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<document id="D098D9282300C9ACCCF251DEB4ADFBAA" ID-CLB-Dataset="24063" ID-DOI="10.1080/00222933.2019.1692088" ID-GBIF-Dataset="e6443eb6-bfcc-44a6-a547-7af9a94d4a7d" ID-ISSN="1464-5262" ID-Zenodo-Dep="3665519" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1581529324984" checkinUser="valdenar" docAuthor="Nunes, Rafael V. &amp; Vaz-de-Mello, Fernando Z." docDate="2019" docId="C740D6092A3BBB1FFEF2E0DCE7212A9A" docLanguage="en" docName="J_NAT_HIST.53.37–38.2231–2351.pdf.imf" docOrigin="Journal of Natural History 53 (37)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Dichotomius (Cephagonus) frizzasae Nunes &amp; Vaz-de-Mello 2019, new species" docType="treatment" docVersion="18" lastPageNumber="2320" masterDocId="3B79AE712A62BB45FFA2E57EE46C2C23" masterDocTitle="Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)" masterLastPageNumber="2351" masterPageNumber="2231" pageNumber="2319" updateTime="1698782751744" updateUser="ExternalLinkService">
<mods:mods id="D38CC6364E51F512F7D5EE3F7EED85D0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="91E3C94BFB378A970A2EEC727F92D0F3">
<mods:title id="53716B24DC72C454413E93010AB99013">Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)</mods:title>
</mods:titleInfo>
<mods:name id="6BF9CFF39D83401108152D36F2D26EEF" type="personal">
<mods:role id="E9DCE2E49E56A6C0D38AFD26D1D76518">
<mods:roleTerm id="DA23CA1D79EFAF4D3325823851AFAE4C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BBCDB47BF8F3E44D76D0D11CB5F720CE">Nunes, Rafael V.</mods:namePart>
</mods:name>
<mods:name id="A0CAA97AC6E96A43B35F989E42AE1767" type="personal">
<mods:role id="2C35FBC5746F564B7517AB0942A15B29">
<mods:roleTerm id="36ACBF43ABBA7F595CB6680D3F923720">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="62D72BC40018166A38BA2F57756E2688">Vaz-de-Mello, Fernando Z.</mods:namePart>
</mods:name>
<mods:typeOfResource id="558925836C5BBC42E9D6184FA9145C35">text</mods:typeOfResource>
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<mods:titleInfo id="542A592B70C09935C185DFFC794F9F3F">
<mods:title id="7412E2421C79741D82EE75ACBA1B8C59">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="46B5BDEBEC8EAD941399252CA9E12655">
<mods:date id="49EE98E2BEB552AFE85863AE34981548">2019</mods:date>
<mods:detail id="225704C92C50CEEA5A7E692CA2C0D555" type="pubDate">
<mods:number id="C31DB3EEFFC6D7A8271F1DAAA95A4900">2020-01-13</mods:number>
</mods:detail>
<mods:detail id="6146AE83CD143B5C8E52C82F46BEDBC6" type="volume">
<mods:number id="60E1B4E4CEBD6F3450B11A26361F9C03">53</mods:number>
</mods:detail>
<mods:detail id="E41222AF638770709694161372CCD16F" type="issue">
<mods:number id="050C8EC8ECB150BE01BC7CDF9B713BA4">37</mods:number>
</mods:detail>
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<mods:start id="2DB7AB4950633232B61D3BEAF3529869">2231</mods:start>
<mods:end id="547D2F1BF8FD6F167775E3ACBFB54C7B">2351</mods:end>
</mods:extent>
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<mods:classification id="BEC1CB2CFDD8922633E51056A7189195">journal article</mods:classification>
<mods:identifier id="CD7E3FB21397BF52FB44AE289FF06C8D" type="CLB-Dataset">24063</mods:identifier>
<mods:identifier id="762D1B0536E319E5FA681B58D984D2D6" type="DOI">10.1080/00222933.2019.1692088</mods:identifier>
<mods:identifier id="61135B0B856BD9DD934A91A4785E60CB" type="GBIF-Dataset">e6443eb6-bfcc-44a6-a547-7af9a94d4a7d</mods:identifier>
<mods:identifier id="684A152029ADD85DE26D68FCBA043511" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="B3E3B2C99E7326DD197C14C4E99DDCE6" type="Zenodo-Dep">3665519</mods:identifier>
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<treatment id="C740D6092A3BBB1FFEF2E0DCE7212A9A" ID-DOI="http://doi.org/10.5281/zenodo.3671928" ID-GBIF-Taxon="162116272" ID-Zenodo-Dep="3671928" LSID="urn:lsid:plazi:treatment:C740D6092A3BBB1FFEF2E0DCE7212A9A" httpUri="http://treatment.plazi.org/id/C740D6092A3BBB1FFEF2E0DCE7212A9A" lastPageId="90" lastPageNumber="2320" pageId="89" pageNumber="2319">
<subSubSection id="07F334942A3BBB1CFEF2E0DCE68029FD" pageId="89" pageNumber="2319" type="nomenclature">
<paragraph id="4F56671F2A3BBB1CFEF2E0DCE7BF2998" blockId="89.[336,979,1442,1502]" box="[336,979,1442,1467]" pageId="89" pageNumber="2319">
<heading id="141ED0732A3BBB1CFEF2E0DCE7BF2998" box="[336,979,1442,1467]" centered="true" fontSize="11" level="2" pageId="89" pageNumber="2319" reason="2">
3.14.:
<taxonomicName id="88E91C9C2A3BBB1CFE36E0DCE7582998" ID-CoL="6CRRK" authority="Nunes &amp; Vaz-de-Mello, 2019" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[404,820,1442,1467]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="2319" phylum="Arthropoda" rank="species" species="frizzasae" status="sp. nov." subGenus="Cephagonus">
<emphasis id="7D9DBB0D2A3BBB1CFE36E0DCE6482998" box="[404,548,1442,1467]" italics="true" pageId="89" pageNumber="2319">Dichotomius</emphasis>
(
<emphasis id="7D9DBB0D2A3BBB1CFD96E0DCE6AE2998" box="[564,706,1442,1467]" italics="true" pageId="89" pageNumber="2319">Cephagonus</emphasis>
)
<emphasis id="7D9DBB0D2A3BBB1CFD73E0DCE7582998" box="[721,820,1442,1467]" italics="true" pageId="89" pageNumber="2319">frizzasae</emphasis>
</taxonomicName>
<emphasis id="7D9DBB0D2A3BBB1CFC9EE0DCE7BF2998" bold="true" box="[828,979,1442,1467]" pageId="89" pageNumber="2319">
<taxonomicNameLabel id="66AE06762A3BBB1CFC9EE0DCE7BF2998" box="[828,979,1442,1467]" pageId="89" pageNumber="2319" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4F56671F2A3BBB1CFD95E0BBE68029FD" blockId="89.[336,979,1442,1502]" box="[567,748,1477,1502]" pageId="89" pageNumber="2319">
<heading id="141ED0732A3BBB1CFD95E0BBE68029FD" box="[567,748,1477,1502]" centered="true" fontSize="11" level="2" pageId="89" pageNumber="2319" reason="5">
(
<figureCitation id="D7D27B9A2A3BBB1CFD9DE0BBE6C329FD" box="[575,687,1477,1502]" captionStart="Figure 37" captionStartId="52.[160,225,1535,1558]" captionTargetBox="[163,1145,148,1503]" captionTargetId="figure@52.[163,1153,142,1503]" captionTargetPageId="52" captionText="Figure 37. D. edmondsi: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view. D. frizzasae: (e) male dorsal habitus; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view. D. gardneri (i) male dorsal habitus; (j) paramera dorsal view; (k) paramera ventral view; (l) paramera lateral view." figureDoi="http://doi.org/10.5281/zenodo.3665589" httpUri="https://zenodo.org/record/3665589/files/figure.png" pageId="89" pageNumber="2319">Figure 37</figureCitation>
(e-h))
</heading>
</paragraph>
</subSubSection>
<subSubSection id="07F334942A3BBB1FFF02E362E7E82CCF" lastPageId="90" lastPageNumber="2320" pageId="89" pageNumber="2319" type="diagnosis">
<paragraph id="4F56671F2A3BBB1FFF02E362E7E82CCF" blockId="89.[160,1156,1564,1727]" lastBlockId="90.[160,1156,142,236]" lastPageId="90" lastPageNumber="2320" pageId="89" pageNumber="2319">
<emphasis id="7D9DBB0D2A3BBB1CFF02E362E5762A16" bold="true" box="[160,282,1564,1589]" italics="true" pageId="89" pageNumber="2319">Diagnosis.</emphasis>
separated from other in the
<taxonomicName id="88E91C9C2A3BBB1CFD2AE363E7772A16" authorityName="Felsche" authorityYear="1901" box="[648,795,1565,1589]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="2319" phylum="Arthropoda" rank="species" species="quadraticeps">
<emphasis id="7D9DBB0D2A3BBB1CFD2AE363E7772A16" box="[648,795,1565,1589]" italics="true" pageId="89" pageNumber="2319">quadraticeps</emphasis>
</taxonomicName>
group by the following combined characters: clypeo-genal angulation in larger males acute and forming a teeth (like
<emphasis id="7D9DBB0D2A3BBB1CFEF0E31CE7482A59" box="[338,804,1634,1658]" italics="true" pageId="89" pageNumber="2319">
<taxonomicName id="88E91C9C2A3BBB1CFEF0E31CE5802A59" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[338,492,1634,1658]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="2319" phylum="Arthropoda" rank="species" species="edmondsi">D. edmondsi</taxonomicName>
,
<taxonomicName id="88E91C9C2A3BBB1CFDA0E31CE6F62A59" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[514,666,1634,1658]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="2319" phylum="Arthropoda" rank="species" species="paschoali">D. paschoali</taxonomicName>
,
<taxonomicName id="88E91C9C2A3BBB1CFD12E31CE7482A59" authorityName="Harold" authorityYear="1869" box="[688,804,1634,1658]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="2319" phylum="Arthropoda" rank="species" species="reichei">D. reichei</taxonomicName>
</emphasis>
); males pronotal disc with a central pair of lobes on anterior portion (idem); males cephalic horn or tubercle rounded apically and paramera in dorsal and ventral view strongly narrowed medially (as
<taxonomicName id="88E91C9C2A38BB1FFE9CE5F0E5BC2C85" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[318,464,142,166]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="2320" phylum="Arthropoda" rank="species" species="paschoali">
<emphasis id="7D9DBB0D2A38BB1FFE9CE5F0E5BC2C85" box="[318,464,142,166]" italics="true" pageId="90" pageNumber="2320">D. paschoali</emphasis>
</taxonomicName>
and
<taxonomicName id="88E91C9C2A38BB1FFDB5E5F0E61B2C85" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[535,631,142,166]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="2320" phylum="Arthropoda" rank="species" species="emas">
<emphasis id="7D9DBB0D2A38BB1FFDB5E5F0E61B2C85" box="[535,631,142,166]" italics="true" pageId="90" pageNumber="2320">D. emas</emphasis>
</taxonomicName>
). Externallyı this species is very similar to
<emphasis id="7D9DBB0D2A38BB1FFF02E5CFE6D12CEA" box="[160,701,177,201]" italics="true" pageId="90" pageNumber="2320">
<taxonomicName id="88E91C9C2A38BB1FFF02E5CFE5722CEA" authorityName="Felsche" authorityYear="1901" box="[160,286,177,201]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="2320" phylum="Arthropoda" rank="species" species="fissiceps">D. fissiceps</taxonomicName>
,
<taxonomicName id="88E91C9C2A38BB1FFE8EE5CFE5F62CEA" authorityName="Harold" authorityYear="1869" box="[300,410,177,201]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="2320" phylum="Arthropoda" rank="species" species="reichei">D. reichei</taxonomicName>
,
<taxonomicName id="88E91C9C2A38BB1FFE0AE5CFE65D2CEA" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[424,561,177,201]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="2320" phylum="Arthropoda" rank="species" species="barbarae">D. barbarae</taxonomicName>
,
<taxonomicName id="88E91C9C2A38BB1FFDE2E5CFE6D12CEA" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[576,701,177,201]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="2320" phylum="Arthropoda" rank="species" species="furtadoi">D. furtadoi</taxonomicName>
</emphasis>
and
<taxonomicName id="88E91C9C2A38BB1FFD5EE5CFE7E52CEA" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[764,905,177,201]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="2320" phylum="Arthropoda" rank="species" species="paschoali">
<emphasis id="7D9DBB0D2A38BB1FFD5EE5CFE7E52CEA" box="[764,905,177,201]" italics="true" pageId="90" pageNumber="2320">D. paschoali</emphasis>
</taxonomicName>
and is differentiated by type-locality and paramera characters (key and diagnosis).
</paragraph>
</subSubSection>
<subSubSection id="07F334942A38BB1FFF1CE467E7E72D55" pageId="90" pageNumber="2320" type="materials_examined">
<paragraph id="4F56671F2A38BB1FFF1CE467E0372D77" blockId="90.[190,1155,281,374]" pageId="90" pageNumber="2320">
<emphasis id="7D9DBB0D2A38BB1FFF1CE467E4A02D0E" bold="true" box="[190,204,281,301]" pageId="90" pageNumber="2320">●</emphasis>
<materialsCitation id="FF816D422A38BB1FFF7AE467E0372D77" ID-GBIF-Occurrence="2570409570" collectingDate="2009-11-16" collectingDateMax="2009-11-31" collectingDateMin="2009-11-16" collectionCode="CEMT" country="Brazil" location="Aguas Emendadas" municipality="Distrito Federal. Brasilia. Planaltina" pageId="90" pageNumber="2320" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="9052D9BD2A38BB1FFF7AE467E53E2D12" box="[216,338,281,305]" pageId="90" pageNumber="2320" type="holotype">HOLOTYPE</typeStatus>
(
<specimenCount id="59EFAC962A38BB1FFEC9E467E5CA2D12" box="[363,422,281,305]" pageId="90" pageNumber="2320" type="male">male</specimenCount>
):
<collectingCountry id="37FE278F2A38BB1FFE61E467E67F2D12" box="[451,531,281,305]" name="Brazil" pageId="90" pageNumber="2320">BRASIL</collectingCountry>
:
<collectingMunicipality id="AF32FD652A38BB1FFD8AE467E7B32D12" box="[552,991,281,305]" pageId="90" pageNumber="2320">
<emphasis id="7D9DBB0D2A38BB1FFD8AE467E6832D12" bold="true" box="[552,751,281,305]" pageId="90" pageNumber="2320">Distrito Federal</emphasis>
. Brasília. Planaltina
</collectingMunicipality>
. ESEC
<location id="4A3631C42A38BB1FFB9CE467E5312D77" LSID="urn:lsid:plazi:treatment:C740D6092A3BBB1FFEF2E0DCE7212A9A:4A3631C42A38BB1FFB9CE467E5312D77" country="Brazil" municipality="Distrito Federal. Brasilia. Planaltina" name="Aguas Emendadas" pageId="90" pageNumber="2320">Águas Emendadas</location>
.
<geoCoordinate id="2ADD01D82A38BB1FFEC5E442E61C2D77" box="[359,624,315,340]" pageId="90" pageNumber="2320">
15³32
<emphasis id="7D9DBB0D2A38BB1FFE05E445E5C12D70" bold="true" box="[423,429,315,339]" italics="true" pageId="90" pageNumber="2320">ʹ</emphasis>
31
<emphasis id="7D9DBB0D2A38BB1FFE6AE445E5BD2D70" box="[456,465,315,339]" italics="true" pageId="90" pageNumber="2320">”</emphasis>
Sı 47³36
<emphasis id="7D9DBB0D2A38BB1FFD8EE445E65E2D70" bold="true" box="[556,562,315,339]" italics="true" pageId="90" pageNumber="2320">ʹ</emphasis>
49
<emphasis id="7D9DBB0D2A38BB1FFDEFE445E63A2D70" box="[589,598,315,339]" italics="true" pageId="90" pageNumber="2320">”</emphasis>
W
</geoCoordinate>
.
<location id="4A3631C42A38BB1FFDD8E442E6D72D77" LSID="urn:lsid:plazi:treatment:C740D6092A3BBB1FFEF2E0DCE7212A9A:4A3631C42A38BB1FFDD8E442E6D72D77" box="[634,699,316,340]" country="Brazil" municipality="Distrito Federal. Brasilia. Planaltina" name="Pitfall" pageId="90" pageNumber="2320">Pitfall</location>
.
<date id="3B5741DF2A38BB1FFD64E442E70D2D77" box="[710,865,316,340]" pageId="90" pageNumber="2320" value="2009-11-16" valueMax="2009-11-31" valueMin="2009-11-16">
<collectingDate id="2B13B8372A38BB1FFD64E442E70D2D77" box="[710,865,316,340]" pageId="90" pageNumber="2320" value="2009-11-16" valueMax="2009-11-31" valueMin="2009-11-16">16-31-xi-2009</collectingDate>
</date>
. MR
<location id="4A3631C42A38BB1FFC34E442E78C2D77" LSID="urn:lsid:plazi:treatment:C740D6092A3BBB1FFEF2E0DCE7212A9A:4A3631C42A38BB1FFC34E442E78C2D77" box="[918,992,316,340]" country="Brazil" municipality="Distrito Federal. Brasilia. Planaltina" name="Frizzas" pageId="90" pageNumber="2320">Frizzas</location>
[at
<collectionCode id="29F8FFDA2A38BB1FFBAFE442E03E2D77" box="[1037,1106,316,340]" pageId="90" pageNumber="2320">CEMT</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph id="4F56671F2A38BB1FFF1CE421E7E72D55" blockId="90.[190,1155,281,374]" box="[190,907,350,374]" pageId="90" pageNumber="2320">
<emphasis id="7D9DBB0D2A38BB1FFF1CE421E4A02D50" bold="true" box="[190,204,351,371]" pageId="90" pageNumber="2320">●</emphasis>
<typeStatus id="9052D9BD2A38BB1FFF7AE420E5302D55" box="[216,348,350,374]" pageId="90" pageNumber="2320" type="paratype">PARATYPES</typeStatus>
: same data of
<typeStatus id="9052D9BD2A38BB1FFDA2E420E60B2D55" box="[512,615,350,374]" pageId="90" pageNumber="2320" type="holotype">Holotype</typeStatus>
[46 ƋƋı
<specimenCount id="59EFAC962A38BB1FFD6FE420E7742D55" box="[717,792,350,374]" pageId="90" pageNumber="2320" type="female">20 ♀♀</specimenCount>
at
<collectionCode id="29F8FFDA2A38BB1FFC9FE420E7EE2D55" box="[829,898,350,374]" pageId="90" pageNumber="2320">CEMT</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="07F334942A38BB1FFF02E4CDE65A29C8" pageId="90" pageNumber="2320" type="description">
<paragraph id="4F56671F2A38BB1FFF02E4CDE5E62818" blockId="90.[160,1156,435,1083]" pageId="90" pageNumber="2320">
<emphasis id="7D9DBB0D2A38BB1FFF02E4CDE5992DEF" bold="true" box="[160,501,435,460]" italics="true" pageId="90" pageNumber="2320">Description (Holotypeı male).</emphasis>
BL: 20 mmı PW:
<quantity id="8811CAFA2A38BB1FFD11E4CDE76F2DE8" box="[691,771,435,459]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="90" pageNumber="2320" unit="mm" value="13.0">13 mm</quantity>
. Colour: blackı shiny.
<emphasis id="7D9DBB0D2A38BB1FFC4EE4CDE0462DE8" bold="true" box="[1004,1066,435,459]" pageId="90" pageNumber="2320">Head</emphasis>
: frontoclypealı genal and inter-ocular surfaces smooth. Gena borders slightly curved inward. Clypeo-genal angle acute. Cephalic carina longer than wideı similar height to pronotumı conical with rounded apex.
<emphasis id="7D9DBB0D2A38BB1FFE77E765E6212E10" bold="true" box="[469,589,539,563]" pageId="90" pageNumber="2320">Pronotum</emphasis>
: pronotal disc shinyı lacking punctures and having a central pair of lobes. Region adjacent to eyes with a row of ocellate punctures. Posterior margin having 1
<emphasis id="7D9DBB0D2A38BB1FFEC1E71FE51C2E5A" box="[355,368,609,633]" italics="true" pageId="90" pageNumber="2320">–</emphasis>
2 rows of ocellate punctures.
<emphasis id="7D9DBB0D2A38BB1FFD75E71FE71B2E5A" bold="true" box="[727,887,609,633]" pageId="90" pageNumber="2320">Metasternum</emphasis>
: sides and margins of anterior lobe covered by long hair. Anterior portion of anterior lobe glabrous. Setigerous punctures equally spaced at sides.
<emphasis id="7D9DBB0D2A38BB1FFD63E7D8E76A2E9D" bold="true" box="[705,774,678,702]" pageId="90" pageNumber="2320">Elytra</emphasis>
: striae deeply impressed having fine punctures spaced by twice their diameter. Interstriae slightly convexı disc with black colourı lacking microsculptureı with very fine punctures (20 x).
<emphasis id="7D9DBB0D2A38BB1FFC09E795E04C2F20" bold="true" box="[939,1056,747,771]" pageId="90" pageNumber="2320">Abdomen</emphasis>
: sides of each ventrite excavated near anterior portionı with groups of ocellate punctures- some punctures setigerous with very short hair (under 35x magnification). Ventrite 1 with 8
<emphasis id="7D9DBB0D2A38BB1FFBF8E64FE00B2F6A" box="[1114,1127,817,841]" italics="true" pageId="90" pageNumber="2320">–</emphasis>
10 setae at each side. Puncture of each ventrite interrupted medially as well as groove along the posterior margin of sixth ventrite.
<emphasis id="7D9DBB0D2A38BB1FFDCFE608E68F2FAD" bold="true" box="[621,739,886,910]" pageId="90" pageNumber="2320">Aedeagus</emphasis>
(
<figureCitation id="D7D27B9A2A38BB1FFD5AE608E7062FAD" box="[760,874,886,910]" captionStart="Figure 37" captionStartId="52.[160,225,1535,1558]" captionTargetBox="[163,1145,148,1503]" captionTargetId="figure@52.[163,1153,142,1503]" captionTargetPageId="52" captionText="Figure 37. D. edmondsi: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view. D. frizzasae: (e) male dorsal habitus; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view. D. gardneri (i) male dorsal habitus; (j) paramera dorsal view; (k) paramera ventral view; (l) paramera lateral view." figureDoi="http://doi.org/10.5281/zenodo.3665589" httpUri="https://zenodo.org/record/3665589/files/figure.png" pageId="90" pageNumber="2320">Figure 37</figureCitation>
(f-h)): dorsallyı paramere strongly narrowed medially. Apexes roundedı parallel; longitudinal excavation absentı produced in a pair of knobs. Laterallyı abrupt declivity from basis towards apex; longitudinal sinuous carina separating ventral of dorsal excavations. Ventrallyı basis with three tuberclesı one central and two lateral ones. Subgenital plate rounded apically and emarginated basally.
</paragraph>
<paragraph id="4F56671F2A38BB1FFF02E106E65A29C8" blockId="90.[160,1156,1144,1515]" pageId="90" pageNumber="2320">
<emphasis id="7D9DBB0D2A38BB1FFF02E106E5AB28B2" bold="true" box="[160,455,1144,1169]" italics="true" pageId="90" pageNumber="2320">Morphological variation.</emphasis>
males BL: 12
<emphasis id="7D9DBB0D2A38BB1FFDCCE107E61728B2" box="[622,635,1145,1169]" italics="true" pageId="90" pageNumber="2320">–</emphasis>
20 mmı PW:
<quantity id="8811CAFA2A38BB1FFCB5E107E7E528B2" box="[791,905,1145,1169]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.3" metricValueMin="0.7" pageId="90" pageNumber="2320" unit="mm" value="10.0" valueMax="13.0" valueMin="7.0">
7
<emphasis id="7D9DBB0D2A38BB1FFC87E107E75E28B2" box="[805,818,1145,1169]" italics="true" pageId="90" pageNumber="2320">–</emphasis>
13 mm
</quantity>
. Male genitalia comparison allows us to identify three phenotypes. Larger males (described above) bear pronotal lobes and tall cephalic horns. Smaller males have simply convex pronotum with deeply impressed fine puncture on disc (10x)ı and the cephalic horn is produced in a conical simple tubercle. Females BL: 16
<emphasis id="7D9DBB0D2A38BB1FFD15E07DE6A82938" box="[695,708,1283,1307]" italics="true" pageId="90" pageNumber="2320">–</emphasis>
19 mmı PW: 8
<emphasis id="7D9DBB0D2A38BB1FFCD1E07DE7EC2938" box="[883,896,1283,1307]" italics="true" pageId="90" pageNumber="2320">–</emphasis>
12 mmı distinguished from males as follows: Head: fronto-clypeal surface striated. Coarse puncture present posterior to cephalic carina. Cephalic carina transversalı bifurcated at apex forming a weak pair of tubercles. Pronotum: simply convexı lacking lobesı borders produced into carinas. Elytra: striae deeply impressedı punctures separated by twice their diameter. Blue sheen present near basis. Abdomen: sixth ventrite bearing a pair of weak tubercles on central portion.
</paragraph>
</subSubSection>
<subSubSection id="07F334942A38BB1FFF02E356E0EF2A62" box="[160,1155,1576,1601]" pageId="90" pageNumber="2320" type="distribution">
<paragraph id="4F56671F2A38BB1FFF02E356E0EF2A62" blockId="90.[160,1155,1576,1601]" box="[160,1155,1576,1601]" pageId="90" pageNumber="2320">
<emphasis id="7D9DBB0D2A38BB1FFF02E356E55F2A62" bold="true" box="[160,307,1576,1601]" italics="true" pageId="90" pageNumber="2320">Distribution.</emphasis>
municipality of Brasíliaı administrative region of Planaltinaı
<collectingCountry id="37FE278F2A38BB1FFC65E357E06E2A62" box="[967,1026,1577,1601]" name="Brazil" pageId="90" pageNumber="2320">Brazil</collectingCountry>
(
<figureCitation id="D7D27B9A2A38BB1FFBB2E357E01B2A62" box="[1040,1143,1577,1601]" captionStart="Figure 36" captionStartId="51.[160,225,717,740]" captionTargetBox="[164,1151,142,685]" captionTargetId="figure@51.[163,1153,142,686]" captionTargetPageId="51" captionText="Figure 36. Distribution map of D. ricardosilvaiı D. marcoscarvalhoiı D. furtadoiı D. barbaraeı D. tissianiaeı D. eması D. frizzasae and D. candangoi." figureDoi="http://doi.org/10.5281/zenodo.3665587" httpUri="https://zenodo.org/record/3665587/files/figure.png" pageId="90" pageNumber="2320">Figure 36</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="07F334942A38BB1FFF02E303E7212A9A" pageId="90" pageNumber="2320" type="etymology">
<paragraph id="4F56671F2A38BB1FFF02E303E7212A9A" blockId="90.[160,1155,1661,1721]" pageId="90" pageNumber="2320">
<emphasis id="7D9DBB0D2A38BB1FFF02E303E54F2AB5" bold="true" box="[160,291,1661,1686]" italics="true" pageId="90" pageNumber="2320">Etymology.</emphasis>
named after our friendı the entomologist Marina Regina Frizzas (Universidade de Brasíliaı
<collectingCountry id="37FE278F2A38BB1FFE62E3DFE66C2A9A" box="[448,512,1697,1721]" name="Brazil" pageId="90" pageNumber="2320">Brasil</collectingCountry>
)ı who collected the
<typeStatus id="9052D9BD2A38BB1FFD40E3DFE7252A9A" box="[738,841,1697,1721]" pageId="90" pageNumber="2320" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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