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<document id="07CC1084FAEAD2EAF05D6BEAD0291FA6" ID-CLB-Dataset="24063" ID-DOI="10.1080/00222933.2019.1692088" ID-GBIF-Dataset="e6443eb6-bfcc-44a6-a547-7af9a94d4a7d" ID-ISSN="1464-5262" ID-Zenodo-Dep="3665519" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1581529324984" checkinUser="valdenar" docAuthor="Nunes, Rafael V. &amp; Vaz-de-Mello, Fernando Z." docDate="2019" docId="C740D6092A36BB10FEF7E62BE58928B8" docLanguage="en" docName="J_NAT_HIST.53.37–38.2231–2351.pdf.imf" docOrigin="Journal of Natural History 53 (37)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Dichotomius (Cephagonus) barbarae Nunes &amp; Vaz-de-Mello 2019, new species" docType="treatment" docVersion="18" lastPageNumber="2315" masterDocId="3B79AE712A62BB45FFA2E57EE46C2C23" masterDocTitle="Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)" masterLastPageNumber="2351" masterPageNumber="2231" pageNumber="2314" updateTime="1698782751744" updateUser="ExternalLinkService">
<mods:mods id="5AA856D46C54D00F7FC1E32E77165D4C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="138399232C69A98EFFF807617842C1C1">
<mods:title id="3C78030ABCB32A3752B4FE874C906AB5">Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)</mods:title>
</mods:titleInfo>
<mods:name id="F63603E16DF7AB963FE928B40A41DAA7" type="personal">
<mods:role id="00733FE3275D158EDE91642570657F1D">
<mods:roleTerm id="DBD3FE25B3DF4F2D76E7AE4B8C9BDF99">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="209A14B0D1DCDF53951CA63CCBA63C04">Nunes, Rafael V.</mods:namePart>
</mods:name>
<mods:name id="F6CC6C5913FCEE0628B6D13C0C9AB79D" type="personal">
<mods:role id="D58B16189DD351A81A192652BDB90BAA">
<mods:roleTerm id="D0DD9402E9E1D563B4F25214213F2366">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C617683011F3706EA380582A97AFF0F9">Vaz-de-Mello, Fernando Z.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A8B1341B85E8B811FBEDB42834604170">text</mods:typeOfResource>
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<mods:titleInfo id="B954A4AA0C3FC33A369E307086F37F32">
<mods:title id="FE1344D437DB4A9BF4AC533566303C40">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="888056A768E4D721EEA848D82593887F">
<mods:date id="A306D434AB8889EC534CA9D8A389A939">2019</mods:date>
<mods:detail id="98B34D207D98A4C14376252642350601" type="pubDate">
<mods:number id="8A13A679004C38FF897EA11ABB10FB18">2020-01-13</mods:number>
</mods:detail>
<mods:detail id="1C27689EEEC2B151E9672258A8123BE2" type="volume">
<mods:number id="865C07FE5AC10C58C27FA09930879BB6">53</mods:number>
</mods:detail>
<mods:detail id="85939DD5344A80A9900E3D873EC6D98F" type="issue">
<mods:number id="A39E5627CBFB2CB5141DE62143645950">37</mods:number>
</mods:detail>
<mods:extent id="13C27093D364678F9C8C4355D1A28BAA" unit="page">
<mods:start id="1E1C11D4D9AE41AADFF102C1502223E8">2231</mods:start>
<mods:end id="B5054BF5E655C6E864F6ECD890332115">2351</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="58D0B3999EC79DAA4437F1CF5D3A1E4A">journal article</mods:classification>
<mods:identifier id="DF7190DF5725B0C459D5530CFCFC5428" type="CLB-Dataset">24063</mods:identifier>
<mods:identifier id="0EF93AB78980AC62A936530628DE33AE" type="DOI">10.1080/00222933.2019.1692088</mods:identifier>
<mods:identifier id="3AF173BA724B385CCAB761CA3F628909" type="GBIF-Dataset">e6443eb6-bfcc-44a6-a547-7af9a94d4a7d</mods:identifier>
<mods:identifier id="3DF8AFD9FBE00488B83A90974C5FA303" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="3A360272B648EDEC32D633400DD21229" type="Zenodo-Dep">3665519</mods:identifier>
</mods:mods>
<treatment id="C740D6092A36BB10FEF7E62BE58928B8" ID-DOI="http://doi.org/10.5281/zenodo.3671920" ID-GBIF-Taxon="162116276" ID-Zenodo-Dep="3671920" LSID="urn:lsid:plazi:treatment:C740D6092A36BB10FEF7E62BE58928B8" httpUri="http://treatment.plazi.org/id/C740D6092A36BB10FEF7E62BE58928B8" lastPageId="85" lastPageNumber="2315" pageId="84" pageNumber="2314">
<subSubSection id="07F334942A36BB11FEF7E62BE6882FB3" pageId="84" pageNumber="2314" type="nomenclature">
<paragraph id="4F56671F2A36BB11FEF7E62BE7A22F4D" blockId="84.[341,974,853,912]" box="[341,974,853,878]" pageId="84" pageNumber="2314">
<heading id="141ED0732A36BB11FEF7E62BE7A22F4D" box="[341,974,853,878]" centered="true" fontSize="11" level="2" pageId="84" pageNumber="2314" reason="2">
3.9.:
<taxonomicName id="88E91C9C2A36BB11FE29E62BE75C2F4D" ID-CoL="35N9N" authority="Nunes &amp; Vaz-de-Mello, 2019" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[395,816,853,878]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="2314" phylum="Arthropoda" rank="species" species="barbarae" status="sp. nov." subGenus="Cephagonus">
<emphasis id="7D9DBB0D2A36BB11FE29E62BE6772F4D" box="[395,539,853,878]" italics="true" pageId="84" pageNumber="2314">Dichotomius</emphasis>
(
<emphasis id="7D9DBB0D2A36BB11FD88E62BE6D42F4D" box="[554,696,853,878]" italics="true" pageId="84" pageNumber="2314">Cephagonus</emphasis>
)
<emphasis id="7D9DBB0D2A36BB11FD6AE62BE75C2F4D" box="[712,816,853,878]" italics="true" pageId="84" pageNumber="2314">barbarae</emphasis>
</taxonomicName>
<emphasis id="7D9DBB0D2A36BB11FC95E62BE7A22F4D" bold="true" box="[823,974,853,878]" pageId="84" pageNumber="2314">
<taxonomicNameLabel id="66AE06762A36BB11FC95E62BE7A22F4D" box="[823,974,853,878]" pageId="84" pageNumber="2314" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4F56671F2A36BB11FD9DE609E6882FB3" blockId="84.[341,974,853,912]" box="[575,740,887,912]" pageId="84" pageNumber="2314">
<heading id="141ED0732A36BB11FD9DE609E6882FB3" box="[575,740,887,912]" centered="true" fontSize="11" level="2" pageId="84" pageNumber="2314" reason="5">
(
<figureCitation id="D7D27B9A2A36BB11FDE5E609E6D42FB3" box="[583,696,887,912]" captionStart="Figure 33" captionStartId="48.[160,225,1500,1523]" captionTargetBox="[163,1153,142,1469]" captionTargetId="figure@48.[163,1153,142,1469]" captionTargetPageId="48" captionText="Figure 33. D. arnaudi: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view (grey arrow indicates the position of longitudinal excavation on lateral view). D. asenjoi: (e) male dorsal habitus; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view. D. barbarae (i) male dorsal habitus; (j) paramera dorsal view (black arrow indicating slight medial constriction); (k) paramera ventral view; (l) paramera lateral view (black arrow indicates the surface of longitudinal excavation, which is smooth with fine punctures (see D. paschoali) and the differentiation of both species)." figureDoi="http://doi.org/10.5281/zenodo.3665581" httpUri="https://zenodo.org/record/3665581/files/figure.png" pageId="84" pageNumber="2314">Figure 33</figureCitation>
(i-l))
</heading>
</paragraph>
</subSubSection>
<subSubSection id="07F334942A36BB11FF02E6B0E61828B7" pageId="84" pageNumber="2314" type="diagnosis">
<paragraph id="4F56671F2A36BB11FF02E6B0E61828B7" blockId="84.[160,1156,974,1172]" pageId="84" pageNumber="2314">
<emphasis id="7D9DBB0D2A36BB11FF02E6B0E5762FC4" bold="true" box="[160,282,974,999]" italics="true" pageId="84" pageNumber="2314">Diagnosis.</emphasis>
this species is separated from other in the group by the following combined characters: larger males pronotum with a single central lobe on anterior portion of disc; elytral striae and puncture deeply impressed; males paramera with sub-apical spine (as
<taxonomicName id="88E91C9C2A36BB11FF02E149E57B286C" authorityName="Felsche" authorityYear="1901" box="[160,279,1079,1103]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="2314" phylum="Arthropoda" rank="species" species="fissiceps">
<emphasis id="7D9DBB0D2A36BB11FF02E149E57B286C" box="[160,279,1079,1103]" italics="true" pageId="84" pageNumber="2314">D. fissiceps</emphasis>
</taxonomicName>
) and paramera not narrowed posterior do sub-apical spine on dorsal view. Externallyı this species is very similar to
<taxonomicName id="88E91C9C2A36BB11FDC6E124E6A42851" authorityName="Harold" authorityYear="1869" box="[612,712,1114,1138]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="2314" phylum="Arthropoda" rank="species" species="reichei">
<emphasis id="7D9DBB0D2A36BB11FDC6E124E6A42851" box="[612,712,1114,1138]" italics="true" pageId="84" pageNumber="2314">D. reichei</emphasis>
</taxonomicName>
and
<taxonomicName id="88E91C9C2A36BB11FCA1E124E7142851" authorityName="Felsche" authorityYear="1901" box="[771,888,1114,1138]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="2314" phylum="Arthropoda" rank="species" species="fissiceps">
<emphasis id="7D9DBB0D2A36BB11FCA1E124E7142851" box="[771,888,1114,1138]" italics="true" pageId="84" pageNumber="2314">D. fissiceps</emphasis>
</taxonomicName>
andı like bothı external characters are subject to strong variation.
</paragraph>
</subSubSection>
<subSubSection id="07F334942A36BB11FF1CE1BCE7E529A4" pageId="84" pageNumber="2314" type="materials_examined">
<paragraph id="4F56671F2A36BB11FF1CE1BCE01628DF" blockId="84.[190,1157,1218,1415]" pageId="84" pageNumber="2314">
<emphasis id="7D9DBB0D2A36BB11FF1CE1BCE4A028F5" bold="true" box="[190,204,1218,1238]" pageId="84" pageNumber="2314">●</emphasis>
<materialsCitation id="FF816D422A36BB11FF7AE1BCE01D28DF" ID-GBIF-Occurrence="2570409558" collectingDate="2015-03-15" collectionCode="CEMT" country="Brazil" location="Gallery" municipality="Chapada dos Guimaraes. Vale do Jamaca" pageId="84" pageNumber="2314" specimenCount="761" specimenCount-male="1" stateProvince="Mato Grosso" typeStatus="holotype">
<typeStatus id="9052D9BD2A36BB11FF7AE1BCE53E28F9" box="[216,338,1218,1242]" pageId="84" pageNumber="2314" type="holotype">HOLOTYPE</typeStatus>
(
<specimenCount id="59EFAC962A36BB11FEC2E1BCE5F728F9" box="[352,411,1218,1242]" pageId="84" pageNumber="2314" type="male">male</specimenCount>
):
<collectingCountry id="37FE278F2A36BB11FE0FE1BCE59128F9" box="[429,509,1218,1242]" name="Brazil" pageId="84" pageNumber="2314">BRASIL</collectingCountry>
:
<emphasis id="7D9DBB0D2A36BB11FDAAE1BCE6CC28F9" bold="true" box="[520,672,1218,1242]" pageId="84" pageNumber="2314">
<collectingRegion id="8D2DA9FD2A36BB11FDAAE1BCE6CC28F9" box="[520,672,1218,1242]" country="Brazil" name="Mato Grosso" pageId="84" pageNumber="2314">Mato Grosso</collectingRegion>
</emphasis>
.
<collectingMunicipality id="AF32FD652A36BB11FD0CE1BCE0ED28F9" box="[686,1153,1218,1242]" pageId="84" pageNumber="2314">Chapada dos Guimarães. Vale do Jamacá</collectingMunicipality>
.
<geoCoordinate id="2ADD01D82A36BB11FF7AE19AE58428DF" box="[216,488,1252,1276]" pageId="84" pageNumber="2314">
15³27
<emphasis id="7D9DBB0D2A36BB11FEBAE19AE57228DF" bold="true" box="[280,286,1252,1276]" italics="true" pageId="84" pageNumber="2314">ʹ</emphasis>
51
<emphasis id="7D9DBB0D2A36BB11FE9BE19AE52928DF" box="[313,325,1252,1276]" italics="true" pageId="84" pageNumber="2314">
<emphasis id="7D9DBB0D2A36BB11FE9BE19AE55328DF" bold="true" box="[313,319,1252,1276]" italics="true" pageId="84" pageNumber="2314">ʹ</emphasis>
’
</emphasis>
Sı 55³42
<emphasis id="7D9DBB0D2A36BB11FE3DE19AE5C928DF" bold="true" box="[415,421,1252,1276]" italics="true" pageId="84" pageNumber="2314">ʹ</emphasis>
45
<emphasis id="7D9DBB0D2A36BB11FE62E19AE5A728DF" box="[448,459,1252,1276]" italics="true" pageId="84" pageNumber="2314">
<emphasis id="7D9DBB0D2A36BB11FE62E19AE5AA28DF" bold="true" box="[448,454,1252,1276]" italics="true" pageId="84" pageNumber="2314">ʹ</emphasis>
’
</emphasis>
Wı
</geoCoordinate>
<specimenCount id="59EFAC962A36BB11FE4DE19AE65028DF" box="[495,572,1252,1276]" pageId="84" pageNumber="2314" type="generic">
<quantity id="8811CAFA2A36BB11FE4DE19AE65428DF" box="[495,568,1252,1276]" metricMagnitude="2" metricUnit="m" metricValue="7.6" pageId="84" pageNumber="2314" unit="m" value="760.0">760 m</quantity>
.
</specimenCount>
<location id="4A3631C42A36BB11FDE1E19AE6FC28DF" LSID="urn:lsid:plazi:treatment:C740D6092A36BB10FEF7E62BE58928B8:4A3631C42A36BB11FDE1E19AE6FC28DF" box="[579,656,1252,1276]" country="Brazil" municipality="Chapada dos Guimaraes. Vale do Jamaca" name="Gallery" pageId="84" pageNumber="2314" stateProvince="Mato Grosso">Gallery</location>
forest. FIT.
<date id="3B5741DF2A36BB11FCB0E19AE7E628DF" box="[786,906,1252,1276]" pageId="84" pageNumber="2314" value="2015-03-15">
<collectingDate id="2B13B8372A36BB11FCB0E19AE7E628DF" box="[786,906,1252,1276]" pageId="84" pageNumber="2314" value="2015-03-15">15-iii-2015</collectingDate>
</date>
. RV
<location id="4A3631C42A36BB11FC1BE19AE06C28DF" LSID="urn:lsid:plazi:treatment:C740D6092A36BB10FEF7E62BE58928B8:4A3631C42A36BB11FC1BE19AE06C28DF" box="[953,1024,1252,1276]" country="Brazil" municipality="Chapada dos Guimaraes. Vale do Jamaca" name="Nunes" pageId="84" pageNumber="2314" stateProvince="Mato Grosso">Nunes</location>
[at
<collectionCode id="29F8FFDA2A36BB11FB8EE19AE01D28DF" box="[1068,1137,1252,1276]" pageId="84" pageNumber="2314">CEMT</collectionCode>
</materialsCitation>
].
</paragraph>
<paragraph id="4F56671F2A36BB11FF1CE076E7E529A4" blockId="84.[190,1157,1218,1415]" pageId="84" pageNumber="2314">
<emphasis id="7D9DBB0D2A36BB11FF1CE076E4A0293F" bold="true" box="[190,204,1288,1308]" pageId="84" pageNumber="2314">●</emphasis>
<typeStatus id="9052D9BD2A36BB11FF7AE079E530293C" box="[216,348,1287,1311]" pageId="84" pageNumber="2314" type="paratype">PARATYPES</typeStatus>
: same data of
<typeStatus id="9052D9BD2A36BB11FDA0E079E605293C" box="[514,617,1287,1311]" pageId="84" pageNumber="2314" type="holotype">Holotype</typeStatus>
[1 Ƌı
<specimenCount id="59EFAC962A36BB11FD0DE079E6B5293C" box="[687,729,1287,1311]" pageId="84" pageNumber="2314" type="female">1 ♀</specimenCount>
at
<collectionCode id="29F8FFDA2A36BB11FD5CE079E72F293C" box="[766,835,1287,1311]" pageId="84" pageNumber="2314">CEMT</collectionCode>
]; same data of
<typeStatus id="9052D9BD2A36BB11FC4CE079E039293C" box="[1006,1109,1287,1311]" pageId="84" pageNumber="2314" type="holotype">Holotype</typeStatus>
but
<date id="3B5741DF2A36BB11FF7AE054E5502961" box="[216,316,1322,1346]" pageId="84" pageNumber="2314" value="2013-02-09">9-ii-2013</date>
. GM Daniel [2 ƋƋ at
<collectionCode id="29F8FFDA2A36BB11FD96E054E6152961" box="[564,633,1322,1346]" pageId="84" pageNumber="2314">CEMT</collectionCode>
]; same municipality at Casa do Mel. Light.
<date id="3B5741DF2A36BB11FBFDE054E5632947" pageId="84" pageNumber="2314" value="2015-12">Xii- 2015</date>
[
<specimenCount id="59EFAC962A36BB11FE80E032E5202947" box="[290,332,1356,1380]" pageId="84" pageNumber="2314" type="female">1 ♀</specimenCount>
at
<collectionCode id="29F8FFDA2A36BB11FED5E032E5D02947" box="[375,444,1356,1380]" pageId="84" pageNumber="2314">CEMT</collectionCode>
]; Cuiabá.
<collectionCode id="29F8FFDA2A36BB11FD88E032E6152947" box="[554,633,1356,1380]" pageId="84" pageNumber="2314">PARNA</collectionCode>
Chapada dos Guimarães.
<date id="3B5741DF2A36BB11FC08E032E0492947" box="[938,1061,1356,1380]" pageId="84" pageNumber="2314" value="2008-12-13">13-xii-2008</date>
[1 Ƌ at
<collectionCode id="29F8FFDA2A36BB11FF7AE011E57129A4" box="[216,285,1391,1415]" pageId="84" pageNumber="2314">CEMT</collectionCode>
]; same data but
<date id="3B5741DF2A36BB11FE74E011E65C29A4" box="[470,560,1391,1415]" pageId="84" pageNumber="2314" value="2015-12">xii-2015</date>
. Light. A Frolov [
<specimenCount id="59EFAC962A36BB11FD4DE011E77B29A4" box="[751,791,1391,1415]" pageId="84" pageNumber="2314" type="female">1 ♀</specimenCount>
at
<collectionCode id="29F8FFDA2A36BB11FC99E011E7EC29A4" box="[827,896,1391,1415]" pageId="84" pageNumber="2314">CEMT</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="07F334942A36BB10FF02E0CAE06E2F8C" lastPageId="85" lastPageNumber="2315" pageId="84" pageNumber="2314" type="description">
<paragraph id="4F56671F2A36BB10FF02E0CAE7DB2DFD" blockId="84.[160,1156,1460,1727]" lastBlockId="85.[160,1156,142,478]" lastPageId="85" lastPageNumber="2315" pageId="84" pageNumber="2314">
<emphasis id="7D9DBB0D2A36BB11FF02E0CAE59929EE" bold="true" box="[160,501,1460,1485]" italics="true" pageId="84" pageNumber="2314">Description (Holotypeı male).</emphasis>
BL: 17 mmı PW:
<quantity id="8811CAFA2A36BB11FD11E0CBE76F29EE" box="[691,771,1461,1485]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="84" pageNumber="2314" unit="mm" value="10.0">10 mm</quantity>
. Colour: blackı shiny.
<emphasis id="7D9DBB0D2A36BB11FC4EE0CBE04629EE" bold="true" box="[1004,1066,1461,1485]" pageId="84" pageNumber="2314">Head</emphasis>
: frontoclypealı genal and inter-ocular surfaces smooth. Gena borders slightly curved inward. Cephalic carina longer than wideı conical with rounded apex.
<emphasis id="7D9DBB0D2A36BB11FCCAE084E78C2A31" bold="true" box="[872,992,1530,1554]" pageId="84" pageNumber="2314">Pronotum</emphasis>
: pronotal disc shinyı lacking punctures having a round central lobe on anterior portion. Internal eye side with coarse deeply impressed punctures. Posterior margin with two rows of ocellate puncturesı interrupted medially.
<emphasis id="7D9DBB0D2A36BB11FDB6E31CE6D82A59" bold="true" box="[532,692,1634,1658]" pageId="84" pageNumber="2314">Metasternum</emphasis>
: sides and margins of anterior lobe covered by long hair. Anterior portion of anterior lobe glabrous but bearing coarse and ocellate punctures. Setigerous punctures equally spaced at sides.
<emphasis id="7D9DBB0D2A36BB11FC39E3D9E78D2A9C" bold="true" box="[923,993,1703,1727]" pageId="84" pageNumber="2314">Elytra</emphasis>
: striae deeply impressed having fine punctures spaced by one time their diameter. First interstriae (near elytral suture) convex in relation to other interstriae which are flat. Interstriae disc lacking microsculpture with very fine puncture (20 x).
<emphasis id="7D9DBB0D2A37BB10FD3EE5AAE77D2CCF" bold="true" box="[668,785,212,236]" pageId="85" pageNumber="2315">Abdomen</emphasis>
: sides of each ventrite excavatedı glabrous. Ocellate punctures concentrated at sides. Puncture of each ventrite interrupted medially as well as groove along the posterior margin of sixth ventrite.
<emphasis id="7D9DBB0D2A37BB10FC1AE467E0422D12" bold="true" box="[952,1070,281,305]" pageId="85" pageNumber="2315">Aedeagus</emphasis>
(
<figureCitation id="D7D27B9A2A37BB10FB9EE467E4D22D77" captionStart="Figure 33" captionStartId="48.[160,225,1500,1523]" captionTargetBox="[163,1153,142,1469]" captionTargetId="figure@48.[163,1153,142,1469]" captionTargetPageId="48" captionText="Figure 33. D. arnaudi: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view (grey arrow indicates the position of longitudinal excavation on lateral view). D. asenjoi: (e) male dorsal habitus; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view. D. barbarae (i) male dorsal habitus; (j) paramera dorsal view (black arrow indicating slight medial constriction); (k) paramera ventral view; (l) paramera lateral view (black arrow indicates the surface of longitudinal excavation, which is smooth with fine punctures (see D. paschoali) and the differentiation of both species)." figureDoi="http://doi.org/10.5281/zenodo.3665581" httpUri="https://zenodo.org/record/3665581/files/figure.png" pageId="85" pageNumber="2315">Figure 33</figureCitation>
(j-l)): dorsallyı paramere apexes roundedı parallel; longitudinal excavation present taking the whole paramera length; basal-rounded excavation present; sub-apical spines presentı not narrowed posterior to spines. Laterallyı abrupt declivity from basis towards apex; having wide excavation. Ventrallyı basis with a single central tubercle; lateralrounded excavations present; subgenital plate emarginate posteriorly.
</paragraph>
<paragraph id="4F56671F2A37BB10FF02E767E06E2F8C" blockId="85.[160,1156,537,943]" pageId="85" pageNumber="2315">
<emphasis id="7D9DBB0D2A37BB10FF02E767E5AB2E11" bold="true" box="[160,455,537,562]" italics="true" pageId="85" pageNumber="2315">Morphological variation.</emphasis>
males BL: 14
<emphasis id="7D9DBB0D2A37BB10FDC7E767E61E2E12" box="[613,626,537,561]" italics="true" pageId="85" pageNumber="2315">–</emphasis>
17 mmı PW:
<quantity id="8811CAFA2A37BB10FCA6E764E71E2E11" box="[772,882,537,562]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="85" pageNumber="2315" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">
8
<emphasis id="7D9DBB0D2A37BB10FCB0E767E7732E12" box="[786,799,537,561]" italics="true" pageId="85" pageNumber="2315">–</emphasis>
10 mm
</quantity>
. Male genitalia comparison allows us to identify three phenotypes. Larger males (re-described above) bear a strong and rounded pronotal lobe and well-developed cephalic horns with similar height of pronotum. Smaller males pronotum is simply convex while cephalic horn is produced in low transversal cephalic carina. Howeverı both male phenotypes and females have similar puncture and hair distribution pattern. Females BL: 14
<emphasis id="7D9DBB0D2A37BB10FC2BE7B9E7FA2EFC" box="[905,918,711,735]" italics="true" pageId="85" pageNumber="2315">–</emphasis>
18 mmı PW:
<quantity id="8811CAFA2A37BB10FB80E7B9E0EC2EFC" box="[1058,1152,711,735]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="85" pageNumber="2315" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">
7
<emphasis id="7D9DBB0D2A37BB10FB8DE7B9E0502EFC" box="[1071,1084,711,735]" italics="true" pageId="85" pageNumber="2315">–</emphasis>
9 mm
</quantity>
. Distinguished from males as follows: Head: fronto-clypeal surface striated. Coarse puncture present posterior to cephalic carina near both eyes. Cephalic carina lowı not bifurcated at apex. Pronotum: simply convexı lacking lobesı ocellate punctures with same pattern found on males but deeply impressed. Elytra: striae deeply impressedı punctures separated by twice their diameter. Blue sheen present near basis. Abdomen: sixth ventrite four times larger than fifth and bearing a weak protuberance at mid-portion.
</paragraph>
</subSubSection>
<subSubSection id="07F334942A37BB10FF02E697E50E2806" pageId="85" pageNumber="2315" type="distribution">
<paragraph id="4F56671F2A37BB10FF02E697E50E2806" blockId="85.[160,1155,1001,1061]" pageId="85" pageNumber="2315">
<emphasis id="7D9DBB0D2A37BB10FF02E697E55F2821" bold="true" box="[160,307,1001,1026]" italics="true" pageId="85" pageNumber="2315">Distribution.</emphasis>
municipalities of Chapada dos Guimarães and Cuiabáı state of Mato Grossoı
<collectingCountry id="37FE278F2A37BB10FF02E173E4B02806" box="[160,220,1037,1061]" name="Brazil" pageId="85" pageNumber="2315">Brazil</collectingCountry>
(
<figureCitation id="D7D27B9A2A37BB10FF49E173E5382806" box="[235,340,1037,1061]" captionStart="Figure 34" captionStartId="49.[160,225,1006,1029]" captionTargetBox="[162,1151,142,975]" captionTargetId="figure@49.[162,1153,142,975]" captionTargetPageId="49" captionText="Figure 34. Distribution map of D. gardneriı D. aielloaeı D. asenjoiı D. arnaudiı D. Clypeatus and D. edmondsi." figureDoi="http://doi.org/10.5281/zenodo.3665583" httpUri="https://zenodo.org/record/3665583/files/figure.png" pageId="85" pageNumber="2315">Figure 34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="07F334942A37BB10FF02E11EE58928B8" pageId="85" pageNumber="2315" type="etymology">
<paragraph id="4F56671F2A37BB10FF02E11EE58928B8" blockId="85.[160,1155,1120,1179]" pageId="85" pageNumber="2315">
<emphasis id="7D9DBB0D2A37BB10FF02E11EE54F285A" bold="true" box="[160,291,1120,1145]" italics="true" pageId="85" pageNumber="2315">Etymology.</emphasis>
named after Bárbara Fernandes Beckerı who showed me the beauties of the
<typeStatus id="9052D9BD2A37BB10FF02E1FDE4BE28B8" box="[160,210,1155,1179]" pageId="85" pageNumber="2315">type</typeStatus>
locality and of life itself.
</paragraph>
</subSubSection>
</treatment>
</document>
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