181 lines
19 KiB
XML
181 lines
19 KiB
XML
<document id="C24AE3DCE78457250483AA03C0D06339" ID-CLB-Dataset="26920" ID-DOI="10.11646/zootaxa.4599.1.1" ID-GBIF-Dataset="ea727a9b-324b-4c6b-a625-bc7cdd204473" ID-ISSN="1175-5326" ID-Zenodo-Dep="2668058" ID-ZooBank="7CDD24FF-5BA3-4D06-898D-A3E220515D4C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557127209543" checkinUser="plazi" docAuthor="Longino, John T." docDate="2019" docId="5B56E872FFB54A7EFF328870FDBCEAEC" docLanguage="en" docName="zootaxa.4599.1.1.pdf.imf" docOrigin="Zootaxa 4599 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pheidole plebecula Forel 1899" docType="treatment" docVersion="21" lastPageNumber="54" masterDocId="A76F900AFF804A4BFFA58B6FFFF4EF5B" masterDocTitle="Pheidole (Hymenoptera, Formicidae) of Middle American Wet Forest" masterLastPageNumber="126" masterPageNumber="1" pageNumber="54" updateTime="1698726215157" updateUser="ExternalLinkService">
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<mods:mods id="4F2D00C157664816301F659B7483517A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1442048DFE5FAAADA74BB8F38628D9DF">
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<mods:title id="8379E1BF7BD48E17C04D6B20C01F1B67">Pheidole (Hymenoptera, Formicidae) of Middle American Wet Forest</mods:title>
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<mods:name id="FDC8EEEE676BCFDB4B78ECB9BD4BCB8F" type="personal">
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<mods:role id="034AFC930D7CC34E96948ABC8CF21FE2">
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<mods:roleTerm id="CB7E489871F1187FE1289EEBEE138D3E">Author</mods:roleTerm>
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<mods:namePart id="14DE40A535D5A32899559AA907F0169B">Longino, John T.</mods:namePart>
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<mods:typeOfResource id="F6317E3F891046B42864A79BA7008D76">text</mods:typeOfResource>
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<mods:titleInfo id="41DA7E8EB9F5FDCCC2780B3960D77AA9">
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<mods:title id="CF5BB20A46B406FBD474CE7377DBD6D5">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="D2092F6D6E9F8E449C10251CCC17F0D3">
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<mods:date id="6D0820D72C1C6BCB08C5DC43AD652162">2019</mods:date>
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<mods:detail id="78C7FEAFAA3CE5BE44FBBFA9A5430757" type="pubDate">
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<mods:number id="C5068566BDED1F1986464ADC52256240">2019-05-06</mods:number>
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<mods:detail id="C4F22F185418ABFC0AA6CB226CFC5824" type="volume">
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<mods:number id="4CEF50803D7BFEDA7AFA65B5F3389ACF">4599</mods:number>
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<mods:detail id="9505C2D15E24D5DBEA52A417A0F75920" type="issue">
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<mods:start id="E47BB722B653C463067CBAD787682049">1</mods:start>
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<mods:end id="40D7EE3C1FDADDB0D6CEAA723940EF4D">126</mods:end>
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<mods:classification id="3E666EFC165C3E214D4D845F745A278F">journal article</mods:classification>
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<mods:identifier id="F31E74AE114B3A3300BD97809A591478" type="CLB-Dataset">26920</mods:identifier>
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<mods:identifier id="8A59CF82735AB495DF338BA14B639DE4" type="DOI">10.11646/zootaxa.4599.1.1</mods:identifier>
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<mods:identifier id="6B23EE8F748D678A1936DBBCA8F40C6F" type="GBIF-Dataset">ea727a9b-324b-4c6b-a625-bc7cdd204473</mods:identifier>
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<mods:identifier id="AFAE62AC51F868C4C3B74F59A18AF640" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="293D624F45B411B679001B5F3531EB4E" type="Zenodo-Dep">2668058</mods:identifier>
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<mods:identifier id="6F46F8CFDEBB028877DF58C9CC4E887F" type="ZooBank">7CDD24FF-5BA3-4D06-898D-A3E220515D4C</mods:identifier>
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</mods:mods>
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<treatment id="5B56E872FFB54A7EFF328870FDBCEAEC" ID-DOI="http://doi.org/10.5281/zenodo.3513587" ID-GBIF-Taxon="157062582" ID-Zenodo-Dep="3513587" LSID="urn:lsid:plazi:treatment:5B56E872FFB54A7EFF328870FDBCEAEC" httpUri="http://treatment.plazi.org/id/5B56E872FFB54A7EFF328870FDBCEAEC" lastPageNumber="54" pageId="53" pageNumber="54">
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<subSubSection id="9BE50AEFFFB54A7EFF328870FE88EC62" box="[151,380,799,825]" pageId="53" pageNumber="54" type="nomenclature">
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<paragraph id="D3405964FFB54A7EFF328870FE88EC62" blockId="53.[151,380,799,825]" box="[151,380,799,825]" pageId="53" pageNumber="54">
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<heading id="8808EE08FFB54A7EFF328870FE88EC62" bold="true" box="[151,380,799,825]" fontSize="11" level="1" pageId="53" pageNumber="54" reason="1">
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<taxonomicName id="14FF22E7FFB54A7EFF328870FE88EC62" ID-CoL="4FYJ7" authorityName="Forel" authorityYear="1899" box="[151,380,799,825]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="plebecula">
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<emphasis id="E18B8576FFB54A7EFF328870FE88EC62" bold="true" box="[151,380,799,825]" italics="true" pageId="53" pageNumber="54">Pheidole plebecula</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFB54A7EFF328809FE7DEB08" pageId="53" pageNumber="54" type="reference_group">
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<paragraph id="D3405964FFB54A7EFF328809FD5DECC0" blockId="53.[151,1436,870,1107]" pageId="53" pageNumber="54">
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<treatmentCitationGroup id="F3EF7E4AFFB54A7EFF328809FD5DECC0" pageId="53" pageNumber="54">
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<taxonomicName id="14FF22E7FFB54A7EFF328809FE0AEC27" ID-CoL="4FYJ7" authority="Forel, 1899: 68" authorityName="Forel" authorityPageNumber="68" authorityYear="1899" box="[151,510,870,892]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="plebecula">
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<emphasis id="E18B8576FFB54A7EFF328809FEAEEC27" box="[151,346,870,892]" italics="true" pageId="53" pageNumber="54">Pheidole plebecula</emphasis>
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<treatmentCitation id="525E7F75FFB54A7EFEC58809FE0AEC27" author="Forel, A." box="[352,510,870,892]" page="68" pageId="53" pageNumber="54" year="1899">
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<bibRefCitation id="B76E2495FFB54A7EFEC58809FE2CEC27" author="Forel, A." box="[352,472,870,892]" pageId="53" pageNumber="54" pagination="57 - 80" refId="ref51457" refString="Forel, A. (1899) Formicidae. [part c]. In: Insecta. Hymenoptera. Formicidae. R. H. Porter, Dulau & Co., London, pp. 57 - 80." type="book chapter" year="1899">Forel, 1899</bibRefCitation>
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: 68
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</treatmentCitation>
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</taxonomicName>
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.
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<typeStatus id="0C44E7C6FFB54A7EFDAD8809FD92EC27" box="[520,614,870,892]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
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major worker:
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<collectingCountry id="ABE819F4FFB54A7EFCA78809FC87EC27" box="[770,883,870,892]" name="Costa Rica" pageId="53" pageNumber="54">Costa Rica</collectingCountry>
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,
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<collectingRegion id="113B9786FFB54A7EFCD88809FC27EC27" box="[893,979,870,892]" country="Costa Rica" name="Alajuela" pageId="53" pageNumber="54">Alajuela</collectingRegion>
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(Alfaro) [MSNG, unique specimen identifier CASENT0904359] (AntWeb image examined).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="D3405964FFB54A7EFF3288CBFBE7EC83" blockId="53.[151,1436,870,1107]" pageId="53" pageNumber="54">
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<treatmentCitationGroup id="F3EF7E4AFFB54A7EFF3288CBFBE7EC83" pageId="53" pageNumber="54">
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<taxonomicName id="14FF22E7FFB54A7EFF3288CBFDE6ECE1" authority="Wilson, 2003: 240" authorityName="Wilson" authorityPageNumber="240" authorityYear="2003" box="[151,530,932,954]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="texticeps">
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<emphasis id="E18B8576FFB54A7EFF3288CBFEA4ECE1" box="[151,336,932,954]" italics="true" pageId="53" pageNumber="54">Pheidole texticeps</emphasis>
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<treatmentCitation id="525E7F75FFB54A7EFEF088CBFDE6ECE1" ID-DOI="http://doi.org/10.5281/zenodo.3509040" ID-Zenodo-Dep="3509040" author="Wilson, E. O." box="[341,530,932,954]" httpUri="http://treatment.plazi.org/id/A0D5030A63B5AD775B74B2B5E5530E07" page="240" pageId="53" pageNumber="54" year="2003">
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<bibRefCitation id="B76E2495FFB54A7EFEF088CBFE2BECE1" author="Wilson, E. O." box="[341,479,932,954]" pageId="53" pageNumber="54" refId="ref52858" refString="Wilson, E. O. (2003) Pheidole in the New World: A Dominant, Hyperdiverse Ant Genus. Cambridge, Mass, Harvard University Press, 794 pp." type="book" year="2003">Wilson, 2003</bibRefCitation>
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: 240
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</treatmentCitation>
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</taxonomicName>
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, figs.
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<typeStatus id="0C44E7C6FFB54A7EFDE888CBFD5FECE1" box="[589,683,932,954]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
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major worker and associated
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<typeStatus id="0C44E7C6FFB54A7EFC7D88CAFBDBECE1" box="[984,1071,933,954]" pageId="53" pageNumber="54" type="paratype">paratype</typeStatus>
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minor worker:
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<collectingCountry id="ABE819F4FFB54A7EFB6E88CBFAC8ECE1" box="[1227,1340,932,954]" name="Costa Rica" pageId="53" pageNumber="54">Costa Rica</collectingCountry>
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,
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<collectingRegion id="113B9786FFB54A7EFAE088CBFA6CECE1" box="[1349,1432,932,954]" country="Costa Rica" name="Heredia" pageId="53" pageNumber="54">Heredia</collectingRegion>
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,
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<location id="D6200FBFFFB54A7EFF6288ACFE14EC83" LSID="urn:lsid:plazi:treatment:5B56E872FFB54A7EFF328870FDBCEAEC:D6200FBFFFB54A7EFF6288ACFE14EC83" box="[199,480,962,984]" country="Costa Rica" name="La Selva Biological Station" pageId="53" pageNumber="54" stateProvince="Heredia">La Selva Biological Station</location>
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, 1988 (
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<collectorName id="7E0A3CB2FFB54A7EFD8988ADFD72EC83" box="[556,646,962,984]" pageId="53" pageNumber="54">S. Cover</collectorName>
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) [MCZC] (examined).
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<emphasis id="E18B8576FFB54A7EFCD788ADFBF9EC83" bold="true" box="[882,1037,962,984]" pageId="53" pageNumber="54">New Synonym</emphasis>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="D3405964FFB54A7EFF32888EFE7DEB08" blockId="53.[151,1436,870,1107]" pageId="53" pageNumber="54">
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<treatmentCitationGroup id="F3EF7E4AFFB54A7EFF32888EFE7DEB08" infered="true" pageId="53" pageNumber="54">
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<taxonomicName id="14FF22E7FFB54A7EFF32888EFDCFECAC" authority="Wilson, 2003: 213" authorityName="Wilson" authorityPageNumber="213" authorityYear="2003" box="[151,571,993,1015]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="perdiligens">
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<emphasis id="E18B8576FFB54A7EFF32888EFE98ECAC" box="[151,364,993,1015]" italics="true" pageId="53" pageNumber="54">Pheidole perdiligens</emphasis>
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<treatmentCitation id="525E7F75FFB54A7EFED3888EFDCFECAC" ID-DOI="http://doi.org/10.5281/zenodo.3499964" ID-Zenodo-Dep="3499964" author="Wilson, E. O." box="[374,571,993,1015]" httpUri="http://treatment.plazi.org/id/5D6A26333A00928E3A0774F89B2CAC24" page="213" pageId="53" pageNumber="54" year="2003">
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<bibRefCitation id="B76E2495FFB54A7EFED3888EFDF1ECAC" author="Wilson, E. O." box="[374,517,993,1015]" pageId="53" pageNumber="54" refId="ref52858" refString="Wilson, E. O. (2003) Pheidole in the New World: A Dominant, Hyperdiverse Ant Genus. Cambridge, Mass, Harvard University Press, 794 pp." type="book" year="2003">Wilson, 2003</bibRefCitation>
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: 213
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</treatmentCitation>
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</taxonomicName>
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, figs.
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<typeStatus id="0C44E7C6FFB54A7EFD24888EFD2BECAC" box="[641,735,993,1015]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
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major worker and associated
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<typeStatus id="0C44E7C6FFB54A7EFB87888DFB8DECAC" box="[1058,1145,994,1015]" pageId="53" pageNumber="54" type="paratype">paratype</typeStatus>
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minor worker:
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<collectingCountry id="ABE819F4FFB54A7EFA86888EFA6CECAC" box="[1315,1432,993,1015]" name="Costa Rica" pageId="53" pageNumber="54">Costa Rica</collectingCountry>
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,
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<collectingRegion id="113B9786FFB54A7EFF628F6EFECDEB4D" box="[199,313,1025,1046]" country="Costa Rica" name="Puntarenas" pageId="53" pageNumber="54">Puntarenas</collectingRegion>
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, Corcovado National Park, Osa Peninsula, Sirena,
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<date id="A7417FA4FFB54A7EFC928F6FFC43EB4D" box="[823,951,1024,1046]" pageId="53" pageNumber="54" value="1979-02-28">28 Feb 1979</date>
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, secondary rainforest, <1/
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<quantity id="1407F481FFB54A7EFB1F8F6FFB17EB4D" box="[1210,1251,1024,1046]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="53" pageNumber="54" unit="mi" value="2.0">2mi</quantity>
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from coast, rotten branch on forest floor (S. P. Cover CR312) [MCZC] (examined).
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<emphasis id="E18B8576FFB54A7EFC248F71FBEFEB6F" bold="true" box="[897,1051,1054,1076]" pageId="53" pageNumber="54">New synonym</emphasis>
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. Synonymized under
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<taxonomicName id="14FF22E7FFB54A7EFAA18F4FFA8DEB6E" authorityName="Wilson" authorityYear="2003" box="[1284,1401,1056,1077]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="texticeps">
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<emphasis id="E18B8576FFB54A7EFAA18F4FFA8DEB6E" box="[1284,1401,1056,1077]" italics="true" pageId="53" pageNumber="54">P. texticeps</emphasis>
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</taxonomicName>
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by
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<treatmentCitation id="525E7F75FFB54A7EFF628F52FE71EB08" author="Longino, J. T." box="[199,389,1085,1107]" page="85" pageId="53" pageNumber="54" year="2009">
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<bibRefCitation id="B76E2495FFB54A7EFF628F52FEABEB08" author="Longino, J. T." box="[199,351,1085,1107]" pageId="53" pageNumber="54" pagination="1 - 90" refId="ref51720" refString="Longino, J. T. (2009) Additions to the taxonomy of New World Pheidole (Hymenoptera: Formicidae). Zootaxa, 2181, 1 - 90." type="journal article" year="2009">Longino, 2009</bibRefCitation>
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: 85
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFB54A7EFF328F10FDBCEAEC" pageId="53" pageNumber="54" type="discussion">
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<paragraph id="D3405964FFB54A7EFF328F10FAA4EBE7" blockId="53.[151,1436,1151,1464]" pageId="53" pageNumber="54">
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<emphasis id="E18B8576FFB54A7EFF328F10FEECEBC3" bold="true" box="[151,280,1151,1176]" pageId="53" pageNumber="54">Comments</emphasis>
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.
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<bibRefCitation id="B76E2495FFB54A7EFE838F10FE3CEBC3" author="Wilson, E. O." box="[294,456,1151,1176]" pageId="53" pageNumber="54" refId="ref52858" refString="Wilson, E. O. (2003) Pheidole in the New World: A Dominant, Hyperdiverse Ant Genus. Cambridge, Mass, Harvard University Press, 794 pp." type="book" year="2003">Wilson (2003)</bibRefCitation>
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was unable to locate the
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<typeStatus id="0C44E7C6FFB54A7EFD4D8FEFFCECEBC3" box="[744,792,1152,1176]" pageId="53" pageNumber="54">type</typeStatus>
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of
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<taxonomicName id="14FF22E7FFB54A7EFC9A8FEEFC3DEBC3" authorityName="Forel" authorityYear="1899" box="[831,969,1151,1176]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="plebecula">
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<emphasis id="E18B8576FFB54A7EFC9A8FEEFC3DEBC3" box="[831,969,1151,1176]" italics="true" pageId="53" pageNumber="54">P. plebecula</emphasis>
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</taxonomicName>
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and left it
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<emphasis id="E18B8576FFB54A7EFBE08FEEFB15EBC3" box="[1093,1249,1151,1176]" italics="true" pageId="53" pageNumber="54">incertae sedis</emphasis>
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. The
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<typeStatus id="0C44E7C6FFB54A7EFA878FEFFAA9EBC3" box="[1314,1373,1152,1176]" pageId="53" pageNumber="54">types</typeStatus>
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were located and imaged for AntWeb by the Fisher lab. The images easily match Wilson's concept of
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<taxonomicName id="14FF22E7FFB54A7EFB698FCAFABDEBE7" authorityName="Wilson" authorityYear="2003" box="[1228,1353,1189,1212]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="texticeps">
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<emphasis id="E18B8576FFB54A7EFB698FCAFABDEBE7" box="[1228,1353,1189,1212]" italics="true" pageId="53" pageNumber="54">P. texticeps</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="D3405964FFB54A7EFF628FA8FDBCEAEC" blockId="53.[151,1436,1151,1464]" pageId="53" pageNumber="54">
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Local populations of
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<taxonomicName id="14FF22E7FFB54A7EFE138FA6FDCBEBBB" authorityName="Forel" authorityYear="1899" box="[438,575,1223,1248]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="plebecula">
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<emphasis id="E18B8576FFB54A7EFE138FA6FDCBEBBB" box="[438,575,1223,1248]" italics="true" pageId="53" pageNumber="54">P. plebecula</emphasis>
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</taxonomicName>
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vary greatly in color and sculpture. Color varies from yellow orange to dark brown, and to varying extent bicolored, with lighter head and mesosoma and darker gaster. The face of the major varies from being entirely foveolate/rugulose to completely smooth and shining. Pilosity can vary as well, with most populations having multiple pairs of stiff setae on the mesosomal dorsum, but some with no dorsal setae. One of the variants, from Volcán Atitlán,
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<collectingCountry id="ABE819F4FFB54A7EFD948E38FD59EA2B" box="[561,685,1367,1392]" name="Guatemala" pageId="53" pageNumber="54">Guatemala</collectingCountry>
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, was initially separated as the morphospecies JTL199. In
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<bibRefCitation id="B76E2495FFB54A7EFA948E37FED7EACF" author="Economo, E. P. & Huang, J. - P. & Fischer, G. & Sarnat, E. M. & Narula, N. & Janda, M. & Guenard, B. & Longino, J. T. & Knowles, L. L." pageId="53" pageNumber="54" pagination="456 - 470" refId="ref51110" refString="Economo, E. P., Huang, J. - P., Fischer, G., Sarnat, E. M., Narula, N., Janda, M., Guenard, B., Longino, J. T. & Knowles, L. L. (2019) Evolution of the latitudinal diversity gradient in the hyperdiverse ant genus Pheidole. Global Ecology and Biogeography, 28, 456 - 470. https: // doi. org / 10.1111 / geb. 12867" type="journal article" year="2019">
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Economo
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<emphasis id="E18B8576FFB54A7EFF328E12FF24EACE" box="[151,208,1404,1429]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
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(2019)
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</bibRefCitation>
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the two species
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<taxonomicName id="14FF22E7FFB54A7EFE448E12FD94EACF" authorityName="Wilson" authorityYear="2003" box="[481,608,1405,1428]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="texticeps">
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<emphasis id="E18B8576FFB54A7EFE448E12FD94EACF" box="[481,608,1405,1428]" italics="true" pageId="53" pageNumber="54">P. texticeps</emphasis>
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</taxonomicName>
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and
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<emphasis id="E18B8576FFB54A7EFD3D8E12FD52EACF" box="[664,678,1405,1428]" italics="true" pageId="53" pageNumber="54">P</emphasis>
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. JTL199 appear together, with little genetic divergence. These can now both be identified as
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<taxonomicName id="14FF22E7FFB54A7EFE1C8ECEFDB5EAE3" authorityName="Forel" authorityYear="1899" box="[441,577,1439,1464]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="plebecula">
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<emphasis id="E18B8576FFB54A7EFE1C8ECEFDB5EAE3" box="[441,577,1439,1464]" italics="true" pageId="53" pageNumber="54">P. plebecula</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |