214 lines
17 KiB
XML
214 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.326.5344" ID-GBIF-Dataset="e18bae20-fa52-4e93-8738-a1d1a5ee9137" ID-PMC="PMC3764535" ID-Pensoft-Pub="1313-2970-326-27" ID-PubMed="24039534" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-326-27" ModsDocOrigin="ZooKeys 326" ModsDocTitle="On Levymanus, a remarkable new spider genus from Israel, with notes on the Chediminae (Araneae, Palpimanidae)" checkinTime="1451247006434" checkinUser="pensoft" docAuthor="Zonstein, Sergei & Marusik, Yuri M." docDate="2013" docId="449E83C2B1F153C7EBB8D660B72360EF" docLanguage="en" docName="ZooKeys 326: 27-45" docOrigin="ZooKeys 326" docSource="http://dx.doi.org/10.3897/zookeys.326.5344" docTitle="Levymanus gershomi Zonstein & Marusik, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="33" masterDocId="FE4132753556C47C7843346BFFA7884C" masterDocTitle="On Levymanus, a remarkable new spider genus from Israel, with notes on the Chediminae (Araneae, Palpimanidae)" masterLastPageNumber="45" masterPageNumber="27" pageNumber="30" updateTime="1668156356640" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>On Levymanus, a remarkable new spider genus from Israel, with notes on the Chediminae (Araneae, Palpimanidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zonstein, Sergei</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Marusik, Yuri M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>326</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>27</mods:start>
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<mods:end>45</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.326.5344</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.326.5344</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-326-27</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152047327" LSID="urn:lsid:zoobank.org:act:BBABE77C-15AE-494D-8A61-6309D6541855" httpUri="http://treatment.plazi.org/id/449E83C2B1F153C7EBB8D660B72360EF" lastPageId="6" lastPageNumber="33" pageId="3" pageNumber="30">
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<subSubSection pageId="3" pageNumber="30" type="nomenclature">
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<paragraph pageId="3" pageNumber="30">
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<taxonomicName LSID="http://zoobank.org/BBABE77C-15AE-494D-8A61-6309D6541855" class="Arachnida" family="Palpimanidae" genus="Levymanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Levymanus gershomi" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="gershomi">
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<pageBreakToken pageId="3" pageNumber="30" start="start">Levymanus</pageBreakToken>
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gershomi
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</taxonomicName>
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<taxonomicNameLabel pageId="3" pageNumber="30">sp. n.</taxonomicNameLabel>
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Figs 1-2, 10-50
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="30" type="types">
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<paragraph pageId="3" pageNumber="30">Types.</paragraph>
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<paragraph pageId="3" pageNumber="30">
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Male holotype, 3 ♂ and 2 ♀ paratypes from the vicinity of Qetura (Ktura), Arava Valley, Israel (
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="29.966667">29°58'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="35.05">35°03'E</geoCoordinate>
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), 8 May 2003, coll. E. Topel, deposited in TAU (holotype and most paratypes) and ZMMU (few paratypes).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="30" type="diagnosis">
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<paragraph pageId="3" pageNumber="30">Diagnosis.</paragraph>
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<paragraph pageId="3" pageNumber="30">As for the genus.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="32" pageId="3" pageNumber="30" type="description">
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<paragraph pageId="3" pageNumber="30">Description.</paragraph>
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<paragraph pageId="3" pageNumber="30">
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Male (holotype): Total length 2.55; carapace, sternum and labium intensive carmine-red; chelicerae, palps (entirely), coxa and femur I light reddish orange, other segments of leg I and entire legs
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<normalizedToken originalValue="II–IV">II-IV</normalizedToken>
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pale yellowish red; abdomen milk-white with intensive reddish orange dorsal scutum. Carapace (Fig. 10): 1.10 long, 0.76 wide. Diameters of AME, ALE, PLE, PME: 0.10, 0.02, 0.02, 0.02. Interdistances:
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<normalizedToken originalValue="AME–AME">AME-AME</normalizedToken>
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0.08,
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<normalizedToken originalValue="ALE–AME">ALE-AME</normalizedToken>
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0.05,
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<normalizedToken originalValue="ALE–PLE">ALE-PLE</normalizedToken>
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<0.01,
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<normalizedToken originalValue="PLE–PME">PLE-PME</normalizedToken>
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0.11,
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<normalizedToken originalValue="PME–PME">PME-PME</normalizedToken>
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0.14. Chelicerae as shown in Figs 17-18. Sternum (Fig. 11) 0.85 long, 0.65 wide; labium 0.21 long, 0.18 wide at base. Measurements of palp and leg segments as shown in Table 1. Scopulae and tarsal claws as shown in Figs 23, 26-31 and 22, 24, 25, 36, 37 respectively. Scarcely distributed scopular hairs approximately as long as metatarsus and tarsus width (Figs 26-31) At least metatarsus III with comb of setae (Fig. 35). Tarsi
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<normalizedToken originalValue="I–IV">I-IV</normalizedToken>
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with claw tufts, better developed on tarsi
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<normalizedToken originalValue="I–II">I-II</normalizedToken>
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(Figs 22, 24). Tarsal claws
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<normalizedToken originalValue="I–II">I-II</normalizedToken>
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with few teeth (Figs 22, 25),
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<normalizedToken originalValue="III–IV">III-IV</normalizedToken>
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with more and longer teeth (Figs 36, 37). Spinnerets, pedicel tube, and ventral parts of abdominal scutum as shown in Figs 32-34.
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</paragraph>
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<caption pageId="3" pageNumber="30">
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<paragraph pageId="3" pageNumber="30">
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Figures 1-9. Spiders belonging to the subfamily
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<taxonomicName lsidName="" pageId="3" pageNumber="30" rank="subFamily" subFamily="Chediminae">Chediminae</taxonomicName>
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; habitus in lateral (1, 2, 6), dorsal (3-5, 7, 9) and ventral (8) view. 1, 2
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<taxonomicName class="Arachnida" family="Palpimanidae" genus="Levymanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Levymanus gershomi" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="gershomi">Levymanus gershomi</taxonomicName>
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sp. n., holotype male and paratype female, respectively 3
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<taxonomicName class="Arachnida" family="Palpimanidae" genus="Boagrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Boagrius" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">Boagrius</taxonomicName>
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sp. aff. incisus, female 4
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<taxonomicName class="Arachnida" family="Palpimanidae" genus="Scelidocteus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scelidocteus" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">Scelidocteus</taxonomicName>
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sp. aff. vuattouxi, male 5, 6
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<taxonomicName class="Arachnida" family="Palpimanidae" genus="Sarascelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarascelis chaperi" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="chaperi">Sarascelis chaperi</taxonomicName>
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, female and male, respectively 7
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<taxonomicName class="Arachnida" family="Palpimanidae" genus="Scelidomachus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scelidomachus socotranus" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="socotranus">Scelidomachus socotranus</taxonomicName>
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, conspecific male 8, 9
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<taxonomicName class="Arachnida" family="Palpimanidae" genus="Steriphopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Steriphopus macleayi" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="macleayi">Steriphopus macleayi</taxonomicName>
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, holotype male (scale bar = 1 mm).
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</paragraph>
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</caption>
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<caption pageId="3" pageNumber="30">
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<paragraph pageId="3" pageNumber="30">
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Figures 10-13.
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<taxonomicName class="Arachnida" family="Palpimanidae" genus="Levymanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Levymanus gershomi" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="gershomi">Levymanus gershomi</taxonomicName>
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sp. n., male (10, 11) and female paratypes (12, 13). 10, 12 Carapace, dorsal view 11, 13 Sternum, labium, maxillae and chelicerae; ventral view (scale bar = 0.2 mm).
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</paragraph>
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</caption>
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<caption pageId="3" pageNumber="30">
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<paragraph pageId="3" pageNumber="30">
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Figures 14-16.
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<taxonomicName class="Arachnida" family="Palpimanidae" genus="Levymanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Levymanus gershomi" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="gershomi">Levymanus gershomi</taxonomicName>
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sp. n., male (14) and female paratype (15, 16). 14, 15 Habitus, lateral view 16 Carapace and chelicerae, frontal view (scale bar = 0.5 mm for 14, 15; 0.2 mm for 16).
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</paragraph>
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</caption>
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<caption pageId="3" pageNumber="30">
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<paragraph pageId="3" pageNumber="30">
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Figures 17-19.
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<taxonomicName class="Arachnida" family="Palpimanidae" genus="Levymanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Levymanus gershomi" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="gershomi">Levymanus gershomi</taxonomicName>
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sp. n., male (17, 18) and female paratype (18). 17, 18 right chelicera, frontal (17) and prolateral (18) view 19 right palp, femur to tarsus, retrolateral view. (scale bar = 0.05 mm for 17, 18; 0.1 mm for 19).
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</paragraph>
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</caption>
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<caption pageId="3" pageNumber="30">
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<paragraph pageId="3" pageNumber="30">
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Figures 20-31.
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<taxonomicName class="Arachnida" family="Palpimanidae" genus="Levymanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Levymanus gershomi" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="gershomi">Levymanus gershomi</taxonomicName>
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sp. n., paratype male, leg I, prolateral view. 20 femur 21 patella 22 tarsus, showing claws with teeth 23 tibia, showing spatulate hair 24 tarsus, showing claws and claw
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<normalizedToken originalValue="‘pillow’">'pillow'</normalizedToken>
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25 tarsus, showing claws and
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<normalizedToken originalValue="‘pillow’">'pillow'</normalizedToken>
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26 tarsus, showing scopula 27 tibia, showing scopula 28 whole leg, showing scopula and enlarged patella 29, 30 metatarsus I, showing scopula 31 tibia to tarsus, showing scopula (scale bar = 0.1 mm for 20-23, 25-27, 30; 0.05 mm for 24; 0.2 mm for 29, 31).
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</paragraph>
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</caption>
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<caption pageId="3" pageNumber="30">
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<paragraph pageId="3" pageNumber="30">
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Figures 32-37.
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<taxonomicName class="Arachnida" family="Palpimanidae" genus="Levymanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Levymanus gershomi" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="gershomi">Levymanus gershomi</taxonomicName>
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sp. n., paratype male. 32 spinnerets, ventral view 33 epigastral scutum, ventral view 34 abdomen, ventral view 35 distal part of metatarsus III, showing metatarsal comb, ventral view 36 distal part of tarsus III, showing onichum and claws, ventral view 37 same, prolateral view (scale bar = 0.01 mm for 32, 35-37; 0.1 mm for 33, 34).
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</paragraph>
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</caption>
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<paragraph lastPageId="4" lastPageNumber="31" pageId="3" pageNumber="30">
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Palp (Figs 38-48): femur short and swollen, 2 times longer than wide, subequal in length to tibia and slightly shorter than cymbium. Patella globular. Tibia without apophyses, strongly swollen, 1.5 times longer than wide, 1.3 times wider than femur. Cymbium narrow, shorter than bulb, without outgrowths, with two clusters of hairs on prolateral side (Figs 39, 46). Bulb as wide (in widest part) as long (not counting embolic division and tegular process), tegulum with strong and long retrolateral tegular process (
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<pageBreakToken pageId="4" pageNumber="31" start="start">Tp</pageBreakToken>
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); base of process with conical outgrowth (Co) and deep furrow (Tf); embolus fused with other sclerites of the embolic division (Ed). Embolic division, attached to tegulum by a flexible membrane, and bears embolus (Em) with retrolateral outgrowth (Eo), and lamella (La) located between embolus and cymbium.
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</paragraph>
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<caption pageId="4" pageNumber="31">
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<paragraph pageId="4" pageNumber="31">
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Figures 38-46.
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<taxonomicName class="Arachnida" family="Palpimanidae" genus="Levymanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Levymanus gershomi" order="Araneae" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="gershomi">Levymanus gershomi</taxonomicName>
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sp. n., palp of paratype male 38 palp, ventral view 39 whole palp, prolateral view 40 same, apical view 41 terminal part of bulbus, ventral view 42, 43 same, prolateral-ventral view 44 same, ventral-retrolateral view 45 palp, retrolateral view 46 same, apical-retrolateral view. 39, 44-46 palp with embolic division sunken into tegulum (scale bar = 0.1 mm for 38, 39; 0.05 mm for 40-46).
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</paragraph>
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</caption>
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<caption pageId="4" pageNumber="31">
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<paragraph pageId="4" pageNumber="31">
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Figures 47-50.
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<taxonomicName class="Arachnida" family="Palpimanidae" genus="Levymanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Levymanus gershomi" order="Araneae" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="gershomi">Levymanus gershomi</taxonomicName>
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sp. n., male (47, 48) and female paratypes (49, 50). 47, 48 palp, ventral view 49, 50 epigyne; ventral view (scale bar = 0.2 mm).
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</paragraph>
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</caption>
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<paragraph lastPageId="5" lastPageNumber="32" pageId="4" pageNumber="31">
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Female (paratype): coloration as in male. Total length 2.93. Carapace (Fig. 12): 1.34 long, 0.92 wide. Diameters of AME, ALE, PLE, PME: 0.10, 0.03, 0.02, 0.02. Interdistances:
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<normalizedToken originalValue="AME–AME">AME-AME</normalizedToken>
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0.08,
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<normalizedToken originalValue="ALE–AME">ALE-AME</normalizedToken>
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0.05,
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<normalizedToken originalValue="ALE–PLE">ALE-PLE</normalizedToken>
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<0.01,
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<normalizedToken originalValue="PLE–PME">PLE-PME</normalizedToken>
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0.10,
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<normalizedToken originalValue="PME–PME">PME-PME</normalizedToken>
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0.12. Sternum (Fig. 13) 0.94 long, 0.80 wide; labium 0.22 long, 0.24
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<pageBreakToken pageId="5" pageNumber="32" start="start">wide</pageBreakToken>
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at base. Measurements of palp and leg segments as shown in Table 1. Spermathecae weakly sclerotised, round, touching each other, each spermatheca with a pair of accessory glands (Figs 49, 50). Due to the weak sclerotisation the outline of the spermathecae and their ducts are not very clear.
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</paragraph>
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<caption ID-Table-UUID="586DFC7B1932F6957D8F399BE683BA87" httpUri="http://table.plazi.org/id/586DFC7B1932F6957D8F399BE683BA87" pageId="5" pageNumber="32">
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<paragraph pageId="5" pageNumber="32">
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Table 1.
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<taxonomicName class="Arachnida" family="Palpimanidae" genus="Levymanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Levymanus gershomi" order="Araneae" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="gershomi">Levymanus gershomi</taxonomicName>
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, gen. et sp. n., palp and leg measurements (in mm). Male holotype and female paratype (in parentheses).
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</paragraph>
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</caption>
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<paragraph pageId="5" pageNumber="32">
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<table pageId="5" pageNumber="32">
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<tr pageId="5" pageNumber="32">
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<th colspan="1" pageId="5" pageNumber="32" rowspan="1">Palp</th>
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<th colspan="1" pageId="5" pageNumber="32" rowspan="1">Leg I</th>
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<th colspan="1" pageId="5" pageNumber="32" rowspan="1">Leg II</th>
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<th colspan="1" pageId="5" pageNumber="32" rowspan="1">Leg III</th>
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<th colspan="1" pageId="5" pageNumber="32" rowspan="1">Leg IV</th>
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</tr>
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</table>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="32" type="variability">
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<paragraph pageId="5" pageNumber="32">Variability.</paragraph>
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<paragraph pageId="5" pageNumber="32">Carapace length in males 1.01-1.12, in females 1.22-1.34. Coloration varies very slightly, recently moulted specimens are lighter.</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="33" type="distribution">
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<paragraph pageId="6" pageNumber="33">
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<pageBreakToken pageId="6" pageNumber="33" start="start">Distribution</pageBreakToken>
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and habitat.
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</paragraph>
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<paragraph pageId="6" pageNumber="33">The species is known only from the type locality (Qetura) represented by an extra-arid stony desert at 200-500 m above sea level. All specimens were collected with pitfall traps.</paragraph>
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</subSubSection>
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</treatment>
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</document> |