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<document id="B7F86B5B56C0422A12A918E58AB44628" ID-CLB-Dataset="24063" ID-DOI="10.1080/00222933.2019.1692088" ID-GBIF-Dataset="e6443eb6-bfcc-44a6-a547-7af9a94d4a7d" ID-ISSN="1464-5262" ID-Zenodo-Dep="3665519" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1581529324984" checkinUser="valdenar" docAuthor="Nunes, Rafael V. &amp; Vaz-de-Mello, Fernando Z." docDate="2019" docId="C740D6092A3FBB1BFEC5E4CAE04D2D77" docLanguage="en" docName="J_NAT_HIST.53.37–38.2231–2351.pdf.imf" docOrigin="Journal of Natural History 53 (37)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Dichotomius (Cephagonus) emas Nunes &amp; Vaz-de-Mello 2019, new species" docType="treatment" docVersion="18" lastPageNumber="2324" masterDocId="3B79AE712A62BB45FFA2E57EE46C2C23" masterDocTitle="Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)" masterLastPageNumber="2351" masterPageNumber="2231" pageNumber="2323" updateTime="1698782751744" updateUser="ExternalLinkService">
<mods:mods id="83B8B28322510AFA356565791E2F204B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B32991D22F61A05D9B281D12B42E2652">
<mods:title id="BF468736F9AF92F1E4AABF9F063392FB">Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)</mods:title>
</mods:titleInfo>
<mods:name id="E2F03721B3B42831BAE0BB83B2F163A6" type="personal">
<mods:role id="C8D7003623341F25BE115376A6F3CA74">
<mods:roleTerm id="E73C5F2B5DDA6A65232E3D9E80C4312D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0BAEE40AC69FC3D149267C1B71E2554D">Nunes, Rafael V.</mods:namePart>
</mods:name>
<mods:name id="020FCA0C57CB689A4D2903823CFFD41D" type="personal">
<mods:role id="93885BB218D0BE7BCA1E1FE6B970D3C2">
<mods:roleTerm id="53402CE69A5E1D557CC02D72B2B2548F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CFE9DD3776607A7A7B1D25BA12721C1F">Vaz-de-Mello, Fernando Z.</mods:namePart>
</mods:name>
<mods:typeOfResource id="CA92847F1CC53D5D0B71B741028619E7">text</mods:typeOfResource>
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<mods:titleInfo id="0D227A95197C5382078F8EC9C553A3D7">
<mods:title id="41C8539FFE3FB2A732C3A427A0DD42CE">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="CB221701ADAC97C4751E6E633C65D10B">
<mods:date id="087A92C60F82F434E219CE2B4D4C9405">2019</mods:date>
<mods:detail id="42EB3597580322E2BD42845F647D6475" type="pubDate">
<mods:number id="EB81CA5A68FDA132E4F61E4464739BCE">2020-01-13</mods:number>
</mods:detail>
<mods:detail id="EE6A79CC1BFF5F4B19453086007EBA40" type="volume">
<mods:number id="D6BF4EA66BD333B7BA42149E342FED56">53</mods:number>
</mods:detail>
<mods:detail id="97D3B9264EACD86F062D609FF0D2DD68" type="issue">
<mods:number id="D20C970E15B4B9BBB5399D963669EE70">37</mods:number>
</mods:detail>
<mods:extent id="52B5156351DBC9800DE954F36C836EF0" unit="page">
<mods:start id="5F52598591247BEA323869A216E90603">2231</mods:start>
<mods:end id="165B3F70C2C40C88730ED5DBF1050FC4">2351</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D077A33BB3C52D48ECAF52BE125F7D9D">journal article</mods:classification>
<mods:identifier id="F6859C1FE1254AEB8C2F234EFDBA3E39" type="CLB-Dataset">24063</mods:identifier>
<mods:identifier id="3221C79ECF82A4D1E319385E63972720" type="DOI">10.1080/00222933.2019.1692088</mods:identifier>
<mods:identifier id="376E43A033D29051DBC32777F024D833" type="GBIF-Dataset">e6443eb6-bfcc-44a6-a547-7af9a94d4a7d</mods:identifier>
<mods:identifier id="A8473E8E7BC261E57612FAE4B3EE179F" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="BA2F14C3AFB4A0E8319FB54B95CEB2F0" type="Zenodo-Dep">3665519</mods:identifier>
</mods:mods>
<treatment id="C740D6092A3FBB1BFEC5E4CAE04D2D77" ID-DOI="http://doi.org/10.5281/zenodo.3671936" ID-GBIF-Taxon="162116268" ID-Zenodo-Dep="3671936" LSID="urn:lsid:plazi:treatment:C740D6092A3FBB1BFEC5E4CAE04D2D77" httpUri="http://treatment.plazi.org/id/C740D6092A3FBB1BFEC5E4CAE04D2D77" lastPageId="94" lastPageNumber="2324" pageId="93" pageNumber="2323">
<subSubSection id="07F334942A3FBB18FEC5E4CAE6802DD3" pageId="93" pageNumber="2323" type="nomenclature">
<paragraph id="4F56671F2A3FBB18FEC5E4CAE7D02DEE" blockId="93.[359,956,436,496]" box="[359,956,436,461]" pageId="93" pageNumber="2323">
<heading id="141ED0732A3FBB18FEC5E4CAE7D02DEE" box="[359,956,436,461]" centered="true" fontSize="11" level="2" pageId="93" pageNumber="2323" reason="5">
3.17.:
<taxonomicName id="88E91C9C2A3FBB18FE0EE4CAE7492DEE" ID-CoL="6D3RJ" authority="Nunes &amp; Vaz-de-Mello, 2019" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[428,805,436,461]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="2323" phylum="Arthropoda" rank="species" species="emas" status="sp. nov." subGenus="Cephagonus">
<emphasis id="7D9DBB0D2A3FBB18FE0EE4CAE7492DEE" box="[428,805,436,461]" italics="true" pageId="93" pageNumber="2323">Dichotomius (Cephagonus) emas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="66AE06762A3FBB18FC8FE4CAE7D02DEE" box="[813,956,436,461]" pageId="93" pageNumber="2323" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="4F56671F2A3FBB18FD95E4A9E6802DD3" blockId="93.[359,956,436,496]" box="[567,748,471,496]" pageId="93" pageNumber="2323">
<heading id="141ED0732A3FBB18FD95E4A9E6802DD3" box="[567,748,471,496]" centered="true" fontSize="11" level="2" pageId="93" pageNumber="2323" reason="5">
(
<figureCitation id="D7D27B9A2A3FBB18FD9DE4A9E6C32DD3" box="[575,687,471,496]" captionStart="Figure 39" captionStartId="54.[160,225,1518,1541]" captionTargetBox="[163,1152,142,1487]" captionTargetId="figure@54.[163,1153,142,1487]" captionTargetPageId="54" captionText="Figure 39. D. emas: (a) male dorsal habitus; (b) paramera dorsal view (black arrow indicating strong medial constriction on paramera side); (c) paramera ventral view (black arrow indicating strong medial constriction on paramera side); (d) paramera lateral view. D. marcoscarvalhoi: (e) male dorsal habitus; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view. D. paschoali (i) male dorsal habitus; (j) paramera dorsal view; (k) paramera ventral view; (l) paramera lateral view (black arrow indicating basal excavation)." figureDoi="http://doi.org/10.5281/zenodo.3665593" httpUri="https://zenodo.org/record/3665593/files/figure.png" pageId="93" pageNumber="2323">Figure 39</figureCitation>
(a-d))
</heading>
</paragraph>
</subSubSection>
<subSubSection id="07F334942A3FBB18FF02E750E0622EF1" pageId="93" pageNumber="2323" type="diagnosis">
<paragraph id="4F56671F2A3FBB18FF02E750E0622EF1" blockId="93.[160,1155,558,722]" pageId="93" pageNumber="2323">
<emphasis id="7D9DBB0D2A3FBB18FF02E750E5762E64" bold="true" box="[160,282,558,583]" italics="true" pageId="93" pageNumber="2323">Diagnosis.</emphasis>
separated from other in the
<taxonomicName id="88E91C9C2A3FBB18FDCCE751E6962E64" authorityName="Felsche" authorityYear="1901" box="[622,762,559,583]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="2323" phylum="Arthropoda" rank="species" species="quadraticeps">
<emphasis id="7D9DBB0D2A3FBB18FDCCE751E6962E64" box="[622,762,559,583]" italics="true" pageId="93" pageNumber="2323">quadraticeps</emphasis>
</taxonomicName>
group by the following combined characters: clypeo-genal angulation in larger males acute and forming a teeth (like
<emphasis id="7D9DBB0D2A3FBB18FF02E70AE6402EAF" box="[160,556,628,652]" italics="true" pageId="93" pageNumber="2323">
<taxonomicName id="88E91C9C2A3FBB18FF02E70AE5452EAF" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[160,297,628,652]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="2323" phylum="Arthropoda" rank="species" species="edmondsi">D. edmondsi</taxonomicName>
,
<taxonomicName id="88E91C9C2A3FBB18FE97E70AE5D02EAF" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[309,444,628,652]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="2323" phylum="Arthropoda" rank="species" species="paschoali">D. paschoali</taxonomicName>
,
<taxonomicName id="88E91C9C2A3FBB18FE6AE70AE6402EAF" authorityName="Harold" authorityYear="1869" box="[456,556,628,652]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="2323" phylum="Arthropoda" rank="species" species="reichei">D. reichei</taxonomicName>
</emphasis>
); males pronotal disc with a central knob on anterior portion (idem); males cephalic horn or tubercle rounded apically and paramera in dorsal and ventral view strongly narrowed medially (as
<taxonomicName id="88E91C9C2A3FBB18FD64E7C4E7272EF1" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[710,843,698,722]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="2323" phylum="Arthropoda" rank="species" species="paschoali">
<emphasis id="7D9DBB0D2A3FBB18FD64E7C4E7272EF1" box="[710,843,698,722]" italics="true" pageId="93" pageNumber="2323">D. paschoali</emphasis>
</taxonomicName>
and
<taxonomicName id="88E91C9C2A3FBB18FC27E7C4E06D2EF1" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[901,1025,698,722]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="2323" phylum="Arthropoda" rank="species" species="frizzasae">
<emphasis id="7D9DBB0D2A3FBB18FC27E7C4E06D2EF1" box="[901,1025,698,722]" italics="true" pageId="93" pageNumber="2323">D. frizzasae</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="07F334942A3FBB18FF1CE781E7352F7F" pageId="93" pageNumber="2323" type="materials_examined">
<paragraph id="4F56671F2A3FBB18FF1CE781E5D32F19" blockId="93.[190,1155,767,860]" pageId="93" pageNumber="2323">
<emphasis id="7D9DBB0D2A3FBB18FF1CE781E4A02F30" bold="true" box="[190,204,767,787]" pageId="93" pageNumber="2323">●</emphasis>
<materialsCitation id="FF816D422A3FBB18FF7AE781E5D32F19" ID-GBIF-Occurrence="2570409580" collectionCode="PARNA, CEMT" country="Brazil" location="Emas" municipality="Mineiros" pageId="93" pageNumber="2323" specimenCount="821" specimenCount-male="1" stateProvince="Goias" typeStatus="holotype">
<typeStatus id="9052D9BD2A3FBB18FF7AE781E53E2F34" box="[216,338,767,791]" pageId="93" pageNumber="2323" type="holotype">HOLOTYPE</typeStatus>
(
<specimenCount id="59EFAC962A3FBB18FEC3E781E5F02F34" box="[353,412,767,791]" pageId="93" pageNumber="2323" type="male">male</specimenCount>
):
<collectingCountry id="37FE278F2A3FBB18FE0DE781E5932F34" box="[431,511,767,791]" name="Brazil" pageId="93" pageNumber="2323">BRASIL</collectingCountry>
:
<emphasis id="7D9DBB0D2A3FBB18FDA8E781E6212F34" bold="true" box="[522,589,767,791]" pageId="93" pageNumber="2323">
<collectingRegion id="8D2DA9FD2A3FBB18FDA8E781E6212F34" box="[522,589,767,791]" country="Brazil" name="Goias" pageId="93" pageNumber="2323">Goiás</collectingRegion>
</emphasis>
.
<collectingMunicipality id="AF32FD652A3FBB18FDF9E781E6D22F34" box="[603,702,767,791]" pageId="93" pageNumber="2323">Mineiros</collectingMunicipality>
.
<collectionCode id="29F8FFDA2A3FBB18FD6BE781E7742F34" box="[713,792,767,791]" pageId="93" pageNumber="2323">PARNA</collectionCode>
<location id="4A3631C42A3FBB18FC83E781E7332F34" LSID="urn:lsid:plazi:treatment:C740D6092A3FBB1BFEC5E4CAE04D2D77:4A3631C42A3FBB18FC83E781E7332F34" box="[801,863,767,791]" country="Brazil" municipality="Mineiros" name="Emas" pageId="93" pageNumber="2323" stateProvince="Goias">Emas</location>
.
<geoCoordinate id="2ADD01D82A3FBB18FCC8E781E0412F34" box="[874,1069,767,791]" pageId="93" pageNumber="2323">
17³53
<emphasis id="7D9DBB0D2A3FBB18FC08E781E7DC2F34" box="[938,944,767,791]" italics="true" pageId="93" pageNumber="2323">’</emphasis>
Sı 53³59
<emphasis id="7D9DBB0D2A3FBB18FBA9E781E07D2F34" bold="true" box="[1035,1041,767,791]" italics="true" pageId="93" pageNumber="2323">ʹ</emphasis>
Wı
</geoCoordinate>
<specimenCount id="59EFAC962A3FBB18FB94E781E0EF2F34" box="[1078,1155,767,791]" pageId="93" pageNumber="2323" type="generic">
<quantity id="8811CAFA2A3FBB18FB94E781E0132F34" box="[1078,1151,767,791]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="93" pageNumber="2323" unit="m" value="820.0">820 m</quantity>
.
</specimenCount>
MF
<location id="4A3631C42A3FBB18FEA0E65CE5292F19" LSID="urn:lsid:plazi:treatment:C740D6092A3FBB1BFEC5E4CAE04D2D77:4A3631C42A3FBB18FEA0E65CE5292F19" box="[258,325,802,826]" country="Brazil" municipality="Mineiros" name="Souza" pageId="93" pageNumber="2323" stateProvince="Goias">Souza</location>
[at
<collectionCode id="29F8FFDA2A3FBB18FED3E65CE5DA2F19" box="[369,438,802,826]" pageId="93" pageNumber="2323">CEMT</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph id="4F56671F2A3FBB18FF1CE63BE7352F7F" blockId="93.[190,1155,767,860]" box="[190,857,836,860]" pageId="93" pageNumber="2323">
<emphasis id="7D9DBB0D2A3FBB18FF1CE63BE4A02F7A" bold="true" box="[190,204,837,857]" pageId="93" pageNumber="2323">●</emphasis>
<typeStatus id="9052D9BD2A3FBB18FF7AE63AE5302F7F" box="[216,348,836,860]" pageId="93" pageNumber="2323" type="paratype">PARATYPES</typeStatus>
: same data of
<typeStatus id="9052D9BD2A3FBB18FDA2E63AE6082F7F" box="[512,612,836,860]" pageId="93" pageNumber="2323" type="holotype">holotype</typeStatus>
[1 Ƌı
<specimenCount id="59EFAC962A3FBB18FD0BE63AE68A2F7F" box="[681,742,836,860]" pageId="93" pageNumber="2323" type="female">2 ♀♀</specimenCount>
at
<collectionCode id="29F8FFDA2A3FBB18FCA9E63AE73C2F7F" box="[779,848,836,860]" pageId="93" pageNumber="2323">CEMT</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="07F334942A3FBB18FF02E6F7E5222A9C" pageId="93" pageNumber="2323" type="description">
<paragraph id="4F56671F2A3FBB18FF02E6F7E68B2989" blockId="93.[160,1156,905,1450]" pageId="93" pageNumber="2323">
<emphasis id="7D9DBB0D2A3FBB18FF02E6F7E5412F81" bold="true" box="[160,301,905,930]" italics="true" pageId="93" pageNumber="2323">Description.</emphasis>
colour: blackı shiny.
<emphasis id="7D9DBB0D2A3FBB18FD8AE6F4E60A2F81" bold="true" box="[552,614,906,930]" pageId="93" pageNumber="2323">Head</emphasis>
: fronto-clypealı genal and inter-ocular surfaces smooth. clypeo-genal angle straight. Cephalic carina longer than wideı conical with rounded apex.
<emphasis id="7D9DBB0D2A3FBB18FEF3E6B1E5A52FC4" bold="true" box="[337,457,975,999]" pageId="93" pageNumber="2323">Pronotum</emphasis>
: disc shinyı lacking punctures and having a central anterior knob with a weakly emargination at each side of the knob. Posterior margin with a row of ocellate punctures interrupted medially.
<emphasis id="7D9DBB0D2A3FBB18FDD8E16AE776280F" bold="true" box="[634,794,1044,1068]" pageId="93" pageNumber="2323">Metasternum</emphasis>
: sides and margins of anterior lobe covered by long hair. Anterior portion of anterior lobe glabrous but bearing ocellate punctures. Setigerous punctures equally spaced at sides.
<emphasis id="7D9DBB0D2A3FBB18FC84E124E7002851" bold="true" box="[806,876,1114,1138]" pageId="93" pageNumber="2323">Elytra</emphasis>
: striae deeply impressed having coarse punctures spaced by their diameter. Interstriae convexı disc lacking microsculpture with very fine punctures (20 x).
<emphasis id="7D9DBB0D2A3FBB18FDC5E1E1E6B02894" bold="true" box="[615,732,1183,1207]" pageId="93" pageNumber="2323">Abdomen</emphasis>
: ventrite 1 with groups of 5
<emphasis id="7D9DBB0D2A3FBB18FBAEE1E1E0752894" box="[1036,1049,1183,1207]" italics="true" pageId="93" pageNumber="2323">–</emphasis>
8 setae at each side. Puncture of each ventrite interrupted medially as well as groove along the posterior margin of sixth ventrite.
<emphasis id="7D9DBB0D2A3FBB18FD96E19AE6C628DF" bold="true" box="[564,682,1252,1276]" pageId="93" pageNumber="2323">Aedeagus</emphasis>
(
<figureCitation id="D7D27B9A2A3FBB18FD1FE19AE74128DF" box="[701,813,1252,1276]" captionStart="Figure 39" captionStartId="54.[160,225,1518,1541]" captionTargetBox="[163,1152,142,1487]" captionTargetId="figure@54.[163,1153,142,1487]" captionTargetPageId="54" captionText="Figure 39. D. emas: (a) male dorsal habitus; (b) paramera dorsal view (black arrow indicating strong medial constriction on paramera side); (c) paramera ventral view (black arrow indicating strong medial constriction on paramera side); (d) paramera lateral view. D. marcoscarvalhoi: (e) male dorsal habitus; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view. D. paschoali (i) male dorsal habitus; (j) paramera dorsal view; (k) paramera ventral view; (l) paramera lateral view (black arrow indicating basal excavation)." figureDoi="http://doi.org/10.5281/zenodo.3665593" httpUri="https://zenodo.org/record/3665593/files/figure.png" pageId="93" pageNumber="2323">Figure 39</figureCitation>
(b-d)): dorsallyı paramere narrowed medially
<emphasis id="7D9DBB0D2A3FBB18FEF0E079E533293C" box="[338,351,1287,1311]" italics="true" pageId="93" pageNumber="2323">–</emphasis>
emargination truncated
<emphasis id="7D9DBB0D2A3FBB18FDD6E079E615293C" box="[628,633,1287,1311]" italics="true" pageId="93" pageNumber="2323">‘</emphasis>
V
<emphasis id="7D9DBB0D2A3FBB18FD2BE079E6E3293C" box="[649,655,1287,1311]" italics="true" pageId="93" pageNumber="2323">’</emphasis>
shaped. Apexes roundedı parallel; sub-apical spine present; longitudinal excavation taking half of paramera length. Basal excavation narrowı delimited by basal knobs. Laterallyı abrupt declivity from basis towards apex. Ventrallyı basis with a single central tubercle and two lateral-rounded excavations; subgenital plate emarginated basallyı rounded apically.
</paragraph>
<paragraph id="4F56671F2A3FBB18FF02E0A8E5222A9C" blockId="93.[160,1155,1494,1727]" pageId="93" pageNumber="2323">
<emphasis id="7D9DBB0D2A3FBB18FF02E0A8E5A829CC" bold="true" box="[160,452,1494,1519]" italics="true" pageId="93" pageNumber="2323">Morphological variation.</emphasis>
male BL: 16 mmı PW:
<quantity id="8811CAFA2A3FBB18FD62E0A9E76F29CC" box="[704,771,1495,1519]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="93" pageNumber="2323" unit="mm" value="9.0">9 mm</quantity>
. Female BL: 16 mmı PW:
<quantity id="8811CAFA2A3FBB18FBB1E0A9E03E29CC" box="[1043,1106,1495,1519]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="93" pageNumber="2323" unit="mm" value="9.0">9 mm</quantity>
and distinguished from males as follows: Head: fronto-clypeal surface striated. Coarse puncture present posterior to cephalic carina. Cephalic carina transversalı bifurcated at apex forming a weak pair of tubercles. Pronotum: simply convexı lacking lobesı borders carinate. Elytra: striae weakly impressedı punctures separated by their diameter. Blue sheen present near basis. Abdomen: sixth ventrite bearing a pair of weak tubercles at central portion.
</paragraph>
</subSubSection>
<subSubSection id="07F334942A3CBB1BFF02E5F0E5722CE9" pageId="94" pageNumber="2324" type="distribution">
<paragraph id="4F56671F2A3CBB1BFF02E5F0E5722CE9" blockId="94.[160,1155,142,202]" pageId="94" pageNumber="2324">
<emphasis id="7D9DBB0D2A3CBB1BFF02E5F0E55F2C84" bold="true" box="[160,307,142,167]" italics="true" pageId="94" pageNumber="2324">Distribution.</emphasis>
Brazilı state of
<collectingRegion id="8D2DA9FD2A3CBB1BFE4EE5F1E6462C84" box="[492,554,143,167]" country="Brazil" name="Goias" pageId="94" pageNumber="2324">Goiás</collectingRegion>
in the municipality of Mineiros (Emas National Park) (
<figureCitation id="D7D27B9A2A3CBB1BFF05E5CCE57D2CE9" box="[167,273,178,202]" captionStart="Figure 36" captionStartId="51.[160,225,717,740]" captionTargetBox="[164,1151,142,685]" captionTargetId="figure@51.[163,1153,142,686]" captionTargetPageId="51" captionText="Figure 36. Distribution map of D. ricardosilvaiı D. marcoscarvalhoiı D. furtadoiı D. barbaraeı D. tissianiaeı D. eması D. frizzasae and D. candangoi." figureDoi="http://doi.org/10.5281/zenodo.3665587" httpUri="https://zenodo.org/record/3665587/files/figure.png" pageId="94" pageNumber="2324">Figure 36</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="07F334942A3CBB1BFF02E588E04D2D77" pageId="94" pageNumber="2324" type="etymology">
<paragraph id="4F56671F2A3CBB1BFF02E588E04D2D77" blockId="94.[160,1155,246,340]" pageId="94" pageNumber="2324">
<emphasis id="7D9DBB0D2A3CBB1BFF02E588E54F2D2C" bold="true" box="[160,291,246,271]" italics="true" pageId="94" pageNumber="2324">Etymology.</emphasis>
eması noun in appositionı refers to Emas National Park (Mineirosı Goiásı
<collectingCountry id="37FE278F2A3CBB1BFF02E464E48D2D11" box="[160,225,282,306]" name="Brazil" pageId="94" pageNumber="2324">Brazil</collectingCountry>
)ı
<typeStatus id="9052D9BD2A3CBB1BFF53E464E54F2D11" box="[241,291,282,306]" pageId="94" pageNumber="2324">type</typeStatus>
locality and where a considerable portion of Cerrado is preserved. This new species is one of the many endemic species of Taquari Plateauı in central
<collectingCountry id="37FE278F2A3CBB1BFC7CE442E0712D77" box="[990,1053,316,340]" name="Brazil" pageId="94" pageNumber="2324">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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