treatments-xml/data/C7/40/D6/C740D6092A32BB14FEBEE168E49B2843.xml
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<document id="E13CA467B53AFA7E93B4268979B7918F" ID-CLB-Dataset="24063" ID-DOI="10.1080/00222933.2019.1692088" ID-GBIF-Dataset="e6443eb6-bfcc-44a6-a547-7af9a94d4a7d" ID-ISSN="1464-5262" ID-Zenodo-Dep="3665519" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1581529324984" checkinUser="valdenar" docAuthor="Nunes, Rafael V. &amp; Vaz-de-Mello, Fernando Z." docDate="2019" docId="C740D6092A32BB14FEBEE168E49B2843" docLanguage="en" docName="J_NAT_HIST.53.3738.22312351.pdf.imf" docOrigin="Journal of Natural History 53 (37)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Dichotomius (Cephagonus) clypeatus" docType="treatment" docVersion="18" lastPageNumber="2311" masterDocId="3B79AE712A62BB45FFA2E57EE46C2C23" masterDocTitle="Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)" masterLastPageNumber="2351" masterPageNumber="2231" pageNumber="2310" updateTime="1698782751744" updateUser="ExternalLinkService">
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<mods:title id="09106A1AF11B2A0CC62134E5D5135E43">Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)</mods:title>
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<mods:namePart id="DB790AAFCF5625B572D20D7CE0335744">Nunes, Rafael V.</mods:namePart>
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<treatment id="C740D6092A32BB14FEBEE168E49B2843" ID-DOI="http://doi.org/10.5281/zenodo.3671912" ID-GBIF-Taxon="162116279" ID-Zenodo-Dep="3671912" LSID="urn:lsid:plazi:treatment:C740D6092A32BB14FEBEE168E49B2843" httpUri="http://treatment.plazi.org/id/C740D6092A32BB14FEBEE168E49B2843" lastPageId="81" lastPageNumber="2311" pageId="80" pageNumber="2310">
<subSubSection id="07F334942A32BB15FEBEE168E6832870" pageId="80" pageNumber="2310" type="nomenclature">
<paragraph id="4F56671F2A32BB15FEBEE168E6832870" blockId="80.[284,1031,1046,1107]" pageId="80" pageNumber="2310">
<heading id="141ED0732A32BB15FEBEE168E6832870" centered="true" fontSize="11" level="2" pageId="80" pageNumber="2310" reason="5">
3.5.:
<taxonomicName id="88E91C9C2A32BB15FEF6E168E06B2813" ID-CoL="35N9R" authority="(Luederwaldt 1929)" baseAuthorityName="Luederwaldt" baseAuthorityYear="1929" box="[340,1031,1046,1072]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="2310" phylum="Arthropoda" rank="species" species="clypeatus" subGenus="Cephagonus">
<emphasis id="7D9DBB0D2A32BB15FEF6E168E5862813" box="[340,490,1046,1072]" italics="true" pageId="80" pageNumber="2310">Dichotomius</emphasis>
(
<emphasis id="7D9DBB0D2A32BB15FE59E168E6FC2813" box="[507,656,1046,1072]" italics="true" pageId="80" pageNumber="2310">Cephagonus</emphasis>
)
<emphasis id="7D9DBB0D2A32BB15FD02E168E77C2813" box="[672,784,1046,1072]" italics="true" pageId="80" pageNumber="2310">clypeatus</emphasis>
(
<bibRefCitation id="2B781AEE2A32BB15FCBCE168E7932813" author="Luederwaldt H" box="[798,1023,1046,1072]" pageId="80" pageNumber="2310" pagination="603 - 776" refId="ref53282" refString="Luederwaldt H. 1929. As especies brasileiras do genero. Pinotus. Rev Museu Paulista. 16: 603 - 776." type="journal article" year="1929">Luederwaldt 1929</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="D7D27B9A2A32BB15FD9EE147E6DC2870" box="[572,688,1081,1107]" captionStart="Figure 32" captionStartId="47.[160,225,1646,1669]" captionTargetBox="[166,1153,142,1614]" captionTargetId="figure@47.[163,1153,142,1614]" captionTargetPageId="47" captionText="Figure 32. D. clypeatus: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view; (e) type labels. D. aielloae: (f) Holotype dorsal habitus; (g) detail of male cephalic carina in frontal view; (h) type labels." figureDoi="http://doi.org/10.5281/zenodo.3665579" httpUri="https://zenodo.org/record/3665579/files/figure.png" pageId="80" pageNumber="2310">Figure 32</figureCitation>
(a-e))
</heading>
</paragraph>
</subSubSection>
<subSubSection id="07F334942A32BB15FF02E110E748288A" pageId="80" pageNumber="2310" type="reference_group">
<paragraph id="4F56671F2A32BB15FF02E110E748288A" blockId="80.[160,1088,1134,1193]" pageId="80" pageNumber="2310">
<taxonomicName id="88E91C9C2A32BB15FF02E110E73528A5" authority="Luederwaldt 1929" authorityName="Luederwaldt" authorityYear="1929" box="[160,857,1134,1158]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="2310" phylum="Arthropoda" rank="variety" species="bicuspis" subGenus="Cephagonus" variety="clypeatus">
<emphasis id="7D9DBB0D2A32BB15FF02E110E49C28A5" box="[160,240,1134,1158]" italics="true" pageId="80" pageNumber="2310">Pinotus</emphasis>
(
<emphasis id="7D9DBB0D2A32BB15FF5DE110E5EA28A5" box="[255,390,1134,1158]" italics="true" pageId="80" pageNumber="2310">Cephagonus</emphasis>
)
<emphasis id="7D9DBB0D2A32BB15FE37E110E58628A5" box="[405,490,1134,1158]" italics="true" pageId="80" pageNumber="2310">bicuspis</emphasis>
var.
<emphasis id="7D9DBB0D2A32BB15FD80E110E6EB28A5" box="[546,647,1134,1158]" italics="true" pageId="80" pageNumber="2310">clypeatus</emphasis>
<bibRefCitation id="2B781AEE2A32BB15FD2CE110E73528A5" author="Luederwaldt H" box="[654,857,1134,1158]" pageId="80" pageNumber="2310" pagination="603 - 776" refId="ref53282" refString="Luederwaldt H. 1929. As especies brasileiras do genero. Pinotus. Rev Museu Paulista. 16: 603 - 776." type="journal article" year="1929">Luederwaldt 1929</bibRefCitation>
</taxonomicName>
ı p. 129 (description)
<taxonomicName id="88E91C9C2A32BB15FF02E1EFE596288A" authority="Luederwaldt 1936" authorityName="Luederwaldш" authorityYear="1936" box="[160,506,1169,1193]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="2310" phylum="Arthropoda" rank="species" species="clypeatus" subGenus="Cephagonus">
<emphasis id="7D9DBB0D2A32BB15FF02E1EFE49C288A" box="[160,240,1169,1193]" italics="true" pageId="80" pageNumber="2310">Pinotus</emphasis>
(
<emphasis id="7D9DBB0D2A32BB15FF5DE1EFE5EA288A" box="[255,390,1169,1193]" italics="true" pageId="80" pageNumber="2310">Cephagonus</emphasis>
)
<emphasis id="7D9DBB0D2A32BB15FE37E1EFE596288A" box="[405,506,1169,1193]" italics="true" pageId="80" pageNumber="2310">clypeatus</emphasis>
</taxonomicName>
ı
<bibRefCitation id="2B781AEE2A32BB15FDAAE1EFE6BF288A" author="Luederwaldt H" box="[520,723,1169,1193]" pageId="80" pageNumber="2310" pagination="207 - 216" refId="ref53377" refString="Luederwaldt H. 1936. Notas complementares &quot; As especies Brasileiras do genero Pinotus &quot;. Rev Museu Paulista. 20: 207 - 216." type="journal article" year="1936">
Luederwaldt
<date id="3B5741DF2A32BB15FD3EE1EFE6BF288A" box="[668,723,1169,1193]" pageId="80" pageNumber="2310" value="1936-01">1936</date>
</bibRefCitation>
ı p. 214
</paragraph>
</subSubSection>
<subSubSection id="07F334942A32BB15FF02E1A3E6E42980" pageId="80" pageNumber="2310" type="diagnosis">
<paragraph id="4F56671F2A32BB15FF02E1A3E6E42980" blockId="80.[160,1156,1245,1443]" pageId="80" pageNumber="2310">
<emphasis id="7D9DBB0D2A32BB15FF02E1A3E57628D5" bold="true" box="[160,282,1245,1270]" italics="true" pageId="80" pageNumber="2310">Diagnosis.</emphasis>
this species is separated from other in the group by the following combined characters: males pronotal disc with a central pair of tubercles on anterior portion; elytral striae and punctures strongly impressed (as in
<taxonomicName id="88E91C9C2A32BB15FD16E05DE7762918" baseAuthorityName="Luederwaldti" baseAuthorityYear="1935" box="[692,794,1315,1339]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="2310" phylum="Arthropoda" rank="species" species="ingens">
<emphasis id="7D9DBB0D2A32BB15FD16E05DE7762918" box="[692,794,1315,1339]" italics="true" pageId="80" pageNumber="2310">D. ingens</emphasis>
</taxonomicName>
and
<taxonomicName id="88E91C9C2A32BB15FCF6E05DE7A42918" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[852,968,1315,1339]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="2310" phylum="Arthropoda" rank="species" species="arnaudi">
<emphasis id="7D9DBB0D2A32BB15FCF6E05DE7A42918" box="[852,968,1315,1339]" italics="true" pageId="80" pageNumber="2310">D. arnaudi</emphasis>
</taxonomicName>
); males cephalic horn or tubercle rounded apically (as
<taxonomicName id="88E91C9C2A32BB15FDC2E03BE6BB297E" box="[608,727,1349,1373]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="2310" phylum="Arthropoda" rank="species" species="bicuspis">
<emphasis id="7D9DBB0D2A32BB15FDC2E03BE6BB297E" box="[608,727,1349,1373]" italics="true" pageId="80" pageNumber="2310">D. bicuspis</emphasis>
</taxonomicName>
); genae borders divergentı posterior angle rounded; paramera lacking basal nor longitudinal excavations and sides of elytra subparallel in dorsal view (as in
<taxonomicName id="88E91C9C2A32BB15FDAAE0F5E6102980" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[520,636,1419,1443]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="2310" phylum="Arthropoda" rank="species" species="arnaudi">
<emphasis id="7D9DBB0D2A32BB15FDAAE0F5E6102980" box="[520,636,1419,1443]" italics="true" pageId="80" pageNumber="2310">D. arnaudi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="07F334942A32BB14FF1CE0AFE0582CEA" lastPageId="81" lastPageNumber="2311" pageId="80" pageNumber="2310" type="materials_examined">
<paragraph id="4F56671F2A32BB15FF1CE0AFE69A2A73" blockId="80.[190,1156,1488,1616]" pageId="80" pageNumber="2310">
<emphasis id="7D9DBB0D2A32BB15FF1CE0AFE4A029C6" bold="true" box="[190,204,1489,1509]" pageId="80" pageNumber="2310"></emphasis>
<typeStatus id="9052D9BD2A32BB15FF7AE0AEE53E29CB" box="[216,338,1488,1512]" pageId="80" pageNumber="2310" type="holotype">HOLOTYPE</typeStatus>
(male) (
<figureCitation id="D7D27B9A2A32BB15FE13E0AEE67329CB" box="[433,543,1488,1512]" captionStart="Figure 32" captionStartId="47.[160,225,1646,1669]" captionTargetBox="[166,1153,142,1614]" captionTargetId="figure@47.[163,1153,142,1614]" captionTargetPageId="47" captionText="Figure 32. D. clypeatus: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view; (e) type labels. D. aielloae: (f) Holotype dorsal habitus; (g) detail of male cephalic carina in frontal view; (h) type labels." figureDoi="http://doi.org/10.5281/zenodo.3665579" httpUri="https://zenodo.org/record/3665579/files/figure.png" pageId="80" pageNumber="2310">Figure 32</figureCitation>
(e)): Oxapampa
<collectingCountry id="37FE278F2A32BB15FD70E0AEE76929CB" box="[722,773,1488,1512]" name="Peru" pageId="80" pageNumber="2310">Peru</collectingCountry>
{whiteı printed}/
<taxonomicName id="88E91C9C2A32BB15FC67E0AEE50A2A28" authority="Felsche" authorityName="Felsche" authorityYear="1901" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="2310" phylum="Arthropoda" rank="species" species="quadraticeps">
<emphasis id="7D9DBB0D2A32BB15FC67E0AEE56B2A28" italics="true" pageId="80" pageNumber="2310">Pinotus quadraticeps</emphasis>
Felsche
</taxonomicName>
{whiteı handwritten}/Muséum Paris 1936 Coll. A. Boucomont {white with black marginsı printed}/
<taxonomicName id="88E91C9C2A32BB15FDBDE36BE7F32A0E" authorityName="Luederwaldt" authorityYear="1929" box="[543,927,1557,1581]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="2310" phylum="Arthropoda" rank="variety" species="bicuspis" variety="clypeatus">
<emphasis id="7D9DBB0D2A32BB15FDBDE36BE6A12A0E" box="[543,717,1557,1581]" italics="true" pageId="80" pageNumber="2310">Pinotus bicuspis</emphasis>
Har. var.
<emphasis id="7D9DBB0D2A32BB15FC98E36BE7F32A0E" box="[826,927,1557,1581]" italics="true" pageId="80" pageNumber="2310">clypeatus</emphasis>
</taxonomicName>
. Clypeatus nov. var.
<bibRefCitation id="2B781AEE2A32BB15FF7AE346E5CF2A73" author="Luederwaldt H" box="[216,419,1592,1616]" pageId="80" pageNumber="2310" pagination="297 - 304" refId="ref53332" refString="Luederwaldt H. 1931. Novos subsidios para o conhecimento do genero. Pinotus. Rev Entomol. 1: 297 - 304." type="journal article" year="1931">Luederwaldt 1931</bibRefCitation>
{whiteı handwritten} [MNHN].
</paragraph>
<paragraph id="4F56671F2A32BB15FF02E3FAE0E82A9C" blockId="80.[160,1156,1668,1727]" pageId="80" pageNumber="2310">
<emphasis id="7D9DBB0D2A32BB15FF02E3FAE5EF2ABE" bold="true" box="[160,387,1668,1693]" italics="true" pageId="80" pageNumber="2310">Non-type material.</emphasis>
<collectingCountry id="37FE278F2A32BB15FE30E3FBE6752ABE" box="[402,537,1669,1693]" name="Argentina" pageId="80" pageNumber="2310">ARGENTINA</collectingCountry>
:
<emphasis id="7D9DBB0D2A32BB15FD85E3FBE60F2ABE" bold="true" box="[551,611,1669,1693]" pageId="80" pageNumber="2310">
<collectingRegion id="8D2DA9FD2A32BB15FD85E3FBE60F2ABE" box="[551,611,1669,1693]" country="Argentina" name="Salta" pageId="80" pageNumber="2310">Salta</collectingRegion>
</emphasis>
: Sierra Tartagal.
<quantity id="8811CAFA2A32BB15FC8AE3FBE71D2ABE" box="[808,881,1669,1693]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="80" pageNumber="2310" unit="km" value="35.0">35 km</quantity>
N Tartagal. 22³19
<emphasis id="7D9DBB0D2A32BB15FBE6E3FAE0262ABF" bold="true" box="[1092,1098,1668,1692]" italics="true" pageId="80" pageNumber="2310">ʹ</emphasis>
36
<emphasis id="7D9DBB0D2A32BB15FBC7E3FAE01C2ABF" box="[1125,1136,1668,1692]" italics="true" pageId="80" pageNumber="2310">
<emphasis id="7D9DBB0D2A32BB15FBC7E3FAE0072ABF" bold="true" box="[1125,1131,1668,1692]" italics="true" pageId="80" pageNumber="2310">ʹ</emphasis>
</emphasis>
Sı 63³51
<emphasis id="7D9DBB0D2A32BB15FF42E3D9E48A2A9C" bold="true" box="[224,230,1703,1727]" italics="true" pageId="80" pageNumber="2310">ʹ</emphasis>
00
<emphasis id="7D9DBB0D2A32BB15FEA3E3D9E5612A9C" box="[257,269,1703,1727]" italics="true" pageId="80" pageNumber="2310">
<emphasis id="7D9DBB0D2A32BB15FEA3E3D9E56B2A9C" bold="true" box="[257,263,1703,1727]" italics="true" pageId="80" pageNumber="2310">ʹ</emphasis>
</emphasis>
Wı
<specimenCount id="59EFAC962A32BB15FE8DE3D9E5172A9C" box="[303,379,1703,1727]" pageId="80" pageNumber="2310" type="generic">
<quantity id="8811CAFA2A32BB15FE8DE3D9E51A2A9C" box="[303,374,1703,1727]" metricMagnitude="2" metricUnit="m" metricValue="8.74" pageId="80" pageNumber="2310" unit="m" value="874.0">874 m</quantity>
.
</specimenCount>
<date id="3B5741DF2A32BB15FE23E3D9E59F2A9C" box="[385,499,1703,1727]" pageId="80" pageNumber="2310" value="2006-11-14">14.xi.2006</date>
. Berger-Dostal [1 Ƌ at
<collectionCode id="29F8FFDA2A32BB15FD4FE3D9E75E2A9C" box="[749,818,1703,1727]" pageId="80" pageNumber="2310">CEMT</collectionCode>
];
<collectingCountry id="37FE278F2A32BB15FC9DE3D9E7F72A9C" box="[831,923,1703,1727]" name="Bolivia" pageId="80" pageNumber="2310">BOLIVIA</collectingCountry>
:
<emphasis id="7D9DBB0D2A32BB15FC07E3D9E0492A9C" bold="true" box="[933,1061,1703,1727]" pageId="80" pageNumber="2310">
<collectingRegion id="8D2DA9FD2A32BB15FC07E3D9E0492A9C" box="[933,1061,1703,1727]" country="Bolivia" name="Santa Cruz" pageId="80" pageNumber="2310">Santa Cruz</collectingRegion>
</emphasis>
: Florida.
</paragraph>
<paragraph id="4F56671F2A33BB14FF02E5F0E0582CEA" blockId="81.[160,1156,142,201]" pageId="81" pageNumber="2311">
18³08
<emphasis id="7D9DBB0D2A33BB14FF42E5F0E4892C85" box="[224,229,142,166]" italics="true" pageId="81" pageNumber="2311"></emphasis>
Sı 63³48
<emphasis id="7D9DBB0D2A33BB14FEE2E5F0E52A2C85" bold="true" box="[320,326,142,166]" italics="true" pageId="81" pageNumber="2311">ʹ</emphasis>
Wı
<quantity id="8811CAFA2A33BB14FEC8E5F0E5D62C85" box="[362,442,142,166]" metricMagnitude="3" metricUnit="m" metricValue="1.73" pageId="81" pageNumber="2311" unit="m" value="1730.0">1730m</quantity>
.
<date id="3B5741DF2A33BB14FE67E5F0E6502C85" box="[453,572,142,166]" pageId="81" pageNumber="2311" value="2006-11-19">19-xi-2006</date>
. WD Edmonds &amp; P Reyes [1 Ƌı
<specimenCount id="59EFAC962A33BB14FC3CE5F0E7AB2C85" box="[926,967,142,166]" pageId="81" pageNumber="2311" type="female">1 ♀</specimenCount>
at
<collectionCode id="29F8FFDA2A33BB14FC4FE5F0E05E2C85" box="[1005,1074,142,166]" pageId="81" pageNumber="2311">CEMT</collectionCode>
];
<collectingRegion id="8D2DA9FD2A33BB14FBE3E5F0E0EC2C85" box="[1089,1152,142,166]" country="Bolivia" name="Tarija" pageId="81" pageNumber="2311">Tarija</collectingRegion>
. Entre Riosı Quebrada del Abra.
<date id="3B5741DF2A33BB14FDA2E5CFE61E2CEA" box="[512,626,177,201]" pageId="81" pageNumber="2311" value="1998-02-25">25-ii-1998</date>
. Ayzama &amp; F Guerra [1 Ƌı
<specimenCount id="59EFAC962A33BB14FC3AE5CFE7AD2CEA" box="[920,961,177,201]" pageId="81" pageNumber="2311" type="female">1 ♀</specimenCount>
at
<collectionCode id="29F8FFDA2A33BB14FC44E5CFE0472CEA" box="[998,1067,177,201]" pageId="81" pageNumber="2311">CEMT</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="07F334942A33BB14FF02E585E5212FD1" pageId="81" pageNumber="2311" type="description">
<paragraph id="4F56671F2A33BB14FF02E585E7642EDA" blockId="81.[160,1156,251,761]" pageId="81" pageNumber="2311">
<emphasis id="7D9DBB0D2A33BB14FF02E585E52A2D37" bold="true" box="[160,326,251,276]" italics="true" pageId="81" pageNumber="2311">Redescription.</emphasis>
colour: blackı shiny.
<emphasis id="7D9DBB0D2A33BB14FD9AE582E61A2D37" bold="true" box="[568,630,252,276]" pageId="81" pageNumber="2311">Head</emphasis>
: fronto-clypealı genal and inter-ocular surfaces smooth. Gena borders not parallel. Cephalic carina conical with rounded apex.
<emphasis id="7D9DBB0D2A33BB14FBA7E461E0112D14" bold="true" box="[1029,1149,287,311]" pageId="81" pageNumber="2311">Pronotum</emphasis>
: borders smoothı not serrated. Pronotal disc shinyı lacking punctures and delimited anteriorly by two central weak tubercles.
<emphasis id="7D9DBB0D2A33BB14FD23E41AE74D2D5F" bold="true" box="[641,801,356,380]" pageId="81" pageNumber="2311">Metasternum</emphasis>
: sides and margins of anterior lobe covered by long hair. Anterior portion of anterior lobe glabrous. Setigerous punctures evenly spaced at sides.
<emphasis id="7D9DBB0D2A33BB14FE65E4D7E6602DE2" bold="true" box="[455,524,425,449]" pageId="81" pageNumber="2311">Elytra</emphasis>
: sub-parallel. Striae deeply impressed having fine weakly impressed punctures spaced by one to three times their diameter. Interstriae convexı lacking microsculpture with very fine puncture (20 x).
<emphasis id="7D9DBB0D2A33BB14FCA9E491E7EC2E24" bold="true" box="[779,896,495,519]" pageId="81" pageNumber="2311">Abdomen</emphasis>
: sides of each ventrite excavated. 1st ventrite with a single row of hair along anterior margin. Each side of ventrites 2 to 5 with groups of 1
<emphasis id="7D9DBB0D2A33BB14FD8EE74AE6552E6F" box="[556,569,564,588]" italics="true" pageId="81" pageNumber="2311"></emphasis>
5 ocelatte setigerous punctures. Puncture of each ventrite interrupted medially as well as groove along the posterior margin of sixth ventrite.
<emphasis id="7D9DBB0D2A33BB14FEABE707E5132EB2" bold="true" box="[265,383,633,657]" pageId="81" pageNumber="2311">Aedeagus</emphasis>
(
<figureCitation id="D7D27B9A2A33BB14FE30E707E66F2EB2" box="[402,515,633,657]" captionStart="Figure 32" captionStartId="47.[160,225,1646,1669]" captionTargetBox="[166,1153,142,1614]" captionTargetId="figure@47.[163,1153,142,1614]" captionTargetPageId="47" captionText="Figure 32. D. clypeatus: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view; (e) type labels. D. aielloae: (f) Holotype dorsal habitus; (g) detail of male cephalic carina in frontal view; (h) type labels." figureDoi="http://doi.org/10.5281/zenodo.3665579" httpUri="https://zenodo.org/record/3665579/files/figure.png" pageId="81" pageNumber="2311">Figure 32</figureCitation>
(b-d)). Dorsallyı paramere apexes roundedı convergent; longitudinal and basal excavations lacking; membranous portion taking more than 2/3 of dorsal surface. Laterallyı subtle declivity from basis towards apex. Ventrallyı subgenital plate widely emarginate at apex and rounded at basis.
</paragraph>
<paragraph id="4F56671F2A33BB14FF02E652E5212FD1" blockId="81.[160,1156,812,1010]" pageId="81" pageNumber="2311">
<emphasis id="7D9DBB0D2A33BB14FF02E652E5A42F66" bold="true" box="[160,456,812,837]" italics="true" pageId="81" pageNumber="2311">Morphological variation.</emphasis>
Male BL: 13
<emphasis id="7D9DBB0D2A33BB14FDFDE653E6002F66" box="[607,620,813,837]" italics="true" pageId="81" pageNumber="2311"></emphasis>
15 mmı PW:
<quantity id="8811CAFA2A33BB14FCA0E653E7082F66" box="[770,868,813,837]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="81" pageNumber="2311" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">
7
<emphasis id="7D9DBB0D2A33BB14FCB2E653E7712F66" box="[784,797,813,837]" italics="true" pageId="81" pageNumber="2311"></emphasis>
9 mm
</quantity>
. Female BL: 16 mmı PW:
<quantity id="8811CAFA2A33BB14FF02E631E4B32F44" box="[160,223,847,871]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="81" pageNumber="2311" unit="mm" value="6.0">6 mm</quantity>
and distinguished from males as follows: Head: dorsal surface striated. Clypeo-genal junction very obtuseı rounded. Cephalic carina absent forming at most a central weak knob. Pronotum: simply convex. Elytra: interstriae microsculpture chagrinated. Abdomen: sixth ventrite three times larger than fifthı pygidium swollenı lacking modifications at medial portion.
</paragraph>
</subSubSection>
<subSubSection id="07F334942A33BB14FF02E15AE49B2843" pageId="81" pageNumber="2311" type="distribution">
<paragraph id="4F56671F2A33BB14FF02E15AE49B2843" blockId="81.[160,1155,1060,1120]" pageId="81" pageNumber="2311">
<emphasis id="7D9DBB0D2A33BB14FF02E15AE55F281E" bold="true" box="[160,307,1060,1085]" italics="true" pageId="81" pageNumber="2311">Distribution.</emphasis>
<collectingCountry id="37FE278F2A33BB14FEE1E15BE5DE281E" box="[323,434,1061,1085]" name="Argentina" pageId="81" pageNumber="2311">Argentina</collectingCountry>
(
<collectingRegion id="8D2DA9FD2A33BB14FE61E15BE593281E" box="[451,511,1061,1085]" country="Argentina" name="Salta" pageId="81" pageNumber="2311">Salta</collectingRegion>
)ı
<collectingCountry id="37FE278F2A33BB14FDB1E15BE631281E" box="[531,605,1061,1085]" name="Bolivia" pageId="81" pageNumber="2311">Bolivia</collectingCountry>
(Santa Cruzı
<collectingRegion id="8D2DA9FD2A33BB14FD5EE15BE756281E" box="[764,826,1061,1085]" country="Bolivia" name="Tarija" pageId="81" pageNumber="2311">Tarija</collectingRegion>
) and
<collectingCountry id="37FE278F2A33BB14FC23E15BE7D8281E" box="[897,948,1061,1085]" name="Peru" pageId="81" pageNumber="2311">Peru</collectingCountry>
(Oxapampaı
<typeStatus id="9052D9BD2A33BB14FBF3E15BE0EF281E" box="[1105,1155,1061,1085]" pageId="81" pageNumber="2311">type</typeStatus>
locality)
</paragraph>
</subSubSection>
</treatment>
</document>