC33D34FF27FEB63421639F" author="Schawaller, W. & H. M. Al Dhafer & A. A. Elgharbawy" box="[158,366,379,401]" pageId="4" pageNumber="132" pagination="246 - 250" refId="ref2446" refString="Schawaller, W., H. M. Al Dhafer, and A. A. Elgharbawy. 2014. The genera Philhammus and Cnemeplatia (Coleoptera: Tenebrionidae: Cnemeplatiini) in the Arabian Peninsula, with taxonomic and faunistic remarks on additional species. Zoology in the Middle East 60: 246 - 250." type="journal article" year="2014">
|
||
Schawaller
|
||
<emphasis id="4DD49E01FFC33D34FEB0FEB63477639F" box="[265,312,379,401]" italics="true" pageId="4" pageNumber="132">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
). The aedeagi of most species are unknown, so we cannot determine if the form of the genitalia in
|
||
<taxonomicName id="B8A03990FFC33D34FE9BFE7B34CA63C8" authorityName="SCHAWALLER AND STEINER" authorityYear="2018" box="[290,389,432,454]" class="Insecta" family="Tenebrionidae" genus="Philhammus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="132" phylum="Arthropoda" rank="species" species="ambouli">
|
||
<emphasis id="4DD49E01FFC33D34FE9BFE7B34CA63C8" box="[290,389,432,454]" italics="true" pageId="4" pageNumber="132">P. ambouli</emphasis>
|
||
</taxonomicName>
|
||
is distinctive or diagnostic. The aedeagus of a related North American species,
|
||
<taxonomicName id="B8A03990FFC33D34FF2EFE2F34AB63F5" authority="(Horn)" baseAuthorityName="Horn" baseAuthorityYear="1870" box="[151,484,485,507]" class="Insecta" family="Tenebrionidae" genus="Lepidocnemeplatia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="132" phylum="Arthropoda" rank="species" species="sericea">
|
||
<emphasis id="4DD49E01FFC33D34FF2EFE2F34D963F5" box="[151,406,485,507]" italics="true" pageId="4" pageNumber="132">Lepidocnemeplatia sericea</emphasis>
|
||
(Horn)
|
||
</taxonomicName>
|
||
, has a similar slender form but is not parallel-sided, with the parameres narrowing from the base to the acuminate apex, and the basal piece is nearly as long as the parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="37BA1198FFC33D34FF16FD9A35BB60C1" pageId="4" pageNumber="132" type="etymology">
|
||
<paragraph id="7F1F4213FFC33D34FF16FD9A35BB60C1" blockId="4.[151,639,194,719]" pageId="4" pageNumber="132">
|
||
<emphasis id="4DD49E01FFC33D34FF16FD9A346D606B" bold="true" box="[175,290,592,613]" pageId="4" pageNumber="132">Etymology.</emphasis>
|
||
Used here as a noun in apposition, Ambouli is the Latin-like name for the town near Camp Lemonnier, the type locality south of
|
||
<collectingCountry id="07B70283FFC33D34FD96FD4F37316094" box="[559,638,645,666]" name="Djibouti" pageId="4" pageNumber="132">Djibouti</collectingCountry>
|
||
City, and the river valley where
|
||
<specimenCount id="69A6899AFFC33D34FE6CFD6A371F60BB" box="[469,592,672,693]" pageId="4" pageNumber="132" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
||
was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |