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<document id="7FEBCAEA04E9A380A03CD163C250A414" ID-CLB-Dataset="53409" ID-DOI="10.1093/zoolinnean/zlac100" ID-GBIF-Dataset="194d8698-7037-41e1-8bdc-11cd82e6884c" ID-ISSN="0024-4082" ID-Zenodo-Dep="7981762" ID-ZooBank="E3CF73A6-FCA6-4935-A516-D1E38E49CFB3" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1685085252051" checkinUser="plazi" docAuthor="Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matias A. &amp; Carvalho, Leonardo S." docDate="2023" docId="2366961AA524FFACFC3B6B505D7D69AD" docLanguage="en" docName="zlac100.pdf" docOrigin="Zoological Journal of the Linnean Society 198" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Nerudia centaura Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho 2023, SP. NOV." docType="treatment" docVersion="6" lastPageNumber="556" masterDocId="DF5FEE62A537FFBAFFBE6E0659506F3E" masterDocTitle="High and dry: integrative taxonomy of the Andean spider genus Nerudia (Araneae: Pholcidae)" masterLastPageNumber="591" masterPageNumber="534" pageNumber="553" updateTime="1701998993362" updateUser="plazi">
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<mods:title id="1291EE5265C8907BCFF434C8C39D5212">High and dry: integrative taxonomy of the Andean spider genus Nerudia (Araneae: Pholcidae)</mods:title>
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<mods:namePart id="23F05C2E7CECC02BCC7698DD7D1B7329">Huber, Bernhard A.</mods:namePart>
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<mods:namePart id="38B4121C147B79D174BABC39623B45D4">Meng, Guanliang</mods:namePart>
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<mods:namePart id="F9B9F86B4CBA5AE7E47936B8C3269A40">Král, Jiří</mods:namePart>
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<mods:namePart id="E43FF99ACE5F4427FD402F1EEC686DE9">Ávila Herrera, Ivalú M.</mods:namePart>
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<mods:namePart id="16A5498B06D7810F04CFA36A9DD5C941">Izquierdo, Matias A.</mods:namePart>
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<mods:namePart id="02212DB25191D85B786D42273E7C164D">Carvalho, Leonardo S.</mods:namePart>
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<mods:title id="817DC64A8E4A0593954654BC4D86704B">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="C30DD831E7A131D4A4B001D20C63EDF3">2023</mods:date>
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<mods:number id="3F9904A5DC0CC396773A7AA1081C4E81">198</mods:number>
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<treatment id="2366961AA524FFACFC3B6B505D7D69AD" ID-DOI="http://doi.org/10.5281/zenodo.7981792" ID-GBIF-Taxon="209634228" ID-Zenodo-Dep="7981792" LSID="urn:lsid:plazi:treatment:2366961AA524FFACFC3B6B505D7D69AD" httpUri="http://treatment.plazi.org/id/2366961AA524FFACFC3B6B505D7D69AD" lastPageId="22" lastPageNumber="556" pageId="19" pageNumber="553">
<subSubSection id="E3D57487A524FFA9FC3B6B505C656A50" box="[901,1333,1365,1390]" pageId="19" pageNumber="553" type="nomenclature">
<paragraph id="AB70270CA524FFA9FC3B6B505C656A50" blockId="19.[901,1333,1365,1390]" box="[901,1333,1365,1390]" pageId="19" pageNumber="553">
<heading id="F0389060A524FFA9FC3B6B505C656A50" box="[901,1333,1365,1390]" centered="true" fontSize="9" level="2" pageId="19" pageNumber="553" reason="2">
<taxonomicName id="6CCF5C8FA524FFA9FC3B6B505D266A52" authority="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho, 2023" authorityName="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho" authorityYear="2023" box="[901,1142,1366,1389]" class="Arachnida" family="Pholcidae" genus="Nerudia" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="557" phylum="Arthropoda" rank="species" species="centaura" status="sp. nov.">
<emphasis id="99BBFB1EA524FFA9FC3B6B505D266A52" bold="true" box="[901,1142,1366,1389]" italics="true" pageId="19" pageNumber="553">
<smallCapsWord id="AD96B1D0A524FFA9FC3B6B505AA46A52" baselines="1383,1383" box="[901,1012,1366,1389]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Nerudia" pageId="19" pageNumber="553">NERUDIA</smallCapsWord>
<smallCapsWord id="AD96B1D0A524FFA9FC456B5F5D266A52" baselines="1383" box="[1019,1142,1369,1388]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="centaura" pageId="19" pageNumber="553">CENTAURA</smallCapsWord>
</emphasis>
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<smallCapsWord id="AD96B1D0A524FFA9FBC06B5F5D9B6A53" baselines="1384" box="[1150,1227,1369,1389]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="huber" pageId="19" pageNumber="553">HUBER</smallCapsWord>
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<emphasis id="99BBFB1EA524FFA9FB6C6B5F5DA66A53" bold="true" box="[1234,1270,1365,1389]" pageId="19" pageNumber="553">
<smallCapsWord id="AD96B1D0A524FFA9FB6C6B5F5DBF6A52" baselines="1383" box="[1234,1263,1369,1388]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="sp" pageId="19" pageNumber="553">SP</smallCapsWord>
.
</emphasis>
<smallCapsWord id="AD96B1D0A524FFA9FB436B5F5C7E6A52" baselines="1383" box="[1277,1326,1369,1388]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="nov" pageId="19" pageNumber="553">
<emphasis id="99BBFB1EA524FFA9FB436B5F5C7E6A52" bold="true" box="[1277,1326,1369,1388]" pageId="19" pageNumber="553">NOV</emphasis>
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.
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="E3D57487A524FFA9FC526B7A5CC26AE7" pageId="19" pageNumber="553" type="description">
<paragraph id="AB70270CA524FFA9FC526B7A5D9D6AAB" blockId="19.[1004,1229,1404,1429]" box="[1004,1229,1404,1429]" pageId="19" pageNumber="553">
(
<figureCitation id="33F43B89A524FFA9FC4B6B7A5D216AAB" box="[1013,1137,1404,1429]" captionStart="Figure 1" captionStartId="4.[164,244,1761,1783]" captionTargetBox="[324,1283,651,1731]" captionTargetId="figure-233@4.[323,1283,650,1731]" captionTargetPageId="4" captionText="Figure 1. Nerudia, live specimens from Argentina. A, N. colina sp. nov., female from Jujuy, between San Salvador and Purmamarca (type locality). B, N. poma sp. nov., male from Salta, NW Campo Quijano (type locality). C, D, N. centaura sp. nov., male and female from Catamarca, E Paso de San Francisco (type locality). E, N. trigo sp. nov., female from Salta, SW Alemania. F, N. guirnalda sp. nov., male from Catamarca, El Rodeo (type locality). G, N. ola sp. nov., male from San Juan, San Agustin de Valle Fertil (type locality). H, N. nono sp. nov., male from Cordoba, E Nono (type locality)." figureDoi="http://doi.org/10.5281/zenodo.7981764" httpUri="https://zenodo.org/record/7981764/files/figure.png" pageId="19" pageNumber="553">
<smallCapsWord id="AD96B1D0A524FFA9FC4B6B7A5D7B6AAD" baselines="1423,1422" box="[1013,1067,1404,1429]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Figs" pageId="19" pageNumber="553">FIGS</smallCapsWord>
1C, D
</figureCitation>
,
<figureCitation id="33F43B89A524FFA9FBC06B7A5D946AAB" box="[1150,1220,1404,1429]" captionStart-0="Figure 15" captionStart-1="Figure 16" captionStart-2="Figure 17" captionStart-3="Figure 18" captionStartId-0="20.[163,242,989,1011]" captionStartId-1="21.[146,225,1317,1339]" captionStartId-2="22.[163,241,1096,1118]" captionStartId-3="23.[145,223,1383,1405]" captionTargetBox-0="[325,1278,196,955]" captionTargetBox-1="[309,1261,196,1284]" captionTargetBox-2="[324,1283,196,1065]" captionTargetBox-3="[306,1265,196,1352]" captionTargetId-0="figure-427@20.[323,1283,195,960]" captionTargetId-1="figure-282@21.[305,1265,195,1287]" captionTargetId-2="figure-356@22.[323,1283,195,1066]" captionTargetId-3="figure-204@23.[305,1265,195,1353]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionTargetPageId-2="22" captionTargetPageId-3="23" captionText-0="Figure 15. Nerudia centaura sp. nov.; non-type male from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). Left palp, prolateral (A) and retrolateral (B) views. Scale line: 0.3 mm." captionText-1="Figure 16. Nerudia centaura sp. nov.; non-type male and non-type female from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). A–C, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views; arrow points at dorsal hump at basis of procursus. D–F, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm." captionText-2="Figure 17. Nerudia centaura sp. nov.; non-type female from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). A, abdomen and epigynum, ventral view. B, anterior median tube-like structure in internal genitalia (arrow). C, posterior median structures in internal genitalia; arrow: ‘receptacle’. D, E, cleared genitalia, ventral and dorsal views. Scale lines: 0.2 mm (A, D, E), 0.1 mm (B, C)." captionText-3="Figure 18. Nerudia centaura sp. nov.; female from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Arg216). A, epigynum, ventral view; arrow: possibly entry into tube-like internal structure. B, female ALS and PMS. C, putative glandular pores on female tarsus 3. D, glandular pore and slit sensillum(?) on female tarsus 4. E, tarsal organ on female tarsus 3. F, tip of female right tarsus 1, retrolateral view. G, tip of female right tarsus 3, retrolateral view. H, tip of female left tarsus 4, prolateral view; arrow: comb-hair. Scale lines: 100 µm (A), 10 µm (B, C, F–H), 2 µm (D, E)." figureDoi-0="http://doi.org/10.5281/zenodo.7981810" figureDoi-1="http://doi.org/10.5281/zenodo.7981814" figureDoi-2="http://doi.org/10.5281/zenodo.7981816" figureDoi-3="http://doi.org/10.5281/zenodo.7981820" httpUri-0="https://zenodo.org/record/7981810/files/figure.png" httpUri-1="https://zenodo.org/record/7981814/files/figure.png" httpUri-2="https://zenodo.org/record/7981816/files/figure.png" httpUri-3="https://zenodo.org/record/7981820/files/figure.png" pageId="19" pageNumber="553">15–18</figureCitation>
)
</paragraph>
<paragraph id="AB70270CA524FFA9FC976BA35CC26AE7" blockId="19.[809,1426,1444,1497]" pageId="19" pageNumber="553">
<emphasis id="99BBFB1EA524FFA9FC976BA35D256A84" box="[809,1141,1444,1466]" italics="true" pageId="19" pageNumber="553">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act:
<uuid id="DF691DD9A524FFA9FCCB6BC55CDE6AE7" box="[885,1422,1475,1497]" pageId="19" pageNumber="553">124FC4AB-B97D-445C-AD11-C3604843AED3</uuid>
.
</paragraph>
</subSubSection>
<subSubSection id="E3D57487A524FFAEFC976804584C6B42" lastPageId="20" lastPageNumber="554" pageId="19" pageNumber="553" type="diagnosis">
<paragraph id="AB70270CA524FFAEFC976804584C6B42" blockId="19.[809,1425,1538,1897]" lastBlockId="20.[163,778,1096,1148]" lastPageId="20" lastPageNumber="554" pageId="19" pageNumber="553">
<emphasis id="99BBFB1EA524FFA9FC9768045AE56929" box="[809,949,1538,1559]" italics="true" pageId="19" pageNumber="553">Diagnosis:</emphasis>
Easily distinguished from known congeners by dark coloration of prosoma (
<figureCitation id="33F43B89A524FFA9FAB168265CD26908" box="[1295,1410,1568,1590]" captionStart="Figure 1" captionStartId="4.[164,244,1761,1783]" captionTargetBox="[324,1283,651,1731]" captionTargetId="figure-233@4.[323,1283,650,1731]" captionTargetPageId="4" captionText="Figure 1. Nerudia, live specimens from Argentina. A, N. colina sp. nov., female from Jujuy, between San Salvador and Purmamarca (type locality). B, N. poma sp. nov., male from Salta, NW Campo Quijano (type locality). C, D, N. centaura sp. nov., male and female from Catamarca, E Paso de San Francisco (type locality). E, N. trigo sp. nov., female from Salta, SW Alemania. F, N. guirnalda sp. nov., male from Catamarca, El Rodeo (type locality). G, N. ola sp. nov., male from San Juan, San Agustin de Valle Fertil (type locality). H, N. nono sp. nov., male from Cordoba, E Nono (type locality)." figureDoi="http://doi.org/10.5281/zenodo.7981764" httpUri="https://zenodo.org/record/7981764/files/figure.png" pageId="19" pageNumber="553">Fig. 1C, D</figureCitation>
), by shape of procursus (
<figureCitation id="33F43B89A524FFA9FBFD68395D98696B" box="[1091,1224,1599,1621]" captionStart="Figure 16" captionStartId="21.[146,225,1317,1339]" captionTargetBox="[309,1261,196,1284]" captionTargetId="figure-282@21.[305,1265,195,1287]" captionTargetPageId="21" captionText="Figure 16. Nerudia centaura sp. nov.; non-type male and non-type female from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). A–C, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views; arrow points at dorsal hump at basis of procursus. D–F, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7981814" httpUri="https://zenodo.org/record/7981814/files/figure.png" pageId="19" pageNumber="553">Fig. 16A–C</figureCitation>
; dorsal hump at basis, strongly bent towards dorsal, pointed tip), by bulbal processes (
<figureCitation id="33F43B89A524FFA9FC47687A5D2B69AC" box="[1017,1147,1660,1682]" captionStart="Figure 16" captionStartId="21.[146,225,1317,1339]" captionTargetBox="[309,1261,196,1284]" captionTargetId="figure-282@21.[305,1265,195,1287]" captionTargetPageId="21" captionText="Figure 16. Nerudia centaura sp. nov.; non-type male and non-type female from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). A–C, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views; arrow points at dorsal hump at basis of procursus. D–F, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7981814" httpUri="https://zenodo.org/record/7981814/files/figure.png" pageId="19" pageNumber="553">Fig. 16D–F</figureCitation>
; strong pointed ventral apophysis; embolus slightly shorter, with strong dorsal sclerite), by male chelicerae (
<figureCitation id="33F43B89A524FFA9FB0368BC5C6C69F1" box="[1213,1340,1722,1744]" captionStart="Figure 16" captionStartId="21.[146,225,1317,1339]" captionTargetBox="[309,1261,196,1284]" captionTargetId="figure-282@21.[305,1265,195,1287]" captionTargetPageId="21" captionText="Figure 16. Nerudia centaura sp. nov.; non-type male and non-type female from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). A–C, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views; arrow points at dorsal hump at basis of procursus. D–F, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7981814" httpUri="https://zenodo.org/record/7981814/files/figure.png" pageId="19" pageNumber="553">Fig. 16G, H</figureCitation>
; strong frontal apophyses with flattened tips; stridulatory files on strong lateral protrusion), and by epigynum and female internal genitalia (
<figureCitation id="33F43B89A524FFA9FB2A69105DA46812" box="[1172,1268,1814,1836]" captionStart="Figure 16" captionStartId="21.[146,225,1317,1339]" captionTargetBox="[309,1261,196,1284]" captionTargetId="figure-282@21.[305,1265,195,1287]" captionTargetPageId="21" captionText="Figure 16. Nerudia centaura sp. nov.; non-type male and non-type female from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). A–C, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views; arrow points at dorsal hump at basis of procursus. D–F, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7981814" httpUri="https://zenodo.org/record/7981814/files/figure.png" pageId="19" pageNumber="553">Figs 16I</figureCitation>
,
<figureCitation id="33F43B89A524FFA9FABC69105C4E6815" box="[1282,1310,1814,1835]" captionStart="Figure 17" captionStartId="22.[163,241,1096,1118]" captionTargetBox="[324,1283,196,1065]" captionTargetId="figure-356@22.[323,1283,195,1066]" captionTargetPageId="22" captionText="Figure 17. Nerudia centaura sp. nov.; non-type female from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). A, abdomen and epigynum, ventral view. B, anterior median tube-like structure in internal genitalia (arrow). C, posterior median structures in internal genitalia; arrow: ‘receptacle’. D, E, cleared genitalia, ventral and dorsal views. Scale lines: 0.2 mm (A, D, E), 0.1 mm (B, C)." figureDoi="http://doi.org/10.5281/zenodo.7981816" httpUri="https://zenodo.org/record/7981816/files/figure.png" pageId="19" pageNumber="553">17</figureCitation>
; epigynal plate with large, whitish posterior indentation angular anteriorly; internal genitalia with indistinct posterior ‘receptacle’ and unique anterior tubular membranous structure).
</paragraph>
</subSubSection>
<caption id="FFB07784A523FFAEFF1D6DDB5AA36B2F" ID-DOI="http://doi.org/10.5281/zenodo.7981810" ID-Zenodo-Dep="7981810" httpUri="https://zenodo.org/record/7981810/files/figure.png" pageId="20" pageNumber="554" startId="20.[163,242,989,1011]" targetBox="[325,1278,196,955]" targetPageId="20" targetType="figure">
<paragraph id="AB70270CA523FFAEFF1D6DDB5AA36B2F" blockId="20.[163,1442,989,1041]" pageId="20" pageNumber="554">
<emphasis id="99BBFB1EA523FFAEFF1D6DDB58486CCC" bold="true" box="[163,280,989,1011]" pageId="20" pageNumber="554">Figure 15.</emphasis>
<taxonomicName id="6CCF5C8FA523FFAEFE9A6DD8588D6CCD" authority="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho, 2023" authorityName="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho" authorityYear="2023" box="[292,477,990,1012]" class="Arachnida" family="Pholcidae" genus="Nerudia" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="554" phylum="Arthropoda" rank="species" species="centaura" status="sp. nov.">
<emphasis id="99BBFB1EA523FFAEFE9A6DD8588D6CCD" box="[292,477,990,1012]" italics="true" pageId="20" pageNumber="554">Nerudia centaura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82884665A523FFAEFE5A6DD85B636CCA" box="[484,563,990,1012]" pageId="20" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
; non-type male from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). Left palp, prolateral (A) and retrolateral (B) views. Scale line: 0.3 mm.
</paragraph>
</caption>
<subSubSection id="E3D57487A523FFAEFF1D6AA058326852" pageId="20" pageNumber="554" type="materials_examined">
<paragraph id="AB70270CA523FFAEFF1D6AA058426A6A" blockId="20.[163,779,1188,1364]" pageId="20" pageNumber="554">
<emphasis id="99BBFB1EA523FFAEFF1D6AA0580B6B84" box="[163,347,1189,1211]" italics="true" pageId="20" pageNumber="554">Type material:</emphasis>
<materialsCitation id="1BA72D51A523FFAEFED76AA35A5B6A28" ID-GBIF-Occurrence="4118252373" collectingDate="2019-03-27" collectionCode="LABRE-Ar" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="4180" latitude="-26.9276" location="Paso de San Francisco" longLatPrecision="6" longitude="-68.0709" pageId="20" pageNumber="554" specimenCode="LABRE-Ar 587" specimenCount="1" specimenCount-female="2" specimenCount-male="1" stateProvince="Catamarca" typeStatus="holotype">
<collectingCountry id="D3D8679CA523FFAEFED76AA35B416B84" box="[361,529,1189,1210]" name="Argentina" pageId="20" pageNumber="554">ARGENTINA</collectingCountry>
<emphasis id="99BBFB1EA523FFAEFDA36AA05B796B85" box="[541,553,1190,1211]" italics="true" pageId="20" pageNumber="554">–</emphasis>
<emphasis id="99BBFB1EA523FFAEFD8B6AA35B9D6B85" bold="true" box="[565,717,1189,1211]" pageId="20" pageNumber="554">
<collectingRegion id="690BE9EEA523FFAEFD8B6AA35B9D6B85" box="[565,717,1189,1211]" country="Argentina" name="Catamarca" pageId="20" pageNumber="554">Catamarca</collectingRegion>
</emphasis>
: •
<specimenCount id="BDC9EC85A523FFAEFD446AA25A5A6B85" box="[762,778,1188,1211]" pageId="20" pageNumber="554" type="male">♂</specimenCount>
<typeStatus id="747499AEA523FFAEFF1D6AC2585B6BE4" box="[163,267,1220,1242]" pageId="20" pageNumber="554" type="holotype">holotype</typeStatus>
; ~
<quantity id="6C378AE9A523FFAEFE996AC258236BE4" box="[295,371,1220,1242]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="554" unit="km" value="20.0">
<locationDeviation id="BAC23E2EA523FFAEFE996AC258236BE4" box="[295,371,1220,1242]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="554" unit="km" value="20.0">20 km</locationDeviation>
</quantity>
E
<location id="AE1071D7A523FFAEFE266AC25BF96BE4" LSID="urn:lsid:plazi:treatment:2366961AA524FFACFC3B6B505D7D69AD:AE1071D7A523FFAEFE266AC25BF96BE4" box="[408,681,1220,1242]" country="Argentina" latitude="-26.9276" longLatPrecision="6" longitude="-68.0709" name="Paso de San Francisco" pageId="20" pageNumber="554" stateProvince="Catamarca">Paso de San Francisco</location>
, ‘site 1’;
<geoCoordinate id="CEFB41CBA523FFAEFF1D6AE558726BC6" box="[163,290,1251,1273]" degrees="26.9276" direction="south" orientation="latitude" pageId="20" pageNumber="554" precision="5" value="-26.9276">26.9276° S</geoCoordinate>
,
<geoCoordinate id="CEFB41CBA523FFAEFE8E6AE558E76BC6" box="[304,439,1251,1273]" degrees="68.0709" direction="west" orientation="longitude" pageId="20" pageNumber="554" precision="5" value="-68.0709">68.0709° W</geoCoordinate>
;
<quantity id="6C378AE9A523FFAEFE7D6AE55B4A6BC6" box="[451,538,1251,1272]" metricMagnitude="3" metricUnit="m" metricValue="4.18" pageId="20" pageNumber="554" unit="m" value="4180.0">
<elevation id="20E2C03FA523FFAEFE7D6AE55B4A6BC6" box="[451,538,1251,1272]" metricMagnitude="3" metricUnit="m" metricValue="4.18" pageId="20" pageNumber="554" unit="m" value="4180.0">4180 m</elevation>
</quantity>
a.s.l.;
<date id="DF7101CCA523FFAEFDD46AE55A566BC6" box="[618,774,1251,1272]" pageId="20" pageNumber="554" value="2019-03-27">
<collectingDate id="CF35F824A523FFAEFDD46AE55A566BC6" box="[618,774,1251,1272]" pageId="20" pageNumber="554" value="2019-03-27">27 Mar. 2019</collectingDate>
</date>
;
<collectorName id="063A42DAA523FFAEFF1D6B04587B6A29" box="[163,299,1281,1303]" pageId="20" pageNumber="554">B. A. Huber</collectorName>
and
<collectorName id="063A42DAA523FFAEFED96B045B4B6A29" box="[359,539,1281,1303]" pageId="20" pageNumber="554">M. A. Izquierdo</collectorName>
leg.;
<emphasis id="99BBFB1EA523FFAEFDE46B045A5B6A28" box="[602,779,1281,1303]" italics="true" pageId="20" pageNumber="554">
<specimenCode id="FB698F77A523FFAEFDE46B045A5B6A28" box="[602,779,1281,1303]" collectionCode="LABRE-Ar" pageId="20" pageNumber="554">LABRE-Ar 587</specimenCode>
</emphasis>
</materialsCitation>
•
<materialsCitation id="1BA72D51A523FFAEFF076B26585E6A6A" ID-GBIF-Occurrence="4118252309" collectingDate="2019-03-27" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="4180" latitude="-26.9276" location="Paso de San Francisco" longLatPrecision="6" longitude="-68.0709" pageId="20" pageNumber="554" specimenCode="ZFMK Ar 23890" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Catamarca" typeStatus="paratype">
<specimenCount id="BDC9EC85A523FFAEFF076B26598F6A08" box="[185,223,1311,1334]" pageId="20" pageNumber="554" type="male">1 ♂</specimenCount>
,
<specimenCount id="BDC9EC85A523FFAEFF536B2658706A0B" box="[237,288,1309,1333]" pageId="20" pageNumber="554" type="female">2 ♀♀</specimenCount>
,
<typeStatus id="747499AEA523FFAEFE906B2658F36A0B" box="[302,419,1312,1333]" pageId="20" pageNumber="554" type="paratype">paratypes</typeStatus>
; same data as holotype;
<specimenCode id="FB698F77A523FFAEFD7E6B26585E6A6A" pageId="20" pageNumber="554">ZFMK Ar 23890</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="AB70270CA523FFAEFF1D6B8D58326852" blockId="20.[163,780,1419,1900]" pageId="20" pageNumber="554">
<materialsCitation id="1BA72D51A523FFAEFF1D6B8D58B46AE0" ID-GBIF-Occurrence="4118252319" collectingDate="2019-03-27" collectionCode="ZFMK" collectorName="Arg" country="Argentina" elevation="4180" latitude="-26.9276" location="Catamarca" longLatPrecision="6" longitude="-68.0709" pageId="20" pageNumber="554" specimenCode="ZFMK Arg215" specimenCount="6" specimenCount-female="6" stateProvince="Catamarca">
<emphasis id="99BBFB1EA523FFAEFF1D6B8D5B706A9E" box="[163,544,1419,1440]" italics="true" pageId="20" pageNumber="554">Other material examined:</emphasis>
ARGENTINA
<emphasis id="99BBFB1EA523FFAEFD406B8D5A5A6A9E" box="[766,778,1419,1440]" italics="true" pageId="20" pageNumber="554">–</emphasis>
<emphasis id="99BBFB1EA523FFAEFF1D6BAF587F6A81" bold="true" box="[163,303,1449,1471]" pageId="20" pageNumber="554">Catamarca</emphasis>
: •
<specimenCount id="BDC9EC85A523FFAEFEEE6BAF58D26A80" box="[336,386,1446,1471]" pageId="20" pageNumber="554" type="female">6 ♀♀</specimenCount>
, 3 juvs, in pure ethanol; same data as holotype;
<specimenCode id="FB698F77A523FFAEFE846BCE58B46AE0" box="[314,484,1480,1502]" pageId="20" pageNumber="554">ZFMK Arg215</specimenCode>
</materialsCitation>
•
<materialsCitation id="1BA72D51A523FFAEFDBA6BCE58A36AC2" ID-GBIF-Occurrence="4118252357" collectingDate="2019-03-27" collectionCode="LABRE-Ar" collectorName="Arg" country="Argentina" elevation="4180" latitude="-26.9276" location="Paso de San Francisco" longLatPrecision="6" longitude="-68.0709" pageId="20" pageNumber="554" specimenCode="LABRE-Ar 526" specimenCount="10" specimenCount-female="6" specimenCount-male="4" stateProvince="Catamarca">
<specimenCount id="BDC9EC85A523FFAEFDBA6BCE5B6C6AE0" box="[516,572,1479,1502]" pageId="20" pageNumber="554" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="BDC9EC85A523FFAEFDF56BCE5BD06AE3" box="[587,640,1477,1502]" pageId="20" pageNumber="554" type="female">6 ♀♀</specimenCount>
; same data as holotype;
<specimenCode id="FB698F77A523FFAEFE826BE158A36AC2" box="[316,499,1510,1532]" collectionCode="LABRE-Ar" pageId="20" pageNumber="554">LABRE-Ar 526</specimenCode>
</materialsCitation>
•
<materialsCitation id="1BA72D51A523FFAEFDAA6BE15BE66925" ID-GBIF-Occurrence="4118252320" collectingDate="2019-03-27" collectionCode="LABRE-Ar" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="4180" latitude="-26.9276" location="Paso de San Francisco" longLatPrecision="6" longitude="-68.0709" pageId="20" pageNumber="554" specimenCode="LABRE-Ar 542" specimenCount="3" specimenCount-female="3" stateProvince="Catamarca">
<specimenCount id="BDC9EC85A523FFAEFDAA6BE15B1A6AC2" box="[532,586,1508,1532]" pageId="20" pageNumber="554" type="female">3 ♀♀</specimenCount>
, 7 juvs, in pure ethanol; same data as holotype;
<specimenCode id="FB698F77A523FFAEFDB468035BE66925" box="[522,694,1541,1563]" collectionCode="LABRE-Ar" pageId="20" pageNumber="554">LABRE-Ar 542</specimenCode>
</materialsCitation>
•
<materialsCitation id="1BA72D51A523FFAEFD6F68035BB169AB" ID-GBIF-Occurrence="4118252322" collectingDate="2019-03-27" collectionCode="ZFMK" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="4400" location="Paso de San Francisco" longLatPrecision="6" pageId="20" pageNumber="554" specimenCode="ZFMK Ar 23891" specimenCount="10" specimenCount-female="6" specimenCount-male="4" stateProvince="Catamarca">
<specimenCount id="BDC9EC85A523FFAEFD6F68035A556925" box="[721,773,1540,1563]" pageId="20" pageNumber="554" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="BDC9EC85A523FFAEFF1D682259866907" box="[163,214,1569,1594]" pageId="20" pageNumber="554" type="female">5 ♀♀</specimenCount>
(
<specimenCount id="BDC9EC85A523FFAEFF5B6822581D6904" box="[229,333,1572,1594]" pageId="20" pageNumber="554" type="male">one male</specimenCount>
and
<specimenCount id="BDC9EC85A523FFAEFE3968225B5F6904" box="[391,527,1572,1594]" pageId="20" pageNumber="554" type="female">two females</specimenCount>
used for µ-CT study);
<locationDeviation id="BAC23E2EA523FFAEFF1D684558476966" box="[163,279,1603,1625]" pageId="20" pageNumber="554">
~
<quantity id="6C378AE9A523FFAEFF0F684559AC6967" box="[177,252,1603,1625]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="554" unit="km" value="20.0">20 km</quantity>
E
</locationDeviation>
<location id="AE1071D7A523FFAEFE9E68455B7D6967" LSID="urn:lsid:plazi:treatment:2366961AA524FFACFC3B6B505D7D69AD:AE1071D7A523FFAEFE9E68455B7D6967" box="[288,557,1603,1625]" country="Argentina" latitude="-26.9276" longLatPrecision="6" longitude="-68.0709" name="Paso de San Francisco" pageId="20" pageNumber="554" stateProvince="Catamarca">Paso de San Francisco</location>
, ‘site 2’; 26.936° S, 68.090–68.095° W; 4270–
<quantity id="6C378AE9A523FFAEFE0368675B5F6949" box="[445,527,1633,1655]" metricMagnitude="3" metricUnit="m" metricValue="4.335" metricValueMax="4.4" metricValueMin="4.27" pageId="20" pageNumber="554" unit="m" value="4335.0" valueMax="4400.0" valueMin="4270.0">
<elevation id="20E2C03FA523FFAEFE0368675B5F6949" box="[445,527,1633,1655]" pageId="20" pageNumber="554" unit="m" value="4400">4400 m</elevation>
</quantity>
a.s.l.;
<date id="DF7101CCA523FFAEFDE868675BBA6948" box="[598,746,1633,1655]" pageId="20" pageNumber="554" value="2019-03-27">
<collectingDate id="CF35F824A523FFAEFDE868675BBA6948" box="[598,746,1633,1655]" pageId="20" pageNumber="554" value="2019-03-27">27 Mar. 2019</collectingDate>
</date>
;
<collectorName id="063A42DAA523FFAEFD4A6864585A69A8" pageId="20" pageNumber="554">B. A. Huber</collectorName>
and
<collectorName id="063A42DAA523FFAEFEFC688658BF69A8" box="[322,495,1664,1686]" pageId="20" pageNumber="554">M. A. Izquierdo</collectorName>
leg.;
<specimenCode id="FB698F77A523FFAEFD9768865BB169AB" box="[553,737,1664,1685]" pageId="20" pageNumber="554">ZFMK Ar 23891</specimenCode>
</materialsCitation>
•
<materialsCitation id="1BA72D51A523FFAEFD4368865B2769CC" ID-GBIF-Occurrence="4118252341" collectingDate="2019-03-27" collectionCode="ZFMK" collectorName="Arg" country="Argentina" elevation="4335" latitude="68.09" location="20 km E Paso de San Francisco" longLatPrecision="59" longitude="-68.095" pageId="20" pageNumber="554" specimenCode="ZFMK Arg216" specimenCount="8" specimenCount-female="8" stateProvince="Catamarca">
<specimenCount id="BDC9EC85A523FFAEFD4368865991698A" pageId="20" pageNumber="554" type="female">6 ♀♀</specimenCount>
, 2 juvs, in pure ethanol (
<specimenCount id="BDC9EC85A523FFAEFE5C68995B0F698B" box="[482,607,1695,1717]" pageId="20" pageNumber="554" type="female">one female</specimenCount>
used for SEM;
<specimenCount id="BDC9EC85A523FFAEFF1D68BB586B69ED" box="[163,315,1725,1747]" pageId="20" pageNumber="554" type="female">three female</specimenCount>
prosomata used for molecular study); same data as preceding;
<specimenCode id="FB698F77A523FFAEFE7368DA5B2769CC" box="[461,631,1756,1778]" pageId="20" pageNumber="554">ZFMK Arg216</specimenCode>
</materialsCitation>
•
<materialsCitation id="1BA72D51A523FFAEFD2668DA5B36682E" ID-GBIF-Occurrence="4118252334" collectingDate="2019-03-27" collectionCode="LABRE-Ar" collectorName="Arg" country="Argentina" elevation="4335" latitude="68.09" longLatPrecision="59" longitude="-68.095" pageId="20" pageNumber="554" specimenCode="LABRE-Ar 527" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Catamarca">
<specimenCount id="BDC9EC85A523FFAEFD2668DA5B9F69CC" box="[664,719,1755,1778]" pageId="20" pageNumber="554" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="BDC9EC85A523FFAEFD6168DA5A5569CF" box="[735,773,1753,1777]" pageId="20" pageNumber="554" type="female">1 ♀</specimenCount>
; same data as preceding;
<specimenCode id="FB698F77A523FFAEFE0468FD5B36682E" box="[442,614,1786,1808]" collectionCode="LABRE-Ar" pageId="20" pageNumber="554">LABRE-Ar 527</specimenCode>
</materialsCitation>
•
<materialsCitation id="1BA72D51A523FFAEFD3D68FC599D6870" ID-GBIF-Occurrence="4118252332" collectingDate="2019-03-27" collectionCode="LABRE-Ar" collectorName="Arg" country="Argentina" elevation="4335" latitude="68.09" location="20 km E Paso de San Francisco" longLatPrecision="59" longitude="-68.095" pageId="20" pageNumber="554" specimenCode="LABRE-Ar 551" specimenCount="6" specimenCount-female="6" stateProvince="Catamarca">
<specimenCount id="BDC9EC85A523FFAEFD3D68FC5BE5682E" box="[643,693,1784,1808]" pageId="20" pageNumber="554" type="female">6 ♀♀</specimenCount>
, 3 juvs, in pure ethanol; same data as preceding;
<specimenCode id="FB698F77A523FFAEFD35691F599D6870" collectionCode="LABRE-Ar" pageId="20" pageNumber="554">LABRE-Ar 551</specimenCode>
</materialsCitation>
•
<materialsCitation id="1BA72D51A523FFAEFF52693E580E6852" ID-GBIF-Occurrence="4118252379" collectingDate="2019-03-27" collectionCode="ZFMK" collectorName="Ar" country="Argentina" elevation="4450" latitude="68.09" location="20 km E Paso de San Francisco" longLatPrecision="59" longitude="-68.095" pageId="20" pageNumber="554" specimenCode="ZFMK Ar 23892" specimenCount="1" specimenCount-female="1" stateProvince="Catamarca">
<specimenCount id="BDC9EC85A523FFAEFF52693E58426873" box="[236,274,1845,1869]" pageId="20" pageNumber="554" type="female">1 ♀</specimenCount>
; same data as preceding but
<quantity id="6C378AE9A523FFAEFDD0693E5B956873" box="[622,709,1848,1870]" metricMagnitude="3" metricUnit="m" metricValue="4.45" pageId="20" pageNumber="554" unit="m" value="4450.0">
<elevation id="20E2C03FA523FFAEFDD0693E5B956873" box="[622,709,1848,1870]" metricMagnitude="3" metricUnit="m" metricValue="4.45" pageId="20" pageNumber="554" unit="m" value="4450.0">4450 m</elevation>
</quantity>
a.s.l.;
<specimenCode id="FB698F77A523FFAEFF1D6951580E6852" box="[163,350,1879,1900]" pageId="20" pageNumber="554">ZFMK Ar 23892</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E3D57487A523FFAEFC856A4E5DCD6BA5" pageId="20" pageNumber="554" type="etymology">
<paragraph id="AB70270CA523FFAEFC856A4E5DCD6BA5" blockId="20.[827,1443,1096,1179]" pageId="20" pageNumber="554">
<emphasis id="99BBFB1EA523FFAEFC856A4E5AEA6B63" box="[827,954,1096,1117]" italics="true" pageId="20" pageNumber="554">Etymology:</emphasis>
The species epithet
<taxonomicName id="6CCF5C8FA523FFAEFB146A4F5C5F6B60" authorityName="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho" authorityYear="2023" box="[1194,1295,1097,1118]" class="Arachnida" family="Pholcidae" genus="Nerudia" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="554" phylum="Arthropoda" rank="species" species="centaura">
<emphasis id="99BBFB1EA523FFAEFB146A4F5C5F6B60" box="[1194,1295,1097,1118]" italics="true" pageId="20" pageNumber="554">centaura</emphasis>
</taxonomicName>
(Spanish for ‘female centaur’) is taken from Pablo Neruda’s poem ‘Soneto 22’; noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="E3D57487A523FFACFC856AD258466A7F" lastPageId="22" lastPageNumber="556" pageId="20" pageNumber="554" type="description">
<paragraph id="AB70270CA523FFAEFC856AD25A976BD5" blockId="20.[827,967,1236,1259]" box="[827,967,1236,1259]" pageId="20" pageNumber="554">
<emphasis id="99BBFB1EA523FFAEFC856AD25A976BD5" box="[827,967,1236,1259]" italics="true" pageId="20" pageNumber="554">Description</emphasis>
</paragraph>
<paragraph id="AB70270CA523FFAEFC856B0A5C556A84" blockId="20.[827,1443,1292,1467]" pageId="20" pageNumber="554">
<emphasis id="99BBFB1EA523FFAEFC856B0A5DEE6A1C" box="[827,1214,1292,1314]" italics="true" pageId="20" pageNumber="554">
Male (
<typeStatus id="747499AEA523FFAEFC366B0A5AA06A1F" box="[904,1008,1292,1313]" pageId="20" pageNumber="554" type="holotype">holotype</typeStatus>
). Measurements:
</emphasis>
Total body length 1.9, carapace width 0.77. Distance PME–PME 90 µm; diameter PME 55 µm; distance PME–ALE 30 µm; distance AME–AME 20 µm; diameter AME 40 µm. Leg 1: 4.55 (1.25 + 0.25 + 1.30 + 1.30 + 0.45), tibia 2: 1.10, tibia 3: 1.00, tibia 4: 1.38; tibia 1 L/d: 14.
</paragraph>
<paragraph id="AB70270CA523FFAEFC856BE25D0A6909" blockId="20.[827,1441,1508,1591]" pageId="20" pageNumber="554">
<emphasis id="99BBFB1EA523FFAEFC856BE25D6B6AC7" box="[827,1083,1508,1529]" italics="true" pageId="20" pageNumber="554">Colour (in ethanol):</emphasis>
Prosoma and legs ochre to light brown; legs without dark rings; abdomen monochromous pale grey.
</paragraph>
<paragraph id="AB70270CA523FFAEFC8568665C1A69EC" blockId="20.[827,1442,1632,1746]" pageId="20" pageNumber="554">
<emphasis id="99BBFB1EA523FFAEFC8568665A2E694B" box="[827,894,1632,1653]" italics="true" pageId="20" pageNumber="554">Body:</emphasis>
Habitus as in
<figureCitation id="33F43B89A523FFAEFB8668665DE0694B" box="[1080,1200,1632,1654]" captionStart="Figure 1" captionStartId="4.[164,244,1761,1783]" captionTargetBox="[324,1283,651,1731]" captionTargetId="figure-233@4.[323,1283,650,1731]" captionTargetPageId="4" captionText="Figure 1. Nerudia, live specimens from Argentina. A, N. colina sp. nov., female from Jujuy, between San Salvador and Purmamarca (type locality). B, N. poma sp. nov., male from Salta, NW Campo Quijano (type locality). C, D, N. centaura sp. nov., male and female from Catamarca, E Paso de San Francisco (type locality). E, N. trigo sp. nov., female from Salta, SW Alemania. F, N. guirnalda sp. nov., male from Catamarca, El Rodeo (type locality). G, N. ola sp. nov., male from San Juan, San Agustin de Valle Fertil (type locality). H, N. nono sp. nov., male from Cordoba, E Nono (type locality)." figureDoi="http://doi.org/10.5281/zenodo.7981764" httpUri="https://zenodo.org/record/7981764/files/figure.png" pageId="20" pageNumber="554">Figure 1C</figureCitation>
. Ocular area barely raised. Carapace without thoracic groove. Clypeus unmodified. Sternum wider than long (0.56/0.48), without anterior processes. Abdomen globular.
</paragraph>
<paragraph id="AB70270CA523FFAEFC8568FD5D476853" blockId="20.[827,1443,1787,1901]" pageId="20" pageNumber="554">
<emphasis id="99BBFB1EA523FFAEFC8568FD5A96682E" box="[827,966,1787,1808]" italics="true" pageId="20" pageNumber="554">Chelicerae:</emphasis>
As in
<figureCitation id="33F43B89A523FFAEFB9C68FD5D89682E" box="[1058,1241,1787,1809]" captionStart="Figure 16" captionStartId="21.[146,225,1317,1339]" captionTargetBox="[309,1261,196,1284]" captionTargetId="figure-282@21.[305,1265,195,1287]" captionTargetPageId="21" captionText="Figure 16. Nerudia centaura sp. nov.; non-type male and non-type female from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). A–C, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views; arrow points at dorsal hump at basis of procursus. D–F, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7981814" httpUri="https://zenodo.org/record/7981814/files/figure.png" pageId="20" pageNumber="554">Figure 16G, H</figureCitation>
; pair of frontal apophyses directed downwards, with obtuse tip slightly flattened; stridulatory files on pair of distinct lateral protrusions.
</paragraph>
<caption id="FFB07784A522FFAFFF2C6B235D876AAD" ID-DOI="http://doi.org/10.5281/zenodo.7981814" ID-Zenodo-Dep="7981814" httpUri="https://zenodo.org/record/7981814/files/figure.png" pageId="21" pageNumber="555" startId="21.[146,225,1317,1339]" targetBox="[309,1261,196,1284]" targetPageId="21" targetType="figure">
<paragraph id="AB70270CA522FFAFFF2C6B235D876AAD" blockId="21.[145,1425,1317,1427]" pageId="21" pageNumber="555">
<emphasis id="99BBFB1EA522FFAFFF2C6B2358596A04" bold="true" box="[146,265,1317,1339]" pageId="21" pageNumber="555">Figure 16.</emphasis>
<taxonomicName id="6CCF5C8FA522FFAFFEAD6B23589F6A05" authority="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho, 2023" authorityName="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho" authorityYear="2023" box="[275,463,1317,1339]" class="Arachnida" family="Pholcidae" genus="Nerudia" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="555" phylum="Arthropoda" rank="species" species="centaura" status="sp. nov.">
<emphasis id="99BBFB1EA522FFAFFEAD6B23589F6A05" box="[275,463,1317,1339]" italics="true" pageId="21" pageNumber="555">Nerudia centaura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82884665A522FFAFFE686B235B766A05" box="[470,550,1317,1339]" pageId="21" pageNumber="555" rank="species">sp. nov.</taxonomicNameLabel>
; non-type male and non-type female from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). A–C, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views; arrow points at dorsal hump at basis of procursus. D–F, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm.
</paragraph>
</caption>
<paragraph id="AB70270CA522FFAFFF2F6BC25B8269D0" blockId="21.[145,761,1476,1774]" pageId="21" pageNumber="555">
<emphasis id="99BBFB1EA522FFAFFF2F6BC259886AE7" box="[145,216,1476,1497]" italics="true" pageId="21" pageNumber="555">Palps:</emphasis>
As in
<figureCitation id="33F43B89A522FFAFFE956BC258CE6AE4" box="[299,414,1476,1498]" captionStart="Figure 15" captionStartId="20.[163,242,989,1011]" captionTargetBox="[325,1278,196,955]" captionTargetId="figure-427@20.[323,1283,195,960]" captionTargetPageId="20" captionText="Figure 15. Nerudia centaura sp. nov.; non-type male from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). Left palp, prolateral (A) and retrolateral (B) views. Scale line: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.7981810" httpUri="https://zenodo.org/record/7981810/files/figure.png" pageId="21" pageNumber="555">Figure 15</figureCitation>
; coxa unmodified; trochanter with indistinct ventral projection; femur cylindrical, only slightly widened distally, proximally with indistinct retrolateral hump and prolateral stridulatory pick (modified hair); patella short; tibia globular; procursus (
<figureCitation id="33F43B89A522FFAFFE3068585B45694A" box="[398,533,1630,1652]" captionStart="Figure 16" captionStartId="21.[146,225,1317,1339]" captionTargetBox="[309,1261,196,1284]" captionTargetId="figure-282@21.[305,1265,195,1287]" captionTargetPageId="21" captionText="Figure 16. Nerudia centaura sp. nov.; non-type male and non-type female from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). A–C, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views; arrow points at dorsal hump at basis of procursus. D–F, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7981814" httpUri="https://zenodo.org/record/7981814/files/figure.png" pageId="21" pageNumber="555">Fig. 16A–C</figureCitation>
) simple, in lateral view bent towards dorsal, in dorsal view bent towards prolateral, with pointed tip; genital bulb (
<figureCitation id="33F43B89A522FFAFFDCF689D5BA1698F" box="[625,753,1691,1713]" captionStart="Figure 16" captionStartId="21.[146,225,1317,1339]" captionTargetBox="[309,1261,196,1284]" captionTargetId="figure-282@21.[305,1265,195,1287]" captionTargetPageId="21" captionText="Figure 16. Nerudia centaura sp. nov.; non-type male and non-type female from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). A–C, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views; arrow points at dorsal hump at basis of procursus. D–F, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7981814" httpUri="https://zenodo.org/record/7981814/files/figure.png" pageId="21" pageNumber="555">Fig. 16D–F</figureCitation>
) large, with strong pointed ventral apophysis, embolus partly membranous, dorsally with distinct sclerite.
</paragraph>
<paragraph id="AB70270CA522FFAFFF2F691E5DA16AC7" blockId="21.[145,760,1815,1898]" lastBlockId="21.[809,1424,1476,1529]" pageId="21" pageNumber="555">
<emphasis id="99BBFB1EA522FFAFFF2F691E59816813" box="[145,209,1816,1837]" italics="true" pageId="21" pageNumber="555">Legs:</emphasis>
Without spines and curved hairs; vertical hairs in high densities on tibiae 1–2; retrolateral t r i c h o b o t h r i u m o f t i b i a 1 a t 6 8%; p r o l a t e r a l trichobothrium absent on tibia 1; tarsus 1 with six to seven pseudosegments, distally distinct.
</paragraph>
<paragraph id="AB70270CA522FFAFFC9768245DF16968" blockId="21.[809,1423,1570,1622]" pageId="21" pageNumber="555">
<emphasis id="99BBFB1EA522FFAFFC9768245D506909" box="[809,1024,1570,1591]" italics="true" pageId="21" pageNumber="555">Variation (male):</emphasis>
Tibia
<quantity id="6C378AE9A522FFAFFBE468245DDB6906" box="[1114,1163,1570,1592]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="21" pageNumber="555" unit="in" value="1.0">1 in</quantity>
<specimenCount id="BDC9EC85A522FFAFFB2868245C576906" box="[1174,1287,1570,1592]" pageId="21" pageNumber="555" type="male">12 males</specimenCount>
(including
<typeStatus id="747499AEA522FFAFFC9768465ADD6968" box="[809,909,1600,1622]" pageId="21" pageNumber="555" type="holotype">holotype</typeStatus>
): 1.15–1.33 (mean 1.26).
</paragraph>
<paragraph id="AB70270CA522FFACFC97687958466A7F" blockId="21.[809,1426,1663,1900]" lastBlockId="22.[163,779,1293,1345]" lastPageId="22" lastPageNumber="556" pageId="21" pageNumber="555">
<emphasis id="99BBFB1EA522FFAFFC9768795AD169AA" box="[809,897,1663,1684]" italics="true" pageId="21" pageNumber="555">Female:</emphasis>
In general, similar to male (
<figureCitation id="33F43B89A522FFAFFB7368795C7069AA" box="[1229,1312,1663,1685]" captionStart="Figure 1" captionStartId="4.[164,244,1761,1783]" captionTargetBox="[324,1283,651,1731]" captionTargetId="figure-233@4.[323,1283,650,1731]" captionTargetPageId="4" captionText="Figure 1. Nerudia, live specimens from Argentina. A, N. colina sp. nov., female from Jujuy, between San Salvador and Purmamarca (type locality). B, N. poma sp. nov., male from Salta, NW Campo Quijano (type locality). C, D, N. centaura sp. nov., male and female from Catamarca, E Paso de San Francisco (type locality). E, N. trigo sp. nov., female from Salta, SW Alemania. F, N. guirnalda sp. nov., male from Catamarca, El Rodeo (type locality). G, N. ola sp. nov., male from San Juan, San Agustin de Valle Fertil (type locality). H, N. nono sp. nov., male from Cordoba, E Nono (type locality)." figureDoi="http://doi.org/10.5281/zenodo.7981764" httpUri="https://zenodo.org/record/7981764/files/figure.png" pageId="21" pageNumber="555">Fig. 1D</figureCitation>
) but with usual low density of vertical hairs on tibiae. Tibia
<quantity id="6C378AE9A522FFAFFA3D68985A1069EC" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="21" pageNumber="555" unit="in" value="1.0">1 in</quantity>
<specimenCount id="BDC9EC85A522FFAFFCF968BB5A9669EC" box="[839,966,1724,1746]" pageId="21" pageNumber="555" type="female">22 females</specimenCount>
: 1.07–1.48 (mean 1.28). Epigynum (
<figureCitation id="33F43B89A522FFAFFAD868BA5A0769CE" captionStart="Figure 17" captionStartId="22.[163,241,1096,1118]" captionTargetBox="[324,1283,196,1065]" captionTargetId="figure-356@22.[323,1283,195,1066]" captionTargetPageId="22" captionText="Figure 17. Nerudia centaura sp. nov.; non-type female from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). A, abdomen and epigynum, ventral view. B, anterior median tube-like structure in internal genitalia (arrow). C, posterior median structures in internal genitalia; arrow: ‘receptacle’. D, E, cleared genitalia, ventral and dorsal views. Scale lines: 0.2 mm (A, D, E), 0.1 mm (B, C)." figureDoi="http://doi.org/10.5281/zenodo.7981816" httpUri="https://zenodo.org/record/7981816/files/figure.png" pageId="21" pageNumber="555">Fig. 17A</figureCitation>
) anterior plate weakly protruding, with large, whitish posterior indentation; posterior plate large, simple. Internal genitalia (
<figureCitation id="33F43B89A522FFAFFBC3691E5DB36810" box="[1149,1251,1816,1838]" captionStart="Figure 16" captionStartId="21.[146,225,1317,1339]" captionTargetBox="[309,1261,196,1284]" captionTargetId="figure-282@21.[305,1265,195,1287]" captionTargetPageId="21" captionText="Figure 16. Nerudia centaura sp. nov.; non-type male and non-type female from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). A–C, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views; arrow points at dorsal hump at basis of procursus. D–F, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7981814" httpUri="https://zenodo.org/record/7981814/files/figure.png" pageId="21" pageNumber="555">Figs 16I</figureCitation>
,
<figureCitation id="33F43B89A522FFAFFB4B691F5C166810" box="[1269,1350,1817,1838]" captionStart="Figure 17" captionStartId="22.[163,241,1096,1118]" captionTargetBox="[324,1283,196,1065]" captionTargetId="figure-356@22.[323,1283,195,1066]" captionTargetPageId="22" captionText="Figure 17. Nerudia centaura sp. nov.; non-type female from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). A, abdomen and epigynum, ventral view. B, anterior median tube-like structure in internal genitalia (arrow). C, posterior median structures in internal genitalia; arrow: ‘receptacle’. D, E, cleared genitalia, ventral and dorsal views. Scale lines: 0.2 mm (A, D, E), 0.1 mm (B, C)." figureDoi="http://doi.org/10.5281/zenodo.7981816" httpUri="https://zenodo.org/record/7981816/files/figure.png" pageId="21" pageNumber="555">17B–E</figureCitation>
) with simple ‘receptacle’ (
<figureCitation id="33F43B89A522FFAFFB9F69315DD96872" box="[1057,1161,1847,1869]" captionStart="Figure 17" captionStartId="22.[163,241,1096,1118]" captionTargetBox="[324,1283,196,1065]" captionTargetId="figure-356@22.[323,1283,195,1066]" captionTargetPageId="22" captionText="Figure 17. Nerudia centaura sp. nov.; non-type female from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). A, abdomen and epigynum, ventral view. B, anterior median tube-like structure in internal genitalia (arrow). C, posterior median structures in internal genitalia; arrow: ‘receptacle’. D, E, cleared genitalia, ventral and dorsal views. Scale lines: 0.2 mm (A, D, E), 0.1 mm (B, C)." figureDoi="http://doi.org/10.5281/zenodo.7981816" httpUri="https://zenodo.org/record/7981816/files/figure.png" pageId="21" pageNumber="555">Fig. 17C</figureCitation>
) and unique tubular membranous structure in anterior position (possibly opening towards the outside, not towards the uterus externus).
</paragraph>
</subSubSection>
<caption id="FFB07784A521FFACFF1D6A4E5B7F6B88" ID-DOI="http://doi.org/10.5281/zenodo.7981816" ID-Zenodo-Dep="7981816" httpUri="https://zenodo.org/record/7981816/files/figure.png" pageId="22" pageNumber="556" startId="22.[163,241,1096,1118]" targetBox="[324,1283,196,1065]" targetPageId="22" targetType="figure">
<paragraph id="AB70270CA521FFACFF1D6A4E5B7F6B88" blockId="22.[163,1443,1096,1206]" pageId="22" pageNumber="556">
<emphasis id="99BBFB1EA521FFACFF1D6A4E58476B63" bold="true" box="[163,279,1096,1118]" pageId="22" pageNumber="556">Figure 17.</emphasis>
<taxonomicName id="6CCF5C8FA521FFACFE9C6A4E588B6B60" authority="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho, 2023" authorityName="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho" authorityYear="2023" box="[290,475,1096,1118]" class="Arachnida" family="Pholcidae" genus="Nerudia" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="556" phylum="Arthropoda" rank="species" species="centaura" status="sp. nov.">
<emphasis id="99BBFB1EA521FFACFE9C6A4E588B6B60" box="[290,475,1096,1118]" italics="true" pageId="22" pageNumber="556">Nerudia centaura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82884665A521FFACFE5E6A4E5B7E6B60" box="[480,558,1096,1118]" pageId="22" pageNumber="556" rank="species">sp. nov.</taxonomicNameLabel>
; non-type female from Argentina, Catamarca, 20 km E Paso de San Francisco (ZFMK Ar 23891). A, abdomen and epigynum, ventral view. B, anterior median tube-like structure in internal genitalia (arrow). C, posterior median structures in internal genitalia; arrow: ‘receptacle’. D, E, cleared genitalia, ventral and dorsal views. Scale lines: 0.2 mm (A, D, E), 0.1 mm (B, C).
</paragraph>
</caption>
<subSubSection id="E3D57487A521FFACFF1D6B6D5B4C6AA0" pageId="22" pageNumber="556" type="distribution">
<paragraph id="AB70270CA521FFACFF1D6B6D5B4C6AA0" blockId="22.[163,778,1387,1439]" pageId="22" pageNumber="556">
<emphasis id="99BBFB1EA521FFACFF1D6B6D58646ABE" box="[163,308,1387,1408]" italics="true" pageId="22" pageNumber="556">Distribution:</emphasis>
Known from two neighbouring localities in
<collectingCountry id="D3D8679CA521FFACFF016B8F58646AA1" box="[191,308,1417,1439]" name="Argentina" pageId="22" pageNumber="556">Argentina</collectingCountry>
,
<collectingRegion id="690BE9EEA521FFACFEFE6B8F58EE6AA0" box="[320,446,1417,1438]" country="Argentina" name="Catamarca" pageId="22" pageNumber="556">Catamarca</collectingRegion>
(
<figureCitation id="33F43B89A521FFACFE736B8F5B5C6AA0" box="[461,524,1417,1439]" captionStart="Figure 3" captionStartId="8.[164,242,1848,1870]" captionTargetBox="[324,1283,713,1818]" captionTargetId="figure-278@8.[323,1283,712,1819]" captionTargetPageId="8" captionText="Figure 3. Known distribution of Nerudia in South America (inset) and of individual species. Regions in Chile: AT, Atacama; COQ, Coquimbo. Provinces in Argentina: CA, Catamarca; COR, Córdoba; JU, Jujuy; LR, La Rioja; SA, Salta; SJ, San Juan." figureDoi="http://doi.org/10.5281/zenodo.7981770" httpUri="https://zenodo.org/record/7981770/files/figure.png" pageId="22" pageNumber="556">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E3D57487A521FFACFF1D6BCE5D7D69AD" pageId="22" pageNumber="556" type="biology_ecology">
<paragraph id="AB70270CA521FFACFF1D6BCE5DBF6A82" blockId="22.[163,780,1480,1900]" lastBlockId="22.[827,1443,1293,1683]" pageId="22" pageNumber="556">
<emphasis id="99BBFB1EA521FFACFF1D6BCE58056AE3" box="[163,341,1480,1501]" italics="true" pageId="22" pageNumber="556">Naturalhistory:</emphasis>
Bothlocalitiesweresimilar, dominated by rocks and grasses [possibly
<taxonomicName id="6CCF5C8FA521FFACFDAE6BE158AA6924" authority="(Rugolo &amp; Villav.) Govaerts" authorityName="Govaerts" baseAuthorityName="Rugolo &amp; Villav." class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="22" pageNumber="556" phylum="Tracheophyta" rank="species" species="crispa">
<emphasis id="99BBFB1EA521FFACFDAE6BE15A5A6AC2" box="[528,778,1511,1532]" italics="true" pageId="22" pageNumber="556">Calamagrostis crispa</emphasis>
(Rúgolo &amp; Villav.) Govaerts
</taxonomicName>
; Julieta Carilla, pers. comm.
<date id="DF7101CCA521FFACFF4B682258DD6907" box="[245,397,1572,1593]" pageId="22" pageNumber="556" value="2020-08">August 2020</date>
] (
<figureCitation id="33F43B89A521FFACFE1B68225B576904" box="[421,519,1572,1594]" captionStart="Figure 45" captionStartId="51.[145,226,1236,1258]" captionTargetBox="[306,1265,198,1205]" captionTargetId="figure-292@51.[305,1265,196,1206]" captionTargetPageId="51" captionText="Figure 45. Sample of typical Nerudia habitats in Argentina. A, Jujuy, between San Salvador and Purmamarca (type locality of N. colina sp. nov.). B, Salta, NW of Campo Quijano (type locality of N. poma sp. nov.). C, Catamarca, near Paso de San Francisco (type locality of N. centaura sp. nov.). D, San Juan, W of Las Flores (type locality of N. rocio sp. nov.). E, La Rioja, SE of Aimogasta (N. ola sp. nov.). F, Córdoba, near Nono (type locality of N. nono sp. nov.)." figureDoi="http://doi.org/10.5281/zenodo.7981891" httpUri="https://zenodo.org/record/7981891/files/figure.png" pageId="22" pageNumber="556">Fig. 45C</figureCitation>
). The precipitation in this area is low (~
<quantity id="6C378AE9A521FFACFED06844589C6966" box="[366,460,1602,1624]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="556" unit="mm" value="150.0">150 mm</quantity>
mean annual precipitation) and largely limited to three months per year (~70% in December–February) (https://www.meteoblue.com/). We visited the locality at the end of the ‘humid’ season, in March, and often found humid patches of soil under large rocks, with considerable numbers of small, whitish collembolans. At the lower site (
<quantity id="6C378AE9A521FFACFDD268FD5B91682E" box="[620,705,1787,1808]" metricMagnitude="3" metricUnit="m" metricValue="4.18" pageId="22" pageNumber="556" unit="m" value="4180.0">4180 m</quantity>
a.s.l.), this microhabitat seemed to contain little else than one species each of collembolans, pholcids, linyphiids and ants. At the higher site (
<quantity id="6C378AE9A521FFACFE4A69505BC16852" box="[500,657,1878,1900]" metricMagnitude="3" metricUnit="m" metricValue="4.36" metricValueMax="4.45" metricValueMin="4.27" pageId="22" pageNumber="556" unit="m" value="4360.0" valueMax="4450.0" valueMin="4270.0">4270–4450 m</quantity>
a.s.l.), the diversity seemed to be slightly higher, including, in addition, a species each of theridiids, filistatids and neopteran insects. At both localities, the pholcids were found sitting on the undersides of the rocks. They did not move and were thus easy to catch. Of the
<specimenCount id="BDC9EC85A521FFACFA396B8E5AC36A82" pageId="22" pageNumber="556" type="female">36 females</specimenCount>
seen, only one had an egg-sac.
</paragraph>
<paragraph id="AB70270CA521FFACFCED6BC35D7D69AD" blockId="22.[827,1443,1293,1683]" pageId="22" pageNumber="556">Precise climate data do not exist for these localities, but simulations of meteorological models (https://www. meteoblue.com/) suggest that temperatures fall below 0 °C almost every night of the year, with daily minima below –10 °C for seven months (April–October). In 2019, the temperatures repeatedly dropped below –20 °C in this period.</paragraph>
</subSubSection>
</treatment>
</document>
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