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<document id="D132C287C21A21058D1C3B1C0EFCF826" ID-CLB-Dataset="53409" ID-DOI="10.1093/zoolinnean/zlac100" ID-GBIF-Dataset="194d8698-7037-41e1-8bdc-11cd82e6884c" ID-ISSN="0024-4082" ID-Zenodo-Dep="7981762" ID-ZooBank="E3CF73A6-FCA6-4935-A516-D1E38E49CFB3" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1685085252051" checkinUser="plazi" docAuthor="Huber, Bernhard A., Meng, Guanliang, Král, Jiří, Ávila Herrera, Ivalú M., Izquierdo, Matias A. &amp; Carvalho, Leonardo S." docDate="2023" docId="2366961AA517FF9EFE98683C5C286A97" docLanguage="en" docName="zlac100.pdf" docOrigin="Zoological Journal of the Linnean Society 198" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-.journal_article" docStyleVersion="7" docTitle="Nerudia ola Huber, Meng, Král, Herrera, Izquierdo &amp; Carvalho, 2023, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="570" masterDocId="DF5FEE62A537FFBAFFBE6E0659506F3E" masterDocTitle="High and dry: integrative taxonomy of the Andean spider genus Nerudia (Araneae: Pholcidae)" masterLastPageNumber="591" masterPageNumber="534" pageNumber="566" updateTime="1685365769390" updateUser="ExternalLinkService">
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<mods:title id="A9E6B9B7BBE190C7135E73C8492987D8">High and dry: integrative taxonomy of the Andean spider genus Nerudia (Araneae: Pholcidae)</mods:title>
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<mods:namePart id="6A32FBFAF8C28244E42242FCFBC23EC6">Meng, Guanliang</mods:namePart>
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<mods:namePart id="F725A963B0B585C54F6246CBB9258E25">Král, Jiří</mods:namePart>
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<taxonomicName id="6CCF5C8FA517FF9AFE98683C589A696E" authorityName="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho" authorityYear="2023" box="[294,458,1594,1617]" class="Arachnida" family="Pholcidae" genus="Nerudia" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="584" phylum="Arthropoda" rank="species" species="ola">
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.
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<emphasis id="99BBFB1EA517FF9AFDEE683B5BD1696E" bold="true" box="[592,641,1597,1616]" pageId="32" pageNumber="566">NOV</emphasis>
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.
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<subSubSection id="E3D57487A517FF9AFEC868665A5D6983" pageId="32" pageNumber="566" type="description">
<paragraph id="AB70270CA517FF9AFEC868665B676947" blockId="32.[374,567,1632,1657]" box="[374,567,1632,1657]" pageId="32" pageNumber="566">
(
<figureCitation id="33F43B89A517FF9AFEC06866588C6947" box="[382,476,1632,1657]" captionStart="Figure 1" captionStartId="4.[164,244,1761,1783]" captionTargetBox="[324,1283,651,1731]" captionTargetId="figure-233@4.[323,1283,650,1731]" captionTargetPageId="4" captionText="Figure 1. Nerudia, live specimens from Argentina. A, N. colina sp. nov., female from Jujuy, between San Salvador and Purmamarca (type locality). B, N. poma sp. nov., male from Salta, NW Campo Quijano (type locality). C, D, N. centaura sp. nov., male and female from Catamarca, E Paso de San Francisco (type locality). E, N. trigo sp. nov., female from Salta, SW Alemania. F, N. guirnalda sp. nov., male from Catamarca, El Rodeo (type locality). G, N. ola sp. nov., male from San Juan, San Agustin de Valle Fertil (type locality). H, N. nono sp. nov., male from Cordoba, E Nono (type locality)." figureDoi="http://doi.org/10.5281/zenodo.7981764" httpUri="https://zenodo.org/record/7981764/files/figure.png" pageId="32" pageNumber="566">
<smallCapsWord id="AD96B1D0A517FF9AFEC0686658E46946" baselines="1651,1651" box="[382,436,1632,1657]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Figs" pageId="32" pageNumber="566">FIGS</smallCapsWord>
1G
</figureCitation>
,
<figureCitation id="33F43B89A517FF9AFE5768665B7F6947" box="[489,559,1632,1657]" captionStart-0="Figure 27" captionStart-1="Figure 28" captionStart-2="Figure 29" captionStart-3="Figure 30" captionStartId-0="32.[163,241,1320,1342]" captionStartId-1="33.[146,224,974,996]" captionStartId-2="34.[163,242,1300,1322]" captionStartId-3="35.[145,223,1378,1400]" captionTargetBox-0="[327,1275,196,1282]" captionTargetBox-1="[306,1265,197,944]" captionTargetBox-2="[323,1282,198,1270]" captionTargetBox-3="[306,1265,196,1347]" captionTargetId-0="figure-281@32.[323,1283,195,1290]" captionTargetId-1="figure-546@33.[305,1265,196,945]" captionTargetId-2="figure-325@34.[322,1282,197,1271]" captionTargetId-3="figure-250@35.[305,1265,195,1348]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionTargetPageId-2="34" captionTargetPageId-3="35" captionText-0="Figure 27. Nerudia ola sp. nov.; paratype male and paratype female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Ar 23897). AC, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views. DF, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm." captionText-1="Figure 28. Nerudia ola sp. nov.; paratype female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Ar 23897). A, abdomen and epigynum, ventral view. B, median structures in internal genitalia. C, D, cleared genitalia, ventral and dorsal views. Scale lines: 0.2 mm (A, C, D), 0.1 mm (B)." captionText-2="Figure 29. Nerudia ola sp. nov.; male and female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Arg141). A, female prosoma, frontal view. B, left procursus tip, prolateral-dorsal view. C, stridulatory file on male chelicera. D, lateral view of female chelicera, showing absence of stridulatory file. E, stridulatory pick (modified hair) on male palpal femur. F, right male cheliceral apophysis, frontal view. G, processes on male sternum. H, male gonopore. Scale lines: 100 µm (A, G), 10 µm (B, D, F, H), 20 µm (C), 2 µm (E)." captionText-3="Figure 30. Nerudia ola sp. nov.; male and female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Arg141). A, male right tibia 1, retrolateral view. B, male metatarsus 4 with trichobothrium. C, male palpal tarsal organ. D, tip of right female palp, dorsal view; arrow points at tarsal organ. E, tarsal organ on male tarsus 2. F, epigynum, ventral view. G, male ALS and PMS. H, female ALS. Scale lines: 10 µm (A, B, D, G, H), 2 µm (C, E), 100 µm (F)." figureDoi-0="http://doi.org/10.5281/zenodo.7981848" figureDoi-1="http://doi.org/10.5281/zenodo.7981852" figureDoi-2="http://doi.org/10.5281/zenodo.7981854" figureDoi-3="http://doi.org/10.5281/zenodo.7981858" httpUri-0="https://zenodo.org/record/7981848/files/figure.png" httpUri-1="https://zenodo.org/record/7981852/files/figure.png" httpUri-2="https://zenodo.org/record/7981854/files/figure.png" httpUri-3="https://zenodo.org/record/7981858/files/figure.png" pageId="32" pageNumber="566">2730</figureCitation>
)
</paragraph>
<paragraph id="AB70270CA517FF9AFF1D688F5A5D6983" blockId="32.[163,781,1673,1725]" pageId="32" pageNumber="566">
<emphasis id="99BBFB1EA517FF9AFF1D688F58BF69A0" box="[163,495,1673,1694]" italics="true" pageId="32" pageNumber="566">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act:
<uuid id="DF691DD9A517FF9AFF5168A15A586983" box="[239,776,1703,1725]" pageId="32" pageNumber="566">BF76CDC2-9B0C-4B69-8DB9-DE6925D99DD4</uuid>
.
</paragraph>
</subSubSection>
<subSubSection id="E3D57487A517FF9AFF1D68E058A76824" pageId="32" pageNumber="566" type="reference_group">
<paragraph id="AB70270CA517FF9AFF1D68E058A76824" blockId="32.[163,778,1766,1819]" pageId="32" pageNumber="566">
<taxonomicName id="6CCF5C8FA517FF9AFF1D68E0581869C2" authority="Mich" authorityName="Mich" box="[163,328,1766,1788]" class="Arachnida" family="Pholcidae" genus="Nerudia" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="566" phylum="Arthropoda" rank="genus">
<emphasis id="99BBFB1EA517FF9AFF1D68E0585369C5" box="[163,259,1766,1787]" italics="true" pageId="32" pageNumber="566">Nerudia</emphasis>
Mich
</taxonomicName>
20:
<bibRefCitation id="CF5E5AFDA517FF9AFED168E05B1469C5" author="Eberle J &amp; Dimitrov D &amp; Valdez-Mondragon A &amp; Huber BA" box="[367,580,1766,1788]" pageId="32" pageNumber="566" pagination="141" refId="ref32004" refString="Eberle J, Dimitrov D, Valdez-Mondragon A, Huber BA. 2018. Microhabitat change drives diversification in pholcid spiders. BMC EVolutionary Biology 18: 141." type="journal article" year="2018">
Eberle
<emphasis id="99BBFB1EA517FF9AFE7A68E158AE69C5" box="[452,510,1766,1788]" italics="true" pageId="32" pageNumber="566">et al.</emphasis>
, 2018
</bibRefCitation>
(molecular data: 28S).
<bibRefCitation id="CF5E5AFDA517FF9AFF5D690358FE6824" author="Huber BA &amp; Eberle J &amp; Dimitrov D" box="[227,430,1797,1819]" pageId="32" pageNumber="566" pagination="51 - 101" refId="ref32562" refString="Huber BA, Eberle J, Dimitrov D. 2018. The phylogeny of pholcid spiders: a critical evaluation of relationships suggested by molecular data (Araneae, Pholcidae). ZooKeys 789: 51 - 101." type="journal article" year="2018">
Huber
<emphasis id="99BBFB1EA517FF9AFE8C6903583A6824" box="[306,362,1797,1818]" italics="true" pageId="32" pageNumber="566">et al.</emphasis>
, 2018
</bibRefCitation>
: fig. 2.
</paragraph>
</subSubSection>
<subSubSection id="E3D57487A517FF9AFF1D69425DA569EA" pageId="32" pageNumber="566" type="diagnosis">
<paragraph id="AB70270CA517FF9AFF1D69425DA569EA" blockId="32.[163,780,1859,1912]" lastBlockId="32.[827,1443,1481,1748]" pageId="32" pageNumber="566">
<emphasis id="99BBFB1EA517FF9AFF1D6942587F6867" box="[163,303,1860,1881]" italics="true" pageId="32" pageNumber="566">D i a g n o s i s:</emphasis>
E a s i l y d i s t i n g u i s h e d f r o m k n o w n congeners by armature of male chelicerae (
<figureCitation id="33F43B89A517FF9AFD3869645A556849" box="[646,773,1890,1912]" captionStart="Figure 27" captionStartId="32.[163,241,1320,1342]" captionTargetBox="[327,1275,196,1282]" captionTargetId="figure-281@32.[323,1283,195,1290]" captionTargetPageId="32" captionText="Figure 27. Nerudia ola sp. nov.; paratype male and paratype female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Ar 23897). AC, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views. DF, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7981848" httpUri="https://zenodo.org/record/7981848/files/figure.png" pageId="32" pageNumber="566">Fig. 27G, H</figureCitation>
; distinctive pair of long frontal apophyses) and by shape of procursus (
<figureCitation id="33F43B89A517FF9AFC626BE15D0E6AC2" box="[988,1118,1511,1533]" captionStart="Figure 27" captionStartId="32.[163,241,1320,1342]" captionTargetBox="[327,1275,196,1282]" captionTargetId="figure-281@32.[323,1283,195,1290]" captionTargetPageId="32" captionText="Figure 27. Nerudia ola sp. nov.; paratype male and paratype female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Ar 23897). AC, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views. DF, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7981848" httpUri="https://zenodo.org/record/7981848/files/figure.png" pageId="32" pageNumber="566">Fig. 27AC</figureCitation>
; with bifid tip consisting of slender dorsal process and wider ventral membranous part); also by bulbal processes (
<figureCitation id="33F43B89A517FF9AFB0668235C6D6904" box="[1208,1341,1573,1595]" captionStart="Figure 27" captionStartId="32.[163,241,1320,1342]" captionTargetBox="[327,1275,196,1282]" captionTargetId="figure-281@32.[323,1283,195,1290]" captionTargetPageId="32" captionText="Figure 27. Nerudia ola sp. nov.; paratype male and paratype female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Ar 23897). AC, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views. DF, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7981848" httpUri="https://zenodo.org/record/7981848/files/figure.png" pageId="32" pageNumber="566">Fig. 27DF</figureCitation>
; ventral apophysis distally slender, curved towards ventral, same length as embolus) and by epigynum and female internal genitalia (
<figureCitation id="33F43B89A517FF9AFBAF68875D3E69A8" box="[1041,1134,1665,1687]" captionStart="Figure 27" captionStartId="32.[163,241,1320,1342]" captionTargetBox="[327,1275,196,1282]" captionTargetId="figure-281@32.[323,1283,195,1290]" captionTargetPageId="32" captionText="Figure 27. Nerudia ola sp. nov.; paratype male and paratype female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Ar 23897). AC, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views. DF, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7981848" httpUri="https://zenodo.org/record/7981848/files/figure.png" pageId="32" pageNumber="566">Figs 27I</figureCitation>
,
<figureCitation id="33F43B89A517FF9AFBC468875DC569A8" box="[1146,1173,1665,1686]" captionStart="Figure 28" captionStartId="33.[146,224,974,996]" captionTargetBox="[306,1265,197,944]" captionTargetId="figure-546@33.[305,1265,196,945]" captionTargetPageId="33" captionText="Figure 28. Nerudia ola sp. nov.; paratype female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Ar 23897). A, abdomen and epigynum, ventral view. B, median structures in internal genitalia. C, D, cleared genitalia, ventral and dorsal views. Scale lines: 0.2 mm (A, C, D), 0.1 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.7981852" httpUri="https://zenodo.org/record/7981852/files/figure.png" pageId="32" pageNumber="566">28</figureCitation>
; epigynal plate without posterior indentation; internal genitalia simple, with barely visible transparent receptacle).
</paragraph>
</subSubSection>
<subSubSection id="E3D57487A517FF99FC85690E5C7169CC" lastPageId="35" lastPageNumber="569" pageId="32" pageNumber="566" type="materials_examined">
<paragraph id="AB70270CA517FF9BFC85690E58006B96" blockId="32.[827,1443,1798,1913]" lastBlockId="33.[145,760,1137,1192]" lastPageId="33" lastPageNumber="567" pageId="32" pageNumber="566">
<emphasis id="99BBFB1EA517FF9AFC85690E5AA86822" box="[827,1016,1799,1821]" italics="true" pageId="32" pageNumber="566">Type material:</emphasis>
<materialsCitation id="1BA72D51A517FF9AFBB969015CF26847" collectingDate="2019-03-05" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="880" latitude="-30.6366" location="San Agustin" longLatPrecision="6" longitude="-67.4863" pageId="32" pageNumber="566" specimenCount="1" stateProvince="San Juan" typeStatus="holotype">
<collectingCountry id="D3D8679CA517FF9AFBB969015DE26822" box="[1031,1202,1799,1820]" name="Argentina" pageId="32" pageNumber="566">ARGENTINA</collectingCountry>
<emphasis id="99BBFB1EA517FF9AFB7E690E5D9C6823" box="[1216,1228,1800,1821]" italics="true" pageId="32" pageNumber="566"></emphasis>
<emphasis id="99BBFB1EA517FF9AFB6769015C316822" bold="true" box="[1241,1377,1799,1821]" pageId="32" pageNumber="566">
<collectingRegion id="690BE9EEA517FF9AFB6769015C316822" box="[1241,1377,1799,1821]" pageId="32" pageNumber="566">San Juan</collectingRegion>
</emphasis>
: • ♂
<typeStatus id="747499AEA517FF9AFC8569205AF56802" box="[827,933,1830,1852]" pageId="32" pageNumber="566">holotype</typeStatus>
;
<location id="AE1071D7A517FF9AFC0A69205D1C6802" LSID="urn:lsid:plazi:treatment:2366961AA517FF9EFE98683C5C286A97:AE1071D7A517FF9AFC0A69205D1C6802" box="[948,1100,1830,1852]" country="Argentina" latitude="-30.6366" longLatPrecision="6" longitude="-67.4863" name="San Agustin" pageId="32" pageNumber="566" stateProvince="San Juan">San Agustín</location>
de
<taxonomicName id="6CCF5C8FA517FF9AFBC569205C5A6802" authority="Fertil" authorityName="Fertil" box="[1147,1290,1830,1852]" class="Arachnida" family="Pholcidae" genus="Artema" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="566" phylum="Arthropoda" rank="species" species="valle">Valle Fértil</taxonomicName>
;
<geoCoordinate id="CEFB41CBA517FF9AFAA769205CCD6805" box="[1305,1437,1830,1852]" degrees="30.6366" direction="south" orientation="latitude" pageId="32" pageNumber="566" precision="5" value="-30.6366">30.6366° S</geoCoordinate>
,
<geoCoordinate id="CEFB41CBA517FF9AFC8569435AEB6864" box="[827,955,1861,1883]" degrees="67.4863" direction="west" orientation="longitude" pageId="32" pageNumber="566" precision="5" value="-67.4863">67.4863° W</geoCoordinate>
;
<elevation id="20E2C03FA517FF9AFC7B69435D166865" box="[965,1094,1861,1883]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="32" pageNumber="566" unit="m" value="880.0">
<quantity id="6C378AE9A517FF9AFC7B69435D5A6864" box="[965,1034,1861,1882]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="32" pageNumber="566" unit="m" value="880.0">880 m</quantity>
a.s.l.
</elevation>
; under rocks near river;
<date id="DF7101CCA517FF9AFAE569425A246846" pageId="32" pageNumber="566" value="2019-03-05">
<collectingDate id="CF35F824A517FF9AFAE569425A246846" pageId="32" pageNumber="566" value="2019-03-05">5 Mar. 2019</collectingDate>
</date>
;
<collectorName id="063A42DAA517FF9AFCC169625D536847" box="[895,1027,1891,1913]" pageId="32" pageNumber="566">B. A. Huber</collectorName>
and
<collectorName id="063A42DAA517FF9AFB8269625DBB6847" box="[1084,1259,1891,1913]" pageId="32" pageNumber="566">M. A. Izquierdo</collectorName>
leg.;
<emphasis id="99BBFB1EA517FF9AFA9869625CF26847" box="[1318,1442,1892,1913]" italics="true" pageId="32" pageNumber="566">LABRE-Ar</emphasis>
</materialsCitation>
<emphasis id="99BBFB1EA516FF9BFF2F6A7259EB6BB7" box="[145,187,1140,1161]" italics="true" pageId="33" pageNumber="567">591</emphasis>
<materialsCitation id="1BA72D51A516FF9BFF7D6A73581C6B96" collectingDate="2019-03-05" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="880" latitude="-30.6366" location="San Agustin" longLatPrecision="6" longitude="-67.4863" pageId="33" pageNumber="567" specimenCount="2" specimenCount-female="2" stateProvince="San Juan" typeStatus="paratype">
• 5 ♂♂,
<specimenCount id="BDC9EC85A516FF9BFE9F6A7258066BB7" box="[289,342,1137,1161]" pageId="33" pageNumber="567" type="female">2 ♀♀</specimenCount>
,
<typeStatus id="747499AEA516FF9BFEDB6A73588C6BB4" box="[357,476,1141,1162]" pageId="33" pageNumber="567">paratypes</typeStatus>
; same data as holotype; ZFMK Ar 23897
</materialsCitation>
.
</paragraph>
<caption id="FFB07784A516FF9BFF2C6DC85B9C6B21" ID-DOI="http://doi.org/10.5281/zenodo.7981852" ID-Zenodo-Dep="7981852" httpUri="https://zenodo.org/record/7981852/files/figure.png" pageId="33" pageNumber="567" startId="33.[146,224,974,996]" targetBox="[306,1265,197,944]" targetPageId="33" targetType="figure">
<paragraph id="AB70270CA516FF9BFF2C6DC85B9C6B21" blockId="33.[145,1426,974,1055]" pageId="33" pageNumber="567">
<emphasis id="99BBFB1EA516FF9BFF2C6DC858576CDD" bold="true" box="[146,263,974,996]" pageId="33" pageNumber="567">Figure 28.</emphasis>
<taxonomicName id="6CCF5C8FA516FF9BFEAF6DC858DE6CDA" authority="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho, 2023" authorityName="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho" authorityYear="2023" box="[273,398,974,996]" class="Arachnida" family="Pholcidae" genus="Nerudia" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="567" phylum="Arthropoda" rank="species" species="ola" status="sp. nov.">
<emphasis id="99BBFB1EA516FF9BFEAF6DC858DE6CDA" box="[273,398,974,996]" italics="true" pageId="33" pageNumber="567">Nerudia ola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82884665A516FF9BFE2D6DC858B16CDA" box="[403,481,974,996]" pageId="33" pageNumber="567" rank="species">sp. nov.</taxonomicNameLabel>
; paratype female from Argentina, San Juan, San Agustín de
<taxonomicName id="6CCF5C8FA516FF9BFBE76DC85D9D6CDA" authority="Fertil" authorityName="Fertil" box="[1113,1229,974,996]" class="Arachnida" family="Pholcidae" genus="Artema" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="567" phylum="Arthropoda" rank="species" species="valle">Valle Fértil</taxonomicName>
(ZFMK Ar 23897). A, abdomen and epigynum, ventral view. B, median structures in internal genitalia. C, D, cleared genitalia, ventral and dorsal views. Scale lines: 0.2 mm (A, C, D), 0.1 mm (B).
</paragraph>
</caption>
<paragraph id="AB70270CA516FF9BFF2F6AD45C596A5E" blockId="33.[145,762,1233,1899]" lastBlockId="33.[809,1426,1140,1377]" pageId="33" pageNumber="567">
<materialsCitation id="1BA72D51A516FF9BFF2F6AD458686A5F" collectingDate="2012-01-22" collectingDateMax="2019-03-05" collectingDateMin="2012-01-22" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="865" latitude="-30.2232" location="Mich" longLatPrecision="6" longitude="-67.6942" municipality="Arg" pageId="33" pageNumber="567" specimenCount="40" specimenCount-female="38" specimenCount-male="1" stateProvince="San Juan">
<emphasis id="99BBFB1EA516FF9BFF2F6AD458E66BD9" box="[145,438,1234,1255]" italics="true" pageId="33" pageNumber="567">
<typeStatus id="747499AEA516FF9BFF2F6AD458E06BD9" box="[145,432,1234,1255]" isOtherTypeStatus="true" pageId="33" pageNumber="567">Other material examined</typeStatus>
:
</emphasis>
<collectingCountry id="D3D8679CA516FF9BFE7C6AD45B306BD9" box="[450,608,1234,1255]" name="Argentina" pageId="33" pageNumber="567">ARGENTINA</collectingCountry>
<emphasis id="99BBFB1EA516FF9BFDD86AD45B226BD9" box="[614,626,1234,1255]" italics="true" pageId="33" pageNumber="567"></emphasis>
<emphasis id="99BBFB1EA516FF9BFDC46AD75BA36BD9" bold="true" box="[634,755,1233,1255]" pageId="33" pageNumber="567">
<collectingRegion id="690BE9EEA516FF9BFDC46AD75BA36BD9" box="[634,755,1233,1255]" pageId="33" pageNumber="567">San Juan</collectingRegion>
</emphasis>
: • 12 ♂♂,
<specimenCount id="BDC9EC85A516FF9BFF476AF6587D6A3B" box="[249,301,1261,1285]" pageId="33" pageNumber="567" type="female">8 ♀♀</specimenCount>
, 5 juvs, in pure ethanol (
<specimenCount id="BDC9EC85A516FF9BFDF16AF65B956A38" box="[591,709,1264,1286]" pageId="33" pageNumber="567" type="male">two males</specimenCount>
and
<specimenCount id="BDC9EC85A516FF9BFF2F6B09584F6A1B" box="[145,287,1295,1317]" pageId="33" pageNumber="567" type="female">two females</specimenCount>
used for SEM;
<specimenCount id="BDC9EC85A516FF9BFE626B095B226A1B" box="[476,626,1295,1317]" pageId="33" pageNumber="567" type="female">three female</specimenCount>
prosomata used for molecular study); same data as holotype; ZFMK
<collectingMunicipality id="4B14BD76A516FF9BFF5D6B4A58416A5F" box="[227,273,1356,1377]" pageId="33" pageNumber="567">Arg</collectingMunicipality>
141
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFEFE6B4B58136ABF" collectingDate="2012-01-22" collectingDateMax="2019-03-05" collectingDateMin="2012-01-22" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="865" latitude="-30.2232" location="Mich" longLatPrecision="6" longitude="-67.6942" municipality="Arg" pageId="33" pageNumber="567" specimenCount="40" specimenCount-female="38" specimenCount-male="1" stateProvince="San Juan">
<specimenCount id="BDC9EC85A516FF9BFEE96B4A58DA6A5F" box="[343,394,1353,1377]" pageId="33" pageNumber="567" type="female">9 ♀♀</specimenCount>
, 3 juvs; same data as
<typeStatus id="747499AEA516FF9BFD316B4A5BA46A5C" box="[655,756,1356,1378]" pageId="33" pageNumber="567">holotype</typeStatus>
; LABRE-Ar 560
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFEF56B6D59B86A80" collectingDate="2012-01-22" collectingDateMax="2019-03-05" collectingDateMin="2012-01-22" collectorName="J. M. A. Navarro" country="Argentina" elevation="865" latitude="-30.2232" location="Mich" longLatPrecision="6" longitude="-67.6942" municipality="Arg" pageId="33" pageNumber="567" specimenCount="40" specimenCount-female="38" specimenCount-male="1" stateProvince="San Juan">
• 1 ♂, in pure ethanol; same locality as
<typeStatus id="747499AEA516FF9BFF0F6B8C58446A9E" box="[177,276,1418,1440]" pageId="33" pageNumber="567">holotype</typeStatus>
;
<date id="DF7101CCA516FF9BFEA06B8C58E06AA1" box="[286,432,1418,1439]" pageId="33" pageNumber="567" value="2012-01-22">
<collectingDate id="CF35F824A516FF9BFEA06B8C58E06AA1" box="[286,432,1418,1439]" pageId="33" pageNumber="567" value="2012-01-22">22 Jan. 2012</collectingDate>
</date>
;
<collectorName id="063A42DAA516FF9BFE056B8C5B246AA1" box="[443,628,1418,1439]" pageId="33" pageNumber="567">J. M. A. Navarro</collectorName>
leg.; ZFMK
<location id="AE1071D7A516FF9BFF2F6BAE599D6A80" LSID="urn:lsid:plazi:treatment:2366961AA517FF9EFE98683C5C286A97:AE1071D7A516FF9BFF2F6BAE599D6A80" box="[145,205,1448,1470]" country="Argentina" latitude="-30.2232" longLatPrecision="6" longitude="-67.6942" municipality="Arg" name="Mich" pageId="33" pageNumber="567" stateProvince="San Juan">Mich</location>
20
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFF4F6BAF59866927" collectingDate="2012-01-22" collectingDateMax="2019-03-05" collectingDateMin="2012-01-22" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="865" latitude="-30.2232" location="Mich" longLatPrecision="6" longitude="-67.6942" municipality="Arg" pageId="33" pageNumber="567" specimenCount="40" specimenCount-female="38" specimenCount-male="1" stateProvince="San Juan">
• 3 ♂♂,
<specimenCount id="BDC9EC85A516FF9BFEEF6BAE58D66A83" box="[337,390,1445,1469]" pageId="33" pageNumber="567" type="female">2 ♀♀</specimenCount>
; ~
<locationDeviation id="BAC23E2EA516FF9BFE1B6BAE5B366A80" box="[421,614,1448,1470]" location="Astica" pageId="33" pageNumber="567">
<quantity id="6C378AE9A516FF9BFE1B6BAE58A86A80" box="[421,504,1448,1470]" metricMagnitude="3" metricUnit="m" metricValue="7.5" pageId="33" pageNumber="567" unit="km" value="7.5">7.5 km</quantity>
S Astica
</locationDeviation>
;
<geoCoordinate id="CEFB41CBA516FF9BFDCD6BAE5BA46A83" box="[627,756,1448,1469]" degrees="31.0223" direction="south" orientation="latitude" pageId="33" pageNumber="567" precision="5" value="-31.0223">31.0223° S</geoCoordinate>
,
<geoCoordinate id="CEFB41CBA516FF9BFF2F6BC158416AE2" box="[145,273,1479,1501]" degrees="67.2976" direction="west" orientation="longitude" pageId="33" pageNumber="567" precision="5" value="-67.2976">67.2976° W</geoCoordinate>
;
<elevation id="20E2C03FA516FF9BFEA56BC158CC6AE3" box="[283,412,1479,1501]" metricMagnitude="2" metricUnit="m" metricValue="8.65" pageId="33" pageNumber="567" unit="m" value="865.0">
<quantity id="6C378AE9A516FF9BFEA56BC158306AE2" box="[283,352,1479,1501]" metricMagnitude="2" metricUnit="m" metricValue="8.65" pageId="33" pageNumber="567" unit="m" value="865.0">865 m</quantity>
a.s.l.
</elevation>
; under rocks near river;
<date id="DF7101CCA516FF9BFD0F6BC1599B6AC5" pageId="33" pageNumber="567" value="2019-03-04">
<collectingDate id="CF35F824A516FF9BFD0F6BC1599B6AC5" pageId="33" pageNumber="567" value="2019-03-04">4 Mar. 2019</collectingDate>
</date>
;
<collectorName id="063A42DAA516FF9BFF686BE0580E6AC5" box="[214,350,1509,1531]" pageId="33" pageNumber="567">B. A. Huber</collectorName>
and
<collectorName id="063A42DAA516FF9BFE276BE05B1B6AC5" box="[409,587,1509,1531]" pageId="33" pageNumber="567">M. A. Izquierdo</collectorName>
leg.; ZFMK Ar 23898
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFF65680358B56907" collectingDate="2012-01-22" collectingDateMax="2019-03-05" collectingDateMin="2012-01-22" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="865" latitude="-30.2232" location="Mich" longLatPrecision="6" longitude="-67.6942" municipality="Arg" pageId="33" pageNumber="567" specimenCount="40" specimenCount-female="38" specimenCount-male="1" stateProvince="San Juan">
• 1 ♂,
<specimenCount id="BDC9EC85A516FF9BFE9E680258026927" box="[288,338,1537,1561]" pageId="33" pageNumber="567" type="female">3 ♀♀</specimenCount>
, 5 juvs, in pure ethanol; same data as preceding; ZFMK Arg135, 136
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFE52682558EB6969" collectingDate="2012-01-22" collectingDateMax="2019-03-05" collectingDateMin="2012-01-22" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="865" latitude="-30.2232" location="Mich" longLatPrecision="6" longitude="-67.6942" municipality="Arg" pageId="33" pageNumber="567" specimenCount="40" specimenCount-female="38" specimenCount-male="1" stateProvince="San Juan">
• 1 ♂,
<specimenCount id="BDC9EC85A516FF9BFD8D68255B076906" box="[563,599,1568,1592]" pageId="33" pageNumber="567" type="female">1 ♀</specimenCount>
; same data as preceding; LABRE-Ar 528
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFE7C684459A369AB" collectingDate="2012-01-22" collectingDateMax="2019-03-05" collectingDateMin="2012-01-22" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="865" latitude="-30.2232" location="Mich" longLatPrecision="6" longitude="-67.6942" municipality="Arg" pageId="33" pageNumber="567" specimenCount="40" specimenCount-female="38" specimenCount-male="1" stateProvince="San Juan">
• 5 ♂♂,
<specimenCount id="BDC9EC85A516FF9BFDA568445B1E6969" box="[539,590,1599,1623]" pageId="33" pageNumber="567" type="female">7 ♀♀</specimenCount>
,
<specimenCount id="BDC9EC85A516FF9BFDE268445BCB6969" box="[604,667,1601,1623]" pageId="33" pageNumber="567" type="generic">1 juv.</specimenCount>
, in pure ethanol; same data as preceding; LABRE-Ar 529, 547, 548, 552
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFF456879580169CE" collectingDate="2012-01-22" collectingDateMax="2019-03-05" collectingDateMin="2012-01-22" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="865" latitude="-30.2232" location="Mich" longLatPrecision="6" longitude="-67.6942" municipality="Arg" pageId="33" pageNumber="567" specimenCount="40" specimenCount-female="38" specimenCount-male="1" stateProvince="San Juan">
• 1 ♂,
<specimenCount id="BDC9EC85A516FF9BFEF96879583C69AA" box="[327,364,1660,1684]" pageId="33" pageNumber="567" type="female">1 ♀</specimenCount>
; Parque Provincial Ischigualasto;
<geoCoordinate id="CEFB41CBA516FF9BFF2F68985842698D" box="[145,274,1694,1715]" degrees="30.1839" direction="south" orientation="latitude" pageId="33" pageNumber="567" precision="5" value="-30.1839">30.1839° S</geoCoordinate>
,
<geoCoordinate id="CEFB41CBA516FF9BFE9F689B58FB698D" box="[289,427,1693,1715]" degrees="67.9026" direction="west" orientation="longitude" pageId="33" pageNumber="567" precision="5" value="-67.9026">67.9026° W</geoCoordinate>
;
<elevation id="20E2C03FA516FF9BFE06689B5B03698D" box="[440,595,1693,1715]" metricMagnitude="3" metricUnit="m" metricValue="1.425" pageId="33" pageNumber="567" unit="m" value="1425.0">
<quantity id="6C378AE9A516FF9BFE06689B5B41698D" box="[440,529,1693,1715]" metricMagnitude="3" metricUnit="m" metricValue="1.425" pageId="33" pageNumber="567" unit="m" value="1425.0">1425 m</quantity>
a.s.l.
</elevation>
; under rocks;
<date id="DF7101CCA516FF9BFF2F68BA587469EF" box="[145,292,1724,1746]" pageId="33" pageNumber="567" value="2019-03-05">
<collectingDate id="CF35F824A516FF9BFF2F68BA587469EF" box="[145,292,1724,1746]" pageId="33" pageNumber="567" value="2019-03-05">5 Mar. 2019</collectingDate>
</date>
;
<collectorName id="063A42DAA516FF9BFE8C68BB589169EC" box="[306,449,1724,1746]" pageId="33" pageNumber="567">B. A. Huber</collectorName>
and
<collectorName id="063A42DAA516FF9BFDBC68BB5BEE69EC" box="[514,702,1724,1746]" pageId="33" pageNumber="567">M. A. Izquierdo</collectorName>
leg.; ZFMK Ar 23899
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFEE468DD58896831" collectingDate="2012-01-22" collectingDateMax="2019-03-05" collectingDateMin="2012-01-22" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="865" latitude="-30.2232" location="Mich" longLatPrecision="6" longitude="-67.6942" municipality="Arg" pageId="33" pageNumber="567" specimenCount="40" specimenCount-female="38" specimenCount-male="1" stateProvince="San Juan">
<specimenCount id="BDC9EC85A516FF9BFECC68DD58F669CE" box="[370,422,1752,1776]" pageId="33" pageNumber="567" type="female">2 ♀♀</specimenCount>
, in pure ethanol; same data as preceding; ZFMK Arg144
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFE5C68FC58B06810" collectingDate="2012-01-22" collectingDateMax="2019-03-05" collectingDateMin="2012-01-22" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="865" latitude="-30.2232" location="Mich" longLatPrecision="6" longitude="-67.6942" municipality="Arg" pageId="33" pageNumber="567" specimenCount="40" specimenCount-female="38" specimenCount-male="1" stateProvince="San Juan">
<specimenCount id="BDC9EC85A516FF9BFE4768FC5B4E6831" box="[505,542,1783,1807]" pageId="33" pageNumber="567" type="female">1 ♀</specimenCount>
, 2 juvs; same data as preceding; LABRE-Ar 531
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFE59691F5BE36873" collectingDate="2012-01-22" collectingDateMax="2019-03-05" collectingDateMin="2012-01-22" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="865" latitude="-30.2232" location="Mich" longLatPrecision="6" longitude="-67.6942" municipality="Arg" pageId="33" pageNumber="567" specimenCount="40" specimenCount-female="38" specimenCount-male="1" stateProvince="San Juan">
<specimenCount id="BDC9EC85A516FF9BFE43691E5B726813" box="[509,546,1813,1837]" pageId="33" pageNumber="567" type="female">1 ♀</specimenCount>
with eggs, in pure ethanol; same data as preceding; LABRE-Ar 554
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFD0469315DE56BF9" collectingDate="2012-01-22" collectingDateMax="2019-03-05" collectingDateMin="2012-01-22" collectorName="M. Izquierdo; F. Bollatti; A. Albin &amp; C. Mattoni" country="Argentina" elevation="865" latitude="-30.2232" location="Mich" longLatPrecision="6" longitude="-67.6942" municipality="Arg" pageId="33" pageNumber="567" specimenCount="40" specimenCount-female="38" specimenCount-male="1" stateProvince="San Juan">
• 1 ♂; Parque Provincial Ischigualasto;
<geoCoordinate id="CEFB41CBA516FF9BFDB069505BD96855" box="[526,649,1878,1899]" degrees="30.1821" direction="south" orientation="latitude" pageId="33" pageNumber="567" precision="5" value="-30.1821">30.1821° S</geoCoordinate>
,
<geoCoordinate id="CEFB41CBA516FF9BFD2B69535A126BB4" box="[661,834,1141,1899]" degrees="67.9010" direction="west" orientation="longitude" pageId="33" pageNumber="567" precision="5" value="-67.901">67.9010° W</geoCoordinate>
;
<elevation id="20E2C03FA516FF9BFCF06A725AB76BB4" box="[846,999,1140,1162]" metricMagnitude="3" metricUnit="m" metricValue="1.425" pageId="33" pageNumber="567" unit="m" value="1425.0">
<quantity id="6C378AE9A516FF9BFCF06A725AF66BB4" box="[846,934,1140,1162]" metricMagnitude="3" metricUnit="m" metricValue="1.425" pageId="33" pageNumber="567" unit="m" value="1425.0">1425 m</quantity>
a.s.l.
</elevation>
; in dry riverbed, hand collecting at night;
<date id="DF7101CCA516FF9BFCCD6A955D576B96" box="[883,1031,1171,1192]" pageId="33" pageNumber="567" value="2018-12-19">
<collectingDate id="CF35F824A516FF9BFCCD6A955D576B96" box="[883,1031,1171,1192]" pageId="33" pageNumber="567" value="2018-12-19">19 Dec. 2018</collectingDate>
</date>
;
<collectorName id="063A42DAA516FF9BFBAC6A955DF56B97" box="[1042,1189,1171,1193]" pageId="33" pageNumber="567">M. Izquierdo</collectorName>
,
<collectorName id="063A42DAA516FF9BFB0E6A955C746B97" box="[1200,1316,1171,1193]" pageId="33" pageNumber="567">F. Bollatti</collectorName>
,
<collectorName id="063A42DAA516FF9BFA906A955CDD6B97" box="[1326,1421,1171,1193]" pageId="33" pageNumber="567">A. Albin</collectorName>
, and
<collectorName id="063A42DAA516FF9BFCE76AB45A816BF9" box="[857,977,1201,1223]" pageId="33" pageNumber="567">C. Mattoni</collectorName>
leg.; LABRE-Ar 445
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFB046AB45C1A6BDB" collectingDate="2012-01-22" collectingDateMax="2019-03-05" collectingDateMin="2012-01-22" collectorName="M. Izquierdo; F. Bollatti; A. Albin &amp; C. Mattoni" country="Argentina" elevation="865" latitude="-30.2232" location="Mich" longLatPrecision="6" longitude="-67.6942" municipality="Arg" pageId="33" pageNumber="567" specimenCount="40" specimenCount-female="38" specimenCount-male="1" stateProvince="San Juan">• 1 ♂; same data as preceding; LABRE-Ar 444, prep. code MAI-4709</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFAEF6AD75C556A5E" collectingDate="2012-01-22" collectingDateMax="2019-03-05" collectingDateMin="2012-01-22" collectorName="M. Izquierdo; F. Bollatti; A. Albin &amp; C. Mattoni" country="Argentina" elevation="865" latitude="-30.2232" location="Mich" longLatPrecision="6" longitude="-67.6942" municipality="Arg" pageId="33" pageNumber="567" specimenCount="40" specimenCount-female="38" specimenCount-male="1" stateProvince="San Juan">
<specimenCount id="BDC9EC85A516FF9BFAD96AD65CDB6BDB" box="[1383,1419,1229,1253]" pageId="33" pageNumber="567" type="female">1 ♀</specimenCount>
;
<collectorName id="063A42DAA516FF9BFC976AE95AF16A3B" box="[809,929,1263,1285]" pageId="33" pageNumber="567">Baldecitos</collectorName>
;
<geoCoordinate id="CEFB41CBA516FF9BFC156AE95D746A3A" box="[939,1060,1263,1284]" degrees="30.2232" direction="south" orientation="latitude" pageId="33" pageNumber="567" precision="5" value="-30.2232">30.2232° S</geoCoordinate>
,
<geoCoordinate id="CEFB41CBA516FF9BFB916AE95DFF6A3A" box="[1071,1199,1263,1285]" degrees="67.6942" direction="west" orientation="longitude" pageId="33" pageNumber="567" precision="5" value="-67.6942">67.6942° W</geoCoordinate>
;
<elevation id="20E2C03FA516FF9BFB076AE95C186A3B" box="[1209,1352,1263,1285]" metricMagnitude="3" metricUnit="m" metricValue="1.255" pageId="33" pageNumber="567" unit="m" value="1255.0">
<quantity id="6C378AE9A516FF9BFB076AE95C5D6A3A" box="[1209,1293,1263,1285]" metricMagnitude="3" metricUnit="m" metricValue="1.255" pageId="33" pageNumber="567" unit="m" value="1255.0">1255 m</quantity>
a.s.l.
</elevation>
; stone wall of buildings, hand collecting at night;
<date id="DF7101CCA516FF9BFA876B085A326A7F" pageId="33" pageNumber="567" value="2018-12-21">
<collectingDate id="CF35F824A516FF9BFA876B085A326A7F" pageId="33" pageNumber="567" value="2018-12-21">21 Dec. 2018</collectingDate>
</date>
;
<collectorName id="063A42DAA516FF9BFCD56B2B5AAD6A7C" box="[875,1021,1324,1346]" pageId="33" pageNumber="567">M. Izquierdo</collectorName>
,
<collectorName id="063A42DAA516FF9BFBB86B2B5D286A7C" box="[1030,1144,1324,1346]" pageId="33" pageNumber="567">F. Bollatti</collectorName>
,
<collectorName id="063A42DAA516FF9BFB3F6B2A5D8E6A7C" box="[1153,1246,1324,1346]" pageId="33" pageNumber="567">A. Albin</collectorName>
, and
<collectorName id="063A42DAA516FF9BFAA76B2A5CC06A7C" box="[1305,1424,1324,1346]" pageId="33" pageNumber="567">C. Mattoni</collectorName>
leg.; LABRE-Ar 448, prep. code MAI-4710
</materialsCitation>
.
</paragraph>
<paragraph id="AB70270CA516FF99FC976B8F59BB698A" blockId="33.[809,1426,1417,1899]" lastBlockId="35.[145,762,1542,1716]" lastPageId="35" lastPageNumber="569" pageId="33" pageNumber="567">
<emphasis id="99BBFB1EA516FF9BFC976B8F5C336AA0" box="[809,1379,1417,1439]" italics="true" pageId="33" pageNumber="567">Assigned tentatiƲely (see Male Ʋariation beloae):</emphasis>
<materialsCitation id="1BA72D51A516FF9BFAD16B8C5CC16AC5" collectingDate="2019-03-08" collectorName="Cuesta" elevation="1960" latitude="-29.3511" location="La Rioja" longLatPrecision="6" longitude="-67.7924" pageId="33" pageNumber="567" specimenCount="1" specimenCount-female="1" stateProvince="La Rioja">
<emphasis id="99BBFB1EA516FF9BFAD16B8C5A3C6A80" bold="true" pageId="33" pageNumber="567">
<collectingRegion id="690BE9EEA516FF9BFAD16B8C5A3C6A80" country="Argentina" name="La Rioja" pageId="33" pageNumber="567">La Rioja</collectingRegion>
</emphasis>
: • 1 ♂,
<specimenCount id="BDC9EC85A516FF9BFC076BAE5A8B6A83" box="[953,987,1445,1469]" pageId="33" pageNumber="567" type="female">1 ♀</specimenCount>
; Cuesta de Miranda, site 1;
<geoCoordinate id="CEFB41CBA516FF9BFAAB6BAE5CDC6A83" box="[1301,1420,1448,1470]" degrees="29.3511" direction="south" orientation="latitude" pageId="33" pageNumber="567" precision="5" value="-29.3511">29.3511° S</geoCoordinate>
,
<geoCoordinate id="CEFB41CBA516FF9BFC976BC15AE06AE2" box="[809,944,1479,1501]" degrees="67.7924" direction="west" orientation="longitude" pageId="33" pageNumber="567" precision="5" value="-67.7924">67.7924° W</geoCoordinate>
);
<elevation id="20E2C03FA516FF9BFC7A6BC15D0A6AE3" box="[964,1114,1479,1501]" metricMagnitude="3" metricUnit="m" metricValue="1.96" pageId="33" pageNumber="567" unit="m" value="1960.0">
<quantity id="6C378AE9A516FF9BFC7A6BC15D4A6AE2" box="[964,1050,1479,1501]" metricMagnitude="3" metricUnit="m" metricValue="1.96" pageId="33" pageNumber="567" unit="m" value="1960.0">1960 m</quantity>
a.s.l.
</elevation>
; under rocks;
<date id="DF7101CCA516FF9BFABE6BC15CDD6AE2" box="[1280,1421,1479,1500]" pageId="33" pageNumber="567" value="2019-03-08">
<collectingDate id="CF35F824A516FF9BFABE6BC15CDD6AE2" box="[1280,1421,1479,1500]" pageId="33" pageNumber="567" value="2019-03-08">8 Mar. 2019</collectingDate>
</date>
;
<collectorName id="063A42DAA516FF9BFC976BE05AFF6AC5" box="[809,943,1509,1531]" pageId="33" pageNumber="567">B. A. Huber</collectorName>
and
<collectorName id="063A42DAA516FF9BFC576BE05DC96AC5" box="[1001,1177,1509,1531]" pageId="33" pageNumber="567">M. A. Izquierdo</collectorName>
leg.; ZFMK
<collectorName id="063A42DAA516FF9BFA996BE05C146AC5" box="[1319,1348,1510,1531]" pageId="33" pageNumber="567">Ar</collectorName>
23900
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFC9768035A2B6907" collectingDate="2019-03-08" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="1960" latitude="-29.3511" location="La Rioja" longLatPrecision="6" longitude="-67.7924" pageId="33" pageNumber="567" specimenCount="3" specimenCount-female="3" stateProvince="Miranda">
<specimenCount id="BDC9EC85A516FF9BFC8068025A206927" box="[830,880,1537,1561]" pageId="33" pageNumber="567" type="female">2 ♀♀</specimenCount>
, in pure ethanol; same data as preceding; ZFMK
<collectorName id="063A42DAA516FF9BFC9768255A046907" box="[809,852,1571,1593]" pageId="33" pageNumber="567">Arg</collectorName>
152
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFC3E68255CC1694B" collectingDate="2019-03-08" collectingDateMax="2019-03-10" collectingDateMin="2019-03-08" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="1960" lastPageId="33" lastPageNumber="568" latitude="-29.3511" location="La Rioja" longLatPrecision="6" longitude="-67.7924" pageId="33" pageNumber="567" specimenCount="3" specimenCount-female="3" stateProvince="Miranda">
• 2 ♂♂,
<specimenCount id="BDC9EC85A516FF9BFC6C68255AA46906" box="[978,1012,1568,1592]" pageId="33" pageNumber="567" type="female">1 ♀</specimenCount>
;
<collectorName id="063A42DAA516FF9BFC4168255D1E6906" box="[1023,1102,1571,1592]" pageId="33" pageNumber="567">Cuesta</collectorName>
de
<collectingRegion id="690BE9EEA516FF9BFBCD68255D876907" box="[1139,1239,1571,1593]" country="Venezuela" name="Miranda" pageId="33" pageNumber="567">Miranda</collectingRegion>
, site 2;
<geoCoordinate id="CEFB41CBA516FF9BFA9168255A696969" degrees="29.3468" direction="south" orientation="latitude" pageId="33" pageNumber="567" precision="5" value="-29.3468">29.3468° S</geoCoordinate>
,
<geoCoordinate id="CEFB41CBA516FF9BFCFB68475A966969" box="[837,966,1601,1623]" degrees="67.7205" direction="west" orientation="longitude" pageId="33" pageNumber="567" precision="5" value="-67.7205">67.7205° W</geoCoordinate>
;
<elevation id="20E2C03FA516FF9BFC6F68475D336969" box="[977,1123,1601,1623]" metricMagnitude="3" metricUnit="m" metricValue="1.66" pageId="33" pageNumber="567" unit="m" value="1660.0">
<quantity id="6C378AE9A516FF9BFC6F68475D766969" box="[977,1062,1601,1623]" metricMagnitude="3" metricUnit="m" metricValue="1.66" pageId="33" pageNumber="567" unit="m" value="1660.0">1660 m</quantity>
a.s.l.
</elevation>
; under rocks;
<date id="DF7101CCA516FF9BFABA68445CDD6969" box="[1284,1421,1602,1623]" pageId="33" pageNumber="567" value="2019-03-08">
<collectingDate id="CF35F824A516FF9BFABA68445CDD6969" box="[1284,1421,1602,1623]" pageId="33" pageNumber="567" value="2019-03-08">8 Mar. 2019</collectingDate>
</date>
;
<collectorName id="063A42DAA516FF9BFC9768675AFF6948" box="[809,943,1632,1654]" pageId="33" pageNumber="567">B. A. Huber</collectorName>
and
<collectorName id="063A42DAA516FF9BFC5768675DC96948" box="[1001,1177,1632,1654]" pageId="33" pageNumber="567">M. A. Izquierdo</collectorName>
leg.; ZFMK
<location id="AE1071D7A516FF9BFA9968665C14694B" LSID="urn:lsid:plazi:treatment:2366961AA517FF9EFE98683C5C286A97:AE1071D7A516FF9BFA9968665C14694B" box="[1319,1348,1632,1653]" latitude="-28.9015" longLatPrecision="6" longitude="-66.6538" name="Ar" pageId="33" pageNumber="567" stateProvince="Miranda">Ar</location>
23901
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFC9768795A2B698D" collectingDate="2019-03-08" collectingDateMax="2019-03-10" collectingDateMin="2019-03-08" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="1660" lastPageId="33" lastPageNumber="568" latitude="-28.9015" location="Ar" longLatPrecision="6" longitude="-66.6538" pageId="33" pageNumber="567" specimenCount="16" specimenCount-female="15" stateProvince="Miranda">
<specimenCount id="BDC9EC85A516FF9BFC8068795A2069AA" box="[830,880,1660,1684]" pageId="33" pageNumber="567" type="female">2 ♀♀</specimenCount>
, in pure ethanol; same data as preceding; ZFMK Arg154
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFC3F68985A0369EC" collectingDate="2019-03-08" collectingDateMax="2019-03-10" collectingDateMin="2019-03-08" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="1660" lastPageId="33" lastPageNumber="568" latitude="-28.9015" location="Ar" longLatPrecision="6" longitude="-66.6538" pageId="33" pageNumber="567" specimenCount="16" specimenCount-female="15" stateProvince="Miranda">
• 1 ♂,
<specimenCount id="BDC9EC85A516FF9BFC78689B5AA7698C" box="[966,1015,1690,1714]" pageId="33" pageNumber="567" type="female">2 ♀♀</specimenCount>
; same data as preceding; LABRE-Ar 533
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFCE268BA5A8469CF" collectingDate="2019-03-08" collectingDateMax="2019-03-10" collectingDateMin="2019-03-08" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="1660" lastPageId="33" lastPageNumber="568" latitude="-28.9015" location="Ar" longLatPrecision="6" longitude="-66.6538" pageId="33" pageNumber="567" specimenCount="16" specimenCount-female="15" stateProvince="Miranda">
<specimenCount id="BDC9EC85A516FF9BFCCA68BA5AC969EF" box="[884,921,1721,1745]" pageId="33" pageNumber="567" type="female">1 ♀</specimenCount>
, in pure ethanol; same data as preceding; LABRE-Ar 553
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFC6468DD5C1A6813" collectingDate="2019-03-08" collectingDateMax="2019-03-10" collectingDateMin="2019-03-08" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="1660" lastPageId="33" lastPageNumber="568" latitude="-28.9015" location="Ar" longLatPrecision="6" longitude="-66.6538" pageId="33" pageNumber="567" specimenCount="16" specimenCount-female="15" stateProvince="Miranda">
<specimenCount id="BDC9EC85A516FF9BFC5168DD5D4269CE" box="[1007,1042,1752,1776]" pageId="33" pageNumber="567" type="female">1 ♀</specimenCount>
;
<locationDeviation id="BAC23E2EA516FF9BFBA068DD5CDC69CF" box="[1054,1420,1755,1777]" firstLocation="Chilecito" pageId="33" pageNumber="567" secondLocation="Famatina">between Chilecito and Famatina</locationDeviation>
;
<geoCoordinate id="CEFB41CBA516FF9BFC9768FF5AF16830" box="[809,929,1785,1806]" degrees="29.0027" direction="south" orientation="latitude" pageId="33" pageNumber="567" precision="5" value="-29.0027">29.0027° S</geoCoordinate>
,
<geoCoordinate id="CEFB41CBA516FF9BFC1268FF5D7B6831" box="[940,1067,1785,1807]" degrees="67.4855" direction="west" orientation="longitude" pageId="33" pageNumber="567" precision="5" value="-67.4855">67.4855° W</geoCoordinate>
;
<elevation id="20E2C03FA516FF9BFB8B68FF5D926831" box="[1077,1218,1785,1807]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="33" pageNumber="567" unit="m" value="1300.0">
<quantity id="6C378AE9A516FF9BFB8B68FF5DD76831" box="[1077,1159,1785,1807]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="33" pageNumber="567" unit="m" value="1300.0">1300 m</quantity>
a.s.l.
</elevation>
;
<date id="DF7101CCA516FF9BFB7068FF5C036830" box="[1230,1363,1785,1807]" pageId="33" pageNumber="567" value="2019-03-09">
<collectingDate id="CF35F824A516FF9BFB7068FF5C036830" box="[1230,1363,1785,1807]" pageId="33" pageNumber="567" value="2019-03-09">9 Mar. 2019</collectingDate>
</date>
;
<collectorName id="063A42DAA516FF9BFAE368FC5A216810" pageId="33" pageNumber="567">B. A. Huber</collectorName>
and
<collectorName id="063A42DAA516FF9BFC14691E5D076810" box="[938,1111,1816,1838]" pageId="33" pageNumber="567">M. A. Izquierdo</collectorName>
leg.; ZFMK Ar 23902
</materialsCitation>
<materialsCitation id="1BA72D51A516FF9BFAEF691E5DB26873" collectingDate="2019-03-08" collectingDateMax="2019-03-10" collectingDateMin="2019-03-08" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="1660" lastPageId="33" lastPageNumber="568" latitude="-28.9015" location="Ar" longLatPrecision="6" longitude="-66.6538" pageId="33" pageNumber="567" specimenCount="16" specimenCount-female="15" stateProvince="Miranda">
<specimenCount id="BDC9EC85A516FF9BFAD9691E5CDB6813" box="[1383,1419,1813,1837]" pageId="33" pageNumber="567" type="female">1 ♀</specimenCount>
; same data as preceding; LABRE-Ar 541
</materialsCitation>
<materialsCitation id="1BA72D51A516FF98FB596931599D6AC7" collectingDate="2019-03-08" collectingDateMax="2019-03-10" collectingDateMin="2019-03-08" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="1660" lastPageId="34" lastPageNumber="568" latitude="-28.9015" location="Ar" longLatPrecision="6" longitude="-66.6538" pageId="33" pageNumber="567" specimenCount="16" specimenCount-female="15" stateProvince="Miranda">
<specimenCount id="BDC9EC85A516FF9BFB4569315C4C6872" box="[1275,1308,1844,1868]" pageId="33" pageNumber="567" type="female">1 ♀</specimenCount>
with eggs, in pure ethanol; same data as preceding; LABRE-Ar 546
</materialsCitation>
<materialsCitation id="1BA72D51A515FF98FF6B6BE55BE9690B" collectingDate="2019-03-08" collectingDateMax="2019-03-10" collectingDateMin="2019-03-08" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="1660" lastPageId="34" lastPageNumber="568" latitude="-28.9015" location="Ar" longLatPrecision="6" longitude="-66.6538" pageId="34" pageNumber="567" specimenCount="16" specimenCount-female="15" stateProvince="Miranda">
• 4 ♂♂,
<specimenCount id="BDC9EC85A515FF98FE8D6BE558386AC6" box="[307,360,1504,1528]" pageId="34" pageNumber="568" type="female">2 ♀♀</specimenCount>
;
<collectorName id="063A42DAA515FF98FEC96BE55B706AC7" box="[375,544,1507,1529]" pageId="34" pageNumber="568">SE Aimogasta</collectorName>
, site 2;
<geoCoordinate id="CEFB41CBA515FF98FD396BE55A566AC6" box="[647,774,1507,1529]" degrees="28.9015" direction="south" orientation="latitude" pageId="34" pageNumber="568" precision="5" value="-28.9015">28.9015° S</geoCoordinate>
,
<geoCoordinate id="CEFB41CBA515FF98FF1D680758716929" box="[163,289,1537,1559]" degrees="66.6538" direction="west" orientation="longitude" pageId="34" pageNumber="568" precision="5" value="-66.6538">66.6538° W</geoCoordinate>
;
<elevation id="20E2C03FA515FF98FE97680758F76929" box="[297,423,1537,1559]" metricMagnitude="2" metricUnit="m" metricValue="7.55" pageId="34" pageNumber="568" unit="m" value="755.0">
<quantity id="6C378AE9A515FF98FE976807583D6929" box="[297,365,1537,1559]" metricMagnitude="2" metricUnit="m" metricValue="7.55" pageId="34" pageNumber="568" unit="m" value="755.0">755 m</quantity>
a.s.l.
</elevation>
; under rocks;
<date id="DF7101CCA515FF98FD8168045B9F6929" box="[575,719,1538,1559]" pageId="34" pageNumber="568" value="2019-03-10">
<collectingDate id="CF35F824A515FF98FD8168045B9F6929" box="[575,719,1538,1559]" pageId="34" pageNumber="568" value="2019-03-10">10 Mar. 2019</collectingDate>
</date>
;
<collectorName id="063A42DAA515FF98FD66680459BB6908" pageId="34" pageNumber="568">B. A. Huber</collectorName>
and
<collectorName id="063A42DAA515FF98FE9F6827589B6908" box="[289,459,1568,1590]" pageId="34" pageNumber="568">M. A. Izquierdo</collectorName>
leg.; ZFMK
<collectorName id="063A42DAA515FF98FDED68265B20690B" box="[595,624,1568,1589]" pageId="34" pageNumber="568">Ar</collectorName>
23903
</materialsCitation>
<materialsCitation id="1BA72D51A515FF98FD0168275A5A696B" collectingDate="2019-03-08" collectingDateMax="2019-03-10" collectingDateMin="2019-03-08" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="1660" lastPageId="34" lastPageNumber="568" latitude="-28.9015" location="Ar" longLatPrecision="6" longitude="-66.6538" pageId="34" pageNumber="567" specimenCount="16" specimenCount-female="15" stateProvince="Miranda">
<specimenCount id="BDC9EC85A515FF98FD6A68265A55690B" box="[724,773,1565,1589]" pageId="34" pageNumber="568" type="female">3 ♀♀</specimenCount>
, in pure ethanol; same data as preceding; ZFMK
<collectorName id="063A42DAA515FF98FD0668395BB3696B" box="[696,739,1599,1621]" pageId="34" pageNumber="568">Arg</collectorName>
162
</materialsCitation>
<materialsCitation id="1BA72D51A515FF98FF1D6858599D69AC" collectingDate="2019-03-08" collectingDateMax="2019-03-10" collectingDateMin="2019-03-08" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="1660" lastPageId="34" lastPageNumber="568" latitude="-28.9015" location="Ar" longLatPrecision="6" longitude="-66.6538" pageId="34" pageNumber="567" specimenCount="16" specimenCount-female="15" stateProvince="Miranda">
• 1 ♂,
<specimenCount id="BDC9EC85A515FF98FF5068585842694D" box="[238,274,1627,1651]" pageId="34" pageNumber="568" type="female">1 ♀</specimenCount>
,
<specimenCount id="BDC9EC85A515FF98FE9E68585831694D" box="[288,353,1629,1651]" pageId="34" pageNumber="568" type="generic">1 juv.</specimenCount>
; same data as preceding; LABRE-Ar 532
</materialsCitation>
<materialsCitation id="1BA72D51A515FF98FF68687B5803698F" collectingDate="2019-03-08" collectingDateMax="2019-03-10" collectingDateMin="2019-03-08" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="1660" lastPageId="34" lastPageNumber="568" latitude="-28.9015" location="Ar" longLatPrecision="6" longitude="-66.6538" pageId="34" pageNumber="567" specimenCount="16" specimenCount-female="15" stateProvince="Miranda">
<specimenCount id="BDC9EC85A515FF98FF50687A584369AF" box="[238,275,1657,1681]" pageId="34" pageNumber="568" type="female">1 ♀</specimenCount>
, in pure ethanol; same data as preceding; LABRE-Ar 567
</materialsCitation>
.
<materialsCitation id="1BA72D51A515FF98FEE3689D580D6815" collectingDate="2019-03-25" collectingDateMax="2019-03-29" collectingDateMin="2019-03-25" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="640" lastPageId="34" lastPageNumber="569" latitude="-27.1559" location="Balneario El Caolin" longLatPrecision="6" longitude="-66.6925" pageId="34" pageNumber="568" specimenCount="49" specimenCount-female="44" specimenCount-male="3" stateProvince="Catamarca">
<collectingRegion id="690BE9EEA515FF98FEE3689D58B9698F" box="[349,489,1691,1713]" country="Argentina" name="Catamarca" pageId="34" pageNumber="568">
<emphasis id="99BBFB1EA515FF98FEE3689D58B9698F" bold="true" box="[349,489,1691,1713]" pageId="34" pageNumber="568">Catamarca</emphasis>
</collectingRegion>
: • 2 ♂♂,
<specimenCount id="BDC9EC85A515FF98FDF1689D5BD2698E" box="[591,642,1688,1712]" pageId="34" pageNumber="568" type="female">4 ♀♀</specimenCount>
; ~
<locationDeviation id="BAC23E2EA515FF98FD20689D587769F1" location="Chumbicha" pageId="34" pageNumber="568">
<quantity id="6C378AE9A515FF98FD20689D5B88698F" box="[670,728,1691,1713]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="568" unit="km" value="5.0">5 km</quantity>
NW Chumbicha
</locationDeviation>
, near
<collectorName id="063A42DAA515FF98FED568BF5B1C69F1" box="[363,588,1721,1743]" pageId="34" pageNumber="568">
<location id="AE1071D7A515FF98FED568BF5B1C69F1" LSID="urn:lsid:plazi:treatment:2366961AA517FF9EFE98683C5C286A97:AE1071D7A515FF98FED568BF5B1C69F1" box="[363,588,1721,1743]" latitude="-27.1559" longLatPrecision="6" longitude="-66.6925" name="Balneario El Caolin" pageId="34" pageNumber="568" stateProvince="Catamarca">Balneario El Caolín</location>
</collectorName>
, site 2;
<geoCoordinate id="CEFB41CBA515FF98FD1768BC59E369D3" degrees="28.8109" direction="south" orientation="latitude" pageId="34" pageNumber="568" precision="5" value="-28.8109">28.8109° S</geoCoordinate>
,
<geoCoordinate id="CEFB41CBA515FF98FF7E68DE581569D0" box="[192,325,1752,1774]" degrees="66.2500" direction="west" orientation="longitude" pageId="34" pageNumber="568" precision="5" value="-66.25">66.2500° W</geoCoordinate>
;
<elevation id="20E2C03FA515FF98FEEC68DE588969D0" box="[338,473,1752,1774]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="34" pageNumber="568" unit="m" value="640.0">
<quantity id="6C378AE9A515FF98FEEC68DE58CA69D0" box="[338,410,1752,1774]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="34" pageNumber="568" unit="m" value="640.0">640 m</quantity>
a.s.l.
</elevation>
; steep rock field in forest;
<date id="DF7101CCA515FF98FF1D68F158306832" box="[163,352,1783,1804]" pageId="34" pageNumber="568" value="2019-03-28" valueMax="2019-03-29" valueMin="2019-03-28">
<collectingDate id="CF35F824A515FF98FF1D68F158306832" box="[163,352,1783,1804]" pageId="34" pageNumber="568" value="2019-03-28" valueMax="2019-03-29" valueMin="2019-03-28">2829 Mar. 2019</collectingDate>
</date>
;
<collectorName id="063A42DAA515FF98FED268F158BF6833" box="[364,495,1783,1805]" pageId="34" pageNumber="568">B. A. Huber</collectorName>
and
<collectorName id="063A42DAA515FF98FD9768F15B876833" box="[553,727,1783,1805]" pageId="34" pageNumber="568">M. A. Izquierdo</collectorName>
leg.; ZFMK Ar 23904
</materialsCitation>
<materialsCitation id="1BA72D51A515FF98FEDA691058A26856" collectingDate="2019-03-25" collectingDateMax="2019-03-29" collectingDateMin="2019-03-25" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="640" lastPageId="34" lastPageNumber="569" latitude="-27.1559" location="Balneario El Caolin" longLatPrecision="6" longitude="-66.6925" pageId="34" pageNumber="568" specimenCount="49" specimenCount-female="44" specimenCount-male="3" stateProvince="Catamarca">
<specimenCount id="BDC9EC85A515FF98FEC5691058FE6815" box="[379,430,1811,1835]" pageId="34" pageNumber="568" type="female">7 ♀♀</specimenCount>
, 3 juvs, in pure ethanol (
<specimenCount id="BDC9EC85A515FF98FD70691359BF6874" pageId="34" pageNumber="568" type="female">three female</specimenCount>
prosomata used for molecular study); same data as preceding; ZFMK Arg218
</materialsCitation>
<materialsCitation id="1BA72D51A515FF98FE4469555D5E6929" collectingDate="2019-03-25" collectingDateMax="2019-03-29" collectingDateMin="2019-03-25" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="640" lastPageId="34" lastPageNumber="569" latitude="-27.1559" location="Balneario El Caolin" longLatPrecision="6" longitude="-66.6925" pageId="34" pageNumber="568" specimenCount="49" specimenCount-female="44" specimenCount-male="3" stateProvince="Catamarca">
• 2 ♂♂,
<specimenCount id="BDC9EC85A515FF98FDE769555BDE6856" box="[601,654,1872,1897]" pageId="34" pageNumber="568" type="female">6 ♀♀</specimenCount>
, 8 juvs, in pure ethanol; same data as preceding; LABRE-Ar 530, 544, 561, 563, 566
</materialsCitation>
<materialsCitation id="1BA72D51A515FF98FBA868045ABC694D" collectingDate="2019-03-25" collectingDateMax="2019-03-29" collectingDateMin="2019-03-25" collectorName="M. A. Izquierdo &amp; B. A. Huber" elevation="640" lastPageId="34" lastPageNumber="569" latitude="-27.1559" location="Balneario El Caolin" longLatPrecision="6" longitude="-66.6925" pageId="34" pageNumber="568" specimenCount="49" specimenCount-female="44" specimenCount-male="3" stateProvince="Catamarca">
<specimenCount id="BDC9EC85A515FF98FB9368045D316929" box="[1069,1121,1535,1559]" pageId="34" pageNumber="568" type="female">3 ♀♀</specimenCount>
with eggs, in pure ethanol; same locality as preceding, site 1;
<geoCoordinate id="CEFB41CBA515FF98FB0068265C66690B" box="[1214,1334,1568,1590]" degrees="28.8152" direction="south" orientation="latitude" pageId="34" pageNumber="568" precision="5" value="-28.8152">28.8152° S</geoCoordinate>
,
<geoCoordinate id="CEFB41CBA515FF98FAFF68265A03696A" degrees="66.2478" direction="west" orientation="longitude" pageId="34" pageNumber="568" precision="5" value="-66.2478">66.2478° W</geoCoordinate>
;
<elevation id="20E2C03FA515FF98FCE168395AB6696B" box="[863,998,1599,1621]" metricMagnitude="2" metricUnit="m" metricValue="6.05" pageId="34" pageNumber="568" unit="m" value="605.0">
<quantity id="6C378AE9A515FF98FCE168395AF7696A" box="[863,935,1599,1621]" metricMagnitude="2" metricUnit="m" metricValue="6.05" pageId="34" pageNumber="568" unit="m" value="605.0">605 m</quantity>
a.s.l.
</elevation>
;
<collectorName id="063A42DAA515FF98FC4A68395DF8696B" box="[1012,1192,1599,1621]" pageId="34" pageNumber="568">M. A. Izquierdo</collectorName>
and
<collectorName id="063A42DAA515FF98FB5D68395C3C696B" box="[1251,1388,1599,1621]" pageId="34" pageNumber="568">B. A. Huber</collectorName>
leg.; LABRE-Ar 556
</materialsCitation>
<materialsCitation id="1BA72D51A515FF98FC4A68585D9A69AC" collectingDate="2019-03-25" collectingDateMax="2019-03-29" collectingDateMin="2019-03-25" collectorName="M. A. Izquierdo &amp; B. A. Huber" elevation="640" lastPageId="34" lastPageNumber="569" latitude="-27.1559" location="Balneario El Caolin" longLatPrecision="6" longitude="-66.6925" pageId="34" pageNumber="568" specimenCount="49" specimenCount-female="44" specimenCount-male="3" stateProvince="Catamarca">
<specimenCount id="BDC9EC85A515FF98FBB2685B5D10694C" box="[1036,1088,1626,1650]" pageId="34" pageNumber="568" type="female">2 ♀♀</specimenCount>
, 8 juvs, in pure ethanol; same data as preceding; LABRE-Ar564
</materialsCitation>
<materialsCitation id="1BA72D51A515FF98FB6D687A5AD26832" collectingDate="2019-03-25" collectingDateMax="2019-03-29" collectingDateMin="2019-03-25" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="640" lastPageId="34" lastPageNumber="569" latitude="-27.1559" location="Balneario El Caolin" longLatPrecision="6" longitude="-66.6925" pageId="34" pageNumber="568" specimenCount="49" specimenCount-female="44" specimenCount-male="3" stateProvince="Catamarca">
• 3 ♂♂,
<specimenCount id="BDC9EC85A515FF98FA8D687A5C3869AF" box="[1331,1384,1657,1681]" pageId="34" pageNumber="568" type="female">2 ♀♀</specimenCount>
(
<specimenCount id="BDC9EC85A515FF98FAC7687A5A23698F" pageId="34" pageNumber="568" type="male">one male</specimenCount>
used for µ-CT study); ~
<quantity id="6C378AE9A515FF98FB3B689D5D9D698F" box="[1157,1229,1691,1713]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="568" unit="km" value="10.0">10 km</quantity>
N
<collectingRegion id="690BE9EEA515FF98FB4E689D5C64698F" box="[1264,1332,1691,1713]" country="Germany" name="Berlin" pageId="34" pageNumber="568">Belén</collectingRegion>
;
<geoCoordinate id="CEFB41CBA515FF98FA81689D5A1B69F0" degrees="27.5641" direction="south" orientation="latitude" pageId="34" pageNumber="568" precision="5" value="-27.5641">27.5641° S</geoCoordinate>
,
<geoCoordinate id="CEFB41CBA515FF98FCE968BF5A8A69F1" box="[855,986,1721,1743]" degrees="67.0058" direction="west" orientation="longitude" pageId="34" pageNumber="568" precision="5" value="-67.0058">67.0058° W</geoCoordinate>
;
<elevation id="20E2C03FA515FF98FC5868BF5D2969F1" box="[998,1145,1721,1743]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="34" pageNumber="568" unit="m" value="1380.0">
<quantity id="6C378AE9A515FF98FC5868BF5D6B69F1" box="[998,1083,1721,1743]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="34" pageNumber="568" unit="m" value="1380.0">1380 m</quantity>
a.s.l.
</elevation>
; in pile of stones;
<date id="DF7101CCA515FF98FAF268BF5A2569D3" pageId="34" pageNumber="568" value="2019-03-25">
<collectingDate id="CF35F824A515FF98FAF268BF5A2569D3" pageId="34" pageNumber="568" value="2019-03-25">25 Mar. 2019</collectingDate>
</date>
;
<collectorName id="063A42DAA515FF98FC3E68DE5D5869D0" box="[896,1032,1752,1774]" pageId="34" pageNumber="568">B. A. Huber</collectorName>
and
<collectorName id="063A42DAA515FF98FBFD68DE5DA669D0" box="[1091,1270,1752,1774]" pageId="34" pageNumber="568">M. A. Izquierdo</collectorName>
leg.; ZFMK Ar 23905
</materialsCitation>
<materialsCitation id="1BA72D51A515FF98FC3568F15D356814" collectingDate="2019-03-25" collectingDateMax="2019-03-29" collectingDateMin="2019-03-25" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="640" lastPageId="34" lastPageNumber="569" latitude="-27.1559" location="Balneario El Caolin" longLatPrecision="6" longitude="-66.6925" pageId="34" pageNumber="568" specimenCount="49" specimenCount-female="44" specimenCount-male="3" stateProvince="Catamarca">
<specimenCount id="BDC9EC85A515FF98FC1A68F05A896832" box="[932,985,1780,1804]" pageId="34" pageNumber="568" type="female">5 ♀♀</specimenCount>
,
<specimenCount id="BDC9EC85A515FF98FC5768F15D7C6833" box="[1001,1068,1783,1805]" pageId="34" pageNumber="568" type="generic">1 juv.</specimenCount>
, in pure ethanol; same data as preceding; ZFMK Arg213
</materialsCitation>
<materialsCitation id="1BA72D51A515FF98FBD369105C396874" collectingDate="2019-03-25" collectingDateMax="2019-03-29" collectingDateMin="2019-03-25" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="640" lastPageId="34" lastPageNumber="569" latitude="-27.1559" location="Balneario El Caolin" longLatPrecision="6" longitude="-66.6925" pageId="34" pageNumber="568" specimenCount="49" specimenCount-female="44" specimenCount-male="3" stateProvince="Catamarca">
• 1 ♂,
<specimenCount id="BDC9EC85A515FF98FB0569105DBF6814" box="[1211,1263,1810,1835]" pageId="34" pageNumber="568" type="female">7 ♀♀</specimenCount>
,
<specimenCount id="BDC9EC85A515FF98FB4169135C116815" box="[1279,1345,1813,1835]" pageId="34" pageNumber="568" type="generic">1 juv.</specimenCount>
, in pure ethanol; same data as preceding; LABRE-Ar 545
</materialsCitation>
<materialsCitation id="1BA72D51A515FF99FACF69325BF76946" collectingDate="2019-03-25" collectingDateMax="2019-03-29" collectingDateMin="2019-03-25" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="640" lastPageId="35" lastPageNumber="569" latitude="-27.1559" location="Balneario El Caolin" longLatPrecision="6" longitude="-66.6925" pageId="34" pageNumber="568" specimenCount="49" specimenCount-female="44" specimenCount-male="3" stateProvince="Catamarca">
• 11 ♂♂,
<specimenCount id="BDC9EC85A515FF98FCD469555ACF6856" box="[874,927,1872,1896]" pageId="34" pageNumber="568" type="female">2 ♀♀</specimenCount>
(
<specimenCount id="BDC9EC85A515FF98FC1169555D6C6857" box="[943,1084,1875,1897]" pageId="34" pageNumber="568" type="male">three males</specimenCount>
and
<specimenCount id="BDC9EC85A515FF98FBC669555DA86857" box="[1144,1272,1875,1897]" pageId="34" pageNumber="568" type="female">one female</specimenCount>
used for µ-CT study;
<specimenCount id="BDC9EC85A514FF99FF61680158186922" box="[223,328,1542,1564]" pageId="35" pageNumber="569" type="male">two male</specimenCount>
abdomens used for karyotype study); near Nacimientos;
<geoCoordinate id="CEFB41CBA514FF99FEC6682358AA6904" box="[376,506,1573,1595]" degrees="27.1559" direction="south" orientation="latitude" pageId="35" pageNumber="569" precision="5" value="-27.1559">27.1559° S</geoCoordinate>
,
<geoCoordinate id="CEFB41CBA514FF99FDB768235BC36904" box="[521,659,1573,1595]" degrees="66.6925" direction="west" orientation="longitude" pageId="35" pageNumber="569" precision="5" value="-66.6925">66.6925° W</geoCoordinate>
;
<elevation id="20E2C03FA514FF99FD1E682359996967" metricMagnitude="3" metricUnit="m" metricValue="2.12" pageId="35" pageNumber="569" unit="m" value="2120.0">
<quantity id="6C378AE9A514FF99FD1E68235BA86904" box="[672,760,1573,1594]" metricMagnitude="3" metricUnit="m" metricValue="2.12" pageId="35" pageNumber="569" unit="m" value="2120.0">2120 m</quantity>
a.s.l.
</elevation>
; under rocks on arid slope;
<date id="DF7101CCA514FF99FDA968455BE56966" box="[535,693,1603,1625]" pageId="35" pageNumber="569" value="2019-03-25">
<collectingDate id="CF35F824A514FF99FDA968455BE56966" box="[535,693,1603,1625]" pageId="35" pageNumber="569" value="2019-03-25">25 Mar. 2019</collectingDate>
</date>
;
<collectorName id="063A42DAA514FF99FD7F684259896946" pageId="35" pageNumber="569">B. A. Huber</collectorName>
and
<collectorName id="063A42DAA514FF99FEB1686458E96946" box="[271,441,1634,1656]" pageId="35" pageNumber="569">M. A. Izquierdo</collectorName>
leg.; ZFMK
<collectorName id="063A42DAA514FF99FDFF68645B0E6949" box="[577,606,1634,1655]" pageId="35" pageNumber="569">Ar</collectorName>
23906
</materialsCitation>
<materialsCitation id="1BA72D51A514FF99FD13686459B7698A" collectingDate="2019-03-25" collectingDateMax="2019-03-29" collectingDateMin="2019-03-25" collectorName="B. A. Huber &amp; M. A. Izquierdo" elevation="640" lastPageId="35" lastPageNumber="569" latitude="-27.1559" location="Balneario El Caolin" longLatPrecision="6" longitude="-66.6925" pageId="35" pageNumber="568" specimenCount="49" specimenCount-female="44" specimenCount-male="3" stateProvince="Catamarca">
<specimenCount id="BDC9EC85A514FF99FD7C68645BA36949" box="[706,755,1631,1655]" pageId="35" pageNumber="569" type="female">4 ♀♀</specimenCount>
, 3 juvs, in pure ethanol; same data as preceding; ZFMK
<collectorName id="063A42DAA514FF99FF2F689959EE698A" box="[145,190,1695,1716]" pageId="35" pageNumber="569">Arg</collectorName>
211
</materialsCitation>
.
</paragraph>
<caption id="FFB07784A515FF98FF1D6B125B6C6AA1" ID-DOI="http://doi.org/10.5281/zenodo.7981854" ID-Zenodo-Dep="7981854" httpUri="https://zenodo.org/record/7981854/files/figure.png" pageId="34" pageNumber="568" startId="34.[163,242,1300,1322]" targetBox="[323,1282,198,1270]" targetPageId="34" targetType="figure">
<paragraph id="AB70270CA515FF98FF1D6B125B6C6AA1" blockId="34.[162,1443,1300,1439]" pageId="34" pageNumber="568">
<emphasis id="99BBFB1EA515FF98FF1D6B12584A6A17" bold="true" box="[163,282,1300,1322]" pageId="34" pageNumber="568">Figure 29.</emphasis>
<taxonomicName id="6CCF5C8FA515FF98FE9D6B1258F16A14" authority="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho, 2023" authorityName="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho" authorityYear="2023" box="[291,417,1300,1322]" class="Arachnida" family="Pholcidae" genus="Nerudia" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="568" phylum="Arthropoda" rank="species" species="ola" status="sp. nov.">
<emphasis id="99BBFB1EA515FF98FE9D6B1258F16A14" box="[291,417,1300,1322]" italics="true" pageId="34" pageNumber="568">Nerudia ola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82884665A515FF98FE166B1258A76A14" box="[424,503,1300,1322]" pageId="34" pageNumber="568" rank="species">sp. nov.</taxonomicNameLabel>
; male and female from Argentina, San Juan, San Agustín de
<taxonomicName id="6CCF5C8FA515FF98FBC46B125DA16A14" authority="Fertil" authorityName="Fertil" box="[1146,1265,1300,1322]" class="Arachnida" family="Pholcidae" genus="Artema" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="568" phylum="Arthropoda" rank="species" species="valle">Valle Fértil</taxonomicName>
(ZFMK Arg141). A, female prosoma, frontal view. B, left procursus tip, prolateral-dorsal view. C, stridulatory file on male chelicera. D, lateral view of female chelicera, showing absence of stridulatory file. E, stridulatory pick (modified hair) on male palpal femur. F, right male cheliceral apophysis, frontal view. G, processes on male sternum. H, male gonopore. Scale lines: 100 µm (A, G), 10 µm (B, D, F, H), 20 µm (C), 2 µm (E).
</paragraph>
</caption>
<caption id="FFB07784A514FF99FF2F6B645D606AEE" ID-DOI="http://doi.org/10.5281/zenodo.7981858" ID-Zenodo-Dep="7981858" httpUri="https://zenodo.org/record/7981858/files/figure.png" pageId="35" pageNumber="569" startId="35.[145,223,1378,1400]" targetBox="[306,1265,196,1347]" targetPageId="35" targetType="figure">
<paragraph id="AB70270CA514FF99FF2F6B645D606AEE" blockId="35.[145,1425,1378,1488]" pageId="35" pageNumber="569">
<emphasis id="99BBFB1EA514FF99FF2F6B6458556A49" bold="true" box="[145,261,1378,1400]" pageId="35" pageNumber="569">Figure 30.</emphasis>
<taxonomicName id="6CCF5C8FA514FF99FEAE6B6458DC6A46" authority="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho, 2023" authorityName="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho" authorityYear="2023" box="[272,396,1378,1400]" class="Arachnida" family="Pholcidae" genus="Nerudia" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="569" phylum="Arthropoda" rank="species" species="ola" status="sp. nov.">
<emphasis id="99BBFB1EA514FF99FEAE6B6458DC6A46" box="[272,396,1378,1400]" italics="true" pageId="35" pageNumber="569">Nerudia ola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82884665A514FF99FE2C6B64588F6A46" box="[402,479,1378,1400]" pageId="35" pageNumber="569" rank="species">sp. nov.</taxonomicNameLabel>
; male and female from Argentina, San Juan, San Agustín de
<taxonomicName id="6CCF5C8FA514FF99FBED6B645D976A46" authority="Fertil" authorityName="Fertil" box="[1107,1223,1378,1400]" class="Arachnida" family="Pholcidae" genus="Artema" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="569" phylum="Arthropoda" rank="species" species="valle">Valle Fértil</taxonomicName>
(ZFMK Arg141). A, male right tibia 1, retrolateral view. B, male metatarsus 4 with trichobothrium. C, male palpal tarsal organ. D, tip of right female palp, dorsal view; arrow points at tarsal organ. E, tarsal organ on male tarsus 2. F, epigynum, ventral view. G, male ALS and PMS. H, female ALS. Scale lines: 10 µm (A, B, D, G, H), 2 µm (C, E), 100 µm (F).
</paragraph>
</caption>
<paragraph id="AB70270CA514FF99FF2F68DF5C7169CC" blockId="35.[145,762,1753,1897]" lastBlockId="35.[809,1427,1542,1779]" pageId="35" pageNumber="569">
<emphasis id="99BBFB1EA514FF99FF2F68DF5B0B69D0" box="[145,603,1753,1774]" italics="true" pageId="35" pageNumber="569">Assigned tentatiƲely (only females aƲailable):</emphasis>
<materialsCitation id="1BA72D51A514FF99FDDA68DF59816874" collectingDate="1994-08" collectorName="M. Ramirez &amp; Ar" country="Argentina" elevation="1700" latitude="-29.35" location="Cuesta de Miranda" longLatPrecision="1227" longitude="-67.71667" pageId="35" pageNumber="569" specimenCount="3" specimenCount-female="2" stateProvince="La Rioja">
<collectingCountry id="D3D8679CA514FF99FDDA68DF5BA969D0" box="[612,761,1753,1774]" name="Argentina" pageId="35" pageNumber="569">ARGENTINA</collectingCountry>
<emphasis id="99BBFB1EA514FF99FF2F68FE59CD6833" box="[145,157,1784,1805]" italics="true" pageId="35" pageNumber="569"></emphasis>
<emphasis id="99BBFB1EA514FF99FF1E68F158526833" bold="true" box="[160,258,1783,1805]" pageId="35" pageNumber="569">
<collectingRegion id="690BE9EEA514FF99FF1E68F158526833" box="[160,258,1783,1805]" country="Argentina" name="La Rioja" pageId="35" pageNumber="569">La Rioja</collectingRegion>
</emphasis>
: •
<specimenCount id="BDC9EC85A514FF99FEA368F1581B6832" box="[285,331,1780,1804]" pageId="35" pageNumber="569" type="female">2 ♀♀</specimenCount>
,
<specimenCount id="BDC9EC85A514FF99FEED68F158DA6833" box="[339,394,1783,1805]" pageId="35" pageNumber="569" type="generic">1 juv.</specimenCount>
;
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;
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,
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;
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a.s.l.
</elevation>
;
<date id="DF7101CCA514FF99FEF3691058EE6815" box="[333,446,1814,1835]" pageId="35" pageNumber="569" value="1994-08">
<collectingDate id="CF35F824A514FF99FEF3691058EE6815" box="[333,446,1814,1835]" pageId="35" pageNumber="569" value="1994-08">Aug. 1994</collectingDate>
</date>
;
<collectorName id="063A42DAA514FF99FE7469105B1D6815" box="[458,589,1814,1835]" pageId="35" pageNumber="569">M. Ramírez</collectorName>
leg.; MACN
<collectorName id="063A42DAA514FF99FD6269105BA96815" box="[732,761,1814,1835]" pageId="35" pageNumber="569">Ar</collectorName>
20055
</materialsCitation>
<materialsCitation id="1BA72D51A514FF99FF5669335C526922" collectingDate="2019-03-10" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="915" latitude="-28.8069" location="Aimogasta" longLatPrecision="6" longitude="-66.6635" pageId="35" pageNumber="569" specimenCount="1" specimenCount-female="1" stateProvince="La Rioja">
<specimenCount id="BDC9EC85A514FF99FF56693358596874" box="[232,265,1842,1866]" pageId="35" pageNumber="569" type="female">1 ♀</specimenCount>
, in pure ethanol; SE
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, site 1;
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,
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;
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<quantity id="6C378AE9A514FF99FE946955583D6857" box="[298,365,1875,1897]" metricMagnitude="2" metricUnit="m" metricValue="9.15" pageId="35" pageNumber="569" unit="m" value="915.0">915 m</quantity>
a.s.l.
</elevation>
; under rocks;
<date id="DF7101CCA514FF99FD8A69555BEE6856" box="[564,702,1875,1897]" pageId="35" pageNumber="569" value="2019-03-10">
<collectingDate id="CF35F824A514FF99FD8A69555BEE6856" box="[564,702,1875,1897]" pageId="35" pageNumber="569" value="2019-03-10">10 Mar. 2019</collectingDate>
</date>
;
<collectorName id="063A42DAA514FF99FD7669525A3D6922" box="[712,877,1542,1897]" pageId="35" pageNumber="569">B. A. Huber</collectorName>
and
<collectorName id="063A42DAA514FF99FC2368005D696922" box="[925,1081,1542,1564]" pageId="35" pageNumber="569">M.A. Izquierdo</collectorName>
leg.; ZFMKArg161
</materialsCitation>
.
<materialsCitation id="1BA72D51A514FF99FAB768005D366949" collectingDate="2019-03-26" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="2000" latitude="-27.659" location="14 km W Fiambala" longLatPrecision="6" longitude="-67.7607" pageId="35" pageNumber="569" specimenCount="1" specimenCount-female="1" stateProvince="Catamarca">
<emphasis id="99BBFB1EA514FF99FAB768005CDB6922" bold="true" box="[1289,1419,1542,1564]" pageId="35" pageNumber="569">
<collectingRegion id="690BE9EEA514FF99FAB768005CDB6922" box="[1289,1419,1542,1564]" country="Argentina" name="Catamarca" pageId="35" pageNumber="569">Catamarca</collectingRegion>
</emphasis>
: •
<specimenCount id="BDC9EC85A514FF99FC8068235A316904" box="[830,865,1570,1594]" pageId="35" pageNumber="569" type="female">1 ♀</specimenCount>
; ~
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W Fiambalá
</location>
;
<geoCoordinate id="CEFB41CBA514FF99FBE668225D9B6904" box="[1112,1227,1572,1594]" degrees="27.6590" direction="south" orientation="latitude" pageId="35" pageNumber="569" precision="5" value="-27.659">27.6590° S</geoCoordinate>
,
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;
<elevation id="20E2C03FA514FF99FAE268235A266967" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="569" unit="m" value="2000.0">
<quantity id="6C378AE9A514FF99FAE268235A6F6967" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="569" unit="m" value="2000.0">2000 m</quantity>
a.s.l.
</elevation>
; under rocks;
<date id="DF7101CCA514FF99FBB368455DCB6966" box="[1037,1179,1603,1625]" pageId="35" pageNumber="569" value="2019-03-26">
<collectingDate id="CF35F824A514FF99FBB368455DCB6966" box="[1037,1179,1603,1625]" pageId="35" pageNumber="569" value="2019-03-26">26 Mar. 2019</collectingDate>
</date>
;
<collectorName id="063A42DAA514FF99FB1B68425C736967" box="[1189,1315,1603,1625]" pageId="35" pageNumber="569">B. A. Huber</collectorName>
and
<collectorName id="063A42DAA514FF99FAE668425ADC6946" pageId="35" pageNumber="569">M. A. Izquierdo</collectorName>
leg.; ZFMK
<collectorName id="063A42DAA514FF99FBB968645D736949" box="[1031,1059,1634,1655]" pageId="35" pageNumber="569">Ar</collectorName>
23907
</materialsCitation>
<materialsCitation id="1BA72D51A514FF99FBC368645ADB698A" collectingDate="1994-08" collectorName="M. J. Ramirez &amp; Ar" country="Argentina" elevation="400" latitude="-28.87" location="Chumbicha" longLatPrecision="738" longitude="-66.23" pageId="35" pageNumber="569" specimenCount="2" specimenCount-female="2" stateProvince="Catamarca">
<specimenCount id="BDC9EC85A514FF99FBC368645DFC6949" box="[1149,1196,1631,1655]" pageId="35" pageNumber="569" type="female">2 ♀♀</specimenCount>
;
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;
<geoCoordinate id="CEFB41CBA514FF99FA8968645CDD6949" box="[1335,1421,1634,1655]" degrees="28.87" direction="south" orientation="latitude" pageId="35" pageNumber="569" precision="555" value="-28.87">28.87° S</geoCoordinate>
,
<geoCoordinate id="CEFB41CBA514FF99FC9768875AD769A8" box="[809,903,1665,1687]" degrees="66.23" direction="west" orientation="longitude" pageId="35" pageNumber="569" precision="555" value="-66.23">66.23° W</geoCoordinate>
;
<elevation id="20E2C03FA514FF99FC2E68875D5769A9" box="[912,1031,1665,1687]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="35" pageNumber="569" unit="m" value="400.0">
<quantity id="6C378AE9A514FF99FC2E68875A8269A8" box="[912,978,1665,1686]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="35" pageNumber="569" unit="m" value="400.0">400 m</quantity>
a.s.l.
</elevation>
;
<date id="DF7101CCA514FF99FBAE68875D2B69A8" box="[1040,1147,1665,1686]" pageId="35" pageNumber="569" value="1994-08">
<collectingDate id="CF35F824A514FF99FBAE68875D2B69A8" box="[1040,1147,1665,1686]" pageId="35" pageNumber="569" value="1994-08">Aug. 1994</collectingDate>
</date>
;
<collectorName id="063A42DAA514FF99FB3A68875C4069A8" box="[1156,1296,1665,1686]" pageId="35" pageNumber="569">M.J. Ramírez</collectorName>
leg.; MACN
<collectorName id="063A42DAA514FF99FC9768995A15698A" box="[809,837,1695,1716]" pageId="35" pageNumber="569">Ar</collectorName>
20012
</materialsCitation>
.
<materialsCitation id="1BA72D51A514FF99FC2D68995C4D69CC" collectingDate="2019-03-06" collectorName="B. A. Huber &amp; M. A. Izquierdo" country="Argentina" elevation="2910" latitude="-30.3967" location="35 km W Las Flores" longLatPrecision="6" longitude="-69.5576" pageId="35" pageNumber="569" specimenCount="1" specimenCount-female="1" stateProvince="San Juan">
<emphasis id="99BBFB1EA514FF99FC2D68995D52698A" bold="true" box="[915,1026,1695,1717]" pageId="35" pageNumber="569">
<collectingRegion id="690BE9EEA514FF99FC2D68995D52698A" box="[915,1026,1695,1717]" country="Argentina" name="San Juan" pageId="35" pageNumber="569">San Juan</collectingRegion>
</emphasis>
: •
<specimenCount id="BDC9EC85A514FF99FB9F68995D13698A" box="[1057,1091,1692,1716]" pageId="35" pageNumber="569" type="female">1 ♀</specimenCount>
; ~
<location id="AE1071D7A514FF99FBE468995C7D698B" LSID="urn:lsid:plazi:treatment:2366961AA517FF9EFE98683C5C286A97:AE1071D7A514FF99FBE468995C7D698B" box="[1114,1325,1695,1717]" country="Argentina" latitude="-30.3967" longLatPrecision="6" longitude="-69.5576" name="35 km W Las Flores" pageId="35" pageNumber="569" stateProvince="San Juan">
<quantity id="6C378AE9A514FF99FBE468995DCC698B" box="[1114,1180,1695,1717]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="35" pageNumber="569" unit="km" value="35.0">35 km</quantity>
W Las Flores
</location>
;
<geoCoordinate id="CEFB41CBA514FF99FA8B68995A6869ED" degrees="30.3967" direction="south" orientation="latitude" pageId="35" pageNumber="569" precision="5" value="-30.3967">30.3967° S</geoCoordinate>
,
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;
<elevation id="20E2C03FA514FF99FC7C68B85D1669EA" box="[962,1094,1726,1748]" metricMagnitude="3" metricUnit="m" metricValue="2.91" pageId="35" pageNumber="569" unit="m" value="2910.0">
<quantity id="6C378AE9A514FF99FC7C68B85D4269ED" box="[962,1042,1726,1747]" metricMagnitude="3" metricUnit="m" metricValue="2.91" pageId="35" pageNumber="569" unit="m" value="2910.0">2910 m</quantity>
a.s.l.
</elevation>
; under rocks;
<date id="DF7101CCA514FF99FB6768B85C0669ED" box="[1241,1366,1726,1748]" pageId="35" pageNumber="569" value="2019-03-06">
<collectingDate id="CF35F824A514FF99FB6768B85C0669ED" box="[1241,1366,1726,1748]" pageId="35" pageNumber="569" value="2019-03-06">6 Mar. 2019</collectingDate>
</date>
;
<collectorName id="063A42DAA514FF99FADE68B85A3D69CD" pageId="35" pageNumber="569">B. A. Huber</collectorName>
and
<collectorName id="063A42DAA514FF99FC2168DB5D6D69CD" box="[927,1085,1757,1779]" pageId="35" pageNumber="569">M. A. Izquierdo</collectorName>
leg.; ZFMK
<collectorName id="063A42DAA514FF99FB0468DB5D8669CC" box="[1210,1238,1757,1778]" pageId="35" pageNumber="569">Ar</collectorName>
23908
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E3D57487A514FF99FC9769105AF76857" pageId="35" pageNumber="569" type="etymology">
<paragraph id="AB70270CA514FF99FC9769105AF76857" blockId="35.[809,1425,1814,1897]" pageId="35" pageNumber="569">
<emphasis id="99BBFB1EA514FF99FC9769105AF76815" box="[809,935,1814,1835]" italics="true" pageId="35" pageNumber="569">Etymology:</emphasis>
The species epithet
<taxonomicName id="6CCF5C8FA514FF99FB3169105DE26815" authorityName="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho" authorityYear="2023" box="[1167,1202,1814,1835]" class="Arachnida" family="Pholcidae" genus="Nerudia" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="569" phylum="Arthropoda" rank="species" species="ola">
<emphasis id="99BBFB1EA514FF99FB3169105DE26815" box="[1167,1202,1814,1835]" italics="true" pageId="35" pageNumber="569">ola</emphasis>
</taxonomicName>
(Spanish for wave) is taken from Pablo Nerudas poem Poema 12; noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="E3D57487A513FF9EFF1D6EC05C776D89" pageId="36" pageNumber="570" type="description">
<paragraph id="AB70270CA513FF9EFF1D6EC0587F6FE3" blockId="36.[163,303,198,221]" box="[163,303,198,221]" pageId="36" pageNumber="570">
<emphasis id="99BBFB1EA513FF9EFF1D6EC0587F6FE3" box="[163,303,198,221]" italics="true" pageId="36" pageNumber="570">Description</emphasis>
</paragraph>
<paragraph id="AB70270CA513FF9EFF1D6EFB5B3D6E92" blockId="36.[163,779,253,429]" pageId="36" pageNumber="570">
<emphasis id="99BBFB1EA513FF9EFF1D6EFB5B766E2D" box="[163,550,253,275]" italics="true" pageId="36" pageNumber="570">
Male (
<typeStatus id="747499AEA513FF9EFF4E6EFB58086E2C" box="[240,344,253,274]" pageId="36" pageNumber="570">holotype</typeStatus>
). Measurements:
</emphasis>
Total body length 1.55, carapace width 0.66. Distance
<collectionCode id="CDDEBFC9A513FF9EFDE16F1A5BCB6E0F" box="[607,667,284,305]" pageId="36" pageNumber="570">PME</collectionCode>
PME 70 µm; diameter PME 65 µm; distance
<collectionCode id="CDDEBFC9A513FF9EFD826F3D5B246E6E" box="[572,628,315,336]" pageId="36" pageNumber="570">PME</collectionCode>
ALE 30 µm; distance
<collectionCode id="CDDEBFC9A513FF9EFEB96F5F58126E50" box="[263,322,345,366]" pageId="36" pageNumber="570">AME</collectionCode>
AME 20 µm; diameter AME 40 µm. Leg 1: 5.64 (1.48 + 0.25 + 1.45 + 1.78 + 0.68), tibia 2: 1.18, tibia 3: 1.03, tibia 4: 1.53; tibia 1 L/d: 22.
</paragraph>
<paragraph id="AB70270CA513FF9EFF1D6FCD5B606D02" blockId="36.[163,779,459,573]" pageId="36" pageNumber="570">
<emphasis id="99BBFB1EA513FF9EFF1D6FCD58C16EDE" box="[163,401,459,480]" italics="true" pageId="36" pageNumber="570">Colour (in ethanol):</emphasis>
Prosoma and legs mostly pale ochre-yellow; carapace with light brown Y-mark behind ocular area; sternum whitish; legs without dark rings; abdomen monochromous pale grey.
</paragraph>
<paragraph id="AB70270CA513FF9EFF1D6C655A5C6DCD" blockId="36.[163,780,611,755]" pageId="36" pageNumber="570">
<emphasis id="99BBFB1EA513FF9EFF1D6C6559B56D46" box="[163,229,611,632]" italics="true" pageId="36" pageNumber="570">Body:</emphasis>
Habitus as in
<figureCitation id="33F43B89A513FF9EFE216C655B486D46" box="[415,536,611,633]" captionStart="Figure 1" captionStartId="4.[164,244,1761,1783]" captionTargetBox="[324,1283,651,1731]" captionTargetId="figure-233@4.[323,1283,650,1731]" captionTargetPageId="4" captionText="Figure 1. Nerudia, live specimens from Argentina. A, N. colina sp. nov., female from Jujuy, between San Salvador and Purmamarca (type locality). B, N. poma sp. nov., male from Salta, NW Campo Quijano (type locality). C, D, N. centaura sp. nov., male and female from Catamarca, E Paso de San Francisco (type locality). E, N. trigo sp. nov., female from Salta, SW Alemania. F, N. guirnalda sp. nov., male from Catamarca, El Rodeo (type locality). G, N. ola sp. nov., male from San Juan, San Agustin de Valle Fertil (type locality). H, N. nono sp. nov., male from Cordoba, E Nono (type locality)." figureDoi="http://doi.org/10.5281/zenodo.7981764" httpUri="https://zenodo.org/record/7981764/files/figure.png" pageId="36" pageNumber="570">Figure 1G</figureCitation>
. Ocular area barely raised. Carapace with indistinct thoracic groove. Clypeus unmodified (only rim slightly more sclerotized than in female). Sternum wider than long (0.46/0.38), with pair of distinct anterior processes near coxae 1. Abdomen globular.
</paragraph>
<paragraph id="AB70270CA513FF9EFF1D6D1A58A66CF4" blockId="36.[163,779,796,971]" pageId="36" pageNumber="570">
<emphasis id="99BBFB1EA513FF9EFF1D6D1A587B6C0F" box="[163,299,796,817]" italics="true" pageId="36" pageNumber="570">Chelicerae:</emphasis>
As in
<figureCitation id="33F43B89A513FF9EFE3D6D1A5B666C0F" box="[387,566,796,818]" captionStart="Figure 27" captionStartId="32.[163,241,1320,1342]" captionTargetBox="[327,1275,196,1282]" captionTargetId="figure-281@32.[323,1283,195,1290]" captionTargetPageId="32" captionText="Figure 27. Nerudia ola sp. nov.; paratype male and paratype female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Ar 23897). AC, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views. DF, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7981848" httpUri="https://zenodo.org/record/7981848/files/figure.png" pageId="36" pageNumber="570">Figure 27G, H</figureCitation>
; with distinctive pair of long frontal apophyses, tips slightly bent downwards, obtuse in frontal view, with some ventral hairs directed downwards (
<figureCitation id="33F43B89A513FF9EFE686D7E5B636CB3" box="[470,563,888,910]" captionStart="Figure 29" captionStartId="34.[163,242,1300,1322]" captionTargetBox="[323,1282,198,1270]" captionTargetId="figure-325@34.[322,1282,197,1271]" captionTargetPageId="34" captionText="Figure 29. Nerudia ola sp. nov.; male and female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Arg141). A, female prosoma, frontal view. B, left procursus tip, prolateral-dorsal view. C, stridulatory file on male chelicera. D, lateral view of female chelicera, showing absence of stridulatory file. E, stridulatory pick (modified hair) on male palpal femur. F, right male cheliceral apophysis, frontal view. G, processes on male sternum. H, male gonopore. Scale lines: 100 µm (A, G), 10 µm (B, D, F, H), 20 µm (C), 2 µm (E)." figureDoi="http://doi.org/10.5281/zenodo.7981854" httpUri="https://zenodo.org/record/7981854/files/figure.png" pageId="36" pageNumber="570">Fig. 29F</figureCitation>
); distance between tips of apophyses: 280 µm; stridulatory files on distinct lateral protrusions (
<figureCitation id="33F43B89A513FF9EFE396DB358B66CF4" box="[391,486,949,971]" captionStart="Figure 29" captionStartId="34.[163,242,1300,1322]" captionTargetBox="[323,1282,198,1270]" captionTargetId="figure-325@34.[322,1282,197,1271]" captionTargetPageId="34" captionText="Figure 29. Nerudia ola sp. nov.; male and female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Arg141). A, female prosoma, frontal view. B, left procursus tip, prolateral-dorsal view. C, stridulatory file on male chelicera. D, lateral view of female chelicera, showing absence of stridulatory file. E, stridulatory pick (modified hair) on male palpal femur. F, right male cheliceral apophysis, frontal view. G, processes on male sternum. H, male gonopore. Scale lines: 100 µm (A, G), 10 µm (B, D, F, H), 20 µm (C), 2 µm (E)." figureDoi="http://doi.org/10.5281/zenodo.7981854" httpUri="https://zenodo.org/record/7981854/files/figure.png" pageId="36" pageNumber="570">Fig. 29C</figureCitation>
).
</paragraph>
<paragraph id="AB70270CA513FF9EFF1D6DF25B176BDE" blockId="36.[163,779,1012,1248]" pageId="36" pageNumber="570">
<emphasis id="99BBFB1EA513FF9EFF1D6DF259BA6B37" box="[163,234,1012,1033]" italics="true" pageId="36" pageNumber="570">Palps:</emphasis>
In general, similar to
<taxonomicName id="6CCF5C8FA513FF9EFE466DF35B306B37" authorityName="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho" authorityYear="2023" box="[504,608,1012,1034]" class="Arachnida" family="Pholcidae" genus="Nerudia" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="570" phylum="Arthropoda" rank="species" species="colina">
<emphasis id="99BBFB1EA513FF9EFE466DF35B306B37" box="[504,608,1012,1034]" italics="true" pageId="36" pageNumber="570">N. colina</emphasis>
</taxonomicName>
(cf.
<figureCitation id="33F43B89A513FF9EFD2F6DF25B836B37" box="[657,723,1012,1034]" captionStart="Figure 4" captionStartId="9.[145,224,1819,1841]" captionTargetBox="[306,1261,1042,1781]" captionTargetId="figure-442@9.[305,1265,1039,1789]" captionTargetPageId="9" captionText="Figure 4. Nerudia colina sp. nov.; paratype male from Argentina, Jujuy, between San Salvador and Purmamarca (ZFMK Ar 23883). Left palp, prolateral (A) and retrolateral (B) views. Abbreviations: b, genital bulb; c, coxa; e, embolus; f, femur; p, procursus; pa, patella; ta, tarsus; ti, tibia; tr, trochanter; va, ventral apophysis. Scale line: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.7981776" httpUri="https://zenodo.org/record/7981776/files/figure.png" pageId="36" pageNumber="570">Fig. 4</figureCitation>
) but femur slightly slenderer (length/width 2.27) and tibia slightly thicker (length/width 0.92); procursus simple, in lateral view slightly directed towards dorsal, with bifid tip consisting of slender dorsal process and wider ventral membranous process (
<figureCitation id="33F43B89A513FF9EFE4B6A8B5B216B9C" box="[501,625,1165,1187]" captionStart="Figure 27" captionStartId="32.[163,241,1320,1342]" captionTargetBox="[327,1275,196,1282]" captionTargetId="figure-281@32.[323,1283,195,1290]" captionTargetPageId="32" captionText="Figure 27. Nerudia ola sp. nov.; paratype male and paratype female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Ar 23897). AC, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views. DF, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7981848" httpUri="https://zenodo.org/record/7981848/files/figure.png" pageId="36" pageNumber="570">Fig. 27AC</figureCitation>
); genital bulb similar to
<taxonomicName id="6CCF5C8FA513FF9EFEA16AAB58D96BFF" authorityName="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho" authorityYear="2023" box="[287,393,1196,1218]" class="Arachnida" family="Pholcidae" genus="Nerudia" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="570" phylum="Arthropoda" rank="species" species="colina">
<emphasis id="99BBFB1EA513FF9EFEA16AAB58D96BFF" box="[287,393,1196,1218]" italics="true" pageId="36" pageNumber="570">N. colina</emphasis>
</taxonomicName>
, with slender ventral apophysis curved towards ventral (
<figureCitation id="33F43B89A513FF9EFE046ACC5B686BDE" box="[442,568,1226,1248]" captionStart="Figure 27" captionStartId="32.[163,241,1320,1342]" captionTargetBox="[327,1275,196,1282]" captionTargetId="figure-281@32.[323,1283,195,1290]" captionTargetPageId="32" captionText="Figure 27. Nerudia ola sp. nov.; paratype male and paratype female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Ar 23897). AC, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views. DF, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7981848" httpUri="https://zenodo.org/record/7981848/files/figure.png" pageId="36" pageNumber="570">Fig. 27DF</figureCitation>
).
</paragraph>
<paragraph id="AB70270CA513FF9EFF1D6B0C5B3B6A86" blockId="36.[163,779,1289,1464]" pageId="36" pageNumber="570">
<emphasis id="99BBFB1EA513FF9EFF1D6B0C598E6A21" box="[163,222,1290,1311]" italics="true" pageId="36" pageNumber="570">Legs:</emphasis>
Without spines and curved hairs; with vertical hairs in higher than usual density on proximal half of tibia 1 only, in two dorsal rows (
<figureCitation id="33F43B89A513FF9EFDAF6B405B206A62" box="[529,624,1350,1372]" captionStart="Figure 30" captionStartId="35.[145,223,1378,1400]" captionTargetBox="[306,1265,196,1347]" captionTargetId="figure-250@35.[305,1265,195,1348]" captionTargetPageId="35" captionText="Figure 30. Nerudia ola sp. nov.; male and female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Arg141). A, male right tibia 1, retrolateral view. B, male metatarsus 4 with trichobothrium. C, male palpal tarsal organ. D, tip of right female palp, dorsal view; arrow points at tarsal organ. E, tarsal organ on male tarsus 2. F, epigynum, ventral view. G, male ALS and PMS. H, female ALS. Scale lines: 10 µm (A, B, D, G, H), 2 µm (C, E), 100 µm (F)." figureDoi="http://doi.org/10.5281/zenodo.7981858" httpUri="https://zenodo.org/record/7981858/files/figure.png" pageId="36" pageNumber="570">Fig. 30A</figureCitation>
); retrolateral trichobothrium of tibia 1 at 58%; prolateral trichobothrium absent on tibia 1; tarsus 1 with six to seven pseudosegments, distally distinct.
</paragraph>
<paragraph id="AB70270CA513FF9EFF1D6BE45C366EAD" blockId="36.[163,780,1505,1895]" lastBlockId="36.[827,1443,197,403]" pageId="36" pageNumber="570">
<emphasis id="99BBFB1EA513FF9EFF1D6BE4582A6AC8" box="[163,378,1505,1527]" italics="true" pageId="36" pageNumber="570">Variation (male):</emphasis>
<materialsCitation id="1BA72D51A513FF9EFE396BE75C326EAD" county="The" location="This" municipality="In" pageId="36" pageNumber="570" specimenCount="36" specimenCount-male="36" stateProvince="La Rioja and Catamarca" typeStatus="holotype">
Tibia
<quantity id="6C378AE9A513FF9EFE6D6BE75B556AC9" box="[467,517,1505,1527]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="36" pageNumber="570" unit="in" value="1.0">1 in</quantity>
<specimenCount id="BDC9EC85A513FF9EFDAF6BE75BD16AC9" box="[529,641,1505,1527]" pageId="36" pageNumber="570" type="male">36 males</specimenCount>
(including
<typeStatus id="747499AEA513FF9EFF1D680658556928" box="[163,261,1536,1558]" pageId="36" pageNumber="570">holotype</typeStatus>
): 1.041.60 (mean 1.33).
<collectingCounty id="42115F80A513FF9EFDA668065B136928" box="[536,579,1536,1558]" pageId="36" pageNumber="570">The</collectingCounty>
distance between the tips of the cheliceral apophyses varies considerably (235340 µm), but also within localities (e.g. Nacimientos: 240310 µm).
<collectingMunicipality id="4B14BD76A513FF9EFE6C685A58BB694F" box="[466,491,1628,1649]" pageId="36" pageNumber="570">In</collectingMunicipality>
northern specimens (from
<collectingRegion id="690BE9EEA513FF9EFF1D687D58E969AE" box="[163,441,1658,1680]" isEnumeration="true" pageId="36" pageNumber="570">La Rioja and Catamarca</collectingRegion>
), the cheliceral apophyses are slightly straighter and longer.
<location id="AE1071D7A513FF9EFE52689F5B4E6991" LSID="urn:lsid:plazi:treatment:2366961AA517FF9EFE98683C5C286A97:AE1071D7A513FF9EFE52689F5B4E6991" box="[492,542,1689,1711]" county="The" municipality="In" name="This" pageId="36" pageNumber="570" stateProvince="La Rioja and Catamarca">This</location>
difference is minimal and some males appear intermediate.
<location id="AE1071D7A513FF9EFDD068BE5BD869F3" LSID="urn:lsid:plazi:treatment:2366961AA517FF9EFE98683C5C286A97:AE1071D7A513FF9EFDD068BE5BD869F3" box="[622,648,1720,1741]" county="The" municipality="In" name="In" pageId="36" pageNumber="570" stateProvince="La Rioja and Catamarca">In</location>
specimens from
<location id="AE1071D7A513FF9EFF5A68D0583F69D2" LSID="urn:lsid:plazi:treatment:2366961AA517FF9EFE98683C5C286A97:AE1071D7A513FF9EFF5A68D0583F69D2" box="[228,367,1750,1772]" county="The" municipality="In" name="Chumbicha" pageId="36" pageNumber="570" stateProvince="La Rioja and Catamarca">Chumbicha</location>
, the procursus is slightly shorter.
<location id="AE1071D7A513FF9EFF1D68F358436835" LSID="urn:lsid:plazi:treatment:2366961AA517FF9EFE98683C5C286A97:AE1071D7A513FF9EFF1D68F358436835" box="[163,275,1781,1803]" county="The" municipality="In" name="Abdomen" pageId="36" pageNumber="570" stateProvince="La Rioja and Catamarca">Abdomen</location>
variably dark.
<emphasis id="99BBFB1EA513FF9EFE6B68F35B566834" box="[469,518,1781,1802]" italics="true" pageId="36" pageNumber="570">COI</emphasis>
data indicate a deep split between northern and southern specimens, and a phylogenetic analysis of the molecular data (Supporting Information,
<figureCitation id="33F43B89A513FF9EFE86695758D86858" box="[312,392,1873,1895]" captionStart="Figure 2" captionStartId="5.[143,224,1170,1192]" captionTargetBox="[152,1415,197,1139]" captionTargetId="figure-346@5.[151,1415,195,1140]" captionTargetPageId="5" captionText="Figure 2. NJ COI tree of all sequenced Nerudia and Gertschiola specimens. Species groups on the right are working hypotheses. Photo: female of Nerudia colina sp. nov. For complete tree, and for tree resulting from analysis with IQ-TREE, see Supporting Information, Figures S1, S2." figureDoi="http://doi.org/10.5281/zenodo.7981768" httpUri="https://zenodo.org/record/7981768/files/figure.png" pageId="36" pageNumber="570">Fig. S2</figureCitation>
) even questions the monophyly of the northern + southern clade.
<location id="AE1071D7A513FF9EFB316EC35DA16FE4" LSID="urn:lsid:plazi:treatment:2366961AA517FF9EFE98683C5C286A97:AE1071D7A513FF9EFB316EC35DA16FE4" box="[1167,1265,197,218]" county="The" municipality="In" name="However" pageId="36" pageNumber="570" stateProvince="La Rioja and Catamarca">However</location>
, deep splits also occur among southern specimens and among northern specimens (
<figureCitation id="33F43B89A513FF9EFC786F045D596E29" box="[966,1033,258,280]" captionStart="Figure 2" captionStartId="5.[143,224,1170,1192]" captionTargetBox="[152,1415,197,1139]" captionTargetId="figure-346@5.[151,1415,195,1140]" captionTargetPageId="5" captionText="Figure 2. NJ COI tree of all sequenced Nerudia and Gertschiola specimens. Species groups on the right are working hypotheses. Photo: female of Nerudia colina sp. nov. For complete tree, and for tree resulting from analysis with IQ-TREE, see Supporting Information, Figures S1, S2." figureDoi="http://doi.org/10.5281/zenodo.7981768" httpUri="https://zenodo.org/record/7981768/files/figure.png" pageId="36" pageNumber="570">Fig. 2</figureCitation>
).
<location id="AE1071D7A513FF9EFB9E6F045D6E6E29" LSID="urn:lsid:plazi:treatment:2366961AA517FF9EFE98683C5C286A97:AE1071D7A513FF9EFB9E6F045D6E6E29" box="[1056,1086,258,279]" county="The" municipality="In" name="As" pageId="36" pageNumber="570" stateProvince="La Rioja and Catamarca">As</location>
a result, a preliminary ASAP analysis (Supporting Information,
<figureCitation id="33F43B89A513FF9EFB7F6F275C476E08" box="[1217,1303,289,311]" captionStart="Figure 3" captionStartId="8.[164,242,1848,1870]" captionTargetBox="[324,1283,713,1818]" captionTargetId="figure-278@8.[323,1283,712,1819]" captionTargetPageId="8" captionText="Figure 3. Known distribution of Nerudia in South America (inset) and of individual species. Regions in Chile: AT, Atacama; COQ, Coquimbo. Provinces in Argentina: CA, Catamarca; COR, Córdoba; JU, Jujuy; LR, La Rioja; SA, Salta; SJ, San Juan." figureDoi="http://doi.org/10.5281/zenodo.7981770" httpUri="https://zenodo.org/record/7981770/files/figure.png" pageId="36" pageNumber="570">Figs S3</figureCitation>
, S
<figureCitation id="33F43B89A513FF9EFA8F6F275C106E08" box="[1329,1344,289,310]" captionStart="Figure 4" captionStartId="9.[145,224,1819,1841]" captionTargetBox="[306,1261,1042,1781]" captionTargetId="figure-442@9.[305,1265,1039,1789]" captionTargetPageId="9" captionText="Figure 4. Nerudia colina sp. nov.; paratype male from Argentina, Jujuy, between San Salvador and Purmamarca (ZFMK Ar 23883). Left palp, prolateral (A) and retrolateral (B) views. Abbreviations: b, genital bulb; c, coxa; e, embolus; f, femur; p, procursus; pa, patella; ta, tarsus; ti, tibia; tr, trochanter; va, ventral apophysis. Scale line: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.7981776" httpUri="https://zenodo.org/record/7981776/files/figure.png" pageId="36" pageNumber="570">4</figureCitation>
) favours the existence of up to seven species (lowest scores of 2.0 and 3.0).
<location id="AE1071D7A513FF9EFC1B6F585AB46E4A" LSID="urn:lsid:plazi:treatment:2366961AA517FF9EFE98683C5C286A97:AE1071D7A513FF9EFC1B6F585AB46E4A" box="[933,996,350,372]" county="The" municipality="In" name="What" pageId="36" pageNumber="570" stateProvince="La Rioja and Catamarca">What</location>
is here interpreted as one species may thus in fact be several more or less cryptic species
</materialsCitation>
.
</paragraph>
<paragraph id="AB70270CA513FF9EFC856FAA5C776D89" blockId="36.[827,1443,428,695]" pageId="36" pageNumber="570">
<emphasis id="99BBFB1EA513FF9EFC856FAA5AC96EFF" box="[827,921,428,449]" italics="true" pageId="36" pageNumber="570">Female:</emphasis>
In general, similar to male but sternum without pair of anterior humps and tibia 1 with usual low number of short vertical hairs. Tibia
<quantity id="6C378AE9A513FF9EFB476FEF5C726EC1" box="[1273,1314,489,511]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="36" pageNumber="570" unit="in" value="1.0">1 in</quantity>
<specimenCount id="BDC9EC85A513FF9EFA9B6FEF5CCF6EC1" box="[1317,1439,489,511]" pageId="36" pageNumber="570" type="female">57 females</specimenCount>
: 0.971.55 (mean 1.25). Epigynum (
<figureCitation id="33F43B89A513FF9EFB536C0E5C0A6D23" box="[1261,1370,520,542]" captionStart="Figure 28" captionStartId="33.[146,224,974,996]" captionTargetBox="[306,1265,197,944]" captionTargetId="figure-546@33.[305,1265,196,945]" captionTargetPageId="33" captionText="Figure 28. Nerudia ola sp. nov.; paratype female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Ar 23897). A, abdomen and epigynum, ventral view. B, median structures in internal genitalia. C, D, cleared genitalia, ventral and dorsal views. Scale lines: 0.2 mm (A, C, D), 0.1 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.7981852" httpUri="https://zenodo.org/record/7981852/files/figure.png" pageId="36" pageNumber="570">Figs 28A</figureCitation>
,
<figureCitation id="33F43B89A513FF9EFAD26C0E5CCA6D23" box="[1388,1434,520,541]" captionStart="Figure 30" captionStartId="35.[145,223,1378,1400]" captionTargetBox="[306,1265,196,1347]" captionTargetId="figure-250@35.[305,1265,195,1348]" captionTargetPageId="35" captionText="Figure 30. Nerudia ola sp. nov.; male and female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Arg141). A, male right tibia 1, retrolateral view. B, male metatarsus 4 with trichobothrium. C, male palpal tarsal organ. D, tip of right female palp, dorsal view; arrow points at tarsal organ. E, tarsal organ on male tarsus 2. F, epigynum, ventral view. G, male ALS and PMS. H, female ALS. Scale lines: 10 µm (A, B, D, G, H), 2 µm (C, E), 100 µm (F)." figureDoi="http://doi.org/10.5281/zenodo.7981858" httpUri="https://zenodo.org/record/7981858/files/figure.png" pageId="36" pageNumber="570">30F</figureCitation>
) anterior plate weakly protruding, without posterior indentation; posterior plate large, simple. Internal genitalia (
<figureCitation id="33F43B89A513FF9EFC116C625D5C6D47" box="[943,1036,612,634]" captionStart="Figure 27" captionStartId="32.[163,241,1320,1342]" captionTargetBox="[327,1275,196,1282]" captionTargetId="figure-281@32.[323,1283,195,1290]" captionTargetPageId="32" captionText="Figure 27. Nerudia ola sp. nov.; paratype male and paratype female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Ar 23897). AC, left male palpal tarsus and procursus, prolateral, dorsal, and retrolateral views. DF, left male genital bulb, prolateral, dorsal, and retrolateral views. G, H, male chelicerae, frontal and lateral views. I, cleared female genitalia, dorsal view. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7981848" httpUri="https://zenodo.org/record/7981848/files/figure.png" pageId="36" pageNumber="570">Figs 27I</figureCitation>
,
<figureCitation id="33F43B89A513FF9EFBA76C625D386D47" box="[1049,1128,612,633]" captionStart="Figure 28" captionStartId="33.[146,224,974,996]" captionTargetBox="[306,1265,197,944]" captionTargetId="figure-546@33.[305,1265,196,945]" captionTargetPageId="33" captionText="Figure 28. Nerudia ola sp. nov.; paratype female from Argentina, San Juan, San Agustín de Valle Fértil (ZFMK Ar 23897). A, abdomen and epigynum, ventral view. B, median structures in internal genitalia. C, D, cleared genitalia, ventral and dorsal views. Scale lines: 0.2 mm (A, C, D), 0.1 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.7981852" httpUri="https://zenodo.org/record/7981852/files/figure.png" pageId="36" pageNumber="570">28BD</figureCitation>
) simple, with barely visible transparent receptacle. No difference could be seen between northern and southern specimens.
</paragraph>
</subSubSection>
<subSubSection id="E3D57487A513FF9EFC856CE65D756C6F" pageId="36" pageNumber="570" type="distribution">
<paragraph id="AB70270CA513FF9EFC856CE65D756C6F" blockId="36.[827,1444,736,849]" pageId="36" pageNumber="570">
<emphasis id="99BBFB1EA513FF9EFC856CE65AB26DCB" box="[827,994,736,757]" italics="true" pageId="36" pageNumber="570">Distribution:</emphasis>
Widely distributed in
<collectingCountry id="D3D8679CA513FF9EFAA76CE65CCC6DC8" box="[1305,1436,736,758]" name="Argentina" pageId="36" pageNumber="570">Argentina</collectingCountry>
, Provinces San Juan,
<collectingRegion id="690BE9EEA513FF9EFB926CF95DDC6C2A" box="[1068,1164,766,788]" country="Argentina" name="La Rioja" pageId="36" pageNumber="570">La Rioja</collectingRegion>
and
<collectingRegion id="690BE9EEA513FF9EFB786CF85C156C2D" box="[1222,1349,766,787]" country="Argentina" name="Catamarca" pageId="36" pageNumber="570">Catamarca</collectingRegion>
(
<figureCitation id="33F43B89A513FF9EFAEA6CF85CC56C2D" box="[1364,1429,766,788]" captionStart="Figure 3" captionStartId="8.[164,242,1848,1870]" captionTargetBox="[324,1283,713,1818]" captionTargetId="figure-278@8.[323,1283,712,1819]" captionTargetPageId="8" captionText="Figure 3. Known distribution of Nerudia in South America (inset) and of individual species. Regions in Chile: AT, Atacama; COQ, Coquimbo. Provinces in Argentina: CA, Catamarca; COR, Córdoba; JU, Jujuy; LR, La Rioja; SA, Salta; SJ, San Juan." figureDoi="http://doi.org/10.5281/zenodo.7981770" httpUri="https://zenodo.org/record/7981770/files/figure.png" pageId="36" pageNumber="570">Fig. 3</figureCitation>
). Note that specimens from
<collectingRegion id="690BE9EEA513FF9EFBDA6D1B5D936C0D" box="[1124,1219,797,819]" country="Argentina" name="La Rioja" pageId="36" pageNumber="570">La Rioja</collectingRegion>
and
<collectingRegion id="690BE9EEA513FF9EFB446D1B5C286C0C" box="[1274,1400,797,818]" country="Argentina" name="Catamarca" pageId="36" pageNumber="570">Catamarca</collectingRegion>
are assigned tentatively.
</paragraph>
</subSubSection>
<subSubSection id="E3D57487A513FF9EFC856D8C5C286A97" pageId="36" pageNumber="570" type="biology_ecology">
<paragraph id="AB70270CA513FF9EFC856D8C5C286A97" blockId="36.[827,1443,906,1449]" pageId="36" pageNumber="570">
<emphasis id="99BBFB1EA513FF9EFC856D8C5AA26CA1" box="[827,1010,906,927]" italics="true" pageId="36" pageNumber="570">Natural history:</emphasis>
All specimens were found by turning rocks. The spiders started to run rapidly when disturbed but then stopped suddenly and did barely ever drop from the rock. They were found in a variety of habitats, ranging from a relatively humid block field in a low forest at
<quantity id="6C378AE9A513FF9EFBB26A255D046B07" box="[1036,1108,1059,1081]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="36" pageNumber="570" unit="m" value="640.0">640 m</quantity>
a.s.l. near Chumbicha to an exposed arid hill with dry ravines at
<quantity id="6C378AE9A513FF9EFB656A445C7E6B66" box="[1243,1326,1090,1112]" metricMagnitude="3" metricUnit="m" metricValue="2.12" pageId="36" pageNumber="570" unit="m" value="2120.0">2120 m</quantity>
a.s.l. near Nacimientos.
<taxonomicName id="6CCF5C8FA513FF9EFC6C6A675D206B49" authority="Fertil" authorityName="Fertil" box="[978,1136,1121,1143]" class="Arachnida" family="Pholcidae" genus="Artema" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="570" phylum="Arthropoda" rank="species" species="valle">At Valle Fértil</taxonomicName>
, the spiders seemed to have a patchy distribution, with up to
<specimenCount id="BDC9EC85A513FF9EFB156A795C776BAB" box="[1195,1319,1151,1173]" pageId="36" pageNumber="570" type="adult">eight adult</specimenCount>
specimens on a single large rock in suitable places (with some shade and leaf litter among the rocks). They shared the microhabitat with a superficially similar species of
<taxonomicName id="6CCF5C8FA513FF9EFCE86AFD5D306A31" authority="Simon, 1893" authorityName="Simon" authorityYear="1893" box="[854,1120,1274,1296]" class="Arachnida" family="Pholcidae" genus="Metagonia" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="99BBFB1EA513FF9EFCE86AFD5A9E6A2E" box="[854,974,1275,1296]" italics="true" pageId="36" pageNumber="570">Metagonia</emphasis>
<bibRefCitation id="CF5E5AFDA513FF9EFC6A6AFC5D306A31" author="Simon E" box="[980,1120,1274,1296]" pageId="36" pageNumber="570" refId="ref33979" refString="Simon E. 1893. Histoire Naturelle des Araignees, 2 nd edn. Paris: Encyclopedie Roret." type="book" year="1893">Simon, 1893</bibRefCitation>
</taxonomicName>
(
<emphasis id="99BBFB1EA513FF9EFBD06AFD5DDD6A2E" box="[1134,1165,1275,1296]" italics="true" pageId="36" pageNumber="570">M.</emphasis>
MACN79; undescribed;
<taxonomicName id="6CCF5C8FA513FF9EFC856B1F5AED6A11" authority="Fertil" authorityName="Fertil" box="[827,957,1305,1327]" class="Arachnida" family="Pholcidae" genus="Artema" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="570" phylum="Arthropoda" rank="species" species="valle">Valle Fértil</taxonomicName>
; Chumbicha), with
<taxonomicName id="6CCF5C8FA513FF9EFB226B1F5CF26A10" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1941" box="[1180,1442,1305,1326]" class="Arachnida" family="Pholcidae" genus="Gertschiola" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="570" phylum="Arthropoda" rank="species" species="macrostyla">
<emphasis id="99BBFB1EA513FF9EFB226B1F5CF26A10" box="[1180,1442,1305,1326]" italics="true" pageId="36" pageNumber="570">Gertschiola macrostyla</emphasis>
</taxonomicName>
(Astica;
<taxonomicName id="6CCF5C8FA513FF9EFC296B315D4A6A73" authority="Fertil" authorityName="Fertil" box="[919,1050,1335,1357]" class="Arachnida" family="Pholcidae" genus="Artema" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="570" phylum="Arthropoda" rank="species" species="valle">Valle Fértil</taxonomicName>
; Ischigualasto; SE Aimogasta site 2), with
<taxonomicName id="6CCF5C8FA513FF9EFC1E6B505DAB6A55" authority="Huber, 2000" authorityName="Huber" authorityYear="2000" box="[928,1275,1366,1388]" class="Arachnida" family="Pholcidae" genus="Chibchea" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="570" phylum="Arthropoda" rank="species" species="araona">
<emphasis id="99BBFB1EA513FF9EFC1E6B505D366A52" box="[928,1126,1366,1388]" italics="true" pageId="36" pageNumber="570">Chibchea araona</emphasis>
<bibRefCitation id="CF5E5AFDA513FF9EFBD06B505DAB6A55" author="Huber BA" box="[1134,1275,1366,1388]" pageId="36" pageNumber="570" pagination="1 - 348" refId="ref32245" refString="Huber BA. 2000. New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History 254: 1 - 348." type="journal article" year="2000">Huber, 2000</bibRefCitation>
</taxonomicName>
(?) (Cuesta de
<collectingRegion id="690BE9EEA513FF9EFC856B735AF46AB5" box="[827,932,1397,1419]" country="Venezuela" name="Miranda" pageId="36" pageNumber="570">Miranda</collectingRegion>
, both sites) and, in at least one case, with another species of
<taxonomicName id="6CCF5C8FA513FF9EFBB36B955D3C6A96" box="[1037,1132,1427,1448]" class="Arachnida" family="Pholcidae" genus="Nerudia" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="99BBFB1EA513FF9EFBB36B955D3C6A96" box="[1037,1132,1427,1448]" italics="true" pageId="36" pageNumber="570">Nerudia</emphasis>
</taxonomicName>
(
<taxonomicName id="6CCF5C8FA513FF9EFBC56B925D896A97" authorityName="Huber &amp; Meng &amp; Král &amp; Ávila Herrera &amp; Izquierdo &amp; Carvalho" authorityYear="2023" box="[1147,1241,1428,1449]" class="Arachnida" family="Pholcidae" genus="Nerudia" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="570" phylum="Arthropoda" rank="species" species="poma">
<emphasis id="99BBFB1EA513FF9EFBC56B925D896A97" box="[1147,1241,1428,1449]" italics="true" pageId="36" pageNumber="570">N. poma</emphasis>
</taxonomicName>
; Chumbicha).
</paragraph>
</subSubSection>
</treatment>
</document>