1064 lines
140 KiB
XML
1064 lines
140 KiB
XML
<document id="C667C621DA253AB140D1BC6E8AD00BCD" ID-DOI="10.5281/zenodo.180479" ID-GBIF-Dataset="d1874f7a-7d67-4f0d-8e5e-16c18ececf86" ID-ISSN="1175-5326" ID-Zenodo-Dep="180479" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459808686165" checkinUser="plazi" docAuthor="Lindquist, Evert E. & Moraza, Maria L." docDate="2008" docId="724287C1FF9CFFC63DE8FB402327BD8C" docLanguage="en" docName="zt01685p037.pdf" docOrigin="Zootaxa 1685" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Spadiseius calyptrogynae Lindquist & Moraza, 2008, new species" docType="treatment" docVersion="10" lastPageNumber="21" masterDocId="8E7BFFB9FF94FFD23D7FFF852731BA6E" masterDocTitle="A new genus of flower-dwelling melicharid mites (Acari: Mesostigmata: Ascoidea) phoretic on bats and insects in Costa Rica and Brazil" masterLastPageNumber="37" masterPageNumber="1" pageNumber="9" updateTime="1698220741931" updateUser="plazi">
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<mods:mods id="ED98AFF087140928FF71818A04E75029" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0F63656DEC7B87AA8BDE5AFF8769586D">
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<mods:title id="9072E3CE6CB7304270BEBA579C6736DC">A new genus of flower-dwelling melicharid mites (Acari: Mesostigmata: Ascoidea) phoretic on bats and insects in Costa Rica and Brazil</mods:title>
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</mods:titleInfo>
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<mods:name id="DB09FB93F442B32A27D1185C038BEBB3" type="personal">
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<mods:namePart id="BE1A80FD8A45A3E98E1D412C48567E2A">Lindquist, Evert E.</mods:namePart>
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<mods:name id="6DED0A628DB46BA445010374C9F94875" type="personal">
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<mods:roleTerm id="839679F42BCA9E49581CFBCC5E179454">Author</mods:roleTerm>
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<mods:namePart id="F04EA42C24FE46660627488513801818">Moraza, Maria L.</mods:namePart>
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<mods:typeOfResource id="C20A01FEB13C3D2ECC5DB7AD55417E02">text</mods:typeOfResource>
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<mods:title id="4332560FBD0EE6D2FF739054C3262B3C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="39CA83725E314792BB8750C7F577CBAE">
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<mods:date id="B8633A64F8956CEE761F2851C8C9E610">2008</mods:date>
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<mods:detail id="6208EB7E821E781BEC574AB8E3270F48" type="volume">
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<mods:number id="1FC81CB89B4E060652E7ACF0C38D6B28">1685</mods:number>
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<mods:start id="3CEF402A5C5753E1941FAC7B4AC3E3C3">1</mods:start>
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<mods:end id="5BDD7FDE3706A805F4D11A950DD6AD47">37</mods:end>
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<mods:classification id="B18E6A2C1173E47C80EE8CC600BEE26D">journal article</mods:classification>
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<mods:identifier id="CB3F83FDCC3CBB912DFFE034E0F2E224" type="DOI">10.5281/zenodo.180479</mods:identifier>
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<mods:identifier id="AD11623D0C8B52D38E36B166673449CB" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="3115BBF31737292FCBB6BF0CEC59E59B" type="Zenodo-Dep">180479</mods:identifier>
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<treatment id="724287C1FF9CFFC63DE8FB402327BD8C" ID-DOI="http://doi.org/10.5281/zenodo.5620716" ID-GBIF-Taxon="119353312" ID-Zenodo-Dep="5620716" LSID="urn:lsid:plazi:treatment:724287C1FF9CFFC63DE8FB402327BD8C" httpUri="http://treatment.plazi.org/id/724287C1FF9CFFC63DE8FB402327BD8C" lastPageId="20" lastPageNumber="21" pageId="8" pageNumber="9">
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<subSubSection id="B2F1655CFF9CFFDA3DE8FB40257ABF6C" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="FA5436D7FF9CFFDA3DE8FB40256EBEB1" blockId="8.[151,607,1221,1282]" box="[151,607,1221,1247]" pageId="8" pageNumber="9">
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<heading id="A11C81BBFF9CFFDA3DE8FB40256EBEB1" bold="true" box="[151,607,1221,1247]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
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<emphasis id="C89FEAC5FF9CFFDA3DE8FB40256EBEB1" bold="true" box="[151,607,1221,1247]" pageId="8" pageNumber="9">
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<taxonomicName id="3DEB4D54FF9CFFDA3DE8FB4026F9BEB1" box="[151,456,1221,1247]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="calyptrogynae" status="sp. nov.">
|
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<emphasis id="C89FEAC5FF9CFFDA3DE8FB4026F9BEB1" bold="true" box="[151,456,1221,1247]" italics="true" pageId="8" pageNumber="9">Spadiseius calyptrogynae</emphasis>
|
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</taxonomicName>
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<taxonomicNameLabel id="D3AC57BEFF9CFFDA3CB0FB40256EBEB1" box="[463,607,1221,1247]" pageId="8" pageNumber="9" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="FA5436D7FF9CFFDA3DE8FB6D257ABF6C" blockId="8.[151,607,1221,1282]" box="[151,587,1256,1282]" pageId="8" pageNumber="9">
|
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(
|
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<figureCitation id="62D02A52FF9CFFDA3DE0FB6D263EBF6C" box="[159,271,1256,1282]" captionStart="FIGURES 1 – 9" captionStartId="9.[151,269,1577,1601]" captionTargetBox="[152,1425,197,1576]" captionTargetId="figure@9.[151,1436,197,1643]" captionTargetPageId="9" captionText="FIGURES 1 – 9. Figs. 1 – 5, Spadiseius calyptrogynae n. sp., adult female: 1, subcapitulum; 2, labrum, ventral view; 3, salivary stylus; 4, tritosternum; 5, palpus, ventral view. Fig. 6, S. calyptrogynae n. sp., larva, chelicera, paraxial view. Figs. 7 – 9, Spadiseius spathiphyllae n. sp., adult female: 7, tectum; 8, chelicera, dorso-paraxial view; 9 a, chelicera, movable digit, dorsal view; 9 b, chelicera, movable digit, antiaxial-ventral view (Figs. 2 – 4, same scale)." httpUri="https://zenodo.org/record/180480/files/figure.png" pageId="8" pageNumber="9">Figs. 1–6</figureCitation>
|
||
,
|
||
<figureCitation id="62D02A52FF9CFFDA3C63FB6D2659BF6C" box="[284,360,1256,1282]" captionStart-0="FIGURES 10 – 12" captionStart-1="FIGURES 13 – 15" captionStart-2="FIGURES 16, 17" captionStart-3="FIGURE 18" captionStart-4="FIGURES 19, 20" captionStart-5="FIGURES 21, 22" captionStart-6="FIGURES 23 – 25" captionStart-7="FIGURES 26 – 29" captionStart-8="FIGURES 30 – 34" captionStartId-0="10.[151,269,1581,1605]" captionStartId-1="11.[151,269,1529,1553]" captionStartId-2="13.[151,269,1731,1755]" captionStartId-3="14.[151,255,1744,1768]" captionStartId-4="15.[151,269,1585,1609]" captionStartId-5="16.[151,269,1448,1472]" captionStartId-6="17.[151,269,1573,1597]" captionStartId-7="18.[151,269,1250,1274]" captionStartId-8="19.[151,269,1464,1488]" captionTargetBox-0="[168,1435,385,1580]" captionTargetBox-1="[156,1435,492,1528]" captionTargetBox-2="[151,1435,556,1730]" captionTargetBox-3="[153,1414,370,1743]" captionTargetBox-4="[186,1435,514,1584]" captionTargetBox-5="[264,1317,480,1446]" captionTargetBox-6="[151,1427,403,1572]" captionTargetBox-7="[151,1435,354,1192]" captionTargetBox-8="[164,1434,194,1391]" captionTargetId-0="figure@10.[151,1436,354,1612]" captionTargetId-1="figure@11.[151,1436,394,1627]" captionTargetId-2="figure@13.[151,1436,554,1771]" captionTargetId-3="figure@14.[151,1436,354,1838]" captionTargetId-4="figure@15.[151,1436,514,1681]" captionTargetId-5="figure@16.[264,1323,474,1500]" captionTargetId-6="figure@17.[151,1436,394,1601]" captionTargetId-7="figure@18.[151,1436,354,1270]" captionTargetId-8="figure@19.[151,1436,194,1440]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="13" captionTargetPageId-3="14" captionTargetPageId-4="15" captionTargetPageId-5="16" captionTargetPageId-6="17" captionTargetPageId-7="18" captionTargetPageId-8="19" captionText-0="FIGURES 10 – 12. Spadiseius calyptrogynae n. sp., larva: 10, idiosoma, dorsal aspect; 11, idiosoma, ventral aspect; 12, tectum, variations." captionText-1="FIGURES 13 – 15. Spadiseius calyptrogynae n. sp., protonymph: 13, idiosoma, dorsal aspect; 14, idiosoma, ventral aspect; 15, tectum. Setae added to larval complement denoted without parentheses." captionText-2="FIGURES 16, 17. Spadiseius calyptrogynae n. sp., deutonymph: 16, idiosoma, dorsal aspect; 17, idiosoma, ventral aspect. Setae added to protonymphal complement denoted without parentheses; supernumerary setae denoted by ‘ x’." captionText-3="FIGURE 18. Spadiseius calyptrogynae n. sp., adult female, idiosoma, dorsal aspect. Supernumerary setae denoted by" captionText-4="FIGURES 19, 20. Spadiseius calyptrogynae n. sp., adult female: 19, idiosoma, ventral aspect; 20, spermathecae, detail" captionText-5="FIGURES 21, 22. Spadiseius calyptrogynae n. sp., adult male, “ wartless ” morph: 21, idiosoma, dorsal aspect; 22, idiosoma," captionText-6="FIGURES 23 – 25. Spadiseius calyptrogynae n. sp., adult male: 23, idiosoma, “ warted morph ”, dorsal aspect; 24, peritrematal shield and detail of enlarged gland pore; 25, greatly enlarged detail of peritrematal gland pore." captionText-7="FIGURES 26 – 29. Spadiseius calyptrogynae n. sp., adult male: 26, subcapitulum; 27, chelicera and spermatodactyl, paraxial view; 28, tarsus II, anterolateral view; 29, tarsus IV, dorsal view, without pretarsus. Abbreviation: k, keel." captionText-8="FIGURES 30 – 34. Spadiseius calyptrogynae n. sp., adult female. Figs. 30 – 33, dorsal views of legs excluding tarsi: 30, leg I; 31, leg II; 32, leg III; 33, leg IV; anterolateral (al) and posterolateral (pl) faces indicated by arrows. Fig. 34, tarsus IV, dorsal view." httpUri-0="https://zenodo.org/record/180481/files/figure.png" httpUri-1="https://zenodo.org/record/180482/files/figure.png" httpUri-2="https://zenodo.org/record/180483/files/figure.png" httpUri-3="https://zenodo.org/record/180484/files/figure.png" httpUri-4="https://zenodo.org/record/180485/files/figure.png" httpUri-5="https://zenodo.org/record/180486/files/figure.png" httpUri-6="https://zenodo.org/record/180487/files/figure.png" httpUri-7="https://zenodo.org/record/180488/files/figure.png" httpUri-8="https://zenodo.org/record/180489/files/figure.png" pageId="8" pageNumber="9">10–34</figureCitation>
|
||
,
|
||
<figureCitation id="62D02A52FF9CFFDA3C0BFB6D26F1BF6C" box="[372,448,1256,1282]" captionStart="FIGURES 38 – 41" captionStartId="24.[151,269,1163,1187]" captionTargetBox="[168,1419,313,1131]" captionTargetId="figure@24.[148,1433,288,1233]" captionTargetPageId="24" captionText="FIGURES 38 – 41. Dorsal subapical femoral setae of legs. Figs. 38, 39, larval femora I – III: 38, Spadiseius calyptrogynae n. sp.; 39, Spadiseius spathiphyllae n. sp. Figs. 40, 41, protonymphal femora I – IV: 40, S. calyptrogynae n. sp.; 41, S." httpUri="https://zenodo.org/record/180491/files/figure.png" pageId="8" pageNumber="9">38, 40</figureCitation>
|
||
,
|
||
<figureCitation id="62D02A52FF9CFFDA3CB3FB6D26DDBF6C" box="[460,492,1256,1282]" captionStart="FIGURES 42 – 45" captionStartId="25.[151,269,1816,1840]" captionTargetBox="[154,1431,346,1815]" captionTargetId="figure@25.[151,1436,330,1936]" captionTargetPageId="25" captionText="FIGURES 42 – 45. Figs. 42, 43, Spadiseius spathiphyllae n. sp., deutonymph: 42, idiosoma, dorsal aspect; 43, idiosoma, ventral aspect; supernumerary setae denoted by ‘ x’. Figs. 44, 45, dorsal subapical femoral setae legs I – IV, deutonymph: 44, Spadiseius calyptrogynae n. sp.; 45, Spadiseius spathiphyllae n. sp." httpUri="https://zenodo.org/record/180492/files/figure.png" pageId="8" pageNumber="9">44</figureCitation>
|
||
,
|
||
<figureCitation id="62D02A52FF9CFFDA3C87FB6D2572BF6C" box="[504,579,1256,1282]" captionStart="FIGURES 65 – 69" captionStartId="31.[151,269,1219,1243]" captionTargetBox="[164,1431,194,1163]" captionTargetId="figure@31.[151,1436,194,1283]" captionTargetPageId="31" captionText="FIGURES 65 – 69. Figs. 65 – 67: Spadiseius spathiphyllae n. sp., adult male: 65, tarsus IV, dorsolateral view, without pretarsus; 66, 67, femora IV, dorsal view: 66, “ short-haired ” morph; 67, “ long-haired ” morph. Figs. 68, 69, Spadiseius calyptrogynae n. sp., adult male, femora IV, dorsal view: 68, “ wartless ” morph; 69, “ warted ” morph." httpUri="https://zenodo.org/record/180497/files/figure.png" pageId="8" pageNumber="9">68, 69</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B2F1655CFF9CFFDA3DE8FAB7264DBDA2" pageId="8" pageNumber="9" type="diagnosis">
|
||
<paragraph id="FA5436D7FF9CFFDA3DE8FAB7264DBDA2" blockId="8.[151,1437,1330,1996]" pageId="8" pageNumber="9">
|
||
<emphasis id="C89FEAC5FF9CFFDA3DE8FAB72695BF22" bold="true" box="[151,420,1330,1356]" pageId="8" pageNumber="9">Diagnosis. All instars</emphasis>
|
||
: leg I with apical sclerotized rim of femur, genu, tibia usually smooth, not serrated, ventrally, those of other legs weakly to clearly serrated.
|
||
<emphasis id="C89FEAC5FF9CFFDA3E43FADF24B2BF1A" bold="true" box="[828,899,1370,1396]" pageId="8" pageNumber="9">Adult</emphasis>
|
||
: peritremes of female and male of moderate thickness, and extending anteriorly well beyond humeral setae
|
||
<emphasis id="C89FEAC5FF9CFFDA3E0AFA0624BEBFF2" box="[885,911,1411,1436]" italics="true" pageId="8" pageNumber="9">r3</emphasis>
|
||
to level of setae
|
||
<emphasis id="C89FEAC5FF9CFFDA392BFA06235CBFF2" box="[1108,1133,1411,1436]" italics="true" pageId="8" pageNumber="9">s1</emphasis>
|
||
; hypostomatic setae
|
||
<emphasis id="C89FEAC5FF9CFFDA381CFA0622AABFF2" box="[1379,1435,1411,1436]" italics="true" pageId="8" pageNumber="9">hyp4</emphasis>
|
||
shorter than
|
||
<emphasis id="C89FEAC5FF9CFFDA3C55FA2E269ABFAA" box="[298,427,1451,1476]" italics="true" pageId="8" pageNumber="9">hyp1–hyp3</emphasis>
|
||
.
|
||
<emphasis id="C89FEAC5FF9CFFDA3CC5FA2F256DBFAA" bold="true" box="[442,604,1450,1476]" pageId="8" pageNumber="9">Adult female</emphasis>
|
||
: marginal setae
|
||
<emphasis id="C89FEAC5FF9CFFDA3E62FA2E2451BFAA" box="[797,864,1451,1476]" italics="true" pageId="8" pageNumber="9">r2–r4</emphasis>
|
||
and sometimes
|
||
<emphasis id="C89FEAC5FF9CFFDA3961FA2E2309BFAA" box="[1054,1080,1451,1476]" italics="true" pageId="8" pageNumber="9">r5</emphasis>
|
||
on edge of dorsal shield; subcapitulum with pair of gland-like structures between insertions of setae
|
||
<emphasis id="C89FEAC5FF9CFFDA3E88FA562301BF82" box="[1015,1072,1491,1516]" italics="true" pageId="8" pageNumber="9">hyp1</emphasis>
|
||
, and with setae
|
||
<emphasis id="C89FEAC5FF9CFFDA3988FA562201BF82" box="[1271,1328,1491,1516]" italics="true" pageId="8" pageNumber="9">hyp4</emphasis>
|
||
abruptly swollen basally.
|
||
<emphasis id="C89FEAC5FF9CFFDA3C24FA7F26D6BC7A" bold="true" box="[347,487,1530,1556]" pageId="8" pageNumber="9">Adult male</emphasis>
|
||
: dorsal shield with setae in central region short, simple, often inserted individually on small subcircular warts, and with setae
|
||
<emphasis id="C89FEAC5FF9CFFDA3FC3F9A625E2BC52" box="[700,723,1571,1596]" italics="true" pageId="8" pageNumber="9">j1</emphasis>
|
||
and a variable number of peripheral setae of
|
||
<emphasis id="C89FEAC5FF9CFFDA399AF9A6223EBC52" box="[1253,1295,1571,1596]" italics="true" pageId="8" pageNumber="9">z–Z</emphasis>
|
||
, s–S and
|
||
<emphasis id="C89FEAC5FF9CFFDA38FDF9A62798BC0A" italics="true" pageId="8" pageNumber="9">r– R</emphasis>
|
||
series forming a corona of enlarged, thickened, apically capitate setae; presternal area with pair of small platelets; peritrematal shield with gland pore greatly enlarged at level between coxae II and III; spermatodactyl divided near base into a shorter pointed process and a longer, straight, apically thickened terminus; leg IV lacking spur on basitarsus.
|
||
<emphasis id="C89FEAC5FF9CFFDA3CABF9472546BCB2" bold="true" box="[468,631,1730,1756]" pageId="8" pageNumber="9">Deutonymph</emphasis>
|
||
: coxa IV seta short, half as long as setae on coxae III; legs I–IV with femoral seta
|
||
<emphasis id="C89FEAC5FF9CFFDA3C4EF96E2656BD6A" box="[305,359,1771,1796]" italics="true" pageId="8" pageNumber="9">ad-1</emphasis>
|
||
thicker than adjacent setae, and clearly spatulate on IV but not on I–II and sometimes III.
|
||
<emphasis id="C89FEAC5FF9CFFDA3DE8F8972605BD42" bold="true" box="[151,308,1810,1836]" pageId="8" pageNumber="9">Protonymph</emphasis>
|
||
: pygidial shield usually with poroid-bearing hump or tubercle laterad seta
|
||
<emphasis id="C89FEAC5FF9CFFDA39C1F89623EABD42" box="[1214,1243,1811,1836]" italics="true" pageId="8" pageNumber="9">S5</emphasis>
|
||
; legs I–IV with femoral seta
|
||
<emphasis id="C89FEAC5FF9CFFDA3C4CF8BE2659BD3A" box="[307,360,1851,1876]" italics="true" pageId="8" pageNumber="9">ad-1</emphasis>
|
||
not spatulate but slightly thicker than adjacent setae.
|
||
<emphasis id="C89FEAC5FF9CFFDA3E8EF8BF230CBD3A" bold="true" box="[1009,1085,1850,1876]" pageId="8" pageNumber="9">Larva</emphasis>
|
||
: podonotal shield faintly lineated laterally; pygidial shield sparsely puncate, with medial projection of anterior margin not extending beyond level of setal alveoli
|
||
<emphasis id="C89FEAC5FF9CFFDA3C9AF80E2530BDCA" box="[485,513,1931,1956]" italics="true" pageId="8" pageNumber="9">J2</emphasis>
|
||
; legs I–III with femoral seta
|
||
<emphasis id="C89FEAC5FF9CFFDA3E2DF80E24B9BDCA" box="[850,904,1931,1956]" italics="true" pageId="8" pageNumber="9">ad-1</emphasis>
|
||
not spatulate but less tapered, slightly thicker than adjacent setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="AE94665FFF9DFFDB3DE8F9AC2385BCC6" httpUri="https://zenodo.org/record/180480/files/figure.png" pageId="9" pageNumber="10" targetBox="[152,1425,197,1576]" targetPageId="9">
|
||
<paragraph id="FA5436D7FF9DFFDB3DE8F9AC2385BCC6" blockId="9.[151,1436,1577,1704]" pageId="9" pageNumber="10">
|
||
<emphasis id="C89FEAC5FF9DFFDB3DE8F9AC2675BC2F" bold="true" box="[151,324,1577,1601]" pageId="9" pageNumber="10">FIGURES 1–9.</emphasis>
|
||
Figs. 1–5,
|
||
<taxonomicName id="3DEB4D54FF9DFFDB3CBEF9AF25E2BC2F" box="[449,723,1578,1601]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="calyptrogynae" status="sp. nov.">
|
||
<emphasis id="C89FEAC5FF9DFFDB3CBEF9AF25E2BC2F" box="[449,723,1578,1601]" italics="true" pageId="9" pageNumber="10">Spadiseius calyptrogynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D3AC57BEFF9DFFDB3FA3F9AC2428BC2F" box="[732,793,1577,1601]" pageId="9" pageNumber="10" rank="species">
|
||
<emphasis id="C89FEAC5FF9DFFDB3FA3F9AC2428BC2F" bold="true" box="[732,793,1577,1601]" pageId="9" pageNumber="10">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult female: 1, subcapitulum; 2, labrum, ventral view; 3, salivary stylus; 4, tritosternum; 5, palpus, ventral view. Fig. 6,
|
||
<taxonomicName id="3DEB4D54FF9DFFDB3E39F9C924CABC0D" box="[838,1019,1612,1635]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="calyptrogynae" status="sp. nov.">
|
||
<emphasis id="C89FEAC5FF9DFFDB3E39F9C924CABC0D" box="[838,1019,1612,1635]" italics="true" pageId="9" pageNumber="10">S. calyptrogynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D3AC57BEFF9DFFDB397CF9C92370BC0A" box="[1027,1089,1612,1636]" pageId="9" pageNumber="10" rank="species">
|
||
<emphasis id="C89FEAC5FF9DFFDB397CF9C92370BC0A" bold="true" box="[1027,1089,1612,1636]" pageId="9" pageNumber="10">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, larva, chelicera, paraxial view. Figs. 7–9,
|
||
<taxonomicName id="3DEB4D54FF9DFFDB3C78F9EA253EBCE8" box="[263,527,1647,1670]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="spathiphyllae" status="sp. nov.">
|
||
<emphasis id="C89FEAC5FF9DFFDB3C78F9EA253EBCE8" box="[263,527,1647,1670]" italics="true" pageId="9" pageNumber="10">Spadiseius spathiphyllae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D3AC57BEFF9DFFDB3F68F9EB2562BCE8" box="[535,595,1646,1670]" pageId="9" pageNumber="10" rank="species">
|
||
<emphasis id="C89FEAC5FF9DFFDB3F68F9EB2562BCE8" bold="true" box="[535,595,1646,1670]" pageId="9" pageNumber="10">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult female: 7, tectum; 8, chelicera, dorso-paraxial view; 9a, chelicera, movable digit, dorsal view; 9b, chelicera, movable digit, antiaxial-ventral view (Figs. 2–4, same scale).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="B2F1655CFF9DFFC13DB9F9592436BD6D" lastPageId="19" lastPageNumber="20" pageId="9" pageNumber="10" type="description">
|
||
<paragraph id="FA5436D7FF9DFFD83DB9F9592564BB44" blockId="9.[151,1436,1756,2022]" lastBlockId="10.[151,1437,152,298]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
|
||
<emphasis id="C89FEAC5FF9DFFDB3DB9F9592519BC9B" bold="true" box="[198,552,1756,1782]" pageId="9" pageNumber="10">Description. ADULT FEMALE</emphasis>
|
||
. Dorsal shield 440–540 long, 275–325 at its greatest width at level of setae
|
||
<emphasis id="C89FEAC5FF9DFFDB3DE8F8812780BD73" box="[151,177,1796,1821]" italics="true" pageId="9" pageNumber="10">r4</emphasis>
|
||
(n = 8); shield lightly evenly reticulate over nearly entire surface (
|
||
<figureCitation id="62D02A52FF9DFFDB3EBBF881232ABD70" box="[964,1051,1796,1822]" captionStart="FIGURE 18" captionStartId="14.[151,255,1744,1768]" captionTargetBox="[153,1414,370,1743]" captionTargetId="figure@14.[151,1436,354,1838]" captionTargetPageId="14" captionText="FIGURE 18. Spadiseius calyptrogynae n. sp., adult female, idiosoma, dorsal aspect. Supernumerary setae denoted by" httpUri="https://zenodo.org/record/180484/files/figure.png" pageId="9" pageNumber="10">Fig. 18</figureCitation>
|
||
); podonotal region with longitudinal distance from alveoli of
|
||
<emphasis id="C89FEAC5FF9DFFDB3C81F8A92524BD2B" box="[510,533,1836,1861]" italics="true" pageId="9" pageNumber="10">j1</emphasis>
|
||
to
|
||
<emphasis id="C89FEAC5FF9DFFDB3F42F8A92562BD2B" box="[573,595,1836,1861]" italics="true" pageId="9" pageNumber="10">j5</emphasis>
|
||
slightly longer (about 1.1) than that from
|
||
<emphasis id="C89FEAC5FF9DFFDB3932F8A92352BD2B" box="[1101,1123,1836,1861]" italics="true" pageId="9" pageNumber="10">j5</emphasis>
|
||
to
|
||
<emphasis id="C89FEAC5FF9DFFDB39F3F8A92393BD2B" box="[1164,1186,1836,1861]" italics="true" pageId="9" pageNumber="10">j6</emphasis>
|
||
. Peritrematal shields uniting with dorsal shield at level between setae
|
||
<emphasis id="C89FEAC5FF9DFFDB3FB4F8D1243FBD03" box="[715,782,1876,1901]" italics="true" pageId="9" pageNumber="10">s1–s2</emphasis>
|
||
; peritremes extending to level of setae
|
||
<emphasis id="C89FEAC5FF9DFFDB39A9F8D123C1BD03" box="[1238,1264,1876,1901]" italics="true" pageId="9" pageNumber="10">s1</emphasis>
|
||
. Body dorsum with approximately 52–54 pairs of setae, most inserted on dorsal shield, but variably
|
||
<emphasis id="C89FEAC5FF9DFFDB39F4F8F92397BDFB" box="[1163,1190,1916,1941]" italics="true" pageId="9" pageNumber="10">r5</emphasis>
|
||
and consistenly
|
||
<emphasis id="C89FEAC5FF9DFFDB3813F8F9278EBDD3" italics="true" pageId="9" pageNumber="10">R1– R6,</emphasis>
|
||
and 0 to 3
|
||
<emphasis id="C89FEAC5FF9DFFDB3C3CF8212648BDD3" box="[323,377,1956,1981]" italics="true" pageId="9" pageNumber="10">UR–</emphasis>
|
||
setae inserted on soft cuticle; dorsal shield with 3 to 5 extra pairs of setae amidst
|
||
<emphasis id="C89FEAC5FF9DFFDB384BF821220DBDD3" box="[1332,1340,1956,1981]" italics="true" pageId="9" pageNumber="10">j</emphasis>
|
||
,
|
||
<emphasis id="C89FEAC5FF9DFFDB3833F8212266BDD3" box="[1356,1367,1956,1981]" italics="true" pageId="9" pageNumber="10">z</emphasis>
|
||
and
|
||
<emphasis id="C89FEAC5FF9DFFDB38EFF82122AABDD3" box="[1424,1435,1956,1981]" italics="true" pageId="9" pageNumber="10">s</emphasis>
|
||
series on podonotal region and several paired or asymmetrically unpaired setae amidst
|
||
<emphasis id="C89FEAC5FF9DFFDB39F5F84923A6BD8B" box="[1162,1175,1996,2021]" italics="true" pageId="9" pageNumber="10">J</emphasis>
|
||
series on opisthonotal region. Dorsal setae uniform in simple but slightly curved, scimitar-like form (most of them slightly swollen at base) and moderately short length (17–23),
|
||
<emphasis id="C89FEAC5FF9EFFD83FCEFF4525FFBAB7" box="[689,718,192,217]" italics="true" pageId="10" pageNumber="11">S5</emphasis>
|
||
,
|
||
<emphasis id="C89FEAC5FF9EFFD83FA3FF4525CDBAB7" box="[732,764,192,217]" italics="true" pageId="10" pageNumber="11">Z5</emphasis>
|
||
slightly largest (23–25),
|
||
<emphasis id="C89FEAC5FF9EFFD8395EFF45230DBAB7" box="[1057,1084,192,217]" italics="true" pageId="10" pageNumber="11">J5</emphasis>
|
||
clearly smallest (10). Anterodorsal expansions of exopodal plates above legs I formed as lineated plates (not illustrated, difficult to discern) touching vertex of dorsal shield.
|
||
</paragraph>
|
||
<caption id="AE94665FFF9EFFD83DE8F9A8266CBC06" httpUri="https://zenodo.org/record/180481/files/figure.png" pageId="10" pageNumber="11" targetBox="[168,1435,385,1580]" targetPageId="10">
|
||
<paragraph id="FA5436D7FF9EFFD83DE8F9A8266CBC06" blockId="10.[151,1436,1581,1640]" pageId="10" pageNumber="11">
|
||
<emphasis id="C89FEAC5FF9EFFD83DE8F9A8266FBC2B" bold="true" box="[151,350,1581,1605]" pageId="10" pageNumber="11">FIGURES 10–12.</emphasis>
|
||
<taxonomicName id="3DEB4D54FF9EFFD83C1AF9AB2546BC2B" box="[357,631,1582,1605]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="calyptrogynae" status="sp. nov.">
|
||
<emphasis id="C89FEAC5FF9EFFD83C1AF9AB2546BC2B" box="[357,631,1582,1605]" italics="true" pageId="10" pageNumber="11">Spadiseius calyptrogynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D3AC57BEFF9EFFD83F01F9A8258ABC2B" box="[638,699,1581,1605]" pageId="10" pageNumber="11" rank="species">
|
||
<emphasis id="C89FEAC5FF9EFFD83F01F9A8258ABC2B" bold="true" box="[638,699,1581,1605]" pageId="10" pageNumber="11">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, larva: 10, idiosoma, dorsal aspect; 11, idiosoma, ventral aspect; 12, tectum, variations.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FA5436D7FF9EFFD93DB9F91E242FBB3C" blockId="10.[151,1437,1691,2037]" lastBlockId="11.[151,1436,152,338]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
|
||
Tritosternum (
|
||
<figureCitation id="62D02A52FF9EFFD83C10F91E2687BCDB" box="[367,438,1691,1717]" captionStart="FIGURES 1 – 9" captionStartId="9.[151,269,1577,1601]" captionTargetBox="[152,1425,197,1576]" captionTargetId="figure@9.[151,1436,197,1643]" captionTargetPageId="9" captionText="FIGURES 1 – 9. Figs. 1 – 5, Spadiseius calyptrogynae n. sp., adult female: 1, subcapitulum; 2, labrum, ventral view; 3, salivary stylus; 4, tritosternum; 5, palpus, ventral view. Fig. 6, S. calyptrogynae n. sp., larva, chelicera, paraxial view. Figs. 7 – 9, Spadiseius spathiphyllae n. sp., adult female: 7, tectum; 8, chelicera, dorso-paraxial view; 9 a, chelicera, movable digit, dorsal view; 9 b, chelicera, movable digit, antiaxial-ventral view (Figs. 2 – 4, same scale)." httpUri="https://zenodo.org/record/180480/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
|
||
) with length of trapezoidal base subequal to its basal width (16–17); laciniae free for about 0.6 of total length (70–75 excluding base). Presternal area weakly sclerotized, insensibly fused with sternal shield, and with pair of small platelets amidst lineation. Sternal shield (
|
||
<figureCitation id="62D02A52FF9EFFD8391DF96E238ABD6B" box="[1122,1211,1771,1797]" captionStart="FIGURES 19, 20" captionStartId="15.[151,269,1585,1609]" captionTargetBox="[186,1435,514,1584]" captionTargetId="figure@15.[151,1436,514,1681]" captionTargetPageId="15" captionText="FIGURES 19, 20. Spadiseius calyptrogynae n. sp., adult female: 19, idiosoma, ventral aspect; 20, spermathecae, detail" httpUri="https://zenodo.org/record/180485/files/figure.png" pageId="10" pageNumber="11">Fig. 19</figureCitation>
|
||
) unornamented or faintly lineated along lateral margins, weakly sclerotized over much of surface but faint or desclerotized posterior to level of second poroids, posterior margin faint, slightly concave; setae
|
||
<emphasis id="C89FEAC5FF9EFFD83930F8B92393BD3B" box="[1103,1186,1852,1877]" italics="true" pageId="10" pageNumber="11">st1–st2</emphasis>
|
||
and first and second pairs of poroids inserted on sclerotized portion of shield,
|
||
<emphasis id="C89FEAC5FF9EFFD83E48F8E12468BD13" box="[823,857,1892,1917]" italics="true" pageId="10" pageNumber="11">st3</emphasis>
|
||
inserted on desclerotized portion;
|
||
<emphasis id="C89FEAC5FF9EFFD8398FF8E12273BD13" box="[1264,1346,1892,1917]" italics="true" pageId="10" pageNumber="11">st1–st3</emphasis>
|
||
similar in length (30–35), slightly longer than
|
||
<emphasis id="C89FEAC5FF9EFFD83F27F809254ABDCB" box="[600,635,1932,1957]" italics="true" pageId="10" pageNumber="11">st4</emphasis>
|
||
(25–27) and
|
||
<emphasis id="C89FEAC5FF9EFFD83E6AF8092409BDCB" box="[789,824,1932,1957]" italics="true" pageId="10" pageNumber="11">st5</emphasis>
|
||
(20) on soft cuticle. Endopodal strips faintly formed alongside coxae III and IV. Genital shield unornamented, its lateral margins slightly concave at level of genital setae. Anal shield weakly reticulate-lineate, longer (90) than greatest width at level of anterior edge of anus (75); postanal seta (20–22) slightly longer than para-anal setae (17–18). Ten pairs of opisthogastric setae (
|
||
<emphasis id="C89FEAC5FF9FFFD93DDEFF4526BBBAB7" box="[161,394,192,217]" italics="true" pageId="11" pageNumber="12">JV1–JV5, ZV1–ZV5</emphasis>
|
||
, excluding 1 or 2 pairs of adjacent
|
||
<emphasis id="C89FEAC5FF9FFFD93E58FF45246CBAB7" box="[807,861,192,217]" italics="true" pageId="11" pageNumber="12">UR–</emphasis>
|
||
setae) on soft cuticle around anal shield, all similarly short (16–18) except for slightly larger
|
||
<emphasis id="C89FEAC5FF9FFFD93FDAFF6D25E3BB6F" box="[677,722,232,257]" italics="true" pageId="11" pageNumber="12">JV5</emphasis>
|
||
,
|
||
<emphasis id="C89FEAC5FF9FFFD93F9DFF6D2422BB6F" box="[738,787,232,257]" italics="true" pageId="11" pageNumber="12">ZV5</emphasis>
|
||
(22–24). Metapodal plates undivided, usually elongate triangular, pointed posteriorly, about 25 long, 12 wide. Spermathecal apparatus difficult to discern, with convoluted conduit and weakly sclerotized calyx (
|
||
<figureCitation id="62D02A52FF9FFFD93FC7FEBD2421BB3C" box="[696,784,312,338]" captionStart="FIGURES 19, 20" captionStartId="15.[151,269,1585,1609]" captionTargetBox="[186,1435,514,1584]" captionTargetId="figure@15.[151,1436,514,1681]" captionTargetPageId="15" captionText="FIGURES 19, 20. Spadiseius calyptrogynae n. sp., adult female: 19, idiosoma, ventral aspect; 20, spermathecae, detail" httpUri="https://zenodo.org/record/180485/files/figure.png" pageId="11" pageNumber="12">Fig. 20</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="AE94665FFF9FFFD93DE8FA7C233DBC5D" httpUri="https://zenodo.org/record/180482/files/figure.png" pageId="11" pageNumber="12" targetBox="[156,1435,492,1528]" targetPageId="11">
|
||
<paragraph id="FA5436D7FF9FFFD93DE8FA7C233DBC5D" blockId="11.[151,1436,1529,1587]" pageId="11" pageNumber="12">
|
||
<emphasis id="C89FEAC5FF9FFFD93DE8FA7C2653BC7F" bold="true" box="[151,354,1529,1553]" pageId="11" pageNumber="12">FIGURES 13–15.</emphasis>
|
||
<taxonomicName id="3DEB4D54FF9FFFD93C13FA7F25B1BC7F" box="[364,640,1530,1553]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="calyptrogynae" status="sp. nov.">
|
||
<emphasis id="C89FEAC5FF9FFFD93C13FA7F25B1BC7F" box="[364,640,1530,1553]" italics="true" pageId="11" pageNumber="12">Spadiseius calyptrogynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D3AC57BEFF9FFFD93FF3FA7C25FDBC7F" box="[652,716,1529,1553]" pageId="11" pageNumber="12" rank="species">
|
||
<emphasis id="C89FEAC5FF9FFFD93FF3FA7C25FDBC7F" bold="true" box="[652,716,1529,1553]" pageId="11" pageNumber="12">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, protonymph: 13, idiosoma, dorsal aspect; 14, idiosoma, ventral aspect; 15, tectum. Setae added to larval complement denoted without parentheses.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FA5436D7FF9FFFDE3DB9F9E226F5BB6C" blockId="11.[151,1436,1639,2025]" lastBlockId="12.[151,1437,152,1218]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||
Tectum with anterior margin smooth, sharply pointed medially (as in
|
||
<figureCitation id="62D02A52FF9FFFD93968F9E22353BCEF" box="[1047,1122,1639,1665]" captionStart="FIGURES 1 – 9" captionStartId="9.[151,269,1577,1601]" captionTargetBox="[152,1425,197,1576]" captionTargetId="figure@9.[151,1436,197,1643]" captionTargetPageId="9" captionText="FIGURES 1 – 9. Figs. 1 – 5, Spadiseius calyptrogynae n. sp., adult female: 1, subcapitulum; 2, labrum, ventral view; 3, salivary stylus; 4, tritosternum; 5, palpus, ventral view. Fig. 6, S. calyptrogynae n. sp., larva, chelicera, paraxial view. Figs. 7 – 9, Spadiseius spathiphyllae n. sp., adult female: 7, tectum; 8, chelicera, dorso-paraxial view; 9 a, chelicera, movable digit, dorsal view; 9 b, chelicera, movable digit, antiaxial-ventral view (Figs. 2 – 4, same scale)." httpUri="https://zenodo.org/record/180480/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
|
||
). Fixed chela with 2 or 3 minute teeth beside offset tooth subapically (as in
|
||
<figureCitation id="62D02A52FF9FFFD93F94F90A2472BCC7" box="[747,835,1679,1705]" captionStart="FIGURES 46 – 50" captionStartId="27.[151,269,1773,1797]" captionTargetBox="[186,1409,183,1772]" captionTargetId="figure@27.[186,1411,178,1937]" captionTargetPageId="27" captionText="FIGURES 46 – 50. Spadiseius spathiphyllae n. sp., adult female: 46, idiosoma, dorsal aspect, supernumerary setae denoted by ‘ x’; 47, idiosoma, ventral aspect; 48, spermathecae, detail of calyx; 49, position of spermatheca at base of leg IV;" httpUri="https://zenodo.org/record/180493/files/figure.png" pageId="11" pageNumber="12">Fig. 50</figureCitation>
|
||
); movable chela (length 40) usually weakly bidentate, the more apical tooth hardly discernible; antiaxial face of movable chela with sharply defined ridge along its basal half; ventral mucro slender, attenuate, surpassing basal half of chela (as in
|
||
<figureCitation id="62D02A52FF9FFFD93914F95A23EABC97" box="[1131,1243,1759,1785]" captionStart="FIGURES 1 – 9" captionStartId="9.[151,269,1577,1601]" captionTargetBox="[152,1425,197,1576]" captionTargetId="figure@9.[151,1436,197,1643]" captionTargetPageId="9" captionText="FIGURES 1 – 9. Figs. 1 – 5, Spadiseius calyptrogynae n. sp., adult female: 1, subcapitulum; 2, labrum, ventral view; 3, salivary stylus; 4, tritosternum; 5, palpus, ventral view. Fig. 6, S. calyptrogynae n. sp., larva, chelicera, paraxial view. Figs. 7 – 9, Spadiseius spathiphyllae n. sp., adult female: 7, tectum; 8, chelicera, dorso-paraxial view; 9 a, chelicera, movable digit, dorsal view; 9 b, chelicera, movable digit, antiaxial-ventral view (Figs. 2 – 4, same scale)." httpUri="https://zenodo.org/record/180480/files/figure.png" pageId="11" pageNumber="12">Figs. 8, 9</figureCitation>
|
||
). Corniculi with lateral edges convergent apically but with medial edges nearly parallel such that apices usually separate; internal malae slender, blade-like, shorter than corniculi, their lateral margins smooth (
|
||
<figureCitation id="62D02A52FF9FFFD93924F8AA2391BD27" box="[1115,1184,1839,1865]" captionStart="FIGURES 1 – 9" captionStartId="9.[151,269,1577,1601]" captionTargetBox="[152,1425,197,1576]" captionTargetId="figure@9.[151,1436,197,1643]" captionTargetPageId="9" captionText="FIGURES 1 – 9. Figs. 1 – 5, Spadiseius calyptrogynae n. sp., adult female: 1, subcapitulum; 2, labrum, ventral view; 3, salivary stylus; 4, tritosternum; 5, palpus, ventral view. Fig. 6, S. calyptrogynae n. sp., larva, chelicera, paraxial view. Figs. 7 – 9, Spadiseius spathiphyllae n. sp., adult female: 7, tectum; 8, chelicera, dorso-paraxial view; 9 a, chelicera, movable digit, dorsal view; 9 b, chelicera, movable digit, antiaxial-ventral view (Figs. 2 – 4, same scale)." httpUri="https://zenodo.org/record/180480/files/figure.png" pageId="11" pageNumber="12">Fig. 1</figureCitation>
|
||
). Deutosternum with 7 or 8 transverse rows of denticles, evenly multidenticulate (7–15 denticles), none distinctly widened. Subcapitulum with pair of gland-like structures between bases of anterior subcapitular setae (
|
||
<figureCitation id="62D02A52FF9FFFD939CFF8FA23C6BDF7" box="[1200,1271,1919,1945]" captionStart="FIGURES 1 – 9" captionStartId="9.[151,269,1577,1601]" captionTargetBox="[152,1425,197,1576]" captionTargetId="figure@9.[151,1436,197,1643]" captionTargetPageId="9" captionText="FIGURES 1 – 9. Figs. 1 – 5, Spadiseius calyptrogynae n. sp., adult female: 1, subcapitulum; 2, labrum, ventral view; 3, salivary stylus; 4, tritosternum; 5, palpus, ventral view. Fig. 6, S. calyptrogynae n. sp., larva, chelicera, paraxial view. Figs. 7 – 9, Spadiseius spathiphyllae n. sp., adult female: 7, tectum; 8, chelicera, dorso-paraxial view; 9 a, chelicera, movable digit, dorsal view; 9 b, chelicera, movable digit, antiaxial-ventral view (Figs. 2 – 4, same scale)." httpUri="https://zenodo.org/record/180480/files/figure.png" pageId="11" pageNumber="12">Fig. 1</figureCitation>
|
||
); setae finely tapered, medial pair longest; capitular pair abruptly thickened basally, shorter (12–15) than anterior and medial pairs (27 and 35–40, respectively). Subcapitulum without ridge extending on either side laterally from bases of capitular setae. Palptrochanter setae slender, inner seta slightly longer than outer one; palpgenu with seta
|
||
<emphasis id="C89FEAC5FF98FFDE3DB1FF4527CCBAB7" box="[206,253,192,217]" italics="true" pageId="12" pageNumber="13">al-2</emphasis>
|
||
similar in spatulate form to palpfemoral seta
|
||
<emphasis id="C89FEAC5FF98FFDE3E5AFF45240DBAB7" box="[805,828,192,217]" italics="true" pageId="12" pageNumber="13">al</emphasis>
|
||
; apical sclerotized rim of palpfemur and palpgenu smooth ventrally (
|
||
<figureCitation id="62D02A52FF98FFDE3C13FF6D2685BB6C" box="[364,436,232,258]" captionStart="FIGURES 1 – 9" captionStartId="9.[151,269,1577,1601]" captionTargetBox="[152,1425,197,1576]" captionTargetId="figure@9.[151,1436,197,1643]" captionTargetPageId="9" captionText="FIGURES 1 – 9. Figs. 1 – 5, Spadiseius calyptrogynae n. sp., adult female: 1, subcapitulum; 2, labrum, ventral view; 3, salivary stylus; 4, tritosternum; 5, palpus, ventral view. Fig. 6, S. calyptrogynae n. sp., larva, chelicera, paraxial view. Figs. 7 – 9, Spadiseius spathiphyllae n. sp., adult female: 7, tectum; 8, chelicera, dorso-paraxial view; 9 a, chelicera, movable digit, dorsal view; 9 b, chelicera, movable digit, antiaxial-ventral view (Figs. 2 – 4, same scale)." httpUri="https://zenodo.org/record/180480/files/figure.png" pageId="12" pageNumber="13">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FA5436D7FF98FFDE3DB9FE952207B8D4" blockId="12.[151,1437,152,1218]" pageId="12" pageNumber="13">
|
||
Legs IV the longest (415–480, excluding pretarsus) of legs, about 0.9 as long as dorsal shield; other leg lengths: I 385–430, II 335–375, III 350–400. Tarsi II–IV with pair of apical setal processes (
|
||
<emphasis id="C89FEAC5FF98FFDE39AFFEBD2278BB3F" box="[1232,1353,312,337]" italics="true" pageId="12" pageNumber="13">ad-1, pd-1</emphasis>
|
||
) about one-half as long as length of pretarsi to base of claws (
|
||
<figureCitation id="62D02A52FF98FFDE3E54FEE524B4BB14" box="[811,901,352,378]" captionStart="FIGURES 30 – 34" captionStartId="19.[151,269,1464,1488]" captionTargetBox="[164,1434,194,1391]" captionTargetId="figure@19.[151,1436,194,1440]" captionTargetPageId="19" captionText="FIGURES 30 – 34. Spadiseius calyptrogynae n. sp., adult female. Figs. 30 – 33, dorsal views of legs excluding tarsi: 30, leg I; 31, leg II; 32, leg III; 33, leg IV; anterolateral (al) and posterolateral (pl) faces indicated by arrows. Fig. 34, tarsus IV, dorsal view." httpUri="https://zenodo.org/record/180489/files/figure.png" pageId="12" pageNumber="13">Fig. 34</figureCitation>
|
||
). Leg II slightly thicker than legs III–IV, but thickness and form of its setae not differentiated from those of legs III–IV (
|
||
<figureCitation id="62D02A52FF98FFDE3978FE0D23A6BBCC" box="[1031,1175,392,418]" captionStart="FIGURES 30 – 34" captionStartId="19.[151,269,1464,1488]" captionTargetBox="[164,1434,194,1391]" captionTargetId="figure@19.[151,1436,194,1440]" captionTargetPageId="19" captionText="FIGURES 30 – 34. Spadiseius calyptrogynae n. sp., adult female. Figs. 30 – 33, dorsal views of legs excluding tarsi: 30, leg I; 31, leg II; 32, leg III; 33, leg IV; anterolateral (al) and posterolateral (pl) faces indicated by arrows. Fig. 34, tarsus IV, dorsal view." httpUri="https://zenodo.org/record/180489/files/figure.png" pageId="12" pageNumber="13">Figs. 31–33</figureCitation>
|
||
). Coxae I with several serrated ridges on ventral face, coxae II with 2 or 3 serrated ridges on posteroventral face, coxae III–IV unornamented (other than vertical line caused by small boss below each seta); seta on coxa IV short (11–12), half as long as setae on coxa III (22–24) (
|
||
<figureCitation id="62D02A52FF98FFDE3F39FD8525E4B874" box="[582,725,512,538]" captionStart="FIGURES 30 – 34" captionStartId="19.[151,269,1464,1488]" captionTargetBox="[164,1434,194,1391]" captionTargetId="figure@19.[151,1436,194,1440]" captionTargetPageId="19" captionText="FIGURES 30 – 34. Spadiseius calyptrogynae n. sp., adult female. Figs. 30 – 33, dorsal views of legs excluding tarsi: 30, leg I; 31, leg II; 32, leg III; 33, leg IV; anterolateral (al) and posterolateral (pl) faces indicated by arrows. Fig. 34, tarsus IV, dorsal view." httpUri="https://zenodo.org/record/180489/files/figure.png" pageId="12" pageNumber="13">Figs. 32, 33</figureCitation>
|
||
). Apical sclerotized rim of femur, genu, tibia usually smooth ventrally on leg I, but usually sparsely or irregularly serrated on legs II–IV. Leg IV with trochanter seta
|
||
<emphasis id="C89FEAC5FF98FFDE3832FDAD225BB82F" box="[1357,1386,552,577]" italics="true" pageId="12" pageNumber="13">ad</emphasis>
|
||
and femoral seta
|
||
<emphasis id="C89FEAC5FF98FFDE3C4EFDD52656B807" box="[305,359,592,617]" italics="true" pageId="12" pageNumber="13">ad-2</emphasis>
|
||
stout, poorly tapered. Femora I–IV with dorsal seta
|
||
<emphasis id="C89FEAC5FF98FFDE3EA5FDD52321B807" box="[986,1040,592,617]" italics="true" pageId="12" pageNumber="13">ad-1</emphasis>
|
||
thickened, untapered, rod-like or slightly oar-like in contrast with adjacent simple setae, increasingly longer from I (15–16) to IV (25–27) (
|
||
<figureCitation id="62D02A52FF98FFDE381EFDFD27D4B8D4" captionStart="FIGURES 30 – 34" captionStartId="19.[151,269,1464,1488]" captionTargetBox="[164,1434,194,1391]" captionTargetId="figure@19.[151,1436,194,1440]" captionTargetPageId="19" captionText="FIGURES 30 – 34. Spadiseius calyptrogynae n. sp., adult female. Figs. 30 – 33, dorsal views of legs excluding tarsi: 30, leg I; 31, leg II; 32, leg III; 33, leg IV; anterolateral (al) and posterolateral (pl) faces indicated by arrows. Fig. 34, tarsus IV, dorsal view." httpUri="https://zenodo.org/record/180489/files/figure.png" pageId="12" pageNumber="13">Figs. 30–33</figureCitation>
|
||
); legs without elongated macrosetae. Setation of genua and tibiae I–IV as presented for genus.
|
||
</paragraph>
|
||
<paragraph id="FA5436D7FF98FFDE3DB9FD422224BEAC" blockId="12.[151,1437,152,1218]" pageId="12" pageNumber="13">
|
||
<emphasis id="C89FEAC5FF98FFDE3DB9FD422645B88F" bold="true" box="[198,372,711,737]" pageId="12" pageNumber="13">ADULT MALE.</emphasis>
|
||
Dorsal shield 445–480 long, 335–440 at its greatest width at level of setae
|
||
<emphasis id="C89FEAC5FF98FFDE386BFD4D221EB88F" box="[1300,1327,712,737]" italics="true" pageId="12" pageNumber="13">r4</emphasis>
|
||
(n =10); podonotal region with longitudinal distance from alveoli of
|
||
<emphasis id="C89FEAC5FF98FFDE3E28FD75245FB967" box="[855,878,752,777]" italics="true" pageId="12" pageNumber="13">j1</emphasis>
|
||
to
|
||
<emphasis id="C89FEAC5FF98FFDE3EEBFD75249BB967" box="[916,938,752,777]" italics="true" pageId="12" pageNumber="13">j5</emphasis>
|
||
slightly less (0.8–0.9) than that from
|
||
<emphasis id="C89FEAC5FF98FFDE3818FD75224CB967" box="[1383,1405,752,777]" italics="true" pageId="12" pageNumber="13">j5</emphasis>
|
||
to
|
||
<emphasis id="C89FEAC5FF98FFDE3DE8FC9D279FB95F" box="[151,174,792,817]" italics="true" pageId="12" pageNumber="13">j6</emphasis>
|
||
; shield with uniformly short (7–10) setae, including some hypertrichy, on its central and posterior area, and with a corona of a variable number of enlarged (length to 135), apically capitate or oar-like setae peripherally (
|
||
<figureCitation id="62D02A52FF98FFDE3DE0FCED2634B9EC" box="[159,261,872,898]" captionStart="FIGURES 21, 22" captionStartId="16.[151,269,1448,1472]" captionTargetBox="[264,1317,480,1446]" captionTargetId="figure@16.[264,1323,474,1500]" captionTargetPageId="16" captionText="FIGURES 21, 22. Spadiseius calyptrogynae n. sp., adult male, “ wartless ” morph: 21, idiosoma, dorsal aspect; 22, idiosoma," httpUri="https://zenodo.org/record/180486/files/figure.png" pageId="12" pageNumber="13">Figs. 21</figureCitation>
|
||
,
|
||
<figureCitation id="62D02A52FF98FFDE3C69FCED2606B9EC" box="[278,311,872,898]" captionStart="FIGURES 23 – 25" captionStartId="17.[151,269,1573,1597]" captionTargetBox="[151,1427,403,1572]" captionTargetId="figure@17.[151,1436,394,1601]" captionTargetPageId="17" captionText="FIGURES 23 – 25. Spadiseius calyptrogynae n. sp., adult male: 23, idiosoma, “ warted morph ”, dorsal aspect; 24, peritrematal shield and detail of enlarged gland pore; 25, greatly enlarged detail of peritrematal gland pore." httpUri="https://zenodo.org/record/180487/files/figure.png" pageId="12" pageNumber="13">23</figureCitation>
|
||
); setae
|
||
<emphasis id="C89FEAC5FF98FFDE3CE8FCED269FB9EF" box="[407,430,872,897]" italics="true" pageId="12" pageNumber="13">j1</emphasis>
|
||
consistently untapered, slightly oar-like. Dorsal shield ornamentation and form of peripheral setae polymorphic, with short setae inserted individually on circular wart-like thickenings correlated with more elongated peripheral setae (75–140) on some specimens (
|
||
<figureCitation id="62D02A52FF98FFDE3973FC3D2354B9BC" box="[1036,1125,952,978]" captionStart="FIGURES 23 – 25" captionStartId="17.[151,269,1573,1597]" captionTargetBox="[151,1427,403,1572]" captionTargetId="figure@17.[151,1436,394,1601]" captionTargetPageId="17" captionText="FIGURES 23 – 25. Spadiseius calyptrogynae n. sp., adult male: 23, idiosoma, “ warted morph ”, dorsal aspect; 24, peritrematal shield and detail of enlarged gland pore; 25, greatly enlarged detail of peritrematal gland pore." httpUri="https://zenodo.org/record/180487/files/figure.png" pageId="12" pageNumber="13">Fig. 23</figureCitation>
|
||
), but with none or few of short setae on such thickenings and shield reticulate, correlated with less elongated peripheral setae (30–80) on others (
|
||
<figureCitation id="62D02A52FF98FFDE3C6BFB8D2647BE4C" box="[276,374,1032,1058]" captionStart="FIGURES 21, 22" captionStartId="16.[151,269,1448,1472]" captionTargetBox="[264,1317,480,1446]" captionTargetId="figure@16.[264,1323,474,1500]" captionTargetPageId="16" captionText="FIGURES 21, 22. Spadiseius calyptrogynae n. sp., adult male, “ wartless ” morph: 21, idiosoma, dorsal aspect; 22, idiosoma," httpUri="https://zenodo.org/record/180486/files/figure.png" pageId="12" pageNumber="13">Figs. 21</figureCitation>
|
||
); whether elongated or not,
|
||
<emphasis id="C89FEAC5FF98FFDE3FBEFB8D25EFBE4F" box="[705,734,1032,1057]" italics="true" pageId="12" pageNumber="13">S1</emphasis>
|
||
consistently shorter (0.5–0.7) than adjacent setae
|
||
<emphasis id="C89FEAC5FF98FFDE3854FB8D2274BE4F" box="[1323,1349,1032,1057]" italics="true" pageId="12" pageNumber="13">s6</emphasis>
|
||
and
|
||
<emphasis id="C89FEAC5FF98FFDE3801FB8D22ADBE4F" box="[1406,1436,1032,1057]" italics="true" pageId="12" pageNumber="13">S2</emphasis>
|
||
(
|
||
<tableCitation id="B769036CFF98FFDE3DE1FBB527C6BE24" box="[158,247,1072,1098]" captionStart="TABLE 1" captionStartId="12.[151,239,1271,1295]" captionTargetBox="[159,1399,1328,1761]" captionTargetPageId="12" captionText="TABLE 1. Polymorphic male measurements (µm) - Spadiseius calyptrogynae n. sp." httpUri="http://table.plazi.org/id/AE94665FFF98FFDE3DE8FB722328BF61" pageId="12" pageNumber="13" tableUuid="AE94665FFF98FFDE3DE8FB722328BF61">Table 1</tableCitation>
|
||
). Peritrematal shields uniting with dorsal shield at level between bases of legs II and III, shield gland pore at that level greatly enlarged (
|
||
<figureCitation id="62D02A52FF98FFDE3F4FFBDD258EBE1C" box="[560,703,1112,1138]" captionStart="FIGURES 23 – 25" captionStartId="17.[151,269,1573,1597]" captionTargetBox="[151,1427,403,1572]" captionTargetId="figure@17.[151,1436,394,1601]" captionTargetPageId="17" captionText="FIGURES 23 – 25. Spadiseius calyptrogynae n. sp., adult male: 23, idiosoma, “ warted morph ”, dorsal aspect; 24, peritrematal shield and detail of enlarged gland pore; 25, greatly enlarged detail of peritrematal gland pore." httpUri="https://zenodo.org/record/180487/files/figure.png" pageId="12" pageNumber="13">Figs. 24, 25</figureCitation>
|
||
); peritremes extending to level of setae
|
||
<emphasis id="C89FEAC5FF98FFDE39EBFBDD239FBE1F" box="[1172,1198,1112,1137]" italics="true" pageId="12" pageNumber="13">s1</emphasis>
|
||
. Dorsal shield complement of setae, poroids and gland-pores similar to that in female, marginals
|
||
<emphasis id="C89FEAC5FF98FFDE3943FB052364BEF7" box="[1084,1109,1152,1177]" italics="true" pageId="12" pageNumber="13">r5</emphasis>
|
||
,
|
||
<emphasis id="C89FEAC5FF98FFDE3919FB052387BEF7" box="[1126,1206,1152,1177]" italics="true" pageId="12" pageNumber="13">R1–R2</emphasis>
|
||
,
|
||
<emphasis id="C89FEAC5FF98FFDE39B9FB052227BEF7" box="[1222,1302,1152,1177]" italics="true" pageId="12" pageNumber="13">R4–R6</emphasis>
|
||
, poroid
|
||
<collectingCountry id="82FC7647FF98FFDE3804FB0522ADBEF7" box="[1403,1436,1152,1177]" name="Poland" pageId="12" pageNumber="13">
|
||
<emphasis id="C89FEAC5FF98FFDE3804FB0522ADBEF7" box="[1403,1436,1152,1177]" italics="true" pageId="12" pageNumber="13">Rp</emphasis>
|
||
</collectingCountry>
|
||
and sometimes 1 or 2
|
||
<emphasis id="C89FEAC5FF98FFDE3CE6FB2D26F1BEAF" box="[409,448,1192,1217]" italics="true" pageId="12" pageNumber="13">UR</emphasis>
|
||
setae variably on soft cuticle or on edge of shield;
|
||
<emphasis id="C89FEAC5FF98FFDE3969FB2D2307BEAF" box="[1046,1078,1192,1217]" italics="true" pageId="12" pageNumber="13">R3</emphasis>
|
||
sometimes absent.
|
||
</paragraph>
|
||
<caption id="AE94665FFF98FFDE3DE8FB722328BF61" ID-Table-UUID="AE94665FFF98FFDE3DE8FB722328BF61" box="[151,1049,1271,1295]" httpUri="http://table.plazi.org/id/AE94665FFF98FFDE3DE8FB722328BF61" pageId="12" pageNumber="13" targetBox="[159,1399,1328,1761]" targetIsTable="true" targetPageId="12">
|
||
<paragraph id="FA5436D7FF98FFDE3DE8FB722328BF61" blockId="12.[151,1049,1271,1295]" box="[151,1049,1271,1295]" pageId="12" pageNumber="13">
|
||
<emphasis id="C89FEAC5FF98FFDE3DE8FB722639BF61" bold="true" box="[151,264,1271,1295]" pageId="12" pageNumber="13">TABLE 1.</emphasis>
|
||
Polymorphic male measurements (µm) -
|
||
<taxonomicName id="3DEB4D54FF98FFDE3FB9FB7D24E6BF61" box="[710,983,1272,1295]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="calyptrogynae" status="sp. nov.">
|
||
<emphasis id="C89FEAC5FF98FFDE3FB9FB7D24E6BF61" box="[710,983,1272,1295]" italics="true" pageId="12" pageNumber="13">Spadiseius calyptrogynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D3AC57BEFF98FFDE3EA2FB722328BF61" box="[989,1049,1271,1295]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FA5436D7FF98FFDE3DE0FAB52268BCD1" pageId="12" pageNumber="13">
|
||
<table id="88EBC477FF98002D3DE0FAB52246BC8F" box="[159,1399,1328,1761]" gridcols="11" gridrows="9" pageId="12" pageNumber="13">
|
||
<tr id="44DB3495FF98002D3DE0FAB52246BF07" box="[159,1399,1328,1385]" gridrow="0" pageId="12" pageNumber="13">
|
||
<th id="070A5DE9FF98002D3DE0FAB527CFBF07" box="[159,254,1328,1385]" gridcol="0" gridrow="0" pageId="12" pageNumber="13">Morph</th>
|
||
<th id="070A5DE9FF98002D3C56FAB5265DBF07" box="[297,364,1328,1385]" gridcol="1" gridrow="0" pageId="12" pageNumber="13">shield length</th>
|
||
<th id="070A5DE9FF98002D3CE4FAB52510BF07" box="[411,545,1328,1385]" gridcol="2" gridrow="0" pageId="12" pageNumber="13">shield width / ratio</th>
|
||
<th id="070A5DE9FF98002D3F29FAB525A8BF07" box="[598,665,1328,1385]" gridcol="3" gridrow="0" pageId="12" pageNumber="13">leg IV length</th>
|
||
<th id="070A5DE9FF98002D3FBDFAB525DBBF07" box="[706,746,1328,1385]" gridcol="4" gridrow="0" pageId="12" pageNumber="13">
|
||
<emphasis id="C89FEAC5FF98FFDE3FBDFAB525D0BF29" box="[706,737,1328,1351]" italics="true" pageId="12" pageNumber="13">st5</emphasis>
|
||
</th>
|
||
<th id="070A5DE9FF98002D3E5FFAB5247BBF07" box="[800,842,1328,1385]" gridcol="5" gridrow="0" pageId="12" pageNumber="13">
|
||
<emphasis id="C89FEAC5FF98FFDE3E5FFAB5247BBF29" box="[800,842,1328,1351]" italics="true" pageId="12" pageNumber="13">JV5</emphasis>
|
||
</th>
|
||
<th id="070A5DE9FF98002D3E0FFAB524A9BF07" box="[880,920,1328,1385]" gridcol="6" gridrow="0" pageId="12" pageNumber="13">
|
||
<emphasis id="C89FEAC5FF98FFDE3E0FFAB524B9BF29" box="[880,904,1328,1351]" italics="true" pageId="12" pageNumber="13">s6</emphasis>
|
||
</th>
|
||
<th id="070A5DE9FF98002D3EB1FAB524D8BF07" box="[974,1001,1328,1385]" gridcol="7" gridrow="0" pageId="12" pageNumber="13">
|
||
<emphasis id="C89FEAC5FF98FFDE3EB1FAB524D9BF29" box="[974,1000,1328,1351]" italics="true" pageId="12" pageNumber="13">S1</emphasis>
|
||
</th>
|
||
<th id="070A5DE9FF98002D3961FAB52377BF07" box="[1054,1094,1328,1385]" gridcol="8" gridrow="0" pageId="12" pageNumber="13">
|
||
<emphasis id="C89FEAC5FF98FFDE3961FAB52308BF29" box="[1054,1081,1328,1351]" italics="true" pageId="12" pageNumber="13">S2</emphasis>
|
||
</th>
|
||
<th id="070A5DE9FF98002D3911FAB523A7BF07" box="[1134,1174,1328,1385]" gridcol="9" gridrow="0" pageId="12" pageNumber="13">
|
||
<emphasis id="C89FEAC5FF98FFDE3911FAB523B9BF29" box="[1134,1160,1328,1351]" italics="true" pageId="12" pageNumber="13">S5</emphasis>
|
||
</th>
|
||
<th id="070A5DE9FF98002D39B3FAB52246BF07" box="[1228,1399,1328,1385]" gridcol="10" gridrow="0" pageId="12" pageNumber="13">
|
||
IVFe
|
||
<emphasis id="C89FEAC5FF98FFDE387DFAB52226BF29" box="[1282,1303,1328,1351]" italics="true" pageId="12" pageNumber="13">-v</emphasis>
|
||
IVFe-
|
||
<emphasis id="C89FEAC5FF98FFDE3848FAD72258BF07" box="[1335,1385,1362,1385]" italics="true" pageId="12" pageNumber="13">ad-1</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="44DB3495FF98002D3DE0FAF82246BFFA" box="[159,1399,1405,1428]" gridrow="1" pageId="12" pageNumber="13">
|
||
<th id="070A5DE9FF98002D3DE0FAF827CFBFFA" box="[159,254,1405,1428]" gridcol="0" gridrow="1" pageId="12" pageNumber="13">wartless</th>
|
||
<td id="070A5DE9FF98002D3C56FAF8265DBFFA" box="[297,364,1405,1428]" gridcol="1" gridrow="1" pageId="12" pageNumber="13">440</td>
|
||
<td id="070A5DE9FF98002D3CE4FAF82510BFFA" box="[411,545,1405,1428]" gridcol="2" gridrow="1" pageId="12" pageNumber="13">320 0.72</td>
|
||
<td id="070A5DE9FF98002D3F29FAF825A8BFFA" box="[598,665,1405,1428]" gridcol="3" gridrow="1" pageId="12" pageNumber="13">430</td>
|
||
<td id="070A5DE9FF98002D3FBDFAF825DBBFFA" box="[706,746,1405,1428]" gridcol="4" gridrow="1" pageId="12" pageNumber="13">45</td>
|
||
<td id="070A5DE9FF98002D3E5FFAF8247BBFFA" box="[800,842,1405,1428]" gridcol="5" gridrow="1" pageId="12" pageNumber="13">20</td>
|
||
<td id="070A5DE9FF98002D3E0FFAF824A9BFFA" box="[880,920,1405,1428]" gridcol="6" gridrow="1" pageId="12" pageNumber="13">30</td>
|
||
<td id="070A5DE9FF98002D3EB1FAF824D8BFFA" box="[974,1001,1405,1428]" gridcol="7" gridrow="1" pageId="12" pageNumber="13">20</td>
|
||
<td id="070A5DE9FF98002D3961FAF82377BFFA" box="[1054,1094,1405,1428]" gridcol="8" gridrow="1" pageId="12" pageNumber="13">30</td>
|
||
<td id="070A5DE9FF98002D3911FAF823A7BFFA" box="[1134,1174,1405,1428]" gridcol="9" gridrow="1" pageId="12" pageNumber="13">57</td>
|
||
<td id="070A5DE9FF98002D39B3FAF82246BFFA" box="[1228,1399,1405,1428]" gridcol="10" gridrow="1" pageId="12" pageNumber="13">37 30</td>
|
||
</tr>
|
||
<tr id="44DB3495FF98002D3DE0FA2D2246BFD1" box="[159,1399,1448,1471]" gridrow="2" pageId="12" pageNumber="13">
|
||
<th id="070A5DE9FF98002D3DE0FA2D27CFBFD1" box="[159,254,1448,1471]" gridcol="0" gridrow="2" pageId="12" pageNumber="13">wartless</th>
|
||
<td id="070A5DE9FF98002D3C56FA2D265DBFD1" box="[297,364,1448,1471]" gridcol="1" gridrow="2" pageId="12" pageNumber="13">467</td>
|
||
<td id="070A5DE9FF98002D3CE4FA2D2510BFD1" box="[411,545,1448,1471]" gridcol="2" gridrow="2" pageId="12" pageNumber="13">312 0.67</td>
|
||
<td id="070A5DE9FF98002D3F29FA2D25A8BFD1" box="[598,665,1448,1471]" gridcol="3" gridrow="2" pageId="12" pageNumber="13">450</td>
|
||
<td id="070A5DE9FF98002D3FBDFA2D25DBBFD1" box="[706,746,1448,1471]" gridcol="4" gridrow="2" pageId="12" pageNumber="13">87</td>
|
||
<td id="070A5DE9FF98002D3E5FFA2D247BBFD1" box="[800,842,1448,1471]" gridcol="5" gridrow="2" pageId="12" pageNumber="13">30</td>
|
||
<td id="070A5DE9FF98002D3E0FFA2D24A9BFD1" box="[880,920,1448,1471]" gridcol="6" gridrow="2" pageId="12" pageNumber="13">62</td>
|
||
<td id="070A5DE9FF98002D3EB1FA2D24D8BFD1" box="[974,1001,1448,1471]" gridcol="7" gridrow="2" pageId="12" pageNumber="13">40</td>
|
||
<td id="070A5DE9FF98002D3961FA2D2377BFD1" box="[1054,1094,1448,1471]" gridcol="8" gridrow="2" pageId="12" pageNumber="13">80</td>
|
||
<td id="070A5DE9FF98002D3911FA2D23A7BFD1" box="[1134,1174,1448,1471]" gridcol="9" gridrow="2" pageId="12" pageNumber="13">80</td>
|
||
<td id="070A5DE9FF98002D39B3FA2D2246BFD1" box="[1228,1399,1448,1471]" gridcol="10" gridrow="2" pageId="12" pageNumber="13">52 42</td>
|
||
</tr>
|
||
<tr id="44DB3495FF98002D3DE0FA572246BF87" box="[159,1399,1490,1513]" gridrow="3" pageId="12" pageNumber="13">
|
||
<th id="070A5DE9FF98002D3DE0FA5727CFBF87" box="[159,254,1490,1513]" gridcol="0" gridrow="3" pageId="12" pageNumber="13">warted</th>
|
||
<td id="070A5DE9FF98002D3C56FA57265DBF87" box="[297,364,1490,1513]" gridcol="1" gridrow="3" pageId="12" pageNumber="13">485</td>
|
||
<td id="070A5DE9FF98002D3CE4FA572510BF87" box="[411,545,1490,1513]" gridcol="2" gridrow="3" pageId="12" pageNumber="13">400 0.82</td>
|
||
<td id="070A5DE9FF98002D3F29FA5725A8BF87" box="[598,665,1490,1513]" gridcol="3" gridrow="3" pageId="12" pageNumber="13">492</td>
|
||
<td id="070A5DE9FF98002D3FBDFA5725DBBF87" box="[706,746,1490,1513]" gridcol="4" gridrow="3" pageId="12" pageNumber="13">112</td>
|
||
<td id="070A5DE9FF98002D3E5FFA57247BBF87" box="[800,842,1490,1513]" gridcol="5" gridrow="3" pageId="12" pageNumber="13">60</td>
|
||
<td id="070A5DE9FF98002D3E0FFA5724A9BF87" box="[880,920,1490,1513]" gridcol="6" gridrow="3" pageId="12" pageNumber="13">120</td>
|
||
<td id="070A5DE9FF98002D3EB1FA5724D8BF87" box="[974,1001,1490,1513]" gridcol="7" gridrow="3" pageId="12" pageNumber="13">82</td>
|
||
<td id="070A5DE9FF98002D3961FA572377BF87" box="[1054,1094,1490,1513]" gridcol="8" gridrow="3" pageId="12" pageNumber="13">135</td>
|
||
<td id="070A5DE9FF98002D3911FA5723A7BF87" box="[1134,1174,1490,1513]" gridcol="9" gridrow="3" pageId="12" pageNumber="13">125</td>
|
||
<td id="070A5DE9FF98002D39B3FA572246BF87" box="[1228,1399,1490,1513]" gridcol="10" gridrow="3" pageId="12" pageNumber="13">85 67</td>
|
||
</tr>
|
||
<tr id="44DB3495FF98002D3DE0FA782246BC7A" box="[159,1399,1533,1556]" gridrow="4" pageId="12" pageNumber="13">
|
||
<th id="070A5DE9FF98002D3DE0FA7827CFBC7A" box="[159,254,1533,1556]" gridcol="0" gridrow="4" pageId="12" pageNumber="13">warted</th>
|
||
<td id="070A5DE9FF98002D3C56FA78265DBC7A" box="[297,364,1533,1556]" gridcol="1" gridrow="4" pageId="12" pageNumber="13">460</td>
|
||
<td id="070A5DE9FF98002D3CE4FA782510BC7A" box="[411,545,1533,1556]" gridcol="2" gridrow="4" pageId="12" pageNumber="13">375 0.82</td>
|
||
<td id="070A5DE9FF98002D3F29FA7825A8BC7A" box="[598,665,1533,1556]" gridcol="3" gridrow="4" pageId="12" pageNumber="13">465</td>
|
||
<td id="070A5DE9FF98002D3FBDFA7825DBBC7A" box="[706,746,1533,1556]" gridcol="4" gridrow="4" pageId="12" pageNumber="13">125</td>
|
||
<td id="070A5DE9FF98002D3E5FFA78247BBC7A" box="[800,842,1533,1556]" gridcol="5" gridrow="4" pageId="12" pageNumber="13">50</td>
|
||
<td id="070A5DE9FF98002D3E0FFA7824A9BC7A" box="[880,920,1533,1556]" gridcol="6" gridrow="4" pageId="12" pageNumber="13">100</td>
|
||
<td id="070A5DE9FF98002D3EB1FA7824D8BC7A" box="[974,1001,1533,1556]" gridcol="7" gridrow="4" pageId="12" pageNumber="13">70</td>
|
||
<td id="070A5DE9FF98002D3961FA782377BC7A" box="[1054,1094,1533,1556]" gridcol="8" gridrow="4" pageId="12" pageNumber="13">97</td>
|
||
<td id="070A5DE9FF98002D3911FA7823A7BC7A" box="[1134,1174,1533,1556]" gridcol="9" gridrow="4" pageId="12" pageNumber="13">107</td>
|
||
<td id="070A5DE9FF98002D39B3FA782246BC7A" box="[1228,1399,1533,1556]" gridcol="10" gridrow="4" pageId="12" pageNumber="13">87 55</td>
|
||
</tr>
|
||
<tr id="44DB3495FF98002D3DE0F9AD2246BC51" box="[159,1399,1576,1599]" gridrow="5" pageId="12" pageNumber="13">
|
||
<th id="070A5DE9FF98002D3DE0F9AD27CFBC51" box="[159,254,1576,1599]" gridcol="0" gridrow="5" pageId="12" pageNumber="13">warted</th>
|
||
<td id="070A5DE9FF98002D3C56F9AD265DBC51" box="[297,364,1576,1599]" gridcol="1" gridrow="5" pageId="12" pageNumber="13">485</td>
|
||
<td id="070A5DE9FF98002D3CE4F9AD2510BC51" box="[411,545,1576,1599]" gridcol="2" gridrow="5" pageId="12" pageNumber="13">400 0.82</td>
|
||
<td id="070A5DE9FF98002D3F29F9AD25A8BC51" box="[598,665,1576,1599]" gridcol="3" gridrow="5" pageId="12" pageNumber="13">467</td>
|
||
<td id="070A5DE9FF98002D3FBDF9AD25DBBC51" box="[706,746,1576,1599]" gridcol="4" gridrow="5" pageId="12" pageNumber="13">115</td>
|
||
<td id="070A5DE9FF98002D3E5FF9AD247BBC51" box="[800,842,1576,1599]" gridcol="5" gridrow="5" pageId="12" pageNumber="13">45</td>
|
||
<td id="070A5DE9FF98002D3E0FF9AD24A9BC51" box="[880,920,1576,1599]" gridcol="6" gridrow="5" pageId="12" pageNumber="13">120</td>
|
||
<td id="070A5DE9FF98002D3EB1F9AD24D8BC51" box="[974,1001,1576,1599]" gridcol="7" gridrow="5" pageId="12" pageNumber="13">82</td>
|
||
<td id="070A5DE9FF98002D3961F9AD2377BC51" box="[1054,1094,1576,1599]" gridcol="8" gridrow="5" pageId="12" pageNumber="13">137</td>
|
||
<td id="070A5DE9FF98002D3911F9AD23A7BC51" box="[1134,1174,1576,1599]" gridcol="9" gridrow="5" pageId="12" pageNumber="13">132</td>
|
||
<td id="070A5DE9FF98002D39B3F9AD2246BC51" box="[1228,1399,1576,1599]" gridcol="10" gridrow="5" pageId="12" pageNumber="13">80 57</td>
|
||
</tr>
|
||
<tr id="44DB3495FF98002D3DE0F9D72246BC07" box="[159,1399,1618,1641]" gridrow="6" pageId="12" pageNumber="13">
|
||
<th id="070A5DE9FF98002D3DE0F9D727CFBC07" box="[159,254,1618,1641]" gridcol="0" gridrow="6" pageId="12" pageNumber="13">warted</th>
|
||
<td id="070A5DE9FF98002D3C56F9D7265DBC07" box="[297,364,1618,1641]" gridcol="1" gridrow="6" pageId="12" pageNumber="13">487</td>
|
||
<td id="070A5DE9FF98002D3CE4F9D72510BC07" box="[411,545,1618,1641]" gridcol="2" gridrow="6" pageId="12" pageNumber="13">395 0.81</td>
|
||
<td id="070A5DE9FF98002D3F29F9D725A8BC07" box="[598,665,1618,1641]" gridcol="3" gridrow="6" pageId="12" pageNumber="13">500</td>
|
||
<td id="070A5DE9FF98002D3FBDF9D725DBBC07" box="[706,746,1618,1641]" gridcol="4" gridrow="6" pageId="12" pageNumber="13">100</td>
|
||
<td id="070A5DE9FF98002D3E5FF9D7247BBC07" box="[800,842,1618,1641]" gridcol="5" gridrow="6" pageId="12" pageNumber="13">38</td>
|
||
<td id="070A5DE9FF98002D3E0FF9D724A9BC07" box="[880,920,1618,1641]" gridcol="6" gridrow="6" pageId="12" pageNumber="13">100</td>
|
||
<td id="070A5DE9FF98002D3EB1F9D724D8BC07" box="[974,1001,1618,1641]" gridcol="7" gridrow="6" pageId="12" pageNumber="13">65</td>
|
||
<td id="070A5DE9FF98002D3961F9D72377BC07" box="[1054,1094,1618,1641]" gridcol="8" gridrow="6" pageId="12" pageNumber="13">100</td>
|
||
<td id="070A5DE9FF98002D3911F9D723A7BC07" box="[1134,1174,1618,1641]" gridcol="9" gridrow="6" pageId="12" pageNumber="13">107</td>
|
||
<td id="070A5DE9FF98002D39B3F9D72246BC07" box="[1228,1399,1618,1641]" gridcol="10" gridrow="6" pageId="12" pageNumber="13">80 55</td>
|
||
</tr>
|
||
<tr id="44DB3495FF98002D3DE0F9F82246BCFA" box="[159,1399,1661,1684]" gridrow="7" pageId="12" pageNumber="13">
|
||
<th id="070A5DE9FF98002D3DE0F9F827CFBCFA" box="[159,254,1661,1684]" gridcol="0" gridrow="7" pageId="12" pageNumber="13">warted</th>
|
||
<td id="070A5DE9FF98002D3C56F9F8265DBCFA" box="[297,364,1661,1684]" gridcol="1" gridrow="7" pageId="12" pageNumber="13">480</td>
|
||
<td id="070A5DE9FF98002D3CE4F9F82510BCFA" box="[411,545,1661,1684]" gridcol="2" gridrow="7" pageId="12" pageNumber="13">360 0.75</td>
|
||
<td id="070A5DE9FF98002D3F29F9F825A8BCFA" box="[598,665,1661,1684]" gridcol="3" gridrow="7" pageId="12" pageNumber="13">467</td>
|
||
<td id="070A5DE9FF98002D3FBDF9F825DBBCFA" box="[706,746,1661,1684]" gridcol="4" gridrow="7" pageId="12" pageNumber="13">112</td>
|
||
<td id="070A5DE9FF98002D3E5FF9F8247BBCFA" box="[800,842,1661,1684]" gridcol="5" gridrow="7" pageId="12" pageNumber="13">32</td>
|
||
<td id="070A5DE9FF98002D3E0FF9F824A9BCFA" box="[880,920,1661,1684]" gridcol="6" gridrow="7" pageId="12" pageNumber="13">112</td>
|
||
<td id="070A5DE9FF98002D3EB1F9F824D8BCFA" box="[974,1001,1661,1684]" gridcol="7" gridrow="7" pageId="12" pageNumber="13">65</td>
|
||
<td id="070A5DE9FF98002D3961F9F82377BCFA" box="[1054,1094,1661,1684]" gridcol="8" gridrow="7" pageId="12" pageNumber="13">100</td>
|
||
<td id="070A5DE9FF98002D3911F9F823A7BCFA" box="[1134,1174,1661,1684]" gridcol="9" gridrow="7" pageId="12" pageNumber="13">107</td>
|
||
<td id="070A5DE9FF98002D39B3F9F82246BCFA" box="[1228,1399,1661,1684]" gridcol="10" gridrow="7" pageId="12" pageNumber="13">80 50</td>
|
||
</tr>
|
||
<tr id="44DB3495FF98002D3DE0F92D2246BC8F" box="[159,1399,1704,1761]" gridrow="8" pageId="12" pageNumber="13">
|
||
<th id="070A5DE9FF98002D3DE0F92D27CFBC8F" box="[159,254,1704,1761]" gridcol="0" gridrow="8" pageId="12" pageNumber="13">warted intermed</th>
|
||
<td id="070A5DE9FF98002D3C56F92D265DBC8F" box="[297,364,1704,1761]" gridcol="1" gridrow="8" pageId="12" pageNumber="13">467</td>
|
||
<td id="070A5DE9FF98002D3CE4F92D2510BC8F" box="[411,545,1704,1761]" gridcol="2" gridrow="8" pageId="12" pageNumber="13">350 0.75</td>
|
||
<td id="070A5DE9FF98002D3F29F92D25A8BC8F" box="[598,665,1704,1761]" gridcol="3" gridrow="8" pageId="12" pageNumber="13">445</td>
|
||
<td id="070A5DE9FF98002D3FBDF92D25DBBC8F" box="[706,746,1704,1761]" gridcol="4" gridrow="8" pageId="12" pageNumber="13">90</td>
|
||
<td id="070A5DE9FF98002D3E5FF92D247BBC8F" box="[800,842,1704,1761]" gridcol="5" gridrow="8" pageId="12" pageNumber="13">30</td>
|
||
<td id="070A5DE9FF98002D3E0FF92D24A9BC8F" box="[880,920,1704,1761]" gridcol="6" gridrow="8" pageId="12" pageNumber="13">80</td>
|
||
<td id="070A5DE9FF98002D3EB1F92D24D8BC8F" box="[974,1001,1704,1761]" gridcol="7" gridrow="8" pageId="12" pageNumber="13">50</td>
|
||
<td id="070A5DE9FF98002D3961F92D2377BC8F" box="[1054,1094,1704,1761]" gridcol="8" gridrow="8" pageId="12" pageNumber="13">80</td>
|
||
<td id="070A5DE9FF98002D3911F92D23A7BC8F" box="[1134,1174,1704,1761]" gridcol="9" gridrow="8" pageId="12" pageNumber="13">87</td>
|
||
<td id="070A5DE9FF98002D39B3F92D2246BC8F" box="[1228,1399,1704,1761]" gridcol="10" gridrow="8" pageId="12" pageNumber="13">67 50</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="FA5436D7FF98FFDF3DB9F8B72683BB9C" blockId="12.[151,1437,1842,2028]" lastBlockId="13.[151,1436,152,498]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
|
||
Tritosternum much as in female, but with length of trapezoidal base (13–15) less than its basal width (20– 22), and laciniae free for about 0.8 of total length (65–68 excluding base). Presternal area with pair of small platelets (
|
||
<figureCitation id="62D02A52FF98FFDE3C74F8072656BDF2" box="[267,359,1922,1948]" captionStart="FIGURES 21, 22" captionStartId="16.[151,269,1448,1472]" captionTargetBox="[264,1317,480,1446]" captionTargetId="figure@16.[264,1323,474,1500]" captionTargetPageId="16" captionText="FIGURES 21, 22. Spadiseius calyptrogynae n. sp., adult male, “ wartless ” morph: 21, idiosoma, dorsal aspect; 22, idiosoma," httpUri="https://zenodo.org/record/180486/files/figure.png" pageId="12" pageNumber="13">Fig. 22</figureCitation>
|
||
). Sternogenital shield unornamented; of four or five pairs of setae on shield,
|
||
<emphasis id="C89FEAC5FF98FFDE386CF8062204BDF2" box="[1299,1333,1923,1948]" italics="true" pageId="12" pageNumber="13">st4</emphasis>
|
||
(50–60) clearly longer than
|
||
<emphasis id="C89FEAC5FF98FFDE3C07F82E26FBBDAA" box="[376,458,1963,1988]" italics="true" pageId="12" pageNumber="13">st1–st2</emphasis>
|
||
(15–28) and
|
||
<emphasis id="C89FEAC5FF98FFDE3F1AF82E25B6BDAA" box="[613,647,1963,1988]" italics="true" pageId="12" pageNumber="13">st3</emphasis>
|
||
(20–25),
|
||
<emphasis id="C89FEAC5FF98FFDE3F86F82E242ABDAA" box="[761,795,1963,1988]" italics="true" pageId="12" pageNumber="13">st5</emphasis>
|
||
variably much longer (85–110) and often inserted near paragenital poroids on soft cuticle closely beside invaginated posterolateral corners of shield; posterior margin of shield strongly delineated, broadly rounded. Ventrianal shield unornamented, with weakly delineated anterior and lateral margins variable in outline; shield length (185–190) about 0.6–0.7 its greatest width (280–305) anteriorly where metapodal plates incorporated; anal region more clearly sclerotized, sometimes appearing like a distinct plate surrounded by desclerotized cuticle (
|
||
<figureCitation id="62D02A52FF99FFDF3E4FFE9524B9BB44" box="[816,904,272,298]" captionStart="FIGURES 21, 22" captionStartId="16.[151,269,1448,1472]" captionTargetBox="[264,1317,480,1446]" captionTargetId="figure@16.[264,1323,474,1500]" captionTargetPageId="16" captionText="FIGURES 21, 22. Spadiseius calyptrogynae n. sp., adult male, “ wartless ” morph: 21, idiosoma, dorsal aspect; 22, idiosoma," httpUri="https://zenodo.org/record/180486/files/figure.png" pageId="13" pageNumber="14">Fig. 22</figureCitation>
|
||
); ventrianal shield with circumanal setae and usually 7 pairs of opisthogastric setae (
|
||
<emphasis id="C89FEAC5FF99FFDF3F13FEBD25E6BB3F" box="[620,727,312,337]" italics="true" pageId="13" pageNumber="14">JV1–JV4</emphasis>
|
||
,
|
||
<emphasis id="C89FEAC5FF99FFDF3F98FEBD2466BB3F" box="[743,855,312,337]" italics="true" pageId="13" pageNumber="14">ZV1–ZV3</emphasis>
|
||
);
|
||
<emphasis id="C89FEAC5FF99FFDF3E0CFEBD24D2BB3F" box="[883,995,312,337]" italics="true" pageId="13" pageNumber="14">ZV1–ZV2</emphasis>
|
||
more attenuated and variably longer (15–50), and
|
||
<emphasis id="C89FEAC5FF99FFDF3C4CFEE52651BB17" box="[307,352,352,377]" italics="true" pageId="13" pageNumber="14">JV4</emphasis>
|
||
variably longer (25–80) than in female;
|
||
<emphasis id="C89FEAC5FF99FFDF3E47FEE52458BB17" box="[824,873,352,377]" italics="true" pageId="13" pageNumber="14">ZV3</emphasis>
|
||
variably oarlike (25) or attenuate (63–90); postanal seta variably longer (25–40) than para-anal setae (16–25);
|
||
<emphasis id="C89FEAC5FF99FFDF3EFCFE0D2481BBCF" box="[899,944,392,417]" italics="true" pageId="13" pageNumber="14">JV5</emphasis>
|
||
usually clavate distally (30–60) and variably attenuate (ca 90), inserted on soft cuticle;
|
||
<emphasis id="C89FEAC5FF99FFDF3FB3FE35240DBBA7" box="[716,828,432,457]" italics="true" pageId="13" pageNumber="14">ZV4–ZV5</emphasis>
|
||
absent. Form of peritrematal and exopodal shields posteriorly as in female.
|
||
</paragraph>
|
||
<caption id="AE94665FFF99FFDF3DE8F9462243BC93" httpUri="https://zenodo.org/record/180483/files/figure.png" pageId="13" pageNumber="14" targetBox="[151,1435,556,1730]" targetPageId="13">
|
||
<paragraph id="FA5436D7FF99FFDF3DE8F9462243BC93" blockId="13.[151,1436,1731,1789]" pageId="13" pageNumber="14">
|
||
<emphasis id="C89FEAC5FF99FFDF3DE8F9462654BCB5" bold="true" box="[151,357,1731,1755]" pageId="13" pageNumber="14">FIGURES 16, 17.</emphasis>
|
||
<taxonomicName id="3DEB4D54FF99FFDF3C10F94125B3BCB5" box="[367,642,1732,1755]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="calyptrogynae" status="sp. nov.">
|
||
<emphasis id="C89FEAC5FF99FFDF3C10F94125B3BCB5" box="[367,642,1732,1755]" italics="true" pageId="13" pageNumber="14">Spadiseius calyptrogynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D3AC57BEFF99FFDF3FF2F94625FCBCB5" box="[653,717,1731,1755]" pageId="13" pageNumber="14" rank="species">
|
||
<emphasis id="C89FEAC5FF99FFDF3FF2F94625FCBCB5" bold="true" box="[653,717,1731,1755]" pageId="13" pageNumber="14">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, deutonymph: 16, idiosoma, dorsal aspect; 17, idiosoma, ventral aspect. Setae added to protonymphal complement denoted without parentheses; supernumerary setae denoted by ‘x’.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FA5436D7FF99FFDC3DB9F8B425F3BB44" blockId="13.[151,1436,1841,2027]" lastBlockId="14.[151,1437,152,298]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||
Tectum as in female. Cheliceral shaft abruptly narrowed dorsally at base of fixed digit (
|
||
<figureCitation id="62D02A52FF99FFDF3992F8B42276BD25" box="[1261,1351,1841,1867]" captionStart="FIGURES 26 – 29" captionStartId="18.[151,269,1250,1274]" captionTargetBox="[151,1435,354,1192]" captionTargetId="figure@18.[151,1436,354,1270]" captionTargetPageId="18" captionText="FIGURES 26 – 29. Spadiseius calyptrogynae n. sp., adult male: 26, subcapitulum; 27, chelicera and spermatodactyl, paraxial view; 28, tarsus II, anterolateral view; 29, tarsus IV, dorsal view, without pretarsus. Abbreviation: k, keel." httpUri="https://zenodo.org/record/180488/files/figure.png" pageId="13" pageNumber="14">Fig. 27</figureCitation>
|
||
); fixed digit with a small offset tooth followed by 2–3 minute teeth at level of hyaline pilus dentilis, and with a large rounded tooth (or lobe) basally; spermatodactyl biramous, with a pointed hyaline process above the nearly straight, apically lobed main shaft which extends anteriorly about 28–35 beyond apex of chela (
|
||
<figureCitation id="62D02A52FF99FFDF3986F82C2260BDAD" box="[1273,1361,1961,1987]" captionStart="FIGURES 26 – 29" captionStartId="18.[151,269,1250,1274]" captionTargetBox="[151,1435,354,1192]" captionTargetId="figure@18.[151,1436,354,1270]" captionTargetPageId="18" captionText="FIGURES 26 – 29. Spadiseius calyptrogynae n. sp., adult male: 26, subcapitulum; 27, chelicera and spermatodactyl, paraxial view; 28, tarsus II, anterolateral view; 29, tarsus IV, dorsal view, without pretarsus. Abbreviation: k, keel." httpUri="https://zenodo.org/record/180488/files/figure.png" pageId="13" pageNumber="14">Fig. 27</figureCitation>
|
||
). Corniculi more slender and widely spaced basally than in female; internal malae larger than in female, with fringed lateral margins and bifid apices (
|
||
<figureCitation id="62D02A52FF9AFFDC3F02FF1D25E6BADC" box="[637,727,152,178]" captionStart="FIGURES 26 – 29" captionStartId="18.[151,269,1250,1274]" captionTargetBox="[151,1435,354,1192]" captionTargetId="figure@18.[151,1436,354,1270]" captionTargetPageId="18" captionText="FIGURES 26 – 29. Spadiseius calyptrogynae n. sp., adult male: 26, subcapitulum; 27, chelicera and spermatodactyl, paraxial view; 28, tarsus II, anterolateral view; 29, tarsus IV, dorsal view, without pretarsus. Abbreviation: k, keel." httpUri="https://zenodo.org/record/180488/files/figure.png" pageId="14" pageNumber="15">Fig. 26</figureCitation>
|
||
). Subcapitulum with rows of deutosternal denticles slightly wider and more multidenticulate than in female. Subcapitulum lacking the pair of gland-like structures near setae
|
||
<emphasis id="C89FEAC5FF9AFFDC3DA6FF6D2623BB6F" box="[217,274,232,257]" italics="true" pageId="14" pageNumber="15">
|
||
hyp
|
||
<quantity id="3D139B32FF9AFFDC3C79FF6D2623BB6F" box="[262,274,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="15" unit="in" value="1.0">1</quantity>
|
||
</emphasis>
|
||
in female, and with capitular setae short but not conspicuously swollen basally; other subcapitular and palpal structures similar to those in female.
|
||
</paragraph>
|
||
<caption id="AE94665FFF9AFFDC3DE8F95522AABC89" box="[151,1435,1744,1768]" httpUri="https://zenodo.org/record/180484/files/figure.png" pageId="14" pageNumber="15" targetBox="[153,1414,370,1743]" targetPageId="14">
|
||
<paragraph id="FA5436D7FF9AFFDC3DE8F95522AABC89" blockId="14.[151,1435,1744,1768]" box="[151,1435,1744,1768]" pageId="14" pageNumber="15">
|
||
<emphasis id="C89FEAC5FF9AFFDC3DE8F9552619BC86" bold="true" box="[151,296,1744,1768]" pageId="14" pageNumber="15">FIGURE 18.</emphasis>
|
||
<taxonomicName id="3DEB4D54FF9AFFDC3C4EF9542572BC86" box="[305,579,1745,1768]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="calyptrogynae" status="sp. nov.">
|
||
<emphasis id="C89FEAC5FF9AFFDC3C4EF9542572BC86" box="[305,579,1745,1768]" italics="true" pageId="14" pageNumber="15">Spadiseius calyptrogynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D3AC57BEFF9AFFDC3F34F95525B8BC86" box="[587,649,1744,1768]" pageId="14" pageNumber="15" rank="species">
|
||
<emphasis id="C89FEAC5FF9AFFDC3F34F95525B8BC86" bold="true" box="[587,649,1744,1768]" pageId="14" pageNumber="15">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult female, idiosoma, dorsal aspect. Supernumerary setae denoted by
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FA5436D7FF9AFFDD3DB9F8BB26B9BBA4" blockId="14.[151,1436,1854,2000]" lastBlockId="15.[151,1437,152,458]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
|
||
Relative leg lengths, excluding pretarsi, similar to those of female, I (380–440), II (325–395), III (350– 420), IV (425–510). Coxa IV seta relatively short as in female; apical sclerotized rim of femur, genu, tibia usually smooth ventrally on legs I and sometimes on legs II, but usually serrated on legs III–IV. Tarsi II–IV with pair of apical setal processes (
|
||
<emphasis id="C89FEAC5FF9AFFDC3F49F8332581BDA1" box="[566,688,1974,1999]" italics="true" pageId="14" pageNumber="15">ad-1, pd-1</emphasis>
|
||
) nearly half as long as length of pretarsi to base of claws. Legs II curved and thicker relative to legs III–IV than on female, and with setae
|
||
<emphasis id="C89FEAC5FF9BFFDD396DFF1D2377BADF" box="[1042,1094,152,177]" italics="true" pageId="15" pageNumber="16">av-1</emphasis>
|
||
,
|
||
<emphasis id="C89FEAC5FF9BFFDD392AFF1D23B8BADF" box="[1109,1161,152,177]" italics="true" pageId="15" pageNumber="16">av-2</emphasis>
|
||
,
|
||
<emphasis id="C89FEAC5FF9BFFDD39E6FF1D23FDBADF" box="[1177,1228,152,177]" italics="true" pageId="15" pageNumber="16">av-3</emphasis>
|
||
(
|
||
<emphasis id="C89FEAC5FF9BFFDD39A0FF1D2230BADF" box="[1247,1281,152,177]" italics="true" pageId="15" pageNumber="16">mv</emphasis>
|
||
) modified to short, stout, bulbous spines (
|
||
<figureCitation id="62D02A52FF9BFFDD3C98FF452571BAB4" box="[487,576,192,218]" captionStart="FIGURES 26 – 29" captionStartId="18.[151,269,1250,1274]" captionTargetBox="[151,1435,354,1192]" captionTargetId="figure@18.[151,1436,354,1270]" captionTargetPageId="18" captionText="FIGURES 26 – 29. Spadiseius calyptrogynae n. sp., adult male: 26, subcapitulum; 27, chelicera and spermatodactyl, paraxial view; 28, tarsus II, anterolateral view; 29, tarsus IV, dorsal view, without pretarsus. Abbreviation: k, keel." httpUri="https://zenodo.org/record/180488/files/figure.png" pageId="15" pageNumber="16">Fig. 28</figureCitation>
|
||
); legs III–IV also slightly thicker relative to leg I than in female; basitarsus of legs III–IV without spur (
|
||
<figureCitation id="62D02A52FF9BFFDD3F65FF6D2545BB6C" box="[538,628,232,258]" captionStart="FIGURES 26 – 29" captionStartId="18.[151,269,1250,1274]" captionTargetBox="[151,1435,354,1192]" captionTargetId="figure@18.[151,1436,354,1270]" captionTargetPageId="18" captionText="FIGURES 26 – 29. Spadiseius calyptrogynae n. sp., adult male: 26, subcapitulum; 27, chelicera and spermatodactyl, paraxial view; 28, tarsus II, anterolateral view; 29, tarsus IV, dorsal view, without pretarsus. Abbreviation: k, keel." httpUri="https://zenodo.org/record/180488/files/figure.png" pageId="15" pageNumber="16">Fig. 29</figureCitation>
|
||
). Leg IV with trochanter seta
|
||
<emphasis id="C89FEAC5FF9BFFDD3EA2FF6D24CBBB6F" box="[989,1018,232,257]" italics="true" pageId="15" pageNumber="16">ad</emphasis>
|
||
and femoral seta
|
||
<emphasis id="C89FEAC5FF9BFFDD39ADFF6D2236BB6F" box="[1234,1287,232,257]" italics="true" pageId="15" pageNumber="16">ad-2</emphasis>
|
||
attenuate or clavate. Femur I with dorsal seta
|
||
<emphasis id="C89FEAC5FF9BFFDD3F65FE95257EBB47" box="[538,591,272,297]" italics="true" pageId="15" pageNumber="16">ad-1</emphasis>
|
||
variably rod-like (30) or oar-like (45) but slenderer than in female;
|
||
<emphasis id="C89FEAC5FF9BFFDD3819FE9522ADBB47" box="[1382,1436,272,297]" italics="true" pageId="15" pageNumber="16">ad-1</emphasis>
|
||
on femora II–IV variably oar-like (32–65) or attenuated (90–135), either way increasingly longer from II to IV. Legs I–IV with size and form of many dorsal and lateral setae variably polymorphic, oar-like or attenuated on genu, tibia and tarsus, and legs III–IV with some ventral setae variably attenuated on femur, genu, tibia and tarsus (
|
||
<figureCitation id="62D02A52FF9BFFDD3D94FE35264BBBA4" box="[235,378,432,458]" captionStart="FIGURES 65 – 69" captionStartId="31.[151,269,1219,1243]" captionTargetBox="[164,1431,194,1163]" captionTargetId="figure@31.[151,1436,194,1283]" captionTargetPageId="31" captionText="FIGURES 65 – 69. Figs. 65 – 67: Spadiseius spathiphyllae n. sp., adult male: 65, tarsus IV, dorsolateral view, without pretarsus; 66, 67, femora IV, dorsal view: 66, “ short-haired ” morph; 67, “ long-haired ” morph. Figs. 68, 69, Spadiseius calyptrogynae n. sp., adult male, femora IV, dorsal view: 68, “ wartless ” morph; 69, “ warted ” morph." httpUri="https://zenodo.org/record/180497/files/figure.png" pageId="15" pageNumber="16">Figs. 68, 69</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="AE94665FFF9BFFDD3DE8F9B422AABC27" box="[151,1435,1585,1609]" httpUri="https://zenodo.org/record/180485/files/figure.png" pageId="15" pageNumber="16" targetBox="[186,1435,514,1584]" targetPageId="15">
|
||
<paragraph id="FA5436D7FF9BFFDD3DE8F9B422AABC27" blockId="15.[151,1435,1585,1609]" box="[151,1435,1585,1609]" pageId="15" pageNumber="16">
|
||
<emphasis id="C89FEAC5FF9BFFDD3DE8F9B4266EBC27" bold="true" box="[151,351,1585,1609]" pageId="15" pageNumber="16">FIGURES 19, 20.</emphasis>
|
||
<taxonomicName id="3DEB4D54FF9BFFDD3C19F9B72549BC27" box="[358,632,1586,1609]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="calyptrogynae" status="sp. nov.">
|
||
<emphasis id="C89FEAC5FF9BFFDD3C19F9B72549BC27" box="[358,632,1586,1609]" italics="true" pageId="15" pageNumber="16">Spadiseius calyptrogynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D3AC57BEFF9BFFDD3F01F9B4258ABC27" box="[638,699,1585,1609]" pageId="15" pageNumber="16" rank="species">
|
||
<emphasis id="C89FEAC5FF9BFFDD3F01F9B4258ABC27" bold="true" box="[638,699,1585,1609]" pageId="15" pageNumber="16">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult female: 19, idiosoma, ventral aspect; 20, spermathecae, detail
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FA5436D7FF9BFFC23DB9F91A22A4BADC" blockId="15.[151,1436,1695,2001]" lastBlockId="16.[151,1436,152,418]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
|
||
<emphasis id="C89FEAC5FF9BFFDD3DB9F91A264CBCD7" bold="true" box="[198,381,1695,1721]" pageId="15" pageNumber="16">DEUTONYMPH.</emphasis>
|
||
Dorsal shield 405–450 long, 270–310 wide at level of lateral incisions by setae
|
||
<emphasis id="C89FEAC5FF9BFFDD385BF91A220CBCD6" box="[1316,1341,1695,1720]" italics="true" pageId="15" pageNumber="16">s6</emphasis>
|
||
(n = 4); shield lightly evenly reticulate over nearly entire surface (
|
||
<figureCitation id="62D02A52FF9BFFDD3E2CF942249DBC8F" box="[851,940,1735,1761]" captionStart="FIGURES 16, 17" captionStartId="13.[151,269,1731,1755]" captionTargetBox="[151,1435,556,1730]" captionTargetId="figure@13.[151,1436,554,1771]" captionTargetPageId="13" captionText="FIGURES 16, 17. Spadiseius calyptrogynae n. sp., deutonymph: 16, idiosoma, dorsal aspect; 17, idiosoma, ventral aspect. Setae added to protonymphal complement denoted without parentheses; supernumerary setae denoted by ‘ x’." httpUri="https://zenodo.org/record/180483/files/figure.png" pageId="15" pageNumber="16">Fig. 16</figureCitation>
|
||
); podonotal region with longitudinal distance from alveoli of
|
||
<emphasis id="C89FEAC5FF9BFFDD3CEEF96A2699BD66" box="[401,424,1775,1800]" italics="true" pageId="15" pageNumber="16">j1</emphasis>
|
||
to
|
||
<emphasis id="C89FEAC5FF9BFFDD3CB3F96A26D2BD66" box="[460,483,1775,1800]" italics="true" pageId="15" pageNumber="16">j5</emphasis>
|
||
nearly equal (0.9) to that from
|
||
<emphasis id="C89FEAC5FF9BFFDD3E2FF96A2457BD66" box="[848,870,1775,1800]" italics="true" pageId="15" pageNumber="16">j5</emphasis>
|
||
to
|
||
<emphasis id="C89FEAC5FF9BFFDD3EF4F96A2493BD66" box="[907,930,1775,1800]" italics="true" pageId="15" pageNumber="16">j6</emphasis>
|
||
. Dorsal shield with approximately 35 to 40 pairs of setae: about 17–18 pairs (
|
||
<emphasis id="C89FEAC5FF9BFFDD3F5CF8922430BD5E" box="[547,769,1815,1840]" italics="true" pageId="15" pageNumber="16">j1–j6, z2–z6, s3–s6</emphasis>
|
||
, and 2 to 4 pairs of extra setae) on podonotal region, and 18 pairs (
|
||
<emphasis id="C89FEAC5FF9BFFDD3C7BF8BA26CDBD36" box="[260,508,1855,1880]" italics="true" pageId="15" pageNumber="16">J1–J5, Z1–Z5, S1–S5</emphasis>
|
||
, and a few paired or asymmetrically unpaired setae amidst
|
||
<emphasis id="C89FEAC5FF9BFFDD39CEF8BA238FBD36" box="[1201,1214,1855,1880]" italics="true" pageId="15" pageNumber="16">J</emphasis>
|
||
series) on opisthonotal region. Dorsal setae mostly alike in simple form (most of them slightly swollen at base) and short length (10–15) in central area,
|
||
<emphasis id="C89FEAC5FF9BFFDD3CCFF80A257ABDC6" box="[432,587,1935,1960]" italics="true" pageId="15" pageNumber="16">s3–s6, S2–S3</emphasis>
|
||
slightly larger (20–23),
|
||
<emphasis id="C89FEAC5FF9BFFDD3E17F80A24B4BDC6" box="[872,901,1935,1960]" italics="true" pageId="15" pageNumber="16">S1</emphasis>
|
||
only half as long as adjacent setae
|
||
<emphasis id="C89FEAC5FF9BFFDD385BF80A220FBDC6" box="[1316,1342,1935,1960]" italics="true" pageId="15" pageNumber="16">s6</emphasis>
|
||
and
|
||
<emphasis id="C89FEAC5FF9BFFDD3808F80A22A4BDC6" box="[1399,1429,1935,1960]" italics="true" pageId="15" pageNumber="16">S2</emphasis>
|
||
, and
|
||
<emphasis id="C89FEAC5FF9BFFDD3DB6F8322673BDBE" box="[201,322,1975,2000]" italics="true" pageId="15" pageNumber="16">S4, S5, Z5</emphasis>
|
||
poorly tapered, slightly barbed and largest (25–28). Dorsal setae on soft cuticle mostly 10–15 long,
|
||
<emphasis id="C89FEAC5FF84FFC23DA6FF1D27C2BADF" box="[217,243,152,177]" italics="true" pageId="16" pageNumber="17">r3</emphasis>
|
||
and
|
||
<emphasis id="C89FEAC5FF84FFC23C54FF1D267DBADF" box="[299,332,152,177]" italics="true" pageId="16" pageNumber="17">R1</emphasis>
|
||
slightly longest (15–20). Peritremes and their shields extending anteriorly to level of setae
|
||
<emphasis id="C89FEAC5FF84FFC2380BFF1D22BCBADF" box="[1396,1421,152,177]" italics="true" pageId="16" pageNumber="17">s1</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FA5436D7FF84FFC23DB9FF452420BBCC" blockId="16.[151,1436,152,418]" pageId="16" pageNumber="17">
|
||
Tritosternum similar in form to that in adult female. Sternal shield smooth, with narrowly rounded posterior margin (
|
||
<figureCitation id="62D02A52FF84FFC23C53FF6D26B2BB6C" box="[300,387,232,258]" captionStart="FIGURES 16, 17" captionStartId="13.[151,269,1731,1755]" captionTargetBox="[151,1435,556,1730]" captionTargetId="figure@13.[151,1436,554,1771]" captionTargetPageId="13" captionText="FIGURES 16, 17. Spadiseius calyptrogynae n. sp., deutonymph: 16, idiosoma, dorsal aspect; 17, idiosoma, ventral aspect. Setae added to protonymphal complement denoted without parentheses; supernumerary setae denoted by ‘ x’." httpUri="https://zenodo.org/record/180483/files/figure.png" pageId="16" pageNumber="17">Fig. 17</figureCitation>
|
||
); setae
|
||
<emphasis id="C89FEAC5FF84FFC23CA3FF6D26CEBB6F" box="[476,511,232,257]" italics="true" pageId="16" pageNumber="17">st4</emphasis>
|
||
clearly shorter (8–10) than
|
||
<emphasis id="C89FEAC5FF84FFC23E38FF6D24ABBB6F" box="[839,922,232,257]" italics="true" pageId="16" pageNumber="17">st1–st3</emphasis>
|
||
(25–27). Anal shield unornamented, length subequal to greatest width (90–95) at level of anterior edge of anus; postanal seta subequally as long as paraanal setae (15–17). Opisthogaster with 8 to 10 pairs of ventral setae plus 1 or 2 adjacent
|
||
<emphasis id="C89FEAC5FF84FFC239D1FEBD23D5BB3F" box="[1198,1252,312,337]" italics="true" pageId="16" pageNumber="17">UR–</emphasis>
|
||
setae (
|
||
<emphasis id="C89FEAC5FF84FFC23846FEBD2258BB3F" box="[1337,1385,312,337]" italics="true" pageId="16" pageNumber="17">ZV4</emphasis>
|
||
and
|
||
<emphasis id="C89FEAC5FF84FFC23DE8FEE527F9BB17" box="[151,200,352,377]" italics="true" pageId="16" pageNumber="17">ZV5</emphasis>
|
||
sometimes absent), all similarly short (10–12), slender except for slightly larger
|
||
<emphasis id="C89FEAC5FF84FFC239CCFEE523D0BB17" box="[1203,1249,352,377]" italics="true" pageId="16" pageNumber="17">JV5</emphasis>
|
||
(15) (
|
||
<figureCitation id="62D02A52FF84FFC2384DFEE522BCBB14" box="[1330,1421,352,378]" captionStart="FIGURES 16, 17" captionStartId="13.[151,269,1731,1755]" captionTargetBox="[151,1435,556,1730]" captionTargetId="figure@13.[151,1436,554,1771]" captionTargetPageId="13" captionText="FIGURES 16, 17. Spadiseius calyptrogynae n. sp., deutonymph: 16, idiosoma, dorsal aspect; 17, idiosoma, ventral aspect. Setae added to protonymphal complement denoted without parentheses; supernumerary setae denoted by ‘ x’." httpUri="https://zenodo.org/record/180483/files/figure.png" pageId="16" pageNumber="17">Fig. 17</figureCitation>
|
||
). Metapodal plates broadly oval, greatest dimension 15.
|
||
</paragraph>
|
||
<caption id="AE94665FFF84FFC23DE8FA2D27E7BF8C" httpUri="https://zenodo.org/record/180486/files/figure.png" pageId="16" pageNumber="17" targetBox="[264,1317,480,1446]" targetPageId="16">
|
||
<paragraph id="FA5436D7FF84FFC23DE8FA2D27E7BF8C" blockId="16.[151,1436,1448,1506]" pageId="16" pageNumber="17">
|
||
<emphasis id="C89FEAC5FF84FFC23DE8FA2D266FBFAE" bold="true" box="[151,350,1448,1472]" pageId="16" pageNumber="17">FIGURES 21, 22.</emphasis>
|
||
<taxonomicName id="3DEB4D54FF84FFC23C1AFA2D2544BFD1" box="[357,629,1448,1471]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="calyptrogynae" status="sp. nov.">
|
||
<emphasis id="C89FEAC5FF84FFC23C1AFA2D2544BFD1" box="[357,629,1448,1471]" italics="true" pageId="16" pageNumber="17">Spadiseius calyptrogynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D3AC57BEFF84FFC23F02FA2D2588BFAE" box="[637,697,1448,1472]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="C89FEAC5FF84FFC23F02FA2D2588BFAE" bold="true" box="[637,697,1448,1472]" pageId="16" pageNumber="17">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult male, “wartless” morph: 21, idiosoma, dorsal aspect; 22, idiosoma,
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FA5436D7FF84FFC23DB9F99323CEBC36" blockId="16.[151,1436,1558,2024]" pageId="16" pageNumber="17">
|
||
Tectum, chelicerae, ventral gnathosomatic structures and palpi similar to those in adult female, except capitular setae not swollen basally, and without pair of gland-like structures between setae
|
||
<emphasis id="C89FEAC5FF84FFC239C0F9BB23C9BC39" box="[1215,1272,1598,1623]" italics="true" pageId="16" pageNumber="17">hyp1</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FA5436D7FF84FFC23DB9F9E32594BD1E" blockId="16.[151,1436,1558,2024]" pageId="16" pageNumber="17">
|
||
Relative leg lengths, excluding pretarsi, similar to those of female, I (360–380), II (310–325), III (325– 350), IV (380–400). Form of legs and their setae as in adult female, except leg II no thicker than legs III–IV; tarsus I with pedestal bearing pretarsus less defined than on protonymph; tarsi II–IV in dorsoventral view tapered apically beyond insertions of
|
||
<emphasis id="C89FEAC5FF84FFC23F31F95B258ABC99" box="[590,699,1758,1783]" italics="true" pageId="16" pageNumber="17">al-1, pl-1</emphasis>
|
||
. Coxa IV seta short (10), at most half as long as coxa III setae. Dorsal seta
|
||
<emphasis id="C89FEAC5FF84FFC23C5CF8832669BD71" box="[291,344,1798,1823]" italics="true" pageId="16" pageNumber="17">ad-1</emphasis>
|
||
increasingly longer on femora I to IV (10 to 20), thicker, poorly tapered but not rod- or oarlike on femur I and sometimes on femur II, but slightly oar-like on femur III and clearly so, in contrast with adjacent simple setae, on femur IV (
|
||
<figureCitation id="62D02A52FF84FFC23F40F8D325A9BD1E" box="[575,664,1878,1904]" captionStart="FIGURES 42 – 45" captionStartId="25.[151,269,1816,1840]" captionTargetBox="[154,1431,346,1815]" captionTargetId="figure@25.[151,1436,330,1936]" captionTargetPageId="25" captionText="FIGURES 42 – 45. Figs. 42, 43, Spadiseius spathiphyllae n. sp., deutonymph: 42, idiosoma, dorsal aspect; 43, idiosoma, ventral aspect; supernumerary setae denoted by ‘ x’. Figs. 44, 45, dorsal subapical femoral setae legs I – IV, deutonymph: 44, Spadiseius calyptrogynae n. sp.; 45, Spadiseius spathiphyllae n. sp." httpUri="https://zenodo.org/record/180492/files/figure.png" pageId="16" pageNumber="17">Fig. 44</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FA5436D7FF84FFC33DB9F8F8258CBB3C" blockId="16.[151,1436,1558,2024]" lastBlockId="17.[151,1436,152,338]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
|
||
<emphasis id="C89FEAC5FF84FFC23DB9F8F8264DBDF9" bold="true" box="[198,380,1917,1943]" pageId="16" pageNumber="17">PROTONYMPH.</emphasis>
|
||
Idiosoma 305–345 long, about 215–235 wide at level of setae
|
||
<emphasis id="C89FEAC5FF84FFC23927F8FB2340BDF9" box="[1112,1137,1918,1943]" italics="true" pageId="16" pageNumber="17">r5</emphasis>
|
||
(n = 5), with lightly sclerotized, fully reticulated podonotal and pygidial shields (
|
||
<figureCitation id="62D02A52FF84FFC23E4FF82324B6BDAE" box="[816,903,1958,1984]" captionStart="FIGURES 13 – 15" captionStartId="11.[151,269,1529,1553]" captionTargetBox="[156,1435,492,1528]" captionTargetId="figure@11.[151,1436,394,1627]" captionTargetPageId="11" captionText="FIGURES 13 – 15. Spadiseius calyptrogynae n. sp., protonymph: 13, idiosoma, dorsal aspect; 14, idiosoma, ventral aspect; 15, tectum. Setae added to larval complement denoted without parentheses." httpUri="https://zenodo.org/record/180482/files/figure.png" pageId="16" pageNumber="17">Fig. 13</figureCitation>
|
||
); podonotal shield 207–220 long, 170–190 at widest level near posterior margin, with longitudinal distance from alveoli of
|
||
<emphasis id="C89FEAC5FF84FFC2395BF84B230ABD89" box="[1060,1083,1998,2023]" italics="true" pageId="16" pageNumber="17">j1</emphasis>
|
||
to
|
||
<emphasis id="C89FEAC5FF84FFC2391EF84B2349BD89" box="[1121,1144,1998,2023]" italics="true" pageId="16" pageNumber="17">j5</emphasis>
|
||
about 0.8 that from
|
||
<emphasis id="C89FEAC5FF84FFC23818F84B224CBD89" box="[1383,1405,1998,2023]" italics="true" pageId="16" pageNumber="17">j5</emphasis>
|
||
to
|
||
<emphasis id="C89FEAC5FF85FFC33DE8FF1D279FBADF" box="[151,174,152,177]" italics="true" pageId="17" pageNumber="18">j6</emphasis>
|
||
; pygidial shield 72–80 along midlength, 143–175 at widest level near seta
|
||
<emphasis id="C89FEAC5FF85FFC33948FF1D2364BADF" box="[1079,1109,152,177]" italics="true" pageId="17" pageNumber="18">S4</emphasis>
|
||
, with anterior margin shallowly excavated at either side of midline, at level of setae
|
||
<emphasis id="C89FEAC5FF85FFC33E2DFF45245FBAB7" box="[850,878,192,217]" italics="true" pageId="17" pageNumber="18">J2</emphasis>
|
||
, and usually with poroid-bearing protuberance laterad seta
|
||
<emphasis id="C89FEAC5FF85FFC33C5BFF6D2670BB6F" box="[292,321,232,257]" italics="true" pageId="17" pageNumber="18">S5</emphasis>
|
||
. Dorsal body setae mostly of similar short length (12–20) and simple form, opisthonotal setae
|
||
<emphasis id="C89FEAC5FF85FFC33DE8FE952787BB47" box="[151,182,272,297]" italics="true" pageId="17" pageNumber="18">Z5</emphasis>
|
||
slightly longer (22–25),
|
||
<emphasis id="C89FEAC5FF85FFC33CA4FE952562BB47" box="[475,595,272,297]" italics="true" pageId="17" pageNumber="18">S4, S5, Z5</emphasis>
|
||
slightly barbed, clunals
|
||
<emphasis id="C89FEAC5FF85FFC33E0CFE9524BFBB47" box="[883,910,272,297]" italics="true" pageId="17" pageNumber="18">J5</emphasis>
|
||
shortest (5). Peritremes extending anteriorly nearly to level of anterior margins of coxae III.
|
||
</paragraph>
|
||
<caption id="AE94665FFF85FFC33DE8F9A023D3BC31" httpUri="https://zenodo.org/record/180487/files/figure.png" pageId="17" pageNumber="18" targetBox="[151,1427,403,1572]" targetPageId="17">
|
||
<paragraph id="FA5436D7FF85FFC33DE8F9A023D3BC31" blockId="17.[151,1436,1573,1631]" pageId="17" pageNumber="18">
|
||
<emphasis id="C89FEAC5FF85FFC33DE8F9A0266EBC53" bold="true" box="[151,351,1573,1597]" pageId="17" pageNumber="18">FIGURES 23–25.</emphasis>
|
||
<taxonomicName id="3DEB4D54FF85FFC33C18F9A02548BC52" box="[359,633,1573,1596]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="calyptrogynae" status="sp. nov.">
|
||
<emphasis id="C89FEAC5FF85FFC33C18F9A02548BC52" box="[359,633,1573,1596]" italics="true" pageId="17" pageNumber="18">Spadiseius calyptrogynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D3AC57BEFF85FFC33FFEF9A0258FBC53" box="[641,702,1573,1597]" pageId="17" pageNumber="18" rank="species">
|
||
<emphasis id="C89FEAC5FF85FFC33FFEF9A0258FBC53" bold="true" box="[641,702,1573,1597]" pageId="17" pageNumber="18">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult male: 23, idiosoma, “warted morph”, dorsal aspect; 24, peritrematal shield and detail of enlarged gland pore; 25, greatly enlarged detail of peritrematal gland pore.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FA5436D7FF85FFC33DB9F9162319BD1B" blockId="17.[151,1437,1683,2029]" pageId="17" pageNumber="18">
|
||
Tritosternum similar in form to that in deutonymph and adult female, laciniae free for about two-thirds of length (50). Sternal shield (
|
||
<figureCitation id="62D02A52FF85FFC33C99F93E250CBCBB" box="[486,573,1723,1749]" captionStart="FIGURES 13 – 15" captionStartId="11.[151,269,1529,1553]" captionTargetBox="[156,1435,492,1528]" captionTargetId="figure@11.[151,1436,394,1627]" captionTargetPageId="11" captionText="FIGURES 13 – 15. Spadiseius calyptrogynae n. sp., protonymph: 13, idiosoma, dorsal aspect; 14, idiosoma, ventral aspect; 15, tectum. Setae added to larval complement denoted without parentheses." httpUri="https://zenodo.org/record/180482/files/figure.png" pageId="17" pageNumber="18">Fig. 14</figureCitation>
|
||
) smooth; setae
|
||
<emphasis id="C89FEAC5FF85FFC33F85F93E242CBCBA" box="[762,797,1723,1748]" italics="true" pageId="17" pageNumber="18">st5</emphasis>
|
||
inserted on soft cuticle much shorter (6) than
|
||
<emphasis id="C89FEAC5FF85FFC33836F93E22AABCBA" box="[1353,1435,1723,1748]" italics="true" pageId="17" pageNumber="18">st1–st3</emphasis>
|
||
(15) and shorter than opisthogastric setae; paragenital poroids absent. Anal shield unornamented, similar in form and structure to those on deutonymph, length subequal to greatest width (60–65); postanal seta subequally as long as para-anal setae (12). Opisthogastric setae on soft cuticle all similarly short (9–10) except for slightly larger
|
||
<emphasis id="C89FEAC5FF85FFC33C3EF8DE265FBD1A" box="[321,366,1883,1908]" italics="true" pageId="17" pageNumber="18">JV5</emphasis>
|
||
(12). Metapodal plates faint, elliptical (6 length, 10 width).
|
||
</paragraph>
|
||
<paragraph id="FA5436D7FF85FFC33DB9F806226DBDAB" blockId="17.[151,1437,1683,2029]" pageId="17" pageNumber="18">
|
||
Tectum more broadly triangular and bluntly pointed than in deutonymph (
|
||
<figureCitation id="62D02A52FF85FFC33944F80623A2BDF3" box="[1083,1171,1923,1949]" captionStart="FIGURES 13 – 15" captionStartId="11.[151,269,1529,1553]" captionTargetBox="[156,1435,492,1528]" captionTargetId="figure@11.[151,1436,394,1627]" captionTargetPageId="11" captionText="FIGURES 13 – 15. Spadiseius calyptrogynae n. sp., protonymph: 13, idiosoma, dorsal aspect; 14, idiosoma, ventral aspect; 15, tectum. Setae added to larval complement denoted without parentheses." httpUri="https://zenodo.org/record/180482/files/figure.png" pageId="17" pageNumber="18">Fig. 15</figureCitation>
|
||
); other gnathosomatic structures similar to those in deutonymph, except palpi with normal protonymphal complement of setae.
|
||
</paragraph>
|
||
<paragraph id="FA5436D7FF85FFC03DB9F856225DBB44" blockId="17.[151,1437,1683,2029]" lastBlockId="18.[151,1436,152,298]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
|
||
Leg lengths, excluding pretarsi, I (280–305), II (232–268), III (240–275), IV (270–325); pretarsi II–IV with apical setalike processes nearly half as long as length of pretarsi to base of claws. Form of legs and their setae generally as in deutonymph, except tarsus I apically with more clearly defined pedestal (5) bearing pretarsus. Coxa IV seta shorter (11) than coxa III setae (15). Femora I–IV with dorsal seta
|
||
<emphasis id="C89FEAC5FF86FFC039E2FF6D23E2BB6F" box="[1181,1235,232,257]" italics="true" pageId="18" pageNumber="19">ad-1</emphasis>
|
||
similar in length (about 10), slightly more thick and setulose than adjacent setae, but tapered, not rod- or oar-like (
|
||
<figureCitation id="62D02A52FF86FFC03878FE952251BB44" box="[1287,1376,272,298]" captionStart="FIGURES 38 – 41" captionStartId="24.[151,269,1163,1187]" captionTargetBox="[168,1419,313,1131]" captionTargetId="figure@24.[148,1433,288,1233]" captionTargetPageId="24" captionText="FIGURES 38 – 41. Dorsal subapical femoral setae of legs. Figs. 38, 39, larval femora I – III: 38, Spadiseius calyptrogynae n. sp.; 39, Spadiseius spathiphyllae n. sp. Figs. 40, 41, protonymphal femora I – IV: 40, S. calyptrogynae n. sp.; 41, S." httpUri="https://zenodo.org/record/180491/files/figure.png" pageId="18" pageNumber="19">Fig. 40</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="AE94665FFF86FFC03DE8FB672268BF72" httpUri="https://zenodo.org/record/180488/files/figure.png" pageId="18" pageNumber="19" targetBox="[151,1435,354,1192]" targetPageId="18">
|
||
<paragraph id="FA5436D7FF86FFC03DE8FB672268BF72" blockId="18.[151,1436,1250,1308]" pageId="18" pageNumber="19">
|
||
<emphasis id="C89FEAC5FF86FFC03DE8FB672650BE94" bold="true" box="[151,353,1250,1274]" pageId="18" pageNumber="19">FIGURES 26–29.</emphasis>
|
||
<taxonomicName id="3DEB4D54FF86FFC03C14FB67254FBE97" box="[363,638,1250,1273]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="calyptrogynae" status="sp. nov.">
|
||
<emphasis id="C89FEAC5FF86FFC03C14FB67254FBE97" box="[363,638,1250,1273]" italics="true" pageId="18" pageNumber="19">Spadiseius calyptrogynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D3AC57BEFF86FFC03FF6FB6725F9BE94" box="[649,712,1250,1274]" pageId="18" pageNumber="19" rank="species">
|
||
<emphasis id="C89FEAC5FF86FFC03FF6FB6725F9BE94" bold="true" box="[649,712,1250,1274]" pageId="18" pageNumber="19">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult male: 26, subcapitulum; 27, chelicera and spermatodactyl, paraxial view; 28, tarsus II, anterolateral view; 29, tarsus IV, dorsal view, without pretarsus. Abbreviation:
|
||
<emphasis id="C89FEAC5FF86FFC03876FA812229BF72" bold="true" box="[1289,1304,1284,1308]" pageId="18" pageNumber="19">k</emphasis>
|
||
, keel.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FA5436D7FF86FFC03DB9FACA2340BCC4" blockId="18.[151,1436,1359,2026]" pageId="18" pageNumber="19">
|
||
<emphasis id="C89FEAC5FF86FFC03DB9FACA2615BF07" bold="true" box="[198,292,1359,1385]" pageId="18" pageNumber="19">LARVA.</emphasis>
|
||
Idiosoma 290–360 long, 175–215 wide at level of setae
|
||
<emphasis id="C89FEAC5FF86FFC03EC8FAD524E0BF07" box="[951,977,1360,1385]" italics="true" pageId="18" pageNumber="19">s4</emphasis>
|
||
(n = 6), with lightly sclerotized, faintly and incompletely lineated podonotal and pygidial shields (
|
||
<figureCitation id="62D02A52FF86FFC03E36FAFD2490BFFC" box="[841,929,1400,1426]" captionStart="FIGURES 10 – 12" captionStartId="10.[151,269,1581,1605]" captionTargetBox="[168,1435,385,1580]" captionTargetId="figure@10.[151,1436,354,1612]" captionTargetPageId="10" captionText="FIGURES 10 – 12. Spadiseius calyptrogynae n. sp., larva: 10, idiosoma, dorsal aspect; 11, idiosoma, ventral aspect; 12, tectum, variations." httpUri="https://zenodo.org/record/180481/files/figure.png" pageId="18" pageNumber="19">Fig. 10</figureCitation>
|
||
); podonotal shield 185–205 long, 150–155 at widest level near posterior margin, with longitudinal distance from alveoli of
|
||
<emphasis id="C89FEAC5FF86FFC0393AFA25236DBFD7" box="[1093,1116,1440,1465]" italics="true" pageId="18" pageNumber="19">j1</emphasis>
|
||
to
|
||
<emphasis id="C89FEAC5FF86FFC039FDFA2523A8BFD7" box="[1154,1177,1440,1465]" italics="true" pageId="18" pageNumber="19">j5</emphasis>
|
||
0.7 that from
|
||
<emphasis id="C89FEAC5FF86FFC0383EFA252266BFD7" box="[1345,1367,1440,1465]" italics="true" pageId="18" pageNumber="19">j5</emphasis>
|
||
to
|
||
<emphasis id="C89FEAC5FF86FFC03801FA2522A5BFD7" box="[1406,1428,1440,1465]" italics="true" pageId="18" pageNumber="19">j6</emphasis>
|
||
, and with undulate lateral margins constricted behind levels of setae
|
||
<emphasis id="C89FEAC5FF86FFC03EC4FA4D24E4BF8F" box="[955,981,1480,1505]" italics="true" pageId="18" pageNumber="19">z2</emphasis>
|
||
and
|
||
<emphasis id="C89FEAC5FF86FFC0396FFA4D231BBF8F" box="[1040,1066,1480,1505]" italics="true" pageId="18" pageNumber="19">s4</emphasis>
|
||
; pygidial shield sparsely punctate, 65–70 along midlength in dorsal view (but further length of 15–20 folded under ventrally), 145–150 at widest level near anterior margin, with anterior margin deeply invaginated on either side of midline, at level of setal alveoli
|
||
<emphasis id="C89FEAC5FF86FFC03C54F9C52677BC37" box="[299,326,1600,1625]" italics="true" pageId="18" pageNumber="19">J3</emphasis>
|
||
, and posterior margin also deeply invaginated on either side of midline at level of setal alveoli
|
||
<emphasis id="C89FEAC5FF86FFC03DE8F9ED2782BCEF" box="[151,179,1640,1665]" italics="true" pageId="18" pageNumber="19">J5</emphasis>
|
||
(
|
||
<figureCitation id="62D02A52FF86FFC03DBDF9ED2626BCEC" box="[194,279,1640,1666]" captionStart="FIGURES 10 – 12" captionStartId="10.[151,269,1581,1605]" captionTargetBox="[168,1435,385,1580]" captionTargetId="figure@10.[151,1436,354,1612]" captionTargetPageId="10" captionText="FIGURES 10 – 12. Spadiseius calyptrogynae n. sp., larva: 10, idiosoma, dorsal aspect; 11, idiosoma, ventral aspect; 12, tectum, variations." httpUri="https://zenodo.org/record/180481/files/figure.png" pageId="18" pageNumber="19">Fig. 11</figureCitation>
|
||
). Dorsal body setae similarly short (6–9), simple on podonotal shield, slightly shorter (4–5) on soft cuticle, but
|
||
<emphasis id="C89FEAC5FF86FFC03C5FF915265CBCC7" box="[288,365,1680,1705]" italics="true" pageId="18" pageNumber="19">Z3–Z4</emphasis>
|
||
on pygidial shield much longer (37–40), thicker, capitate distally.
|
||
</paragraph>
|
||
<paragraph id="FA5436D7FF86FFC03DB9F93D2541BD1C" blockId="18.[151,1436,1359,2026]" pageId="18" pageNumber="19">
|
||
Tritosternum with base longer (17–25) than wide (9–11); laciniae free for about three-fourths of total length (40 excluding base). Sternal shield smooth, with setae
|
||
<emphasis id="C89FEAC5FF86FFC03E17F965248BBC97" box="[872,954,1760,1785]" italics="true" pageId="18" pageNumber="19">st1–st3</emphasis>
|
||
similar in length (12–14) (
|
||
<figureCitation id="62D02A52FF86FFC03988F965227CBC94" box="[1271,1357,1760,1786]" captionStart="FIGURES 10 – 12" captionStartId="10.[151,269,1581,1605]" captionTargetBox="[168,1435,385,1580]" captionTargetId="figure@10.[151,1436,354,1612]" captionTargetPageId="10" captionText="FIGURES 10 – 12. Spadiseius calyptrogynae n. sp., larva: 10, idiosoma, dorsal aspect; 11, idiosoma, ventral aspect; 12, tectum, variations." httpUri="https://zenodo.org/record/180481/files/figure.png" pageId="18" pageNumber="19">Fig. 11</figureCitation>
|
||
). Anal shield smooth, slightly wider (50–55) than long (45–50); para-anal setae (15–18) longer than postanal seta (12–13). Four pairs of opisthogastric setae (
|
||
<emphasis id="C89FEAC5FF86FFC03FE4F8B525FABD27" box="[667,715,1840,1865]" italics="true" pageId="18" pageNumber="19">ZV2</emphasis>
|
||
present) plus
|
||
<emphasis id="C89FEAC5FF86FFC03E0DF8B524BEBD27" box="[882,911,1840,1865]" italics="true" pageId="18" pageNumber="19">S5</emphasis>
|
||
and
|
||
<emphasis id="C89FEAC5FF86FFC03EB6F8B524D9BD27" box="[969,1000,1840,1865]" italics="true" pageId="18" pageNumber="19">Z5</emphasis>
|
||
on soft cuticle all similarly short (3– 4) except for slightly longer
|
||
<emphasis id="C89FEAC5FF86FFC03C9AF8DD2522BD1F" box="[485,531,1880,1905]" italics="true" pageId="18" pageNumber="19">JV2</emphasis>
|
||
(7–14).
|
||
</paragraph>
|
||
<paragraph id="FA5436D7FF86FFC03DB9F8052690BD84" blockId="18.[151,1436,1359,2026]" pageId="18" pageNumber="19">
|
||
Tectum with anterior margin bluntly unevenly triangular (
|
||
<figureCitation id="62D02A52FF86FFC03E0FF80524F9BDF4" box="[880,968,1920,1946]" captionStart="FIGURES 10 – 12" captionStartId="10.[151,269,1581,1605]" captionTargetBox="[168,1435,385,1580]" captionTargetId="figure@10.[151,1436,354,1612]" captionTargetPageId="10" captionText="FIGURES 10 – 12. Spadiseius calyptrogynae n. sp., larva: 10, idiosoma, dorsal aspect; 11, idiosoma, ventral aspect; 12, tectum, variations." httpUri="https://zenodo.org/record/180481/files/figure.png" pageId="18" pageNumber="19">Fig. 12</figureCitation>
|
||
). Fixed digit of chelicera with dentition and pilus dentilis lobe as in protonymph; movable chela 20–22 long, edentate or with a minute tooth, and with ventral mucro (
|
||
<figureCitation id="62D02A52FF86FFC03C35F85526A3BD84" box="[330,402,2000,2026]" captionStart="FIGURES 1 – 9" captionStartId="9.[151,269,1577,1601]" captionTargetBox="[152,1425,197,1576]" captionTargetId="figure@9.[151,1436,197,1643]" captionTargetPageId="9" captionText="FIGURES 1 – 9. Figs. 1 – 5, Spadiseius calyptrogynae n. sp., adult female: 1, subcapitulum; 2, labrum, ventral view; 3, salivary stylus; 4, tritosternum; 5, palpus, ventral view. Fig. 6, S. calyptrogynae n. sp., larva, chelicera, paraxial view. Figs. 7 – 9, Spadiseius spathiphyllae n. sp., adult female: 7, tectum; 8, chelicera, dorso-paraxial view; 9 a, chelicera, movable digit, dorsal view; 9 b, chelicera, movable digit, antiaxial-ventral view (Figs. 2 – 4, same scale)." httpUri="https://zenodo.org/record/180480/files/figure.png" pageId="18" pageNumber="19">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="AE94665FFF87FFC13DE8FA3D260FBC7B" httpUri="https://zenodo.org/record/180489/files/figure.png" pageId="19" pageNumber="20" targetBox="[164,1434,194,1391]" targetPageId="19">
|
||
<paragraph id="FA5436D7FF87FFC13DE8FA3D260FBC7B" blockId="19.[151,1436,1464,1557]" pageId="19" pageNumber="20">
|
||
<emphasis id="C89FEAC5FF87FFC13DE8FA3D266EBFBE" bold="true" box="[151,351,1464,1488]" pageId="19" pageNumber="20">FIGURES 30–34.</emphasis>
|
||
<taxonomicName id="3DEB4D54FF87FFC13C18FA3C2548BFBE" box="[359,633,1465,1488]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="calyptrogynae" status="sp. nov.">
|
||
<emphasis id="C89FEAC5FF87FFC13C18FA3C2548BFBE" box="[359,633,1465,1488]" italics="true" pageId="19" pageNumber="20">Spadiseius calyptrogynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D3AC57BEFF87FFC13FFEFA3D258EBFBE" box="[641,703,1464,1488]" pageId="19" pageNumber="20" rank="species">
|
||
<emphasis id="C89FEAC5FF87FFC13FFEFA3D258EBFBE" bold="true" box="[641,703,1464,1488]" pageId="19" pageNumber="20">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult female. Figs.30–33, dorsal views of legs excluding tarsi: 30, leg I; 31, leg II; 32, leg III; 33, leg IV; anterolateral (
|
||
<emphasis id="C89FEAC5FF87FFC13FB3FA5E25D0BF9D" bold="true" box="[716,737,1499,1523]" pageId="19" pageNumber="20">al</emphasis>
|
||
) and posterolateral (
|
||
<emphasis id="C89FEAC5FF87FFC13EBFFA5E24E7BF9D" bold="true" box="[960,982,1499,1523]" pageId="19" pageNumber="20">pl</emphasis>
|
||
) faces indicated by arrows. Fig. 34, tarsus IV, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FA5436D7FF87FFC13DB9F9CC2436BD6D" blockId="19.[151,1437,1609,2035]" pageId="19" pageNumber="20">
|
||
Leg lengths, excluding pretarsi: I (255–280), II (192–228), III (210–230); pretarsi II–III with apical setalike processes about half as long as length of pretarsi to base of claws. Form of legs and their setae generally as in protonymph. Tarsus I apically with well defined pedestal (7) bearing pretarsus. Femora I–III with dorsal seta
|
||
<emphasis id="C89FEAC5FF87FFC13DB3F9442630BCB4" box="[204,257,1729,1754]" italics="true" pageId="19" pageNumber="20">ad-1</emphasis>
|
||
(8–9) resembling
|
||
<emphasis id="C89FEAC5FF87FFC13CA6F944253FBCB4" box="[473,526,1729,1754]" italics="true" pageId="19" pageNumber="20">pd-1</emphasis>
|
||
of femur and
|
||
<emphasis id="C89FEAC5FF87FFC13FCBF94425CCBCB4" box="[692,765,1729,1754]" italics="true" pageId="19" pageNumber="20">ad–pd</emphasis>
|
||
setae of genua, i.e., slightly more thick and setulose than other femoral setae, but not rod- or oar-like (
|
||
<figureCitation id="62D02A52FF87FFC13FDDF96C25C8BD6D" box="[674,761,1769,1795]" captionStart="FIGURES 38 – 41" captionStartId="24.[151,269,1163,1187]" captionTargetBox="[168,1419,313,1131]" captionTargetId="figure@24.[148,1433,288,1233]" captionTargetPageId="24" captionText="FIGURES 38 – 41. Dorsal subapical femoral setae of legs. Figs. 38, 39, larval femora I – III: 38, Spadiseius calyptrogynae n. sp.; 39, Spadiseius spathiphyllae n. sp. Figs. 40, 41, protonymphal femora I – IV: 40, S. calyptrogynae n. sp.; 41, S." httpUri="https://zenodo.org/record/180491/files/figure.png" pageId="19" pageNumber="20">Fig. 38</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B2F1655CFF87FFC63DB9F8952560BB14" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="materials_examined">
|
||
<paragraph id="FA5436D7FF87FFC13DB9F89522ADBD9C" blockId="19.[151,1437,1609,2035]" pageId="19" pageNumber="20">
|
||
<emphasis id="C89FEAC5FF87FFC13DB9F8952648BD44" bold="true" box="[198,377,1808,1834]" pageId="19" pageNumber="20">
|
||
<typeStatus id="25508875FF87FFC13DB9F8952630BD44" box="[198,257,1808,1834]" pageId="19" pageNumber="20">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
<collectingCountry id="82FC7647FF87FFC13CFDF8942501BD45" box="[386,560,1809,1835]" name="Costa Rica" pageId="19" pageNumber="20">COSTA RICA</collectingCountry>
|
||
, Heredia Province, La Selva Biological Station, (
|
||
<geoCoordinate id="9FDF5010FF87FFC13913F89423CEBD45" box="[1132,1279,1809,1835]" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="15" value="10.433611">10° 26' 1" N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="9FDF5010FF87FFC13874F89422A5BD45" box="[1291,1428,1809,1835]" direction="west" orientation="longitude" pageId="19" pageNumber="20" precision="15" value="-84.01723">84° 1' 2" W</geoCoordinate>
|
||
) elevation
|
||
<quantity id="3D139B32FF87FFC13C75F8BC26B4BD3D" box="[266,389,1849,1875]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="19" pageNumber="20" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50–150 m</quantity>
|
||
:
|
||
<typeStatus id="25508875FF87FFC13CEEF8BC2520BD3D" box="[401,529,1849,1875]" pageId="19" pageNumber="20" type="holotype">HOLOTYPE</typeStatus>
|
||
: adult female, Research Swamp, ex inflorescences of Palmaceae,
|
||
<taxonomicName id="3DEB4D54FF87FFC13863F8BC265BBD14" class="Liliopsida" family="Arecaceae" genus="Calyptrogyne" kingdom="Plantae" order="Arecales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="species" species="sarapiquensis">
|
||
<emphasis id="C89FEAC5FF87FFC13863F8BC265BBD14" italics="true" pageId="19" pageNumber="20">Calyptrogyne sarapiquensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<date id="8E551017FF87FFC13C07F8E4257BBD15" box="[376,586,1889,1915]" pageId="19" pageNumber="20" value="1994-02-10">10 February 1994</date>
|
||
, coll. S. Cunningham & E.E. Lindquist.
|
||
<typeStatus id="25508875FF87FFC1395BF8E4239FBD15" box="[1060,1198,1889,1915]" pageId="19" pageNumber="20" type="paratype">PARATYPES</typeStatus>
|
||
:
|
||
<specimenCount id="ECEDFD5EFF87FFC139C2F8E4223FBD15" box="[1213,1294,1889,1915]" pageId="19" pageNumber="20" type="adult">4 adult</specimenCount>
|
||
females,
|
||
<specimenCount id="ECEDFD5EFF87FFC13800F8E427E3BDCD" pageId="19" pageNumber="20" type="adult">10 adult</specimenCount>
|
||
males, 6 deutonymphs, 14 protonymphs,
|
||
<specimenCount id="ECEDFD5EFF87FFC13FBBF80C2414BDCD" box="[708,805,1929,1955]" pageId="19" pageNumber="20" type="larva">8 larvae</specimenCount>
|
||
, with same data as
|
||
<typeStatus id="25508875FF87FFC13971F80C2349BDCD" box="[1038,1144,1929,1955]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
;
|
||
<specimenCount id="ECEDFD5EFF87FFC139FAF80C23E9BDCD" box="[1157,1240,1929,1955]" pageId="19" pageNumber="20" type="adult">2 adult</specimenCount>
|
||
females,
|
||
<specimenCount id="ECEDFD5EFF87FFC13834F80C22ACBDCD" box="[1355,1437,1929,1955]" pageId="19" pageNumber="20" type="adult">1 adult</specimenCount>
|
||
male, 6 protonymphs,
|
||
<specimenCount id="ECEDFD5EFF87FFC13CE1F83426CFBDA5" box="[414,510,1969,1995]" pageId="19" pageNumber="20" type="larva">3 larvae</specimenCount>
|
||
, Research Swamp, ex inflorescences of
|
||
<taxonomicName id="3DEB4D54FF87FFC13EABF83423A8BDA4" box="[980,1177,1969,1994]" class="Liliopsida" family="Arecaceae" genus="Calyptrogyne" kingdom="Plantae" order="Arecales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="species" species="sarapiquensis">
|
||
<emphasis id="C89FEAC5FF87FFC13EABF83423A8BDA4" box="[980,1177,1969,1994]" italics="true" pageId="19" pageNumber="20">C. sarapiquensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<date id="8E551017FF87FFC139D8F834226DBDA5" box="[1191,1372,1969,1995]" pageId="19" pageNumber="20" value="1991-03-15">15 March 1991</date>
|
||
, coll. S. Cunningham;
|
||
<specimenCount id="ECEDFD5EFF87FFC13C16F85C26F0BD9D" box="[361,449,2009,2035]" pageId="19" pageNumber="20" type="adult">1 adult</specimenCount>
|
||
female,
|
||
<specimenCount id="ECEDFD5EFF87FFC13F4CF85C25B8BD9D" box="[563,649,2009,2035]" pageId="19" pageNumber="20" type="adult">1 adult</specimenCount>
|
||
male, 2 protonymphs, ex scarab beetle on inflorescence of
|
||
<emphasis id="C89FEAC5FF87FFC138FEF85C22ADBD9C" box="[1409,1436,2009,2034]" italics="true" pageId="19" pageNumber="20">C.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="FA5436D7FF80FFC63DE8FF1D2560BB14" blockId="20.[151,1437,152,2018]" pageId="20" pageNumber="21">
|
||
<taxonomicName id="3DEB4D54FF80FFC63DE8FF1D260BBADF" box="[151,314,152,177]" class="Liliopsida" family="Arecaceae" genus="Calyptrogyne" kingdom="Plantae" order="Arecales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="sarapiquensis">
|
||
<emphasis id="C89FEAC5FF80FFC63DE8FF1D260BBADF" box="[151,314,152,177]" italics="true" pageId="20" pageNumber="21">sarapiquensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<date id="8E551017FF80FFC63C33FF1D2538BADC" box="[332,521,152,178]" pageId="20" pageNumber="21" value="1991-03-27">27 March 1991</date>
|
||
, coll. S. Cunningham;
|
||
<specimenCount id="ECEDFD5EFF80FFC63E5BFF1D2449BADC" box="[804,888,152,178]" pageId="20" pageNumber="21" type="adult">1 adult</specimenCount>
|
||
female, 1 protonymph, Research Swamp, ex elytrum of scarab,
|
||
<taxonomicName id="3DEB4D54FF80FFC63C08FF45257EBAB7" box="[375,591,192,217]" class="Insecta" family="Rutelidae" genus="Lagochile" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="collaris">
|
||
<emphasis id="C89FEAC5FF80FFC63C08FF45257EBAB7" box="[375,591,192,217]" italics="true" pageId="20" pageNumber="21">Lagochile collaris</emphasis>
|
||
</taxonomicName>
|
||
, on inflorescences of
|
||
<taxonomicName id="3DEB4D54FF80FFC63E27FF45232EBAB7" box="[856,1055,192,217]" class="Liliopsida" family="Arecaceae" genus="Calyptrogyne" kingdom="Plantae" order="Arecales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="sarapiquensis">
|
||
<emphasis id="C89FEAC5FF80FFC63E27FF45232EBAB7" box="[856,1055,192,217]" italics="true" pageId="20" pageNumber="21">C. sarapiquensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<date id="8E551017FF80FFC63950FF4523F2BAB4" box="[1071,1219,192,218]" pageId="20" pageNumber="21" value="1994-05-02">2 May 1994</date>
|
||
, coll. S. Cunningham;
|
||
<specimenCount id="ECEDFD5EFF80FFC63DA5FF6D261DBB6C" box="[218,300,232,258]" pageId="20" pageNumber="21" type="adult">4 adult</specimenCount>
|
||
females, ex scarab,
|
||
<taxonomicName id="3DEB4D54FF80FFC63F62FF6D25C5BB6F" box="[541,756,232,257]" class="Insecta" family="Rutelidae" genus="Lagochile" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="collaris">
|
||
<emphasis id="C89FEAC5FF80FFC63F62FF6D25C5BB6F" box="[541,756,232,257]" italics="true" pageId="20" pageNumber="21">Lagochile collaris</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<date id="8E551017FF80FFC63E7BFF6D24E8BB6C" box="[772,985,232,258]" pageId="20" pageNumber="21" value="1996-02-22">22 February 1996</date>
|
||
, coll. M. Paniagua;
|
||
<specimenCount id="ECEDFD5EFF80FFC639B7FF6D2228BB6C" box="[1224,1305,232,258]" pageId="20" pageNumber="21" type="adult">2 adult</specimenCount>
|
||
females, 1 protonymph, ex curculionid beetle,
|
||
<date id="8E551017FF80FFC63F47FE95243BBB44" box="[568,778,272,298]" pageId="20" pageNumber="21" value="1994-02-24">24 February 1994</date>
|
||
, coll. S. Cunningham;
|
||
<specimenCount id="ECEDFD5EFF80FFC63969FE952354BB44" box="[1046,1125,272,298]" pageId="20" pageNumber="21" type="adult">4 adult</specimenCount>
|
||
females,
|
||
<specimenCount id="ECEDFD5EFF80FFC639A9FE952216BB44" box="[1238,1319,272,298]" pageId="20" pageNumber="21" type="adult">9 adult</specimenCount>
|
||
males, ex bats,
|
||
<date id="8E551017FF80FFC63DABFEBD261EBB3C" box="[212,303,312,338]" pageId="20" pageNumber="21" value="1993-01">January</date>
|
||
&
|
||
<date id="8E551017FF80FFC63C2AFEBD2533BB3C" box="[341,514,312,338]" pageId="20" pageNumber="21" value="1993-02">February 1993</date>
|
||
, coll. S. Cunningham;
|
||
<specimenCount id="ECEDFD5EFF80FFC63E71FEBD246FBB3C" box="[782,862,312,338]" pageId="20" pageNumber="21" type="adult">1 adult</specimenCount>
|
||
female,
|
||
<specimenCount id="ECEDFD5EFF80FFC63EBBFEBD2324BB3C" box="[964,1045,312,338]" pageId="20" pageNumber="21" type="adult">2 adult</specimenCount>
|
||
males, 1 protonymph, ex bats(?),
|
||
<date id="8E551017FF80FFC63DE8FEE5267DBB14" box="[151,332,352,378]" pageId="20" pageNumber="21" value="1995-03-20">20 March 1995</date>
|
||
, coll. S. Cunningham.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B2F1655CFF80FFC63DBAFE02230BBB9C" pageId="20" pageNumber="21" type="description">
|
||
<paragraph id="FA5436D7FF80FFC63DBAFE02230BBB9C" blockId="20.[151,1437,152,2018]" pageId="20" pageNumber="21">
|
||
<emphasis id="C89FEAC5FF80FFC63DBAFE022638BBCF" bold="true" box="[197,265,391,417]" pageId="20" pageNumber="21">Note:</emphasis>
|
||
The above data are based on the labels affixed to slide preparations or associated with residual samples preserved in ethanol.
|
||
<taxonomicName id="3DEB4D54FF80FFC63CB8FE352330BBA4" authority="H. Wendl. ex Burret" authorityName="H. Wendl. ex Burret" box="[455,1025,432,458]" class="Liliopsida" family="Arecaceae" genus="Calyptrogyne" kingdom="Plantae" order="Arecales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="sarapiquensis">
|
||
<emphasis id="C89FEAC5FF80FFC63CB8FE35243EBBA7" box="[455,783,432,457]" italics="true" pageId="20" pageNumber="21">Calyptrogyne sarapiquensis</emphasis>
|
||
H. Wendl. ex Burret
|
||
</taxonomicName>
|
||
is presently considered to be a junior synonym of
|
||
<taxonomicName id="3DEB4D54FF80FFC63C2EFE5D240ABB9C" authority="Linden & H. Wendl." authorityName="Linden & H. Wendl." box="[337,827,472,498]" class="Liliopsida" family="Arecaceae" genus="Calyptrogyne" kingdom="Plantae" order="Arecales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="ghiesbreghtiana">
|
||
<emphasis id="C89FEAC5FF80FFC63C2EFE5D2501BB9F" box="[337,560,472,497]" italics="true" pageId="20" pageNumber="21">C. ghiesbreghtiana</emphasis>
|
||
(Linden & H. Wendl.)
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="9E7A4B26FF80FFC63E07FE5D231FBB9C" author="Grayum" box="[888,1070,472,498]" pageId="20" pageNumber="37" refString="Grayum, M. H. (2003 b) Arecaceae. In: Hammel, B. E., Grayum, M. H., Herrera C. & Zamora, N. (Eds.), Manual de Plantas de Costa Rica. Volumen II, Gimnospermas y Monocotiledoneas (Agavaceae-Musaceae). Monographs in Systematic Botany from the Missouri Botanical Garden, St. Louis, Vol. 92, pp. 201 - 293." type="journal article" year="2003" yearSuffix="b">Grayum 2003b</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B2F1655CFF80FFC63DB9FE7A244CB82C" pageId="20" pageNumber="21" type="etymology">
|
||
<paragraph id="FA5436D7FF80FFC63DB9FE7A244CB82C" blockId="20.[151,1437,152,2018]" pageId="20" pageNumber="21">
|
||
<emphasis id="C89FEAC5FF80FFC63DB9FE7A2662B877" bold="true" box="[198,339,511,537]" pageId="20" pageNumber="21">Etymology.</emphasis>
|
||
The specific epithet,
|
||
<taxonomicName id="3DEB4D54FF80FFC63F2BFD8525CAB877" box="[596,763,512,537]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="calyptrogynae">
|
||
<emphasis id="C89FEAC5FF80FFC63F2BFD8525CAB877" box="[596,763,512,537]" italics="true" pageId="20" pageNumber="21">calyptrogynae</emphasis>
|
||
</taxonomicName>
|
||
, is based on the name of the genus of palmaceous plants in whose inflorescences these mites undergo their life histories.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B2F1655CFF80FFC63DB9FDCA2327BD8C" pageId="20" pageNumber="21" type="discussion">
|
||
<paragraph id="FA5436D7FF80FFC63DB9FDCA26C7BE84" blockId="20.[151,1437,152,2018]" pageId="20" pageNumber="21">
|
||
<emphasis id="C89FEAC5FF80FFC63DB9FDCA2671B807" bold="true" box="[198,320,591,617]" pageId="20" pageNumber="21">Remarks.</emphasis>
|
||
Of the five recognized species of small palms of the Neotropical genus
|
||
<taxonomicName id="3DEB4D54FF80FFC639F9FDD52215B807" box="[1158,1316,592,617]" class="Liliopsida" family="Arecaceae" genus="Calyptrogyne" kingdom="Plantae" order="Arecales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="C89FEAC5FF80FFC639F9FDD52215B807" box="[1158,1316,592,617]" italics="true" pageId="20" pageNumber="21">Calyptrogyne</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3DEB4D54FF80FFC6384CFDD52626B8FF" class="Liliopsida" family="Arecaceae" genus="Calyptrogyne" kingdom="Plantae" order="Arecales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="ghiesbreghtiana">
|
||
<emphasis id="C89FEAC5FF80FFC6384CFDD52626B8FF" italics="true" pageId="20" pageNumber="21">C. ghiesbreghtiana</emphasis>
|
||
</taxonomicName>
|
||
is the most widespread and variable species, occuring in both lowland rainforests and mid-elevation cloudforests on both the Atlantic and Pacific coasts, from southern
|
||
<collectingCountry id="82FC7647FF80FFC63E95FD252375B8D4" box="[1002,1092,672,698]" name="Mexico" pageId="20" pageNumber="21">Mexico</collectingCountry>
|
||
to
|
||
<collectingCountry id="82FC7647FF80FFC63915FD2523F6B8D4" box="[1130,1223,672,698]" name="Panama" pageId="20" pageNumber="21">Panama</collectingCountry>
|
||
(
|
||
<bibRefCitation id="9E7A4B26FF80FFC639A7FD2522A1B8D4" author="Grayum" box="[1240,1424,672,698]" pageId="20" pageNumber="37" refString="Grayum, M. H. (2003 b) Arecaceae. In: Hammel, B. E., Grayum, M. H., Herrera C. & Zamora, N. (Eds.), Manual de Plantas de Costa Rica. Volumen II, Gimnospermas y Monocotiledoneas (Agavaceae-Musaceae). Monographs in Systematic Botany from the Missouri Botanical Garden, St. Louis, Vol. 92, pp. 201 - 293." type="journal article" year="2003" yearSuffix="b">Grayum 2003b</bibRefCitation>
|
||
).
|
||
<taxonomicName id="3DEB4D54FF80FFC63DE8FD4D26CBB88F" box="[151,506,712,737]" class="Liliopsida" family="Arecaceae" genus="Calyptrogyne" kingdom="Plantae" order="Arecales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="ghiesbreghtiana">
|
||
<emphasis id="C89FEAC5FF80FFC63DE8FD4D26CBB88F" box="[151,506,712,737]" italics="true" pageId="20" pageNumber="21">Calyptrogyne ghiesbreghtiana</emphasis>
|
||
</taxonomicName>
|
||
is the only well documented case in the palm family
|
||
<taxonomicName id="3DEB4D54FF80FFC6390DFD4D23DDB88C" box="[1138,1260,712,738]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
|
||
of being pollinated, in part at least, by bats (
|
||
<bibRefCitation id="9E7A4B26FF80FFC63C85FD7525E8B964" author="Cunningham" box="[506,729,752,778]" pageId="20" pageNumber="36" refString="Cunningham, S. A. (1995) Ecological constraints on fruit initiation by Calyptrogyne ghiesbreghtiana (Arecaceae): floral herbivory, pollen availability and visitation by pollinating bats. American Journal of Botany, 82, 1527 - 1536." type="journal article" year="1995">Cunningham 1995</bibRefCitation>
|
||
).
|
||
<taxonomicName id="3DEB4D54FF80FFC63F92FD752329B967" box="[749,1048,752,777]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="calyptrogynae">
|
||
<emphasis id="C89FEAC5FF80FFC63F92FD752329B967" box="[749,1048,752,777]" italics="true" pageId="20" pageNumber="21">Spadiseius calyptrogynae</emphasis>
|
||
</taxonomicName>
|
||
is the first flower-dwelling melicharine mite recorded to use pollinating bats for dispersal (
|
||
<bibRefCitation id="9E7A4B26FF80FFC63E2DFC9D23FBB95C" author="Tschapka" box="[850,1226,792,818]" pageId="20" pageNumber="37" refString="Tschapka, M. & Cunningham, S. A. (2004) Flower mites of Calyptrogyne ghiesbreghtiana (Arecaceae): evidence for dispersal using pollinating bats. Biotropica, 36, 377 - 381." type="journal article" year="2004">Tschapka & Cunningham 2004</bibRefCitation>
|
||
). Eight species of bats, including nectar-specialists and frugivores, are known to visit inflorescences of
|
||
<taxonomicName id="3DEB4D54FF80FFC639F8FCC52256B937" box="[1159,1383,832,857]" class="Liliopsida" family="Arecaceae" genus="Calyptrogyne" kingdom="Plantae" order="Arecales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="ghiesbreghtiana">
|
||
<emphasis id="C89FEAC5FF80FFC639F8FCC52256B937" box="[1159,1383,832,857]" italics="true" pageId="20" pageNumber="21">C. ghiesbreghtiana</emphasis>
|
||
</taxonomicName>
|
||
in a given study area of lowland rainforest in
|
||
<collectingCountry id="82FC7647FF80FFC63FEFFCED2427B9EC" box="[656,790,872,898]" name="Costa Rica" pageId="20" pageNumber="21">Costa Rica</collectingCountry>
|
||
, and their visiting behaviors differ greatly. While the glossophagine nectar-specialists visit inflorescences in extremely short hovering flights of less than one second, the usually frugivorous stenodermine bats, including two species of
|
||
<taxonomicName id="3DEB4D54FF80FFC63E96FC3D237DB9BF" box="[1001,1100,952,977]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
|
||
<emphasis id="C89FEAC5FF80FFC63E96FC3D237DB9BF" box="[1001,1100,952,977]" italics="true" pageId="20" pageNumber="21">Artibeus</emphasis>
|
||
</taxonomicName>
|
||
, land and perch on the inflorescences for minutes, occasionally up to 17 minutes. Although the glossophagine bats are much more frequent visitors than
|
||
<taxonomicName id="3DEB4D54FF80FFC63CD6FB8D253DBE4F" box="[425,524,1032,1057]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
|
||
<emphasis id="C89FEAC5FF80FFC63CD6FB8D253DBE4F" box="[425,524,1032,1057]" italics="true" pageId="20" pageNumber="21">Artibeus</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="3DEB4D54FF80FFC63F42FB8D2412BE4F" box="[573,803,1032,1057]" class="Liliopsida" family="Arecaceae" genus="Calyptrogyne" kingdom="Plantae" order="Arecales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="ghiesbreghtiana">
|
||
<emphasis id="C89FEAC5FF80FFC63F42FB8D2412BE4F" box="[573,803,1032,1057]" italics="true" pageId="20" pageNumber="21">C. ghiesbreghtiana</emphasis>
|
||
</taxonomicName>
|
||
, their short hovering flights and lack of a smooth extended walkway like that provided by hummingbird bills evidently do not afford the mites sufficient time and means to mount or dismount (
|
||
<bibRefCitation id="9E7A4B26FF80FFC63F59FBDD24ABBE1C" author="Tschapka" box="[550,922,1112,1138]" pageId="20" pageNumber="37" refString="Tschapka, M. & Cunningham, S. A. (2004) Flower mites of Calyptrogyne ghiesbreghtiana (Arecaceae): evidence for dispersal using pollinating bats. Biotropica, 36, 377 - 381." type="journal article" year="2004">Tschapka & Cunningham 2004</bibRefCitation>
|
||
). In contrast, perching by
|
||
<taxonomicName id="3DEB4D54FF80FFC639B5FBDD221CBE1F" box="[1226,1325,1112,1137]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
|
||
<emphasis id="C89FEAC5FF80FFC639B5FBDD221CBE1F" box="[1226,1325,1112,1137]" italics="true" pageId="20" pageNumber="21">Artibeus</emphasis>
|
||
</taxonomicName>
|
||
bats provides close body contact between them and inflorescences along with longer visit durations. Mites were nearly always found on the surfaces of these bats’ wings, in the same places that most of the
|
||
<taxonomicName id="3DEB4D54FF80FFC639A1FB2D27F2BE87" class="Liliopsida" family="Arecaceae" genus="Calyptrogyne" kingdom="Plantae" order="Arecales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="ghiesbreghtiana">
|
||
<emphasis id="C89FEAC5FF80FFC639A1FB2D27F2BE87" italics="true" pageId="20" pageNumber="21">C. ghiesbreghtiana</emphasis>
|
||
</taxonomicName>
|
||
pollen grains were found.
|
||
</paragraph>
|
||
<paragraph id="FA5436D7FF80FFC63DB9FB7D2614BFB4" blockId="20.[151,1437,152,2018]" pageId="20" pageNumber="21">
|
||
Apart from the diversity of mites that are parasites of bats, few other mites have been noted to be phoretic on bats. Adult females of a tarsonemid,
|
||
<taxonomicName id="3DEB4D54FF80FFC63FFFFAA5233FBF54" authority="Ochoa" authorityName="Ochoa" box="[640,1038,1312,1338]" class="Arachnida" family="Tarsonemidae" genus="Alkithoenemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="cynopterus">
|
||
<emphasis id="C89FEAC5FF80FFC63FFFFAA52489BF57" box="[640,952,1312,1337]" italics="true" pageId="20" pageNumber="21">Alkithoenemus cynopterus</emphasis>
|
||
Ochoa
|
||
</taxonomicName>
|
||
& OConnor, thought to undergo development within the fleshy syconial receptacle of figs, are phoretic on the frugivorous bat,
|
||
<taxonomicName id="3DEB4D54FF80FFC6386AFACD2471BFE4" authority="Kitchener & Maharadatunkamsi. However" authorityName="Kitchener & Maharadatunkamsi. However" class="Mammalia" family="Pteropodidae" genus="Cynopterus" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="nusatenggara">
|
||
<emphasis id="C89FEAC5FF80FFC6386AFACD2609BFE7" italics="true" pageId="20" pageNumber="21">Cynopterus nusatenggara</emphasis>
|
||
Kitchener & Maharadatunkamsi. However
|
||
</taxonomicName>
|
||
, this bat is not a pollinator of figs, and species of
|
||
<taxonomicName id="3DEB4D54FF80FFC63DE8FA1D262FBFDF" box="[151,286,1432,1457]" class="Mammalia" family="Pteropodidae" genus="Cynopterus" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
|
||
<emphasis id="C89FEAC5FF80FFC63DE8FA1D262FBFDF" box="[151,286,1432,1457]" italics="true" pageId="20" pageNumber="21">Cynopterus</emphasis>
|
||
</taxonomicName>
|
||
feed on a wide variety of fruits, even though figs may constitute large parts of their diets (
|
||
<bibRefCitation id="9E7A4B26FF80FFC63832FA1D2629BFB4" author="Ochoa" pageId="20" pageNumber="37" refString="Ochoa, R., OConnor, B. M. & Klompen, S. H. (1996) A new genus and species of Tarsonemidae (Acari: Heterostigmata) associated with a fruit bat, Cynopterus nusatenggara (Chiroptera: Pteropodidae) on Lombok Island, Indonesia. International Journal of Acarology, 22, 209 - 213." type="journal article" year="1996">
|
||
Ochoa
|
||
<emphasis id="C89FEAC5FF80FFC63DE8FA4527E0BFB7" box="[151,209,1472,1497]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
|
||
1996
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FA5436D7FF80FFC63DB9FA6D23EABC9C" blockId="20.[151,1437,152,2018]" pageId="20" pageNumber="21">
|
||
The above collection records support observations by
|
||
<bibRefCitation id="9E7A4B26FF80FFC63E09FA6D235DBC6C" author="Cunningham" box="[886,1132,1512,1538]" pageId="20" pageNumber="36" refString="Cunningham, S. A. (1995) Ecological constraints on fruit initiation by Calyptrogyne ghiesbreghtiana (Arecaceae): floral herbivory, pollen availability and visitation by pollinating bats. American Journal of Botany, 82, 1527 - 1536." type="journal article" year="1995">Cunningham (1995)</bibRefCitation>
|
||
that a scarabeid beetle,
|
||
<taxonomicName id="3DEB4D54FF80FFC63DE8F99526C5BC44" authority="Blanchard" authorityName="Blanchard" box="[151,500,1552,1578]" class="Insecta" family="Rutelidae" genus="Lagochile" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="collaris">
|
||
<emphasis id="C89FEAC5FF80FFC63DE8F995265EBC47" box="[151,367,1552,1577]" italics="true" pageId="20" pageNumber="21">Lagochile collaris</emphasis>
|
||
Blanchard
|
||
</taxonomicName>
|
||
, which feeds destructively on
|
||
<taxonomicName id="3DEB4D54FF80FFC63E20F9952371BC47" box="[863,1088,1552,1577]" class="Liliopsida" family="Arecaceae" genus="Calyptrogyne" kingdom="Plantae" order="Arecales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="ghiesbreghtiana">
|
||
<emphasis id="C89FEAC5FF80FFC63E20F9952371BC47" box="[863,1088,1552,1577]" italics="true" pageId="20" pageNumber="21">C. ghiesbreghtiana</emphasis>
|
||
</taxonomicName>
|
||
flowers, provides alternative transportation for this mite. About two-thirds of the beetles observed on inflorescences carried mites (
|
||
<bibRefCitation id="9E7A4B26FF80FFC63DE1F9E52528BC14" author="Tschapka" box="[158,537,1632,1658]" pageId="20" pageNumber="37" refString="Tschapka, M. & Cunningham, S. A. (2004) Flower mites of Calyptrogyne ghiesbreghtiana (Arecaceae): evidence for dispersal using pollinating bats. Biotropica, 36, 377 - 381." type="journal article" year="2004">Tschapka & Cunningham 2004</bibRefCitation>
|
||
), which locate on external surfaces, not underneath the elytra (Ellen Sperr, personal observation,
|
||
<date id="8E551017FF80FFC63CDBF90D2510BCCC" box="[420,545,1672,1698]" pageId="20" pageNumber="21" value="2007-07">July 2007</date>
|
||
). The single collection record of a few female and protonymphal mites phoretic on an unidentified curculionid beetle, without a more specific identification of the latter, needs confirmation as a phoretic association between these mites and a carrier of this family of beetles.
|
||
</paragraph>
|
||
<paragraph id="FA5436D7FF80FFC63DB9F885232BBD04" blockId="20.[151,1437,152,2018]" pageId="20" pageNumber="21">
|
||
A sample of two fully developed inflorescences of
|
||
<taxonomicName id="3DEB4D54FF80FFC63E51F88523A2BD77" box="[814,1171,1792,1817]" class="Liliopsida" family="Arecaceae" genus="Calyptrogyne" kingdom="Plantae" order="Arecales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="ghiesbreghtiana">
|
||
<emphasis id="C89FEAC5FF80FFC63E51F88523A2BD77" box="[814,1171,1792,1817]" italics="true" pageId="20" pageNumber="21">Calyptrogyne ghiesbreghtiana</emphasis>
|
||
</taxonomicName>
|
||
taken at the La Selva Biological Station,
|
||
<date id="8E551017FF80FFC63C05F8AD257BBD2C" box="[378,586,1832,1858]" pageId="20" pageNumber="21" value="1994-02-10">10 February 1994</date>
|
||
by one of us (EEL), contained between 400 and 500 mites, not accounting for eggs, including an abundance of all developmental instars and adults.
|
||
</paragraph>
|
||
<paragraph id="FA5436D7FF80FFC63DB9F8FD2327BD8C" blockId="20.[151,1437,152,2018]" pageId="20" pageNumber="21">
|
||
The greatly enlarged peritrematal gland pore in adult male
|
||
<taxonomicName id="3DEB4D54FF80FFC63EF1F8FD2364BDFF" box="[910,1109,1912,1937]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="calyptrogynae">
|
||
<emphasis id="C89FEAC5FF80FFC63EF1F8FD2364BDFF" box="[910,1109,1912,1937]" italics="true" pageId="20" pageNumber="21">S. calyptrogynae</emphasis>
|
||
</taxonomicName>
|
||
is a form of sexual dimorphism not observed before among mesostigmatic mites. Adult males available of three other congeneric species (
|
||
<taxonomicName id="3DEB4D54FF80FFC63DAAF84D26A0BD8F" box="[213,401,1992,2017]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="spathiphyllae">
|
||
<emphasis id="C89FEAC5FF80FFC63DAAF84D26A0BD8F" box="[213,401,1992,2017]" italics="true" pageId="20" pageNumber="21">S. spathiphyllae</emphasis>
|
||
</taxonomicName>
|
||
and two of the undescribed species) lack this attribute.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |