treatments-xml/data/C7/40/D6/C740D6092A3EBB18FEF3E4A9E5272D55.xml
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<document id="FB9360A05094D4B27F3D58FA7CC9A772" ID-CLB-Dataset="24063" ID-DOI="10.1080/00222933.2019.1692088" ID-GBIF-Dataset="e6443eb6-bfcc-44a6-a547-7af9a94d4a7d" ID-ISSN="1464-5262" ID-Zenodo-Dep="3665519" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1581529324984" checkinUser="valdenar" docAuthor="Nunes, Rafael V. &amp; Vaz-de-Mello, Fernando Z." docDate="2019" docId="C740D6092A3EBB18FEF3E4A9E5272D55" docLanguage="en" docName="J_NAT_HIST.53.3738.22312351.pdf.imf" docOrigin="Journal of Natural History 53 (37)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Dichotomius (Cephagonus) gardneri Nunes &amp; Vaz-de-Mello 2019, new species" docType="treatment" docVersion="14" lastPageNumber="2323" masterDocId="3B79AE712A62BB45FFA2E57EE46C2C23" masterDocTitle="Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)" masterLastPageNumber="2351" masterPageNumber="2231" pageNumber="2322" updateTime="1698782751744" updateUser="ExternalLinkService">
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<mods:title id="C3DC4B511A03A4D81D2407D9A505A8F4">Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)</mods:title>
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<mods:namePart id="EF90E02A00227469985A8DDDE2A90451">Nunes, Rafael V.</mods:namePart>
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<mods:namePart id="4C690ACFACBED7837278A4D8279E9094">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:title id="CC64C34E89AC7E85EAD25A48553FD408">Journal of Natural History</mods:title>
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<mods:date id="54EB58C62ACCFF73BC57DAA5387A759B">2019</mods:date>
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<treatment id="C740D6092A3EBB18FEF3E4A9E5272D55" ID-DOI="http://doi.org/10.5281/zenodo.3671934" ID-GBIF-Taxon="162136333" ID-Zenodo-Dep="3671934" LSID="urn:lsid:plazi:treatment:C740D6092A3EBB18FEF3E4A9E5272D55" httpUri="http://treatment.plazi.org/id/C740D6092A3EBB18FEF3E4A9E5272D55" lastPageId="93" lastPageNumber="2323" pageId="92" pageNumber="2322">
<subSubSection id="07F334942A3EBB19FEF3E4A9E6882E30" pageId="92" pageNumber="2322" type="nomenclature">
<paragraph id="4F56671F2A3EBB19FEF3E4A9E7BE2DD3" blockId="92.[337,978,471,531]" box="[337,978,471,496]" pageId="92" pageNumber="2322">
<heading id="141ED0732A3EBB19FEF3E4A9E7BE2DD3" box="[337,978,471,496]" centered="true" fontSize="11" level="2" pageId="92" pageNumber="2322" reason="2">
3.16.:
<taxonomicName id="88E91C9C2A3EBB19FE34E4A9E75F2DD3" ID-CoL="6D3RK" authority="Nunes &amp; Vaz-de-Mello, 2019" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[406,819,471,496]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="2322" phylum="Arthropoda" rank="species" species="gardneri" status="sp. nov." subGenus="Cephagonus">
<emphasis id="7D9DBB0D2A3EBB19FE34E4A9E75F2DD3" box="[406,819,471,496]" italics="true" pageId="92" pageNumber="2322">Dichotomius (Cephagonus) gardneri</emphasis>
</taxonomicName>
<emphasis id="7D9DBB0D2A3EBB19FC99E4A9E7BE2DD3" bold="true" box="[827,978,471,496]" pageId="92" pageNumber="2322">
<taxonomicNameLabel id="66AE06762A3EBB19FC99E4A9E7BE2DD3" box="[827,978,471,496]" pageId="92" pageNumber="2322" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4F56671F2A3EBB19FD9DE484E6882E30" blockId="92.[337,978,471,531]" box="[575,740,506,531]" pageId="92" pageNumber="2322">
<heading id="141ED0732A3EBB19FD9DE484E6882E30" box="[575,740,506,531]" centered="true" fontSize="11" level="2" pageId="92" pageNumber="2322" reason="5">
(
<figureCitation id="D7D27B9A2A3EBB19FDE5E484E6D52E30" box="[583,697,506,531]" captionStart="Figure 37" captionStartId="52.[160,225,1535,1558]" captionTargetBox="[163,1145,148,1503]" captionTargetId="figure@52.[163,1153,142,1503]" captionTargetPageId="52" captionText="Figure 37. D. edmondsi: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view. D. frizzasae: (e) male dorsal habitus; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view. D. gardneri (i) male dorsal habitus; (j) paramera dorsal view; (k) paramera ventral view; (l) paramera lateral view." figureDoi="http://doi.org/10.5281/zenodo.3665589" httpUri="https://zenodo.org/record/3665589/files/figure.png" pageId="92" pageNumber="2322">Figure 37</figureCitation>
(i-l))
</heading>
</paragraph>
</subSubSection>
<subSubSection id="07F334942A3EBB19FF02E72FE53F2F34" pageId="92" pageNumber="2322" type="diagnosis">
<paragraph id="4F56671F2A3EBB19FF02E72FE53F2F34" blockId="92.[160,1156,593,791]" pageId="92" pageNumber="2322">
<emphasis id="7D9DBB0D2A3EBB19FF02E72FE5762E49" bold="true" box="[160,282,593,618]" italics="true" pageId="92" pageNumber="2322">Diagnosis.</emphasis>
separated from other in the group by the following combined characters: males pronotum central emargination on anterior portion; males cephalic carina inserted both eyesı transversal with acute apex (similar to
<taxonomicName id="88E91C9C2A3EBB19FC1DE7E9E02E2E8C" authorityName="Nunesi" authorityYear="2016" box="[959,1090,663,687]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="2322" phylum="Arthropoda" rank="species" species="aielloae">
<emphasis id="7D9DBB0D2A3EBB19FC1DE7E9E02E2E8C" box="[959,1090,663,687]" italics="true" pageId="92" pageNumber="2322">D. aielloae</emphasis>
</taxonomicName>
and
<taxonomicName id="88E91C9C2A3EBB19FF02E7C4E5782EF1" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[160,276,698,722]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="2322" phylum="Arthropoda" rank="species" species="asenjoi">
<emphasis id="7D9DBB0D2A3EBB19FF02E7C4E5782EF1" box="[160,276,698,722]" italics="true" pageId="92" pageNumber="2322">D. asenjoi</emphasis>
</taxonomicName>
); elytral striae and puncture deeply impressed; males paramera lacking basal and longitudinal excavations and paramera subgenital plate taking 1/5 of the ventral surface.
</paragraph>
</subSubSection>
<subSubSection id="07F334942A3EBB19FF1CE63BE66E2FE7" pageId="92" pageNumber="2322" type="materials_examined">
<paragraph id="4F56671F2A3EBB19FF1CE63BE6BB2F5C" blockId="92.[190,1155,836,964]" pageId="92" pageNumber="2322">
<emphasis id="7D9DBB0D2A3EBB19FF1CE63BE4A02F7A" bold="true" box="[190,204,837,857]" pageId="92" pageNumber="2322"></emphasis>
<materialsCitation id="FF816D422A3EBB19FF7AE63AE6BB2F5C" ID-GBIF-Occurrence="2570409576" collectingDate="2006-04" collectionCode="T, MIM, CEMT" country="Brazil" location="Fezes" municipality="Para. Monte Dourado" pageId="92" pageNumber="2322" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="9052D9BD2A3EBB19FF7AE63AE53E2F7F" box="[216,338,836,860]" pageId="92" pageNumber="2322" type="holotype">HOLOTYPE</typeStatus>
(
<specimenCount id="59EFAC962A3EBB19FEC3E63AE5F02F7F" box="[353,412,836,860]" pageId="92" pageNumber="2322" type="male">male</specimenCount>
):
<collectingCountry id="37FE278F2A3EBB19FE0DE63AE5932F7F" box="[431,511,836,860]" name="Brazil" pageId="92" pageNumber="2322">BRASIL</collectingCountry>
:
<collectingMunicipality id="AF32FD652A3EBB19FDA8E63AE76A2F7F" box="[522,774,836,860]" pageId="92" pageNumber="2322">
<emphasis id="7D9DBB0D2A3EBB19FDA8E63AE62C2F7F" bold="true" box="[522,576,836,860]" pageId="92" pageNumber="2322">Pará</emphasis>
. Monte Dourado
</collectingMunicipality>
.
<geoCoordinate id="2ADD01D82A3EBB19FCB3E63AE07C2F7F" box="[785,1040,836,860]" pageId="92" pageNumber="2322">
0³60
<emphasis id="7D9DBB0D2A3EBB19FCE6E63AE7262F7F" bold="true" box="[836,842,836,860]" italics="true" pageId="92" pageNumber="2322">ʹ</emphasis>
20
<emphasis id="7D9DBB0D2A3EBB19FCC7E63AE71D2F7F" box="[869,881,836,860]" italics="true" pageId="92" pageNumber="2322">
<emphasis id="7D9DBB0D2A3EBB19FCC7E63AE7072F7F" bold="true" box="[869,875,836,860]" italics="true" pageId="92" pageNumber="2322">ʹ</emphasis>
</emphasis>
Sı 52³59
<emphasis id="7D9DBB0D2A3EBB19FC6EE63AE7BE2F7F" bold="true" box="[972,978,836,860]" italics="true" pageId="92" pageNumber="2322">ʹ</emphasis>
07
<emphasis id="7D9DBB0D2A3EBB19FC4FE63AE79A2F7F" box="[1005,1014,836,860]" italics="true" pageId="92" pageNumber="2322"></emphasis>
W
</geoCoordinate>
.
<location id="4A3631C42A3EBB19FBBEE63AE0312F7F" LSID="urn:lsid:plazi:treatment:C740D6092A3EBB18FEF3E4A9E5272D55:4A3631C42A3EBB19FBBEE63AE0312F7F" box="[1052,1117,836,860]" country="Brazil" municipality="Para. Monte Dourado" name="Fezes" pageId="92" pageNumber="2322">Fezes</location>
.
<date id="3B5741DF2A3EBB19FBCAE63AE57D2F5C" pageId="92" pageNumber="2322" value="2006-04">
<collectingDate id="2B13B8372A3EBB19FBCAE63AE57D2F5C" pageId="92" pageNumber="2322" value="2006-04">iv- 2006</collectingDate>
</date>
.
<collectionCode id="29F8FFDA2A3EBB19FEBEE619E5452F5C" box="[284,297,871,895]" name="Tavera, Department of Geology and Geophysics" pageId="92" pageNumber="2322">T</collectionCode>
<location id="4A3631C42A3EBB19FE92E619E5E72F5C" LSID="urn:lsid:plazi:treatment:C740D6092A3EBB18FEF3E4A9E5272D55:4A3631C42A3EBB19FE92E619E5E72F5C" box="[304,395,871,895]" country="Brazil" municipality="Para. Monte Dourado" name="Gardner" pageId="92" pageNumber="2322">Gardner</location>
&amp;
<collectionCode id="29F8FFDA2A3EBB19FE08E619E5B72F5C" box="[426,475,871,895]" country="Russia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15245" lsid="urn:lsid:biocol.org:col:15245" name="Minusinsk Museum of Local Studies" pageId="92" pageNumber="2322" type="Herbarium">MIM</collectionCode>
<location id="4A3631C42A3EBB19FE41E619E6312F5C" LSID="urn:lsid:plazi:treatment:C740D6092A3EBB18FEF3E4A9E5272D55:4A3631C42A3EBB19FE41E619E6312F5C" box="[483,605,871,895]" country="Brazil" municipality="Para. Monte Dourado" name="Hernandez" pageId="92" pageNumber="2322">Hernandéz</location>
[at
<collectionCode id="29F8FFDA2A3EBB19FD2BE619E6A22F5C" box="[649,718,871,895]" pageId="92" pageNumber="2322">CEMT</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph id="4F56671F2A3EBB19FF1CE6F4E66E2FE7" blockId="92.[190,1155,836,964]" pageId="92" pageNumber="2322">
<emphasis id="7D9DBB0D2A3EBB19FF1CE6F4E4A02FBD" bold="true" box="[190,204,906,926]" pageId="92" pageNumber="2322"></emphasis>
<typeStatus id="9052D9BD2A3EBB19FF7AE6F4E5302F81" box="[216,348,906,930]" pageId="92" pageNumber="2322" type="paratype">PARATYPES</typeStatus>
:
<collectingCountry id="37FE278F2A3EBB19FEC7E6F4E5D92F81" box="[357,437,906,930]" name="Brazil" pageId="92" pageNumber="2322">BRASIL</collectingCountry>
.
<emphasis id="7D9DBB0D2A3EBB19FE1CE6F4E5982F81" bold="true" box="[446,500,906,930]" pageId="92" pageNumber="2322">Pará</emphasis>
. Óbidos. 1878. M Nathan [1 Ƌ at
<collectionCode id="29F8FFDA2A3EBB19FCC7E6F4E7C62F81" box="[869,938,906,930]" pageId="92" pageNumber="2322">CEMT</collectionCode>
]; same municipality but
<date id="3B5741DF2A3EBB19FEA4E6D2E53B2FE7" box="[262,343,940,964]" pageId="92" pageNumber="2322" value="1989-11">xi-1989</date>
[
<specimenCount id="59EFAC962A3EBB19FECAE6D2E5FC2FE7" box="[360,400,940,964]" pageId="92" pageNumber="2322" type="female">1 ♀</specimenCount>
at
<collectionCode id="29F8FFDA2A3EBB19FE16E6D2E5952FE7" box="[436,505,940,964]" pageId="92" pageNumber="2322">CEMT</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="07F334942A3EBB18FF02E68FE6CF2CEA" lastPageId="93" lastPageNumber="2323" pageId="92" pageNumber="2322" type="description">
<paragraph id="4F56671F2A3EBB19FF02E68FE7ED2A31" blockId="92.[160,1156,1009,1554]" pageId="92" pageNumber="2322">
<emphasis id="7D9DBB0D2A3EBB19FF02E68FE5922829" bold="true" box="[160,510,1009,1034]" italics="true" pageId="92" pageNumber="2322">Description (Holotypeı male).</emphasis>
BL: 17 mmı PW:
<quantity id="8811CAFA2A3EBB19FD42E68CE7472829" box="[736,811,1010,1034]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="92" pageNumber="2322" unit="mm" value="9.0">9 mm</quantity>
. Colour: blackı shiny.
<emphasis id="7D9DBB0D2A3EBB19FB9FE68CE0122829" bold="true" box="[1085,1150,1010,1034]" pageId="92" pageNumber="2322">Head</emphasis>
: fronto-clypealı genal and inter-ocular surfaces smooth. Few weakly impressed fine punctures near eyes. clypeo-genal angle roundedı obtuse (100³). Cephalic carina longer than wideı conical with rounded apex.
<emphasis id="7D9DBB0D2A3EBB19FD7EE124E7342851" bold="true" box="[732,856,1114,1138]" pageId="92" pageNumber="2322">Pronotum</emphasis>
: disc shinyı with central emargination
<emphasis id="7D9DBB0D2A3EBB19FEE8E102E53B28B7" box="[330,343,1148,1172]" italics="true" pageId="92" pageNumber="2322"></emphasis>
same width as interocular space. Posterior margin with a row of ocellate punctures.
<emphasis id="7D9DBB0D2A3EBB19FE32E1E1E65A2894" bold="true" box="[400,566,1183,1207]" pageId="92" pageNumber="2322">Metasternum</emphasis>
: sides and margins of anterior lobe covered by long hair. Anterior portion of anterior lobe with ocellate setigerous punctures. Setigerous punctures equally spaced at sides.
<emphasis id="7D9DBB0D2A3EBB19FD72E19AE77528DF" bold="true" box="[720,793,1252,1276]" pageId="92" pageNumber="2322">Elytra</emphasis>
: striae deeply impressed having coarse punctures spaced by their diameter. Interstriae convexı disc lacking microsculpture with very fine punctures (20 x).
<emphasis id="7D9DBB0D2A3EBB19FD48E054E70F2961" bold="true" box="[746,867,1322,1346]" pageId="92" pageNumber="2322">Abdomen</emphasis>
: ventrites with ocellate punctures concentrate at sides. Puncture of each ventrite interrupted medially as well as groove along the posterior margin of sixth ventrite.
<emphasis id="7D9DBB0D2A3EBB19FC21E011E79229A4" bold="true" box="[899,1022,1391,1415]" pageId="92" pageNumber="2322">Aedeagus</emphasis>
(
<figureCitation id="D7D27B9A2A3EBB19FBB0E011E0EF29A4" box="[1042,1155,1391,1415]" captionStart="Figure 37" captionStartId="52.[160,225,1535,1558]" captionTargetBox="[163,1145,148,1503]" captionTargetId="figure@52.[163,1153,142,1503]" captionTargetPageId="52" captionText="Figure 37. D. edmondsi: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view. D. frizzasae: (e) male dorsal habitus; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view. D. gardneri (i) male dorsal habitus; (j) paramera dorsal view; (k) paramera ventral view; (l) paramera lateral view." figureDoi="http://doi.org/10.5281/zenodo.3665589" httpUri="https://zenodo.org/record/3665589/files/figure.png" pageId="92" pageNumber="2322">Figure 37</figureCitation>
(j-l)): dorsallyı paramere apexes roundedı convergent; longitudinal and basal excavations lacking; membranous part taking half of dorsal surface. Laterallyı subtle declivity from basis towards apex; lacking central excavation. Ventrallyı subgenital plate with rounded apexes and taking 1/5 of ventral surface.
</paragraph>
<paragraph id="4F56671F2A3EBB18FF02E340E6CF2CEA" blockId="92.[160,1156,1598,1727]" lastBlockId="93.[160,1155,142,201]" lastPageId="93" lastPageNumber="2323" pageId="92" pageNumber="2322">
<emphasis id="7D9DBB0D2A3EBB19FF02E340E5A52A74" bold="true" box="[160,457,1598,1623]" italics="true" pageId="92" pageNumber="2322">Morphological variation.</emphasis>
male BL is 17 mmı PW:
<quantity id="8811CAFA2A3EBB19FD50E341E70E2A74" box="[754,866,1599,1623]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="92" pageNumber="2322" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">
9
<emphasis id="7D9DBB0D2A3EBB19FCA2E341E7612A74" box="[768,781,1599,1623]" italics="true" pageId="92" pageNumber="2322"></emphasis>
10 mm
</quantity>
. Female BL: 16 mmı PW: 11 mmı distinguished from males as follows: Head: fronto-clypeal surface striated. Surface posterior to cephalic carina smooth. Cephalic carina transversalı with straight apex. Pronotum: simply convexı lacking lobes. Elytra: striae weakly impressedı punctures separated by twice their diameter. Blue sheen present near basis. Abdomen: sixth ventrite bearing a pair of tubercles on central portion.
</paragraph>
</subSubSection>
<subSubSection id="07F334942A3FBB18FF02E58BE6F22D2D" box="[160,670,245,270]" pageId="93" pageNumber="2323" type="distribution">
<paragraph id="4F56671F2A3FBB18FF02E58BE6F22D2D" blockId="93.[160,670,245,270]" box="[160,670,245,270]" pageId="93" pageNumber="2323">
<emphasis id="7D9DBB0D2A3FBB18FF02E58BE55F2D2D" bold="true" box="[160,307,245,270]" italics="true" pageId="93" pageNumber="2323">Distribution.</emphasis>
Brazilı state of
<collectingRegion id="8D2DA9FD2A3FBB18FE4AE588E6752D2D" box="[488,537,246,270]" country="Suriname" name="Para" pageId="93" pageNumber="2323">Pará</collectingRegion>
(
<figureCitation id="D7D27B9A2A3FBB18FD85E588E6FD2D2D" box="[551,657,246,270]" captionStart="Figure 34" captionStartId="49.[160,225,1006,1029]" captionTargetBox="[162,1151,142,975]" captionTargetId="figure@49.[162,1153,142,975]" captionTargetPageId="49" captionText="Figure 34. Distribution map of D. gardneriı D. aielloaeı D. asenjoiı D. arnaudiı D. Clypeatus and D. edmondsi." figureDoi="http://doi.org/10.5281/zenodo.3665583" httpUri="https://zenodo.org/record/3665583/files/figure.png" pageId="93" pageNumber="2323">Figure 34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="07F334942A3FBB18FF02E445E5272D55" pageId="93" pageNumber="2323" type="etymology">
<paragraph id="4F56671F2A3FBB18FF02E445E5272D55" blockId="93.[160,1155,315,374]" pageId="93" pageNumber="2323">
<emphasis id="7D9DBB0D2A3FBB18FF02E445E54F2D77" bold="true" box="[160,291,315,340]" italics="true" pageId="93" pageNumber="2323">Etymology.</emphasis>
named after Toby Gardner (Stockholm Environment Institute)ı who collected the
<typeStatus id="9052D9BD2A3FBB18FF6EE420E4922D55" box="[204,254,350,374]" pageId="93" pageNumber="2323">type</typeStatus>
series.
</paragraph>
</subSubSection>
</treatment>
</document>